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<document id="755CDC6B9CCEC8DC6CDA9361F4D1C599" ID-CLB-Dataset="30562" ID-DOI="10.11646/zootaxa.4392.2.4" ID-GBIF-Dataset="dbcec4ac-cf8f-4215-a445-a63e65e0f02f" ID-ISSN="1175-5326" ID-Zenodo-Dep="1195409" ID-ZooBank="C1A8F758-D41A-404C-A5C4-6CBC476EA324" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1520684441588" checkinUser="plazi" docAuthor="Okanishi, Masanori &amp; Fujita, Toshihiko" docDate="2018" docId="2526AA33FF8A4163D9C2E8BBFECDD147" docLanguage="en" docName="zootaxa.43922.2.4.pdf" docOrigin="Zootaxa 4392 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Astrodendrum sagaminum" docType="treatment" docVersion="8" lastPageNumber="309" masterDocId="D91FD24BFF984177D955EC4FFFEAD327" masterDocTitle="A taxonomic review of the genus Astrodendrum (Echinodermata, Ophiuroidea, Euryalida, Gorgonocephalidae) with description of a new species from Japan" masterLastPageNumber="310" masterPageNumber="289" pageNumber="307" updateTime="1698529305170" updateUser="ExternalLinkService">
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<mods:title id="90D312BE51F3465B8E7DE68735635E2C">A taxonomic review of the genus Astrodendrum (Echinodermata, Ophiuroidea, Euryalida, Gorgonocephalidae) with description of a new species from Japan</mods:title>
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<mods:namePart id="F1B185BA5899E47A2AACB7FE2D9CFE56">Okanishi, Masanori</mods:namePart>
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<treatment id="2526AA33FF8A4163D9C2E8BBFECDD147" ID-DOI="http://doi.org/10.5281/zenodo.5998505" ID-GBIF-Taxon="142212007" ID-Zenodo-Dep="5998505" LSID="urn:lsid:plazi:treatment:2526AA33FF8A4163D9C2E8BBFECDD147" httpUri="http://treatment.plazi.org/id/2526AA33FF8A4163D9C2E8BBFECDD147" lastPageId="20" lastPageNumber="309" pageId="18" pageNumber="307">
<subSubSection id="E59548AEFF8A4165D9C2E8BBFF16D608" pageId="18" pageNumber="307" type="nomenclature">
<paragraph id="AD301B25FF8A4165D9C2E8BBFD50D629" blockId="18.[151,698,1268,1363]" box="[151,698,1268,1294]" pageId="18" pageNumber="307">
<heading id="F678AC49FF8A4165D9C2E8BBFD50D629" bold="true" box="[151,698,1268,1294]" fontSize="11" level="1" pageId="18" pageNumber="307" reason="1">
<taxonomicName id="6A8F60A6FF8A4165D9C2E8BBFD50D629" ID-CoL="J48D" authority="(Doderlein, 1902)" baseAuthorityName="Doderlein" baseAuthorityYear="1902" box="[151,698,1268,1294]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Astrodendrum" kingdom="Animalia" order="Euryalida" pageId="18" pageNumber="307" phylum="Echinodermata" rank="species" species="sagaminum">
<emphasis id="9FFBC737FF8A4165D9C2E8BBFD50D629" bold="true" box="[151,698,1268,1294]" pageId="18" pageNumber="307">
<emphasis id="9FFBC737FF8A4165D9C2E8BBFE30D629" bold="true" box="[151,474,1268,1294]" italics="true" pageId="18" pageNumber="307">Astrodendrum sagaminum</emphasis>
(
<bibRefCitation id="C91E66D4FF8A4165D8BCE8BBFD5BD629" box="[489,689,1268,1294]" pageId="18" pageNumber="307" refId="ref9238" refString="Doderlein, L. (1902) Japanische Euryaliden. Zoologishcer Anzeiger, 25, 320 - 326." type="journal article">Döderlein, 1902</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="AD301B25FF8A4165D9C2E959FF16D608" blockId="18.[151,698,1268,1363]" box="[151,252,1302,1327]" pageId="18" pageNumber="307">
(
<figureCitation id="35B407A0FF8A4165D9CAE959FF1ED608" box="[159,244,1302,1327]" captionStart="FIGURE 12" captionStartId="19.[151,250,1929,1951]" captionTargetBox="[151,1416,991,1893]" captionTargetId="figure@19.[151,1436,988,1905]" captionTargetPageId="19" captionText="FIGURE 12. Astrodendrum sagaminum, one syntype (ZSM 448/1). A, aboral disc and proximal portion of the arms; B, oral disc and proximal portion of the arms; C, oral view of proximal portion of the arms; D, lateral view of the disc and proximal portion of the arms. Arrowheads indicate madreporite (B, D). Abbreviations: AS, arm spine; GS, genital slit." httpUri="https://zenodo.org/record/1195433/files/figure.png" pageId="18" pageNumber="307">Figs 12</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="E59548AEFF8A4165D9C2E974FD6CD674" box="[151,646,1338,1363]" pageId="18" pageNumber="307" type="description">
<paragraph id="AD301B25FF8A4165D9C2E974FD6CD674" blockId="18.[151,698,1268,1363]" box="[151,646,1338,1363]" pageId="18" pageNumber="307">[Japanese Name: Tsuruboso-Tezurumozuru]</paragraph>
</subSubSection>
<subSubSection id="E59548AEFF8A4165D9C2E9CDFB43D6D3" pageId="18" pageNumber="307" type="reference_group">
<paragraph id="AD301B25FF8A4165D9C2E9CDFAF2D6BF" blockId="18.[151,1436,1410,1524]" box="[151,1304,1410,1432]" pageId="18" pageNumber="307">
<taxonomicName id="6A8F60A6FF8A4165D9C2E9CDFD86D6BF" ID-CoL="3H38C" authority="Doderlein, 1902" authorityName="Doderlein" authorityYear="1902" box="[151,620,1410,1432]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Gorgonocephalus" kingdom="Animalia" order="Euryalida" pageId="18" pageNumber="307" phylum="Echinodermata" rank="species" species="sagaminus">
<emphasis id="9FFBC737FF8A4165D9C2E9CDFE54D6BF" box="[151,446,1410,1432]" italics="true" pageId="18" pageNumber="307">Gorgonocephalus sagaminus</emphasis>
<bibRefCitation id="C91E66D4FF8A4165D891E9CDFD86D6BF" box="[452,620,1410,1432]" pageId="18" pageNumber="307" refId="ref9238" refString="Doderlein, L. (1902) Japanische Euryaliden. Zoologishcer Anzeiger, 25, 320 - 326." type="journal article">Döderlein, 1902</bibRefCitation>
</taxonomicName>
. 321322;—H. L. Clark, 1911. 292293; Jangoux
<emphasis id="9FFBC737FF8A4165DD38E9CCFB70D6BF" box="[1133,1178,1410,1432]" italics="true" pageId="18" pageNumber="307">et al</emphasis>
., 1987. 308.
