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<document id="D36B1680C80D2F1C21D50AE07F1BC8D5" ID-CLB-Dataset="20497" ID-DOI="10.5169/seals-787048" ID-GBIF-Dataset="8c468ca9-0ee3-4200-b5a3-6adfd98caaea" ID-ISSN="2624-9170" ID-Zenodo-Dep="6284316" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1637935242462" checkinUser="carolina" docAuthor="Balkenohl, Michael" docDate="2017" docId="2559A84DA84BFFE8D6FFFBD68E7EFC24" docLanguage="en" docName="ContribNatHist.35.1-66.pdf.imf" docOrigin="Contributions to Natural History 35" docStyle="DocumentStyle:D319E9290ED630111D2A11900DAFC11F.4:ContribNatHist.2011-.journal_article" docStyleId="D319E9290ED630111D2A11900DAFC11F" docStyleName="ContribNatHist.2011-.journal_article" docStyleVersion="4" docTitle="Orictites mjoebergi LouWERENS 1964" docType="treatment" docVersion="12" lastPageNumber="29" masterDocId="D960D035A853FFF4D660FF9B8D45FFA1" masterDocTitle="Revision of the genus Orictites ANDREWES, 1931 (Coleoptera, Carabidae, Clivinini)" masterLastPageNumber="68" masterPageNumber="1" pageNumber="25" updateTime="1699470517822" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="DD66C8F82191D668717EE05ACCDAAD03">Revision of the genus Orictites ANDREWES, 1931 (Coleoptera, Carabidae, Clivinini)</mods:title>
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<mods:namePart id="0CAC4F73E9F64C4FAB713B186E11F363">Balkenohl, Michael</mods:namePart>
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<mods:title id="4CBB04E55E2504C0777DC116DDBFF4A1">Contributions to Natural History</mods:title>
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<mods:date id="B76376282C6383C92539DFEE96029522">2017</mods:date>
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<mods:identifier id="BB5FD8B74A9EC75B602B1B5E1FBA6EC2" type="DOI">10.5169/seals-787048</mods:identifier>
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<treatment id="2559A84DA84BFFE8D6FFFBD68E7EFC24" ID-DOI="http://doi.org/10.5281/zenodo.6315574" ID-GBIF-Taxon="193586564" ID-Zenodo-Dep="6315574" LSID="urn:lsid:plazi:treatment:2559A84DA84BFFE8D6FFFBD68E7EFC24" httpUri="http://treatment.plazi.org/id/2559A84DA84BFFE8D6FFFBD68E7EFC24" lastPageId="28" lastPageNumber="29" pageId="24" pageNumber="25">
<subSubSection id="E5EA4AD0A84BFFECD6FFFBD68F09FBC4" box="[159,588,1101,1126]" pageId="24" pageNumber="25" type="nomenclature">
<paragraph id="AD4F195BA84BFFECD6FFFBD68F09FBC4" blockId="24.[159,588,1101,1126]" box="[159,588,1101,1126]" pageId="24" pageNumber="25">
<heading id="F607AE37A84BFFECD6FFFBD68F09FBC4" box="[159,588,1101,1126]" centered="true" fontSize="12" level="2" pageId="24" pageNumber="25" reason="2">
<taxonomicName id="6AF062D8A84BFFECD6FFFBD68F09FBC4" authority="LouWERENS, 1964" authorityName="LouWERENS" authorityYear="1964" box="[159,588,1101,1126]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="mjoebergi">
<emphasis id="9F84C549A84BFFECD6FFFBD68C3CFBC7" bold="true" box="[159,377,1101,1126]" italics="true" pageId="24" pageNumber="25">Orictites mjoebergi</emphasis>
<bibRefCitation id="C96164AAA84BFFECD71FFBD68F09FBC4" author="Louwerens, C. J." box="[383,588,1101,1125]" pageId="24" pageNumber="25" pagination="171 - 189" refId="ref27961" refString="Louwerens, C. J. (1964): An annotated list of the Carabidae, chiefly collected in East Borneo by Dr Eric MjOberg with descriptions of new species. - Entomologisk Tidskrift, Stockholm 85: 171 - 189." type="journal article" year="1964">LouWERENS, 1964</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A84BFFECD6FFFB038C57FB11" box="[159,274,1176,1200]" pageId="24" pageNumber="25" type="discussion">
<paragraph id="AD4F195BA84BFFECD6FFFB038C57FB11" blockId="24.[159,755,1176,1312]" box="[159,274,1176,1200]" pageId="24" pageNumber="25">Catalogue</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A84BFFECD6FFFB268FB6FA81" pageId="24" pageNumber="25" type="reference_group">
<paragraph id="AD4F195BA84BFFECD6FFFB268FCCFB74" blockId="24.[159,755,1176,1312]" box="[159,649,1213,1237]" pageId="24" pageNumber="25">
<treatmentCitationGroup id="8DE03E75A84BFFECD6FFFB268FCCFB74" box="[159,649,1213,1237]" pageId="24" pageNumber="25">
<emphasis id="9F84C549A84BFFECD6FFFB268CC5FB74" box="[159,384,1213,1237]" italics="true" pageId="24" pageNumber="25">
<taxonomicName id="6AF062D8A84BFFECD6FFFB268C3CFB74" authorityName="LouWERENS" authorityYear="1964" box="[159,377,1213,1237]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="mjoebergi">Orictites mjoebergi</taxonomicName>