</paragraph>
<paragraph id="AD301B25FF8A4165D9C2E9EEFB43D6D3" blockId="18.[151,1436,1410,1524]" pageId="18" pageNumber="307">
<taxonomicName id="6A8F60A6FF8A4165D9C2E9EEFDBAD690" ID-CoL="J48D" authority="Doderlein, 1911" authorityName="Doderlein" authorityYear="1911" box="[151,592,1441,1463]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Astrodendrum" kingdom="Animalia" order="Euryalida" pageId="18" pageNumber="307" phylum="Echinodermata" rank="species" species="sagaminum">
<emphasis id="9FFBC737FF8A4165D9C2E9EEFE48D690" box="[151,418,1441,1463]" italics="true" pageId="18" pageNumber="307">Astrodendrum sagaminum</emphasis>
<bibRefCitation id="C91E66D4FF8A4165D8FDE9EEFDBAD690" box="[424,592,1441,1463]" pageId="18" pageNumber="307" refId="ref9257" refString="Doderlein, L. (1911) Beitrage zur Naturgeschichte Ostasiens. Uber japanische und andere Euryalae. Abhandlungen der Bayerischen Akademie der Wissenschaften, II. Suppl. - Bd., 5, 1 - 123." type="journal article">Döderlein, 1911</bibRefCitation>
</taxonomicName>
. 3839, pl. 2 figs 35, pl. 7, fig. 8;—Döderlein, 1927. 32, 92;
<bibRefCitation id="C91E66D4FF8A4165DD92E9EEFA8FD690" box="[1223,1381,1441,1463]" pageId="18" pageNumber="307" refId="ref9078" refString="Bomford, T. L. (1913) A note on certain ophiuroids in the Indian Museum. Records of the Indian Museum, 9, 219 - 225." type="journal article">Bomford, 1913</bibRefCitation>
. 220; H. L. Clark, 1915. 185;
<bibRefCitation id="C91E66D4FF8A4165D8EDE98EFD87D6F1" box="[440,621,1472,1494]" pageId="18" pageNumber="307" refId="ref9496" refString="Matsumoto, H. (1917) A monograph of Japanese Ophiuroidea, arranged according to a new classification. Journal of College of Science Imperial University of Tokyo, 38, 1 - 408." type="journal article">Matsumoto, 1917</bibRefCitation>
. 7374, fig. 21;
<bibRefCitation id="C91E66D4FF8A4165DA44E98FFC56D6F1" box="[785,956,1472,1494]" pageId="18" pageNumber="307" refId="ref9620" refString="Murakami, S. (1944) Report on the ophiurans from off Ogasawara Islands and from off the Yaeyama group, Nippon. Journal of the Department of Agriculture, Kyushu Imperial University, 7, 235 - 257." type="journal article">Murakami, 1944</bibRefCitation>
. 247;
<bibRefCitation id="C91E66D4FF8A4165DAADE98FFB5FD6F1" box="[1016,1205,1472,1494]" pageId="18" pageNumber="307" refId="ref9109" refString="Chang, F. Y., Liao, Y. L. &amp; Wu, B. J. (1962) Euryales (Echinodermata: Ophiuroidea: Euryalida) of China. Acta Zoologica Sinica, 14, 53 - 67." type="journal article">
Chang
<emphasis id="9FFBC737FF8A4165DD14E98EFB87D6F1" box="[1089,1133,1472,1494]" italics="true" pageId="18" pageNumber="307">et al</emphasis>
., 1962
</bibRefCitation>
. 5859, pl. 1, figs 34;
<bibRefCitation id="C91E66D4FF8A4165D992E991FEBFD6D3" box="[199,341,1502,1524]" pageId="18" pageNumber="307" refId="ref9351" refString="Irimura, S. (1991) Ophiuroidea. In: Echinoderms from continental shelf and slope around Japan. Vol 2. Japan Fisheries Resource Conservation Association, Tokyo, pp. 111 - 153." type="book chapter">Irimura, 1991</bibRefCitation>
. 120, 2 plates;
<bibRefCitation id="C91E66D4FF8A4165D8B8E991FD15D6D3" box="[493,767,1502,1524]" pageId="18" pageNumber="307" refId="ref9388" refString="Irimura, S. and Kubodera, T. (1998) Ophiuroidea in the East Chinae Sea. Memoires of the National Science Museum, 30, 135 - 143." type="journal article">Irimura &amp; Kubodera, 1998</bibRefCitation>
. 138; Kroh, 2002. 148, 150151, fig. 41.
</paragraph>
</subSubSection>
<subSubSection id="E59548AEFF8A4165D9C2EA6EFB0AD51D" box="[151,1248,1569,1594]" pageId="18" pageNumber="307" type="materials_examined">
<paragraph id="AD301B25FF8A4165D9C2EA6EFB0AD51D" blockId="18.[151,1436,1569,2026]" box="[151,1248,1569,1594]" pageId="18" pageNumber="307">
<materialsCitation id="1DE71178FF8A4165D9C2EA6EFB0AD51D" ID-GBIF-Occurrence="1834883974" box="[151,1248,1569,1594]" collectionCode="ZSM" collectorName="Haberer" country="Japan" location="Sagami Bay" pageId="18" pageNumber="307" specimenCode="ZSM 448" specimenCount="2" typeStatus="syntype">
<emphasis id="9FFBC737FF8A4165D9C2EA6EFEAFD51D" bold="true" box="[151,325,1569,1594]" pageId="18" pageNumber="307">
<typeStatus id="7234A587FF8A4165D9C2EA6EFF38D51D" box="[151,210,1569,1594]" pageId="18" pageNumber="307">Type</typeStatus>
material.