;
</emphasis>
<treatmentCitation id="2C513F4AA84BFFECD7E6FB268FC4FB74" author="Louwerens, C. J." box="[390,641,1213,1237]" page="172" pageId="24" pageNumber="25" year="1964">
<bibRefCitation id="C96164AAA84BFFECD7E6FB268FC4FB74" author="Louwerens, C. J." box="[390,641,1213,1237]" pageId="24" pageNumber="25" pagination="171 - 189" refId="ref27961" refString="Louwerens, C. J. (1964): An annotated list of the Carabidae, chiefly collected in East Borneo by Dr Eric MjOberg with descriptions of new species. - Entomologisk Tidskrift, Stockholm 85: 171 - 189." type="journal article" year="1964">Louwerens, 1964: 172</bibRefCitation>
</treatmentCitation>
;
</treatmentCitationGroup>
</paragraph>
<paragraph id="AD4F195BA84BFFECD6FFFB798F99FB5B" blockId="24.[159,755,1176,1312]" box="[159,732,1250,1274]" pageId="24" pageNumber="25">
<treatmentCitationGroup id="8DE03E75A84BFFECD6FFFB798F99FB5B" box="[159,732,1250,1274]" pageId="24" pageNumber="25">
<taxonomicName id="6AF062D8A84BFFECD6FFFB798F46FB5B" authority="LouWERENS" authorityName="LouWERENS" authorityYear="1964" box="[159,515,1250,1274]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="mjoebergi">
<emphasis id="9F84C549A84BFFECD6FFFB798C3CFB5B" box="[159,377,1250,1274]" italics="true" pageId="24" pageNumber="25">Orictites mjoebergi</emphasis>
LouWERENS
</taxonomicName>
;
<treatmentCitation id="2C513F4AA84BFFECD471FB798F91FB5B" author="Lorenz, W." box="[529,724,1250,1274]" page="135" pageId="24" pageNumber="25" year="1998">
<bibRefCitation id="C96164AAA84BFFECD471FB798F91FB5B" author="Lorenz, W." box="[529,724,1250,1274]" pageId="24" pageNumber="25" refId="ref28004" refString="Lorenz, W. (1998): Systematic list of extant Ground Beetles of the World (Insecta Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodidae). - Tutzing, printed by the author. 502 pp." type="book" year="1998">Lorenz 1998: 135</bibRefCitation>
</treatmentCitation>
;
</treatmentCitationGroup>
</paragraph>
<paragraph id="AD4F195BA84BFFECD6FFFA938FB6FA81" blockId="24.[159,755,1176,1312]" box="[159,755,1288,1312]" pageId="24" pageNumber="25">
<treatmentCitationGroup id="8DE03E75A84BFFECD6FFFA938FB6FA81" box="[159,755,1288,1312]" pageId="24" pageNumber="25">
<taxonomicName id="6AF062D8A84BFFECD6FFFA938F46FA81" authority="LouWERENS" authorityName="LouWERENS" authorityYear="1964" box="[159,515,1288,1312]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="mjoebergi">
<emphasis id="9F84C549A84BFFECD6FFFA938C3CFA81" box="[159,377,1288,1312]" italics="true" pageId="24" pageNumber="25">Orictites mjoebergi</emphasis>
LouWERENS
</taxonomicName>
;
<treatmentCitation id="2C513F4AA84BFFECD471FA938FAEFA81" author="Balkenohl, M." box="[529,747,1288,1312]" page="18" pageId="24" pageNumber="25" year="2001">
<bibRefCitation id="C96164AAA84BFFECD471FA938FAEFA81" author="Balkenohl, M." box="[529,747,1288,1312]" pageId="24" pageNumber="25" refId="ref27660" refString="Balkenohl, M. (2001): Key and Catalogue of the tribe Clivinini from the Oriental realm, with revisions of the genera Thliboclivina KULT, and Trilophidius JEANNEL (Insecta, Coleoptera, Carabidae, Scarititae, Clivinini). - 83 pp., Sofia, Moscow (Pensoft Series Faunistica 21)." type="book" year="2001">Balkenohl 2001: 18</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A84BFFECD6FFFAC98C09F984" pageId="24" pageNumber="25" type="materials_examined">
<paragraph id="AD4F195BA84BFFECD6FFFAC98C09F984" blockId="24.[159,1043,1362,1573]" pageId="24" pageNumber="25">
Material examined:
<materialsCitation id="1D981306A84BFFECD7EEFAC98C09F984" ID-GBIF-Occurrence="3497007340" collectionCode="NHRS" collectorName="O. Borneo MjOberg" determinerName="C. J. Louwerens" location="Pajau River" municipality="Pajau River" pageId="24" pageNumber="25" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
Type material:
<typeStatus id="724BA7F9A84BFFECD423FAC98FF0FACB" box="[579,693,1362,1386]" pageId="24" pageNumber="25" type="lectotype">Lectotype</typeStatus>
(by present designation).
<specimenCount id="BBF6D2D2A84BFFECD599FAC88948FACB" box="[1017,1037,1363,1386]" pageId="24" pageNumber="25" type="male">♂</specimenCount>
, with labels and data: white, handwritten: “Type
<taxonomicName id="6AF062D8A84BFFECD4AAFAE38E6EFA31" authorityName="ANDREWES" authorityYear="1931" box="[714,811,1400,1424]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">Orictites</taxonomicName>
mjÖbergi sp.n.” and printed “det.