</emphasis>
<specimenCode id="FD29B35EFF8A4165D819EA6DFE51D51D" box="[332,443,1570,1594]" collectionCode="ZSM" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="18" pageNumber="307">ZSM 448</specimenCode>
/1, one
<typeStatus id="7234A587FF8A4165DB5CEA6DFD89D51D" box="[521,611,1570,1594]" pageId="18" pageNumber="307" type="syntype">syntype</typeStatus>
, collected by
<collectorName id="007A7EF3FF8A4165DBABEA6EFCB0D51D" box="[766,858,1569,1594]" pageId="18" pageNumber="307">Haberer</collectorName>
,
<location id="A8504DFEFF8A4165DA30EA6EFC04D51D" LSID="urn:lsid:plazi:treatment:2526AA33FF8A4163D9C2E8BBFECDD147:A8504DFEFF8A4165DA30EA6EFC04D51D" box="[869,1006,1569,1594]" country="Japan" name="Sagami Bay" pageId="18" pageNumber="307">Sagami Bay</location>
,
<collectingCountry id="D5985BB5FF8A4165DAACEA6DFBD6D51E" box="[1017,1084,1570,1593]" name="Japan" pageId="18" pageNumber="307">Japan</collectingCountry>
, 1900 (
<figureCitation id="35B407A0FF8A4165DDDBEA6EFB25D51D" box="[1166,1231,1569,1594]" captionStart="FIGURE 1" captionStartId="4.[151,250,1967,1989]" captionTargetBox="[273,1322,208,1919]" captionTargetId="figure@4.[194,1390,193,1949]" captionTargetPageId="4" captionText="FIGURE 1. Distribution of Astrodendrum spinulosum sp. nov., A. capense, A. elingamita, A. galapagense, A. laevigatum and A. sagaminum. Red symbols indicate type localities." httpUri="https://zenodo.org/record/1195411/files/figure.png" pageId="18" pageNumber="307">Fig. 1</figureCitation>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="E59548AEFF8A4165D992EA0AFB18D5CA" pageId="18" pageNumber="307" type="diagnosis">
<paragraph id="AD301B25FF8A4165D992EA0AFB18D5CA" blockId="18.[151,1436,1569,2026]" pageId="18" pageNumber="307">
<emphasis id="9FFBC737FF8A4165D992EA0AFEABD579" bold="true" box="[199,321,1605,1630]" pageId="18" pageNumber="307">Diagnosis.</emphasis>
Two sizes (larger and smaller) and scattered granular external ossicles on interradial aboral disc; uniform-size and fully in contact granular and conical external ossicles on radial shields; external ossicles conical, separated on interradial lateral disc; external ossicles plate-shaped and granular, both fully in contact on oral disc; bulges of lateral edges of proximal portion of arms absent; single terminal projection on each arm spine of proximal portion of arm; 2 secondary teeth on each hook-shaped arm spine of distal portion of arm.
</paragraph>
</subSubSection>
<subSubSection id="E59548AEFF8A4163D992EAB6FD72D397" lastPageId="20" lastPageNumber="309" pageId="18" pageNumber="307" type="description">
<paragraph id="AD301B25FF8A4164D992EAB6FBDAD11B" blockId="18.[151,1436,1569,2026]" lastBlockId="19.[151,1437,151,933]" lastPageId="19" lastPageNumber="308" pageId="18" pageNumber="307">
<emphasis id="9FFBC737FF8A4165D992EAB6FE1DD435" bold="true" box="[199,503,1785,1810]" pageId="18" pageNumber="307">
Description of a
<typeStatus id="7234A587FF8A4165D8C1EAB6FE18D435" box="[404,498,1785,1810]" pageId="18" pageNumber="307" type="syntype">syntype</typeStatus>
.
</emphasis>
<emphasis id="9FFBC737FF8A4165DB55EAB5FDD1D436" box="[512,571,1786,1809]" italics="true" pageId="18" pageNumber="307">Disc.</emphasis>
Disc five-lobed with notched interradial edges,
<quantity id="6A77B6C0FF8A4165DD3CEAB5FB51D436" box="[1129,1211,1786,1810]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="18" pageNumber="307" unit="mm" value="20.0">20 mm</quantity>
disc diameter (
<figureCitation id="35B407A0FF8A4165DC3AEAB6FF21D411" captionStart="FIGURE 12" captionStartId="19.[151,250,1929,1951]" captionTargetBox="[151,1416,991,1893]" captionTargetId="figure@19.[151,1436,988,1905]" captionTargetPageId="19" captionText="FIGURE 12. Astrodendrum sagaminum, one syntype (ZSM 448/1). A, aboral disc and proximal portion of the arms; B, oral disc and proximal portion of the arms; C, oral view of proximal portion of the arms; D, lateral view of the disc and proximal portion of the arms. Arrowheads indicate madreporite (B, D). Abbreviations: AS, arm spine; GS, genital slit." httpUri="https://zenodo.org/record/1195433/files/figure.png" pageId="18" pageNumber="307">Fig. 12A</figureCitation>
). Aborally, radial shields tumid (
<figureCitation id="35B407A0FF8A4165DB6CEB52FD4CD411" box="[569,678,1821,1846]" captionStart="FIGURE 12" captionStartId="19.[151,250,1929,1951]" captionTargetBox="[151,1416,991,1893]" captionTargetId="figure@19.[151,1436,988,1905]" captionTargetPageId="19" captionText="FIGURE 12. Astrodendrum sagaminum, one syntype (ZSM 448/1). A, aboral disc and proximal portion of the arms; B, oral disc and proximal portion of the arms; C, oral view of proximal portion of the arms; D, lateral view of the disc and proximal portion of the arms. Arrowheads indicate madreporite (B, D). Abbreviations: AS, arm spine; GS, genital slit." httpUri="https://zenodo.org/record/1195433/files/figure.png" pageId="18" pageNumber="307">Figs 12A</figureCitation>
). On aboral disc, interradial spaces and spaces between each pair of radial shields entirely covered by different sizes (larger and smaller) of granular external ossicles, separated and scattered (
<figureCitation id="35B407A0FF8A4165D85EEB2AFE9ED459" box="[267,372,1893,1918]" captionStart="FIGURE 12" captionStartId="19.[151,250,1929,1951]" captionTargetBox="[151,1416,991,1893]" captionTargetId="figure@19.[151,1436,988,1905]" captionTargetPageId="19" captionText="FIGURE 12. Astrodendrum sagaminum, one syntype (ZSM 448/1). A, aboral disc and proximal portion of the arms; B, oral disc and proximal portion of the arms; C, oral view of proximal portion of the arms; D, lateral view of the disc and proximal portion of the arms. Arrowheads indicate madreporite (B, D). Abbreviations: AS, arm spine; GS, genital slit." httpUri="https://zenodo.org/record/1195433/files/figure.png" pageId="18" pageNumber="307">Fig. 12A</figureCitation>