<determinerName id="5893F59FA84BFFECD749FA068C91FA14" box="[297,468,1437,1461]" pageId="24" pageNumber="25">C.J.Louwerens</determinerName>
” / blue: “3491 E91 +” / 2
<superScript id="5A85B413A84BFFECD499FA018E55FA0C" attach="left" box="[761,784,1434,1453]" fontSize="8" pageId="24" pageNumber="25">nd</superScript>
pin with original small paper card: small, white, “
<location id="A82F4F80A84BFFECD7ACFA598F10FA7B" LSID="urn:lsid:plazi:treatment:2559A84DA84BFFE8D6FFFBD68E7EFC24:A82F4F80A84BFFECD7ACFA598F10FA7B" box="[460,597,1474,1498]" municipality="Pajau River" name="Pajau River" pageId="24" pageNumber="25">Pajau River</location>
” / white, “O. Borneo MjÖberg” / small blue: “3492 E91 +” / big, white, “
<taxonomicName id="6AF062D8A84BFFECD452FA738FB1F9A1" authority="Andr." authorityName="ANDREWES" authorityYear="1931" box="[562,756,1512,1536]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">ORICTITES Andr.</taxonomicName>
” / big, white: “MjÖbergi Louw.” (
<collectionCode id="CBE1819EA84BFFECD698F9968C07F984" box="[248,322,1549,1573]" collectionName="Sweden, Stockholm, Naturhistoriska riksmuseet" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="24" pageNumber="25">NHRS</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A84AFFEDD6FEFFC68F00FD04" pageId="25" pageNumber="26" type="discussion">
<paragraph id="AD4F195BA84AFFEDD6FEFFC68F00FD04" blockId="25.[121,1006,93,938]" pageId="25" pageNumber="26">
Remarks: C.J.
<bibRefCitation id="C96164AAA84AFFEDD720FFC68F4AFFD4" author="Louwerens, C. J." box="[320,527,93,117]" pageId="25" pageNumber="26" pagination="171 - 189" refId="ref27961" refString="Louwerens, C. J. (1964): An annotated list of the Carabidae, chiefly collected in East Borneo by Dr Eric MjOberg with descriptions of new species. - Entomologisk Tidskrift, Stockholm 85: 171 - 189." type="journal article" year="1964">Louwerens (1964)</bibRefCitation>
based the description on
<specimenCount id="BBF6D2D2A84AFFEDD521FFC68EA8FFD4" box="[833,1005,93,117]" pageId="25" pageNumber="26" type="generic">two specimens</specimenCount>
from Pajan River. One of them is located in the NHRS (see material examined), the other one could not be located and is obviously lost. The available specimen isdesignated here as
<typeStatus id="724BA7F9A84AFFEDD7C7FF568F57FF44" box="[423,530,205,229]" pageId="25" pageNumber="26" type="lectotype">lectotype</typeStatus>
in the interst of the stability. In the specimen the right wing of the clypeus is slightly more angled (dorsal view) and more fused with the median part. I recognise this character as intraindividual variation. Exactly this asymmetric character is described by
<bibRefCitation id="C96164AAA84AFFEDD511FEA68D80FEDB" author="Louwerens, C. J." pageId="25" pageNumber="26" pagination="171 - 189" refId="ref27961" refString="Louwerens, C. J. (1964): An annotated list of the Carabidae, chiefly collected in East Borneo by Dr Eric MjOberg with descriptions of new species. - Entomologisk Tidskrift, Stockholm 85: 171 - 189." type="journal article" year="1964">Louwerens (1964)</bibRefCitation>
. However, careful investigation revealed that this specimen is a male and not a female as indicated by Louwerens. The inner structures were slightly damaged, obviously by a former mounting attempt. The left antennae misses segments 8–11. It should be mentioned that the sketch provided by Louwerens does not display a realistic picture of the species. In addition,
<bibRefCitation id="C96164AAA84AFFEDD512FE638C45FD94" author="Louwerens, C. J." pageId="25" pageNumber="26" pagination="171 - 189" refId="ref27961" refString="Louwerens, C. J. (1964): An annotated list of the Carabidae, chiefly collected in East Borneo by Dr Eric MjOberg with descriptions of new species. - Entomologisk Tidskrift, Stockholm 85: 171 - 189." type="journal article" year="1964">Louwerens (1964: 173)</bibRefCitation>
describes the presence of five setigerous punctures on interval 3 of the elytra. However, the type specimen possesses four, respectively five setigerous punctures (right/left side). The additional specimens all exhibit four setigerous punctures on either side.
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A84AFFEDD6FEFD298C3EFC0B" pageId="25" pageNumber="26" type="materials_examined">
<paragraph id="AD4F195BA84AFFEDD6FEFD298C3EFC0B" blockId="25.[121,1006,93,938]" pageId="25" pageNumber="26">
Additional material:
<materialsCitation id="1D981306A84AFFEDD7FAFD298EF4FD51" ID-GBIF-Occurrence="3497007317" collectingDate="1991-08-05" collectionCode="SMNS" country="NORD-SUMATRA" elevation="1500" location="Kiefernwald" municipality="Umg. Siulakosa" pageId="25" pageNumber="26" specimenCount="1" specimenCount-male="1">
<specimenCount id="BBF6D2D2A84AFFEDD7FAFD298C8FFD6B" box="[410,458,690,714]" pageId="25" pageNumber="26" type="male">1 ♂</specimenCount>
,
<collectingCountry id="D5E759CBA84AFFEDD7BFFD298FE3FD6B" box="[479,678,690,714]" pageId="25" pageNumber="26">NORD-SUMATRA</collectingCountry>
:
<collectingMunicipality id="4D2B8321A84AFFEDD4D7FD298E37FD6B" box="[695,882,690,714]" name="Umg. Siulakosa" pageId="25" pageNumber="26">Umg. Siulakosa</collectingMunicipality>
<date id="D94E3F9BA84AFFEDD5E0FD298EA3FD6B" box="[896,998,690,714]" pageId="25" pageNumber="26" value="1991-08-05">