). Larger granular external ossicles approximately 150200 µm in length, and smaller external ossicles approximately 50100 µm in length (
<figureCitation id="35B407A0FF8A4165DBC9EBC6FCEFD485" box="[668,773,1929,1954]" captionStart="FIGURE 12" captionStartId="19.[151,250,1929,1951]" captionTargetBox="[151,1416,991,1893]" captionTargetId="figure@19.[151,1436,988,1905]" captionTargetPageId="19" captionText="FIGURE 12. Astrodendrum sagaminum, one syntype (ZSM 448/1). A, aboral disc and proximal portion of the arms; B, oral disc and proximal portion of the arms; C, oral view of proximal portion of the arms; D, lateral view of the disc and proximal portion of the arms. Arrowheads indicate madreporite (B, D). Abbreviations: AS, arm spine; GS, genital slit." httpUri="https://zenodo.org/record/1195433/files/figure.png" pageId="18" pageNumber="307">Fig. 12A</figureCitation>
). Radial shields covered by uniform-size external ossicles, fully in contact, approximately 100 µm in length (
<figureCitation id="35B407A0FF8A4165DB85EBE2FCD3D4E1" box="[720,825,1965,1990]" captionStart="FIGURE 12" captionStartId="19.[151,250,1929,1951]" captionTargetBox="[151,1416,991,1893]" captionTargetId="figure@19.[151,1436,988,1905]" captionTargetPageId="19" captionText="FIGURE 12. Astrodendrum sagaminum, one syntype (ZSM 448/1). A, aboral disc and proximal portion of the arms; B, oral disc and proximal portion of the arms; C, oral view of proximal portion of the arms; D, lateral view of the disc and proximal portion of the arms. Arrowheads indicate madreporite (B, D). Abbreviations: AS, arm spine; GS, genital slit." httpUri="https://zenodo.org/record/1195433/files/figure.png" pageId="18" pageNumber="307">Fig. 12A</figureCitation>
), conical 100 µm high on distal edge of radial shields and granular in the other areas (
<figureCitation id="35B407A0FF8A4165DB64EB9EFD49D4CD" box="[561,675,2001,2026]" captionStart="FIGURE 12" captionStartId="19.[151,250,1929,1951]" captionTargetBox="[151,1416,991,1893]" captionTargetId="figure@19.[151,1436,988,1905]" captionTargetPageId="19" captionText="FIGURE 12. Astrodendrum sagaminum, one syntype (ZSM 448/1). A, aboral disc and proximal portion of the arms; B, oral disc and proximal portion of the arms; C, oral view of proximal portion of the arms; D, lateral view of the disc and proximal portion of the arms. Arrowheads indicate madreporite (B, D). Abbreviations: AS, arm spine; GS, genital slit." httpUri="https://zenodo.org/record/1195433/files/figure.png" pageId="18" pageNumber="307">Fig. 12A</figureCitation>
). Center of disc covered by larger granular external ossicles, approximately 250 µm and smaller granular external ossicles approximately 50100 µm in length, both separated and scattered (
<figureCitation id="35B407A0FF8B4164D815ECF3FE41D3F3" box="[320,427,188,213]" captionStart="FIGURE 12" captionStartId="19.[151,250,1929,1951]" captionTargetBox="[151,1416,991,1893]" captionTargetId="figure@19.[151,1436,988,1905]" captionTargetPageId="19" captionText="FIGURE 12. Astrodendrum sagaminum, one syntype (ZSM 448/1). A, aboral disc and proximal portion of the arms; B, oral disc and proximal portion of the arms; C, oral view of proximal portion of the arms; D, lateral view of the disc and proximal portion of the arms. Arrowheads indicate madreporite (B, D). Abbreviations: AS, arm spine; GS, genital slit." httpUri="https://zenodo.org/record/1195433/files/figure.png" pageId="19" pageNumber="308">Fig. 12A</figureCitation>
). Lateral edge of disc covered by uniform-size conical external ossicles, separated and scattered, approximately 100 µm in length and height (
<figureCitation id="35B407A0FF8B4164DA54EC90FC83D3DF" box="[769,873,223,248]" captionStart="FIGURE 12" captionStartId="19.[151,250,1929,1951]" captionTargetBox="[151,1416,991,1893]" captionTargetId="figure@19.[151,1436,988,1905]" captionTargetPageId="19" captionText="FIGURE 12. Astrodendrum sagaminum, one syntype (ZSM 448/1). A, aboral disc and proximal portion of the arms; B, oral disc and proximal portion of the arms; C, oral view of proximal portion of the arms; D, lateral view of the disc and proximal portion of the arms. Arrowheads indicate madreporite (B, D). Abbreviations: AS, arm spine; GS, genital slit." httpUri="https://zenodo.org/record/1195433/files/figure.png" pageId="19" pageNumber="308">Fig. 12A</figureCitation>
). Radial shields completely concealed by external ossicles, bar-like, approximately
<quantity id="6A77B6C0FF8B4164DB5AED4BFD8AD23B" box="[527,608,260,284]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="19" pageNumber="308" unit="mm" value="10.0">10 mm</quantity>
in length,
<quantity id="6A77B6C0FF8B4164DB88ED4BFCDFD23B" box="[733,821,260,285]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="19" pageNumber="308" unit="mm" value="1.5">1.5 mm</quantity>
in width, and almost reaching disc center (
<figureCitation id="35B407A0FF8B4164DC73ED4BFA7AD23B" box="[1318,1424,260,285]" captionStart="FIGURE 12" captionStartId="19.[151,250,1929,1951]" captionTargetBox="[151,1416,991,1893]" captionTargetId="figure@19.[151,1436,988,1905]" captionTargetPageId="19" captionText="FIGURE 12. Astrodendrum sagaminum, one syntype (ZSM 448/1). A, aboral disc and proximal portion of the arms; B, oral disc and proximal portion of the arms; C, oral view of proximal portion of the arms; D, lateral view of the disc and proximal portion of the arms. Arrowheads indicate madreporite (B, D). Abbreviations: AS, arm spine; GS, genital slit." httpUri="https://zenodo.org/record/1195433/files/figure.png" pageId="19" pageNumber="308">Fig. 12A</figureCitation>
). Oral surface of disc covered by external ossicles, approximately 200300 um in length (
<figureCitation id="35B407A0FF8B4164DD29ED68FB09D267" box="[1148,1251,295,320]" captionStart="FIGURE 12" captionStartId="19.[151,250,1929,1951]" captionTargetBox="[151,1416,991,1893]" captionTargetId="figure@19.[151,1436,988,1905]" captionTargetPageId="19" captionText="FIGURE 12. Astrodendrum sagaminum, one syntype (ZSM 448/1). A, aboral disc and proximal portion of the arms; B, oral disc and proximal portion of the arms; C, oral view of proximal portion of the arms; D, lateral view of the disc and proximal portion of the arms. Arrowheads indicate madreporite (B, D). Abbreviations: AS, arm spine; GS, genital slit." httpUri="https://zenodo.org/record/1195433/files/figure.png" pageId="19" pageNumber="308">Fig. 12B</figureCitation>
), flat, polygonal and fully in contact on oral plate and slightly tumid, round and slightly in contact on adoral plates (
<figureCitation id="35B407A0FF8B4164DDBBED03FABDD243" box="[1262,1367,332,357]" captionStart="FIGURE 12" captionStartId="19.[151,250,1929,1951]" captionTargetBox="[151,1416,991,1893]" captionTargetId="figure@19.[151,1436,988,1905]" captionTargetPageId="19" captionText="FIGURE 12. Astrodendrum sagaminum, one syntype (ZSM 448/1). A, aboral disc and proximal portion of the arms; B, oral disc and proximal portion of the arms; C, oral view of proximal portion of the arms; D, lateral view of the disc and proximal portion of the arms. Arrowheads indicate madreporite (B, D). Abbreviations: AS, arm spine; GS, genital slit." httpUri="https://zenodo.org/record/1195433/files/figure.png" pageId="19" pageNumber="308">Fig. 12A</figureCitation>
). Oral shields, adoral shields, oral plates and ventral arm plates completely concealed by external ossicles. Uniformly flat and spiniform teeth situated on the dental plates and edge of each oral plate, approximately
<quantity id="6A77B6C0FF8B4164DDFCEDDBFADBD28B" box="[1193,1329,404,429]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="19" pageNumber="308" unit="mm" value="0.65" valueMax="0.8" valueMin="0.5">0.50.8 mm</quantity>
in length and approximately
<quantity id="6A77B6C0FF8B4164D82DEDF8FE0AD2F7" box="[376,480,439,464]" metricMagnitude="-1" metricUnit="m" metricValue="5.715" metricValueMax="6.35" metricValueMin="5.08" pageId="19" pageNumber="308" unit="in" value="22.5" valueMax="25.0" valueMin="20.0">2025 in</quantity>
number on each jaw (
<figureCitation id="35B407A0FF8B4164DBA4EDF8FCB6D2F7" box="[753,860,439,464]" captionStart="FIGURE 12" captionStartId="19.[151,250,1929,1951]" captionTargetBox="[151,1416,991,1893]" captionTargetId="figure@19.[151,1436,988,1905]" captionTargetPageId="19" captionText="FIGURE 12. Astrodendrum sagaminum, one syntype (ZSM 448/1). A, aboral disc and proximal portion of the arms; B, oral disc and proximal portion of the arms; C, oral view of proximal portion of the arms; D, lateral view of the disc and proximal portion of the arms. Arrowheads indicate madreporite (B, D). Abbreviations: AS, arm spine; GS, genital slit." httpUri="https://zenodo.org/record/1195433/files/figure.png" pageId="19" pageNumber="308">Fig. 12B</figureCitation>
). Lateral surface of oral disc covered by conical external ossicles, separated and scattered, approximately 80100 µm in length and height (
<figureCitation id="35B407A0FF8B4164DD94ED93FAB9D2D3" box="[1217,1363,476,501]" captionStart="FIGURE 12" captionStartId="19.[151,250,1929,1951]" captionTargetBox="[151,1416,991,1893]" captionTargetId="figure@19.[151,1436,988,1905]" captionTargetPageId="19" captionText="FIGURE 12. Astrodendrum sagaminum, one syntype (ZSM 448/1). A, aboral disc and proximal portion of the arms; B, oral disc and proximal portion of the arms; C, oral view of proximal portion of the arms; D, lateral view of the disc and proximal portion of the arms. Arrowheads indicate madreporite (B, D). Abbreviations: AS, arm spine; GS, genital slit." httpUri="https://zenodo.org/record/1195433/files/figure.png" pageId="19" pageNumber="308">Fig. 12B, D</figureCitation>
). Two genital slits (
<quantity id="6A77B6C0FF8B4164D87EEDB0FE84D130" box="[299,366,511,536]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="19" pageNumber="308" unit="mm" value="5.0">5 mm</quantity>
long and
<quantity id="6A77B6C0FF8B4164D8B7EDB0FDD0D130" box="[482,570,511,536]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="19" pageNumber="308" unit="mm" value="1.5">1.5 mm</quantity>
wide) in each interradius (
<figureCitation id="35B407A0FF8B4164DA3BEDB0FC33D13F" box="[878,985,511,536]" captionStart="FIGURE 12" captionStartId="19.[151,250,1929,1951]" captionTargetBox="[151,1416,991,1893]" captionTargetId="figure@19.[151,1436,988,1905]" captionTargetPageId="19" captionText="FIGURE 12. Astrodendrum sagaminum, one syntype (ZSM 448/1). A, aboral disc and proximal portion of the arms; B, oral disc and proximal portion of the arms; C, oral view of proximal portion of the arms; D, lateral view of the disc and proximal portion of the arms. Arrowheads indicate madreporite (B, D). Abbreviations: AS, arm spine; GS, genital slit." httpUri="https://zenodo.org/record/1195433/files/figure.png" pageId="19" pageNumber="308">Fig. 12D</figureCitation>
). One elliptical madreporite situated on oral interradius, approximately
<quantity id="6A77B6C0FF8B4164D8ADEE6BFDA4D11B" box="[504,590,548,572]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="19" pageNumber="308" unit="mm" value="2.7">2.7 mm</quantity>
in length and 1.0 mm in height (
<figureCitation id="35B407A0FF8B4164DAE9EE6BFBC9D11B" box="[956,1059,548,573]" captionStart="FIGURE 12" captionStartId="19.[151,250,1929,1951]" captionTargetBox="[151,1416,991,1893]" captionTargetId="figure@19.[151,1436,988,1905]" captionTargetPageId="19" captionText="FIGURE 12. Astrodendrum sagaminum, one syntype (ZSM 448/1). A, aboral disc and proximal portion of the arms; B, oral disc and proximal portion of the arms; C, oral view of proximal portion of the arms; D, lateral view of the disc and proximal portion of the arms. Arrowheads indicate madreporite (B, D). Abbreviations: AS, arm spine; GS, genital slit." httpUri="https://zenodo.org/record/1195433/files/figure.png" pageId="19" pageNumber="308">Fig. 12B</figureCitation>
).