<collectingDate id="C90AC673A84AFFEDD5E0FD298EA3FD6B" box="[896,998,690,714]" pageId="25" pageNumber="26" value="1991-08-05">5.8.1991</collectingDate>
</date>
, ERBER / Samosir, ob. Tomok
<quantity id="6A08B4BEA84AFFEDD7ABFD438F63FD51" box="[459,550,728,752]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="25" pageNumber="26" unit="m" value="1500.0">
<elevation id="26DDFE68A84AFFEDD7ABFD438F63FD51" box="[459,550,728,752]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="25" pageNumber="26" unit="m" value="1500.0">1500 m</elevation>
</quantity>
,
<location id="A82F4F80A84AFFEDD452FD438FFBFD51" LSID="urn:lsid:plazi:treatment:2559A84DA84BFFE8D6FFFBD68E7EFC24:A82F4F80A84AFFEDD452FD438FFBFD51" box="[562,702,728,752]" country="NORD-SUMATRA" municipality="Umg. Siulakosa" name="Kiefernwald" pageId="25" pageNumber="26">Kiefernwald</location>
,
<location id="A82F4F80A84AFFEDD4A9FD438E0BFD51" LSID="urn:lsid:plazi:treatment:2559A84DA84BFFE8D6FFFBD68E7EFC24:A82F4F80A84AFFEDD4A9FD438E0BFD51" box="[713,846,728,752]" country="NORD-SUMATRA" municipality="Umg. Siulakosa" name="unter Rinde" pageId="25" pageNumber="26">unter Rinde</location>
(
<collectionCode id="CBE1819EA84AFFEDD53DFD438EEEFD51" box="[861,939,728,752]" collectionName="Germany, Stuttgart, Staatliches Museum fuer Naturkunde" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="25" pageNumber="26">SMNS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="1D981306A84AFFEDD5DDFD438E37FCB4" ID-GBIF-Occurrence="3497007332" collectingDate="1995-03" collectionCode="CBP" collectorName="M. Malindang North Sambuanga" country="Philippines" location="M. Malindang North Sambuanga" municipality="Mindanao" pageId="25" pageNumber="26" specimenCount="1" specimenCount-male="1">
<specimenCount id="BBF6D2D2A84AFFEDD5DDFD438EADFD4E" box="[957,1000,728,752]" pageId="25" pageNumber="26" type="male">1 ♂</specimenCount>
,
<collectingCountry id="D5E759CBA84AFFEDD619FD668D9FFCB4" box="[121,218,765,789]" name="Philippines" pageId="25" pageNumber="26">Filippine</collectingCountry>
<collectingMunicipality id="4D2B8321A84AFFEDD681FD668C17FCB4" box="[225,338,765,789]" country="Philippines" name="Mindanao" pageId="25" pageNumber="26">Mindanao</collectingMunicipality>
<date id="D94E3F9BA84AFFEDD739FD668CEBFCB4" box="[345,430,765,789]" pageId="25" pageNumber="26" value="1995-03">
<collectingDate id="C90AC673A84AFFEDD739FD668CEBFCB4" box="[345,430,765,789]" pageId="25" pageNumber="26" value="1995-03">III.1995</collectingDate>
</date>
<location id="A82F4F80A84AFFEDD7D6FD668E6DFCB4" LSID="urn:lsid:plazi:treatment:2559A84DA84BFFE8D6FFFBD68E7EFC24:A82F4F80A84AFFEDD7D6FD668E6DFCB4" box="[438,808,765,789]" country="Philippines" municipality="Mindanao" name="M. Malindang North Sambuanga" pageId="25" pageNumber="26">M. Malindang North Sambuanga</location>
(
<collectionCode id="CBE1819EA84AFFEDD556FD668E29FCB4" box="[822,876,765,789]" pageId="25" pageNumber="26">CBP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="1D981306A84AFFEDD51DFD668ECCFC9B" ID-GBIF-Occurrence="3497007341" collectingDate="2014-11" collectionCode="CBB" collectorName="N Luzon" country="PHIL- IPPINES" location="N Luzon" pageId="25" pageNumber="26" specimenCount="1" specimenCount-female="1" stateProvince="Ifugao">
<specimenCount id="BBF6D2D2A84AFFEDD51DFD668EE3FCB4" box="[893,934,765,789]" pageId="25" pageNumber="26" type="female">1 ♀</specimenCount>
,
<collectingCountry id="D5E759CBA84AFFEDD5D4FD668D9DFC9B" pageId="25" pageNumber="26">PHILIPPINES</collectingCountry>
,
<location id="A82F4F80A84AFFEDD682FCB98C06FC9B" LSID="urn:lsid:plazi:treatment:2559A84DA84BFFE8D6FFFBD68E7EFC24:A82F4F80A84AFFEDD682FCB98C06FC9B" box="[226,323,802,826]" country="PHIL- IPPINES" name="N Luzon" pageId="25" pageNumber="26" stateProvince="Ifugao">N Luzon</location>
,
<location id="A82F4F80A84AFFEDD72EFCB98CDEFC9B" LSID="urn:lsid:plazi:treatment:2559A84DA84BFFE8D6FFFBD68E7EFC24:A82F4F80A84AFFEDD72EFCB98CDEFC9B" box="[334,411,802,826]" country="PHIL- IPPINES" name="Ifugao" pageId="25" pageNumber="26" stateProvince="Ifugao">Ifugao</location>
,
<location id="A82F4F80A84AFFEDD7C5FCB98CBAFC9B" LSID="urn:lsid:plazi:treatment:2559A84DA84BFFE8D6FFFBD68E7EFC24:A82F4F80A84AFFEDD7C5FCB98CBAFC9B" box="[421,511,802,826]" country="PHIL- IPPINES" name="Banaue" pageId="25" pageNumber="26" stateProvince="Ifugao">Banaue</location>
,
<date id="D94E3F9BA84AFFEDD469FCB98F25FC9B" box="[521,608,802,826]" pageId="25" pageNumber="26" value="2014-11">
<collectingDate id="C90AC673A84AFFEDD469FCB98F25FC9B" box="[521,608,802,826]" pageId="25" pageNumber="26" value="2014-11">XI 2014</collectingDate>
</date>
. local collector leg. (
<collectionCode id="CBE1819EA84AFFEDD52CFCB98EC6FC9B" box="[844,899,802,826]" collectionName="CBB" pageId="25" pageNumber="26">CBB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="1D981306A84AFFEDD5F4FCB98C32FC0B" ID-GBIF-Occurrence="3497007306" collectionCode="BMNH" collectorName="Banninger &amp; H. E. Andrewes" country="Philippines" pageId="25" pageNumber="26" specimenCount="1" stateProvince="Ifugao">
<specimenCount id="BBF6D2D2A84AFFEDD5F4FCB98DEBFCC1" pageId="25" pageNumber="26" type="generic">1 specimen</specimenCount>
: head and pronotum lost, F. C. DRESCHER G. Tangkoeban Prahoe 4000 – 5000 Voet Praeanger, Java
<date id="D94E3F9BA84AFFEDD7D0FCF68F4AFC24" box="[432,527,877,901]" pageId="25" pageNumber="26" value="1933-06">VI. 1933</date>
/ Ex coll.
<collectorName id="00057C8DA84AFFEDD4E5FCF68FBDFC24" box="[645,760,877,901]" pageId="25" pageNumber="26">Bänninger</collectorName>
/
<collectorName id="00057C8DA84AFFEDD574FCF68EF6FC24" box="[788,947,877,901]" pageId="25" pageNumber="26">
H.E.
<taxonomicName id="6AF062D8A84AFFEDD559FCF68EF6FC24" box="[825,947,877,901]" pageId="25" pageNumber="1" rank="subGenus" status="subgen. nov." subGenus="Andrewes">Andrewes</taxonomicName>
</collectorName>
Coll. B.M.1945–97. (
<collectionCode id="CBE1819EA84AFFEDD741FC098C34FC0B" box="[289,369,914,938]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="25" pageNumber="26">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A84AFFE8D619FB998F6FFDFB" lastPageId="28" lastPageNumber="29" pageId="25" pageNumber="26" type="description">
<paragraph id="AD4F195BA84AFFEDD619FB998F19FBBB" blockId="25.[121,604,1026,1051]" box="[121,604,1026,1051]" pageId="25" pageNumber="26">
<emphasis id="9F84C549A84AFFEDD619FB998C5DFBBA" bold="true" box="[121,280,1026,1051]" pageId="25" pageNumber="26">Redescription</emphasis>
(
<figureCitation id="35CB05DEA84AFFEDD744FB998C2DFBBB" box="[292,360,1026,1050]" captionStart="Figs 1–4" captionStartId="7.[121,158,1494,1512]" captionTargetBox="[145,1002,80,1436]" captionTargetPageId="7" captionText="Figs 1–4: Orictites spp., habitus. 1: O. minotaur ANDREWES, 1931, lectotype, male; 2: O. costulipennis BATES, 1892, holotype, male; 3: O. mjoebergi LouWERENS, 1964, lectotype, male; 4: O. desuntsetosus sp. nov., holotype, male." figureDoi="http://doi.org/10.5281/zenodo.6284320" httpUri="https://zenodo.org/record/6284320/files/figure.png" pageId="25" pageNumber="26">Figs 3</figureCitation>
,
<figureCitation id="35CB05DEA84AFFEDD715FB998CD4FBBB" box="[373,401,1026,1050]" captionStart="Figs 21–26" captionStartId="23.[121,158,1467,1485]" captionTargetBox="[120,1005,145,1417]" captionTargetPageId="23" captionText="Figs 21–26: Orictites spp., aedeagus, ventral view. 21: O. mjoebergi LouWERENS, 1964, lectotype, scale bar 0.2 mm; 22: O. desuntsetosus sp. nov., holotype, scale bar 0.15 mm; 23: O. plurisetosus sp. nov., holotype, scale bar 0.2 mm; 24: O. anteriortenuis sp. nov., holotype, scale bar 0.15 mm; 25: O. omnipunctatus sp. nov., holotype, scale bar 0.2 mm; 26: O. ampliosetosus sp. nov., holotype, scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6284350" httpUri="https://zenodo.org/record/6284350/files/figure.png" pageId="25" pageNumber="26">21</figureCitation>
,
<figureCitation id="35CB05DEA84AFFEDD7FEFB998CFAFBBB" box="[414,447,1026,1050]" captionStart="Figs 31–34" captionStartId="34.[159,196,1291,1309]" captionTargetBox="[159,1043,80,1253]" captionText="Figs 31–34: Orictites spp., coxostylus. 31: O. minotaur ANDREWES, 1931, scale bar 0.5 mm; 32: O. costulipennis BATES, 1892, scale bar 0.1 mm. 33: O. mjoebergi LouWERENS, 1964, scale bar 0.1 mm; 34: O. desuntsetosus sp. nov., scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6284358" httpUri="https://zenodo.org/record/6284358/files/figure.png" pageId="25" pageNumber="26">33</figureCitation>
; pp. 8, 24, 35)
</paragraph>
<paragraph id="AD4F195BA84AFFEDD619FBD68FBBFB2B" blockId="25.[121,1006,1101,1573]" pageId="25" pageNumber="26">
Measurements:
<typeStatus id="724BA7F9A84AFFEDD751FBD68CE3FBC4" box="[305,422,1101,1125]" pageId="25" pageNumber="26" type="lectotype">Lectotype</typeStatus>
: Body length
<quantity id="6A08B4BEA84AFFEDD422FBD68FDEFBC4" box="[578,667,1101,1125]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="25" pageNumber="26" unit="mm" value="5.4">5.4 mm</quantity>
, width
<quantity id="6A08B4BEA84AFFEDD48CFBD68E17FBC4" box="[748,850,1101,1125]" metricMagnitude="-3" metricUnit="m" metricValue="1.52" pageId="25" pageNumber="26" unit="mm" value="1.52">1.52 mm</quantity>
; ratio length/ width of pronotum 1.07; ratio length/width of elytra 1.95.