</paragraph>
<paragraph id="AD301B25FF8B4164D992EE07FD78D1EB" blockId="19.[151,1437,151,933]" pageId="19" pageNumber="308">
<emphasis id="9FFBC737FF8B4164D992EE07FEE0D178" box="[199,266,584,607]" italics="true" pageId="19" pageNumber="308">Arms.</emphasis>
Arms 5, branching at least 8 times but number of branches indeterminate because of entanglement of arms (
<figureCitation id="35B407A0FF8B4164D988EE23FEAED1A3" box="[221,324,620,645]" captionStart="FIGURE 12" captionStartId="19.[151,250,1929,1951]" captionTargetBox="[151,1416,991,1893]" captionTargetId="figure@19.[151,1436,988,1905]" captionTargetPageId="19" captionText="FIGURE 12. Astrodendrum sagaminum, one syntype (ZSM 448/1). A, aboral disc and proximal portion of the arms; B, oral disc and proximal portion of the arms; C, oral view of proximal portion of the arms; D, lateral view of the disc and proximal portion of the arms. Arrowheads indicate madreporite (B, D). Abbreviations: AS, arm spine; GS, genital slit." httpUri="https://zenodo.org/record/1195433/files/figure.png" pageId="19" pageNumber="308">Fig. 12C</figureCitation>
). The first branch occurs beyond disc (
<figureCitation id="35B407A0FF8B4164DBADEE23FC95D1A3" box="[760,895,620,645]" captionStart="FIGURE 12" captionStartId="19.[151,250,1929,1951]" captionTargetBox="[151,1416,991,1893]" captionTargetId="figure@19.[151,1436,988,1905]" captionTargetPageId="19" captionText="FIGURE 12. Astrodendrum sagaminum, one syntype (ZSM 448/1). A, aboral disc and proximal portion of the arms; B, oral disc and proximal portion of the arms; C, oral view of proximal portion of the arms; D, lateral view of the disc and proximal portion of the arms. Arrowheads indicate madreporite (B, D). Abbreviations: AS, arm spine; GS, genital slit." httpUri="https://zenodo.org/record/1195433/files/figure.png" pageId="19" pageNumber="308">Fig. 12A, B</figureCitation>
). On the proximal portion of the arm, width and height almost equal,
<quantity id="6A77B6C0FF8B4164D8DAEEC0FE00D180" box="[399,490,655,680]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="19" pageNumber="308" unit="mm" value="4.5">4.5 mm</quantity>
in width. Arms with a concave aboral surface and flattened oral surface. Arms tapering gradually toward arm tip (
<figureCitation id="35B407A0FF8B4164DB4AEEFBFD6CD1EB" box="[543,646,692,717]" captionStart="FIGURE 12" captionStartId="19.[151,250,1929,1951]" captionTargetBox="[151,1416,991,1893]" captionTargetId="figure@19.[151,1436,988,1905]" captionTargetPageId="19" captionText="FIGURE 12. Astrodendrum sagaminum, one syntype (ZSM 448/1). A, aboral disc and proximal portion of the arms; B, oral disc and proximal portion of the arms; C, oral view of proximal portion of the arms; D, lateral view of the disc and proximal portion of the arms. Arrowheads indicate madreporite (B, D). Abbreviations: AS, arm spine; GS, genital slit." httpUri="https://zenodo.org/record/1195433/files/figure.png" pageId="19" pageNumber="308">Fig. 12C</figureCitation>
).
</paragraph>
<paragraph id="AD301B25FF8B4164D992EE97FCBAD083" blockId="19.[151,1437,151,933]" pageId="19" pageNumber="308">
On aboral and lateral surface of proximal portion of arm, each arm segment covered by granular external ossicles, fully in contact (
<figureCitation id="35B407A0FF8B4164D896EEB3FDC7D033" box="[451,557,764,789]" captionStart="FIGURE 12" captionStartId="19.[151,250,1929,1951]" captionTargetBox="[151,1416,991,1893]" captionTargetId="figure@19.[151,1436,988,1905]" captionTargetPageId="19" captionText="FIGURE 12. Astrodendrum sagaminum, one syntype (ZSM 448/1). A, aboral disc and proximal portion of the arms; B, oral disc and proximal portion of the arms; C, oral view of proximal portion of the arms; D, lateral view of the disc and proximal portion of the arms. Arrowheads indicate madreporite (B, D). Abbreviations: AS, arm spine; GS, genital slit." httpUri="https://zenodo.org/record/1195433/files/figure.png" pageId="19" pageNumber="308">Fig. 12C</figureCitation>
). On proximal portion of arms, granules 100150 µm in length (
<figureCitation id="35B407A0FF8B4164DC71EEB3FA65D033" box="[1316,1423,764,789]" captionStart="FIGURE 12" captionStartId="19.[151,250,1929,1951]" captionTargetBox="[151,1416,991,1893]" captionTargetId="figure@19.[151,1436,988,1905]" captionTargetPageId="19" captionText="FIGURE 12. Astrodendrum sagaminum, one syntype (ZSM 448/1). A, aboral disc and proximal portion of the arms; B, oral disc and proximal portion of the arms; C, oral view of proximal portion of the arms; D, lateral view of the disc and proximal portion of the arms. Arrowheads indicate madreporite (B, D). Abbreviations: AS, arm spine; GS, genital slit." httpUri="https://zenodo.org/record/1195433/files/figure.png" pageId="19" pageNumber="308">Fig. 12A</figureCitation>
), gradually decreasing in size toward arm tip. Oral surface of proximal portion of arms covered by flat, polygonal plate-shaped external ossicles, similar to those on oral disc, approximately 200300 µm in length (
<figureCitation id="35B407A0FF8B4164DC71EF0BFA7AD07B" box="[1316,1424,836,861]" captionStart="FIGURE 12" captionStartId="19.[151,250,1929,1951]" captionTargetBox="[151,1416,991,1893]" captionTargetId="figure@19.[151,1436,988,1905]" captionTargetPageId="19" captionText="FIGURE 12. Astrodendrum sagaminum, one syntype (ZSM 448/1). A, aboral disc and proximal portion of the arms; B, oral disc and proximal portion of the arms; C, oral view of proximal portion of the arms; D, lateral view of the disc and proximal portion of the arms. Arrowheads indicate madreporite (B, D). Abbreviations: AS, arm spine; GS, genital slit." httpUri="https://zenodo.org/record/1195433/files/figure.png" pageId="19" pageNumber="308">Fig. 12C</figureCitation>
). Before the first branches, the first to third tentacle pores with no arm spine; single arm spine at fourth pores (
<figureCitation id="35B407A0FF8B4164DC3AEF28FF22D083" captionStart="FIGURE 12" captionStartId="19.[151,250,1929,1951]" captionTargetBox="[151,1416,991,1893]" captionTargetId="figure@19.[151,1436,988,1905]" captionTargetPageId="19" captionText="FIGURE 12. Astrodendrum sagaminum, one syntype (ZSM 448/1). A, aboral disc and proximal portion of the arms; B, oral disc and proximal portion of the arms; C, oral view of proximal portion of the arms; D, lateral view of the disc and proximal portion of the arms. Arrowheads indicate madreporite (B, D). Abbreviations: AS, arm spine; GS, genital slit." httpUri="https://zenodo.org/record/1195433/files/figure.png" pageId="19" pageNumber="308">Fig. 12B</figureCitation>
), and subsequent pores with 2 or 3 arm spines (
<figureCitation id="35B407A0FF8B4164DB89EFC3FCA9D083" box="[732,835,908,933]" captionStart="FIGURE 12" captionStartId="19.[151,250,1929,1951]" captionTargetBox="[151,1416,991,1893]" captionTargetId="figure@19.[151,1436,988,1905]" captionTargetPageId="19" captionText="FIGURE 12. Astrodendrum sagaminum, one syntype (ZSM 448/1). A, aboral disc and proximal portion of the arms; B, oral disc and proximal portion of the arms; C, oral view of proximal portion of the arms; D, lateral view of the disc and proximal portion of the arms. Arrowheads indicate madreporite (B, D). Abbreviations: AS, arm spine; GS, genital slit." httpUri="https://zenodo.org/record/1195433/files/figure.png" pageId="19" pageNumber="308">Fig. 12B</figureCitation>
).
</paragraph>
<caption id="F9F04BADFF8B4164D9C2EBC6FB09D4FB" httpUri="https://zenodo.org/record/1195433/files/figure.png" pageId="19" pageNumber="308" startId="19.[151,250,1929,1951]" targetBox="[151,1416,991,1893]" targetPageId="19">
<paragraph id="AD301B25FF8B4164D9C2EBC6FB09D4FB" blockId="19.[151,1436,1929,2012]" pageId="19" pageNumber="308">
<emphasis id="9FFBC737FF8B4164D9C2EBC6FECBD4B9" bold="true" box="[151,289,1929,1951]" pageId="19" pageNumber="308">FIGURE 12.</emphasis>
<taxonomicName id="6A8F60A6FF8B4164D87DEBC6FDDED4B8" authorityName="Doderlein" authorityYear="1911" box="[296,564,1929,1951]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Astrodendrum" kingdom="Animalia" order="Euryalida" pageId="19" pageNumber="308" phylum="Echinodermata" rank="species" species="sagaminum">
<emphasis id="9FFBC737FF8B4164D87DEBC6FDDED4B8" box="[296,564,1929,1951]" italics="true" pageId="19" pageNumber="308">Astrodendrum sagaminum</emphasis>
</taxonomicName>
, one syntype (ZSM 448/1). A, aboral disc and proximal portion of the arms; B, oral disc and proximal portion of the arms; C, oral view of proximal portion of the arms; D, lateral view of the disc and proximal portion of the arms. Arrowheads indicate madreporite (B, D). Abbreviations: AS, arm spine; GS, genital slit.
</paragraph>
</caption>
<paragraph id="AD301B25FF8C4163D992ECD8FD72D397" blockId="20.[151,1436,151,608]" box="[199,664,151,176]" pageId="20" pageNumber="309">
<emphasis id="9FFBC737FF8C4163D992ECD8FEE7D397" box="[199,269,151,176]" italics="true" pageId="20" pageNumber="309">Color.</emphasis>
Uniformly creamy white (
<figureCitation id="35B407A0FF8C4163DB62ECD8FD60D397" box="[567,650,151,176]" captionStart="FIGURE 12" captionStartId="19.[151,250,1929,1951]" captionTargetBox="[151,1416,991,1893]" captionTargetId="figure@19.[151,1436,988,1905]" captionTargetPageId="19" captionText="FIGURE 12. Astrodendrum sagaminum, one syntype (ZSM 448/1). A, aboral disc and proximal portion of the arms; B, oral disc and proximal portion of the arms; C, oral view of proximal portion of the arms; D, lateral view of the disc and proximal portion of the arms. Arrowheads indicate madreporite (B, D). Abbreviations: AS, arm spine; GS, genital slit." httpUri="https://zenodo.org/record/1195433/files/figure.png" pageId="20" pageNumber="309">Fig. 12</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E59548AEFF8C4163D992ECF3FD35D243" pageId="20" pageNumber="309" type="distribution">
<paragraph id="AD301B25FF8C4163D992ECF3FD35D243" blockId="20.[151,1436,151,608]" pageId="20" pageNumber="309">
<emphasis id="9FFBC737FF8C4163D992ECF3FE8AD3F2" bold="true" box="[199,352,188,213]" pageId="20" pageNumber="309">Distribution.</emphasis>
<collectingCountry id="D5985BB5FF8C4163D83FECF3FE28D3F3" box="[362,450,188,212]" name="Japan" pageId="20" pageNumber="309">JAPAN</collectingCountry>
. Sagami Bay (
<typeStatus id="7234A587FF8C4163DB39ECF3FD74D3F3" box="[620,670,188,212]" pageId="20" pageNumber="309">type</typeStatus>
locality,
<bibRefCitation id="C91E66D4FF8C4163DA5EECF3FC2DD3F3" box="[779,967,188,213]" pageId="20" pageNumber="309" refId="ref9238" refString="Doderlein, L. (1902) Japanische Euryaliden. Zoologishcer Anzeiger, 25, 320 - 326." type="journal article">Döderlein, 1902</bibRefCitation>
;
<bibRefCitation id="C91E66D4FF8C4163DA80ECF3FBE5D3F3" box="[981,1039,188,212]" pageId="20" pageNumber="309" refId="ref9257" refString="Doderlein, L. (1911) Beitrage zur Naturgeschichte Ostasiens. Uber japanische und andere Euryalae. Abhandlungen der Bayerischen Akademie der Wissenschaften, II. Suppl. - Bd., 5, 1 - 123." type="journal article">1911</bibRefCitation>
;
<bibRefCitation id="C91E66D4FF8C4163DD4BECF2FB00D3F3" box="[1054,1258,188,212]" pageId="20" pageNumber="309" refId="ref9496" refString="Matsumoto, H. (1917) A monograph of Japanese Ophiuroidea, arranged according to a new classification. Journal of College of Science Imperial University of Tokyo, 38, 1 - 408." type="journal article">Matsumoto, 1917</bibRefCitation>
;
<bibRefCitation id="C91E66D4FF8C4163DDADECF3FA72D3F3" box="[1272,1432,188,213]" pageId="20" pageNumber="309" refId="ref9327" refString="Irimura, S. (1982) The Brittle-stars of Sagami Bay. Maruzen Co. Ltd., Tokyo, 95 pp." type="book">Irimura, 1982</bibRefCitation>
,
<bibRefCitation id="C91E66D4FF8C4163D9C2ECAFFF38D3DF" box="[151,210,224,248]" pageId="20" pageNumber="309" refId="ref9351" refString="Irimura, S. (1991) Ophiuroidea. In: Echinoderms from continental shelf and slope around Japan. Vol 2. Japan Fisheries Resource Conservation Association, Tokyo, pp. 111 - 153." type="book chapter">1991</bibRefCitation>
; Jangoux
<emphasis id="9FFBC737FF8C4163D814ECAEFE9BD3DF" box="[321,369,223,248]" italics="true" pageId="20" pageNumber="309">et al</emphasis>
., 1987), off eastern
<collectingCountry id="D5985BB5FF8C4163DB04ECAFFD7BD3D0" box="[593,657,224,247]" name="Japan" pageId="20" pageNumber="309">Japan</collectingCountry>
(H. L. Clark, 1915); Suruga Bay (H. L. Clark, 1911; Döderlein, 1927; Kroh, 2002); Ogasawara or Yaeyama Islands (
<bibRefCitation id="C91E66D4FF8C4163DBE1ED4BFB3ED23A" box="[692,1236,260,285]" pageId="20" pageNumber="309" refId="ref9620" refString="Murakami, S. (1944) Report on the ophiurans from off Ogasawara Islands and from off the Yaeyama group, Nippon. Journal of the Department of Agriculture, Kyushu Imperial University, 7, 235 - 257." type="journal article">Murakami, 1944: detailed locality is unknown</bibRefCitation>
). East
<collectingCountry id="D5985BB5FF8C4163DC73ED4BFA83D23A" box="[1318,1385,260,285]" name="China" pageId="20" pageNumber="309">China</collectingCountry>
Sea (
<bibRefCitation id="C91E66D4FF8C4163D9CBED68FE3BD267" box="[158,465,295,320]" pageId="20" pageNumber="309" refId="ref9388" refString="Irimura, S. and Kubodera, T. (1998) Ophiuroidea in the East Chinae Sea. Memoires of the National Science Museum, 30, 135 - 143." type="journal article">Irimura &amp; Kubodera, 1998</bibRefCitation>
).