</paragraph>
<paragraph id="AD4F195BA84AFFEDD6FEFB038F9CFB5B" blockId="25.[121,1006,1101,1573]" pageId="25" pageNumber="26">
Additional material (n = 3): Body length
<quantity id="6A08B4BEA84AFFEDD40BFB038E47FB11" box="[619,770,1176,1200]" metricMagnitude="-3" metricUnit="m" metricValue="5.415" metricValueMax="5.9" metricValueMin="4.93" pageId="25" pageNumber="26" unit="mm" value="5.415" valueMax="5.9" valueMin="4.93">4.93–5.9 mm</quantity>
(
<emphasis id="9F84C549A84AFFEDD572FB0C8E67FB11" bold="true" box="[786,802,1175,1200]" italics="true" pageId="25" pageNumber="26"></emphasis>
=
<quantity id="6A08B4BEA84AFFEDD523FB038ED8FB11" box="[835,925,1176,1200]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="25" pageNumber="26" unit="mm" value="5.4">5.4 mm</quantity>
), width
<quantity id="6A08B4BEA84AFFEDD619FB268C5CFB74" box="[121,281,1213,1237]" metricMagnitude="-3" metricUnit="m" metricValue="1.51" metricValueMax="1.65" metricValueMin="1.37" pageId="25" pageNumber="26" unit="mm" value="1.51" valueMax="1.65" valueMin="1.37">1.37–1.65 mm</quantity>
(
<emphasis id="9F84C549A84AFFEDD74AFB278C7FFB74" bold="true" box="[298,314,1212,1237]" italics="true" pageId="25" pageNumber="26"></emphasis>
=
<quantity id="6A08B4BEA84AFFEDD73BFB268C87FB74" box="[347,450,1213,1237]" metricMagnitude="-3" metricUnit="m" metricValue="1.53" pageId="25" pageNumber="26" unit="mm" value="1.53">1.53 mm</quantity>
); ratio length/width of pronotum 0.96–1.06 (
<emphasis id="9F84C549A84AFFEDD5A4FB278E91FB74" bold="true" box="[964,980,1212,1237]" italics="true" pageId="25" pageNumber="26"></emphasis>
= 1.0); ratio length/width of elytra 1.78–1.97 (
<emphasis id="9F84C549A84AFFEDD40AFB7A8F3FFB5B" bold="true" box="[618,634,1249,1274]" italics="true" pageId="25" pageNumber="26"></emphasis>
= 1.89).
</paragraph>
<paragraph id="AD4F195BA84AFFEDD6FEFA938E69FACB" blockId="25.[121,1006,1101,1573]" pageId="25" pageNumber="26">Colour: Shiny. Black. Elytra red-brown, pronotum darker. Wings of clypeus and supraantennal plates transparent red-brown. Antennae, mouthparts, middle and hind legs yellowish brown, front legs dark red-brown.</paragraph>
<paragraph id="AD4F195BA84AFFEED6FEFAE38C59FA81" blockId="25.[121,1006,1101,1573]" lastBlockId="26.[159,1044,93,1573]" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
Head: A quarter narrower than pronotum. Clypeus limited laterally by small obtuse angled tooth, anterior margin straight, with blunt raised knob between the two clypeal setae, posterior part of knob covered with medium sized punctures, anterior part smooth; wings obtuse-angular, rounded at tip, hollowed out, divided from middle part of clypeus by slight incisions and carinae, divided from supraantennal plates by distinct obtusely angled incisions and indistinct carina, clypeus and wings reflexed margined, supraantennal plates vaulted, elongated as supraorbital carina, smooth, with reflexed rounded margin, frons separated from clypeus by flat transverse depression and a group of punctures formed like a letter V, with nearly parallel running costate frontal carinae at each side posteriorly, with broad furrow between frontal carina and supraorbital carina, sharp furrow between frons and frontal carina. Frons moderately convex, nearly smooth, with small but distinct longitudinal impression at middle. Supraorbital setae located in broad furrows between supraorbital carina and frontal carina, the posterior one arising from a tubercle. Eyes of moderate size, moderately convex, genae enlarged, regularly rounded, not as high as and a third shorter than eyes, enclosing eyes posteriorly. Grooves for reception of antennae short, length 0.4 of eye diameter. Neck constriction marked as distinct step, with band of big punctures, neck laterally densely covered with medium sized punctures. Labrum indistinctly trilobed, appearing straight, 7-setose, ciliate laterally, with isodiametric reticulation. Mandibles half as long as head, flattened, broadened at base, acutely curved at apex, carinae of scrobe complete, both mandibles obtusely angled and somewhat hollowed out dorsally. Maxilla nearly straight, acutely hooked at apex. Apical maxillary palpomere slender, slightly securiform. Apical labial palpomere slender, straight, longer than 2
<superScript id="5A85B413A849FFEED4CBFCDF8F87FCF6" attach="left" box="[683,706,836,855]" fontSize="8" pageId="26" pageNumber="27">nd</superScript>
segment, 2
<superScript id="5A85B413A849FFEED52EFCDF8E20FCF6" attach="left" box="[846,869,836,855]" fontSize="8" pageId="26" pageNumber="27">nd</superScript>
segment bisetose. Ligula with one long seta; paraglossae slender. Ventral surface of neck covered with medium sized punctures and irregular reticulation. Gula smooth. Submentum and mentum distinctly separated; submentum with two pairs of setae; mentum keeled at middle, nearly smooth, with fine and complete reflexed margin, lateral lobes projecting, thought obtusely angled at tips, with two setae near base of each lobe, median tooth forming broad obtuse triangle, bisetose, less projecting than lateral lobes. Paragenae sharp, carina-like, without tooth. Antennae of medium length, reaching over middle of pronotum, scapus carinate, with one seta dorsoapically, scapus and pedicellus with fine reticulation, segments 5–10 transverse (L/
<geoCoordinate id="C8C47F9CA849FFEED4F7FB268FB4FB74" box="[663,753,1213,1237]" degrees="0. 80" direction="west" orientation="longitude" pageId="26" pageNumber="27" precision="555" value="-0.8">W 0.80</geoCoordinate>
), antennomeres subglobose, nearly dull, segments 4–11 densely pubescent, with a few additional long setae.
</paragraph>
<paragraph id="AD4F195BA849FFEFD6A4FAB68DF9FDFB" blockId="26.[159,1044,93,1573]" lastBlockId="27.[121,1006,93,1573]" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
Pronotum: Square (L/
<geoCoordinate id="C8C47F9CA849FFEED7A6FAB68F65FAE4" box="[454,544,1325,1349]" degrees="1. 07" direction="west" orientation="longitude" pageId="26" pageNumber="27" precision="555" value="-1.07">W 1.07</geoCoordinate>
), sides straight but slightly diverging to base, gently rounded before posterior angle. Anterior margin slightly bisinuate. Reflexed lateral margin nearly smooth, with indistinct reticulation (100x), from posterior angle to base running as smooth, slightly convex line. Lateral channel distinctly broadened between the two lateral setigerous punctures, with fine irregular reticulation, with a few medium-sized punctures situated in broadened part of channel, basal channel broad, deep, foveolate. Anterior setigerous puncture located at end of anterior quarter, the posterior one located at level of posterior angle, removed from lateral channel by diameter of pore. Anterior angle slightly but distinctly projecting, rounded, formed by reflexed lateral margin, posterior angle developed as large rounded tooth, projecting laterally. Anterior transverse line sharp, punctured, not adjoining lateral channel. Median line deep, broad, thought with indistinct blunt carina-like vault bilaterally, surpassing level of anterior transverse line but not joining, smaller at base, joining base. Surface with two longitudinal groups of paramedian and paralateral big punctures and with a few fine, irregularly situated punctures, with irregular reticulation laterally, shiny on disc; basal impression developed as small flat furrow; base of disc with indistinct reticulation. Basal part of disc elongated posteriorly and bilaterally at declivity, as tooth-like vault slightly hanging over basal channel (lateral view). Flange raised keel-like (lateral view).
</paragraph>
<paragraph id="AD4F195BA848FFEFD6FEFDF38EA0FBC4" blockId="27.[121,1006,93,1573]" pageId="27" pageNumber="28">
Elytra: Subcylindrical, sides slightly diverging, marginal channel broad, not completely visible from above due to convexity of intervals 7 and 8, with uninterrupted row of big setigerous punctures arising from broad tubercles, with additional row of punctures laterally. Reflexed margin with fine crenulation in anterior quarter, channel broadened at humerus, bending with margin over rounded toothless humerus up to 5
<superScript id="5A85B413A848FFEFD469FC848F59FC93" attach="left" box="[521,540,799,818]" fontSize="8" pageId="27" pageNumber="28">th</superScript>
stria. Basal tubercle big, with setigerous puncture, situated at declivity of first stria. Striae 1–4 free at base, all striae deep, broad, distinctly punctate-striate, striae 1–2, 3–4 and 5–6 joined apically; striae 6 and 7 shortened at humerus. Striole distinct, interval between striole and sutura costiform, raised, basal area around striole moderately impressed. Intervals shiny, distinctly convex, 3
<superScript id="5A85B413A848FFEFD4E8FC418FDEFC4C" attach="left" box="[648,667,986,1005]" fontSize="8" pageId="27" pageNumber="28">rd</superScript>
broader, strikingly raised and with large tubercle at base, 4
<superScript id="5A85B413A848FFEFD7A5FC648C9DFBB3" attach="left" box="[453,472,1023,1042]" fontSize="8" pageId="27" pageNumber="28">th</superScript>
broader and raised at base but not as strongly as 3
<superScript id="5A85B413A848FFEFD6CBFBBF8DFBFB96" attach="none" box="[171,190,1060,1079]" fontSize="8" pageId="27" pageNumber="28">rd</superScript>
, 7
<superScript id="5A85B413A848FFEFD6BDFBBF8DB5FB96" attach="left" box="[221,240,1060,1079]" fontSize="8" pageId="27" pageNumber="28">th</superScript>
running to apex as obtuse carina. 8
<superScript id="5A85B413A848FFEFD4FBFBBF8FEBFB96" attach="left" box="[667,686,1060,1079]" fontSize="8" pageId="27" pageNumber="28">th</superScript>
small, completely carinate. Interval 3 with four/five setigerous punctures adjoining stria 3 (see variation).
</paragraph>
<paragraph id="AD4F195BA848FFEFD6FEFBE98CA6FB2B" blockId="27.[121,1006,93,1573]" box="[158,483,1138,1162]" pageId="27" pageNumber="28">Hind wings: Fully developed.</paragraph>
<paragraph id="AD4F195BA848FFEFD6FEFB038CFEFA7B" blockId="27.[121,1006,93,1573]" pageId="27" pageNumber="28">
Ventral surface: Proepisternum with complete submarginal ridge, with some punctures, with irregular reticulation laterally. Proepipleuron small, with punctures and indistinct reticulation. Epipleuron broadened and with big punctures in basal quarter. Proepisternum somewhat swollen at level of posterior setigerous puncture, prosternite smooth, double keeled at middle, mesosternum smooth. Sternites smooth at middle, each with big punctures at base laterally, 3
<superScript id="5A85B413A848FFEFD736FAEF8C2CFA26" attach="left" box="[342,361,1396,1415]" fontSize="8" pageId="27" pageNumber="28">rd</superScript>
to 5
<superScript id="5A85B413A848FFEFD7C2FAEF8CF0FA26" attach="left" box="[418,437,1396,1415]" fontSize="8" pageId="27" pageNumber="28">th</superScript>
with paralateral ambulatory setae at each side, ventral strigae distinct, sternite 6 smooth at middle, with two widely separated apical setae at each side.
</paragraph>
<paragraph id="AD4F195BA848FFE8D6FEFA738CFBFF61" blockId="27.[121,1006,93,1573]" lastBlockId="28.[159,1044,93,901]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">Legs: Anterior tibia with strong digitation, with two big and one small lateral preapical denticles, apical spine curved distinctly towards ventral and lateral, no dorsal furrow, basal tarsomere elongated, longer than the following four together. Mesotibia with apical spine, furnished with strong seta. All tarsomeres relatively broad.</paragraph>
<paragraph id="AD4F195BA84FFFE8D6A4FF568F90FE91" blockId="28.[159,1044,93,901]" pageId="28" pageNumber="29">
Male genitalia (
<figureCitation id="35CB05DEA84FFFE8D71EFF568C94FF44" box="[382,465,205,229]" captionStart="Figs 21–26" captionStartId="23.[121,158,1467,1485]" captionTargetBox="[120,1005,145,1417]" captionTargetPageId="23" captionText="Figs 21–26: Orictites spp., aedeagus, ventral view. 21: O. mjoebergi LouWERENS, 1964, lectotype, scale bar 0.2 mm; 22: O. desuntsetosus sp. nov., holotype, scale bar 0.15 mm; 23: O. plurisetosus sp. nov., holotype, scale bar 0.2 mm; 24: O. anteriortenuis sp. nov., holotype, scale bar 0.15 mm; 25: O. omnipunctatus sp. nov., holotype, scale bar 0.2 mm; 26: O. ampliosetosus sp. nov., holotype, scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6284350" httpUri="https://zenodo.org/record/6284350/files/figure.png" pageId="28" pageNumber="29">Fig. 21</figureCitation>
): Relatively short, median lobe broad, regularly arcuate at middle, apex twisted apically. Endophallus with bristles laterally. Both parameres slender, slightly twisted, asetose.
</paragraph>
<paragraph id="AD4F195BA84FFFE8D6A4FEA68EC8FE64" blockId="28.[159,1044,93,901]" pageId="28" pageNumber="29">
Female genitalia (
<figureCitation id="35CB05DEA84FFFE8D7EDFEA68C9FFEF4" box="[397,474,317,341]" captionStart="Figs 31–34" captionStartId="34.[159,196,1291,1309]" captionTargetBox="[159,1043,80,1253]" captionText="Figs 31–34: Orictites spp., coxostylus. 31: O. minotaur ANDREWES, 1931, scale bar 0.5 mm; 32: O. costulipennis BATES, 1892, scale bar 0.1 mm. 33: O. mjoebergi LouWERENS, 1964, scale bar 0.1 mm; 34: O. desuntsetosus sp. nov., scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6284358" httpUri="https://zenodo.org/record/6284358/files/figure.png" pageId="28" pageNumber="29">Fig. 33</figureCitation>
): Coxostylus of moderate width, distinctly curved to apex, with two preapical setae close together, dorsally with some minute scars in apical half, with obtuse knob-like tooth dorsally at base, with two short ensiform setae at middle, with four longer nematiform setae.
</paragraph>
<paragraph id="AD4F195BA84FFFE8D6A4FE498F6FFDFB" blockId="28.[159,1044,93,901]" pageId="28" pageNumber="29">
Variation: Bilateral variation is observed in the right clypeal wing, mentioned under remarks. In addition, in the
<typeStatus id="724BA7F9A84FFFE8D4E3FE638FABFDB1" box="[643,750,504,528]" pageId="28" pageNumber="29" type="lectotype">lectotype</typeStatus>
the left elytra bears five setigerous punctures and the right one four. The other specimens all have four setigerous punctures on each side.
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A84FFFE8D6A4FDF38E7EFC24" pageId="28" pageNumber="29" type="diagnosis">
<paragraph id="AD4F195BA84FFFE8D6A4FDF38E7EFC24" blockId="28.[159,1044,93,901]" pageId="28" pageNumber="29">
Diagnosis: A medium-sized almost black species with red-brownish elytra, with distinct knob on the clypeus, humerus without tooth, and four setigerous punctures on interval 3 (but compare remarks). Only in this species and in
<taxonomicName id="6AF062D8A84FFFE8D6FFFD438C17FD51" authorityName="Balkenohl" authorityYear="2017" box="[159,338,728,752]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="anteriorlatus">
<emphasis id="9F84C549A84FFFE8D6FFFD438C17FD51" box="[159,338,728,752]" italics="true" pageId="28" pageNumber="29">O. anteriorlatus</emphasis>
</taxonomicName>
is the basal part of the disc of the pronotum posteriorly elongated bilaterally at the declivity as a tooth-like vault slightly hanging over the basal channel. By contrast to
<taxonomicName id="6AF062D8A84FFFE8D797FCB98FEEFC9B" authorityName="Balkenohl" authorityYear="2017" box="[503,683,802,826]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="anteriorlatus">
<emphasis id="9F84C549A84FFFE8D797FCB98FEEFC9B" box="[503,683,802,826]" italics="true" pageId="28" pageNumber="29">O. anteriorlatus</emphasis>
</taxonomicName>
,
<taxonomicName id="6AF062D8A84FFFE8D4DAFCB98E0AFC9B" authorityName="LouWERENS" authorityYear="1964" box="[698,847,802,826]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="mjoebergi">
<emphasis id="9F84C549A84FFFE8D4DAFCB98E0AFC9B" box="[698,847,802,826]" italics="true" pageId="28" pageNumber="29">O. mjoebergi</emphasis>
</taxonomicName>
exhibits a tubercle at the base of interval 3 of the elytra, only, and the furrow between clypeus and frons consists of punctures in form of the letter V.
</paragraph>
</subSubSection>
</treatment>
</document>
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