<collectingCountry id="D5985BB5FF8C4163D8B0ED67FDABD267" box="[485,577,296,320]" name="China" pageId="20" pageNumber="309">CHINA</collectingCountry>
. East
<collectingCountry id="D5985BB5FF8C4163DBD6ED68FD2CD267" box="[643,710,295,320]" name="China" pageId="20" pageNumber="309">China</collectingCountry>
Sea (
<bibRefCitation id="C91E66D4FF8C4163DA50ED68FC33D267" box="[773,985,295,320]" pageId="20" pageNumber="309" refId="ref9109" refString="Chang, F. Y., Liao, Y. L. &amp; Wu, B. J. (1962) Euryales (Echinodermata: Ophiuroidea: Euryalida) of China. Acta Zoologica Sinica, 14, 53 - 67." type="journal article">
Chang
<emphasis id="9FFBC737FF8C4163DA02ED66FC6DD267" box="[855,903,295,320]" italics="true" pageId="20" pageNumber="309">et al</emphasis>
., 1962
</bibRefCitation>
).
<collectingCountry id="D5985BB5FF8C4163DAB8ED67FB6CD267" box="[1005,1158,296,320]" name="Sri Lanka" pageId="20" pageNumber="309">SRI LANKA</collectingCountry>
. Southwest off Colombo (
<bibRefCitation id="C91E66D4FF8C4163D9CBED03FEA5D243" box="[158,335,332,357]" pageId="20" pageNumber="309" refId="ref9078" refString="Bomford, T. L. (1913) A note on certain ophiuroids in the Indian Museum. Records of the Indian Museum, 9, 219 - 225." type="journal article">Bomford, 1913</bibRefCitation>
). Depth ranges
<quantity id="6A77B6C0FF8C4163D8ABED03FD97D243" box="[510,637,332,356]" metricMagnitude="2" metricUnit="m" metricValue="6.95" metricValueMax="13.0" metricValueMin="0.9" pageId="20" pageNumber="309" unit="m" value="695.0" valueMax="1300.0" valueMin="90.0">901300 m</quantity>
(
<figureCitation id="35B407A0FF8C4163DBD9ED03FD27D243" box="[652,717,332,357]" captionStart="FIGURE 1" captionStartId="4.[151,250,1967,1989]" captionTargetBox="[273,1322,208,1919]" captionTargetId="figure@4.[194,1390,193,1949]" captionTargetPageId="4" captionText="FIGURE 1. Distribution of Astrodendrum spinulosum sp. nov., A. capense, A. elingamita, A. galapagense, A. laevigatum and A. sagaminum. Red symbols indicate type localities." httpUri="https://zenodo.org/record/1195411/files/figure.png" pageId="20" pageNumber="309">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E59548AEFF8C4163D992ED20FECDD147" pageId="20" pageNumber="309" type="discussion">
<paragraph id="AD301B25FF8C4163D992ED20FB2ED2F7" blockId="20.[151,1436,151,608]" pageId="20" pageNumber="309">
<emphasis id="9FFBC737FF8C4163D992ED20FED1D2AF" bold="true" box="[199,315,367,392]" pageId="20" pageNumber="309">Remarks.</emphasis>
<taxonomicName id="6A8F60A6FF8C4163D813ED20FD98D2AF" authorityName="Doderlein" authorityYear="1911" box="[326,626,367,392]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Astrodendrum" kingdom="Animalia" order="Euryalida" pageId="20" pageNumber="309" phylum="Echinodermata" rank="species" species="sagaminum">
<emphasis id="9FFBC737FF8C4163D813ED20FD98D2AF" box="[326,626,367,392]" italics="true" pageId="20" pageNumber="309">Astrodendrum sagaminum</emphasis>
</taxonomicName>
shares with
<taxonomicName id="6A8F60A6FF8C4163DA58ED3FFC5AD2AF" authority="Okanishi &amp; Fujita, 2018" authorityName="Okanishi &amp; Fujita" authorityYear="2018" box="[781,944,367,392]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Astrodendrum" kingdom="Animalia" order="Euryalida" pageId="20" pageNumber="309" phylum="Echinodermata" rank="species" species="spinulosum" status="sp. nov.">
<emphasis id="9FFBC737FF8C4163DA58ED3FFC5AD2AF" box="[781,944,367,392]" italics="true" pageId="20" pageNumber="309">A. spinulosum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84C87A4CFF8C4163DAE9ED20FBF1D2AF" box="[956,1051,367,392]" pageId="20" pageNumber="309" rank="species">
<emphasis id="9FFBC737FF8C4163DAE9ED20FBF1D2AF" bold="true" box="[956,1051,367,392]" pageId="20" pageNumber="309">sp. nov.</emphasis>
</taxonomicNameLabel>
the presence of conical external ossicles, that are, separated and scattered on interradial lateral disc, while those of
<emphasis id="9FFBC737FF8C4163DD65EDDBFAA4D28A" box="[1072,1358,404,429]" italics="true" pageId="20" pageNumber="309">
<taxonomicName id="6A8F60A6FF8C4163DD65EDDBFB46D28B" baseAuthorityName="Mortensen" baseAuthorityYear="1933" box="[1072,1196,404,428]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Astrodendrum" kingdom="Animalia" order="Euryalida" pageId="20" pageNumber="309" phylum="Echinodermata" rank="species" species="capense">A. capense</taxonomicName>
,
<taxonomicName id="6A8F60A6FF8C4163DDE2EDDBFAA4D28A" baseAuthorityName="Baker" baseAuthorityYear="1980" box="[1207,1358,404,429]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Astrodendrum" kingdom="Animalia" order="Euryalida" pageId="20" pageNumber="309" phylum="Echinodermata" rank="species" species="elingamita">A. elingamita</taxonomicName>
</emphasis>
and
<taxonomicName id="6A8F60A6FF8C4163DCD1EDDBFECED2F7" authorityName="A. H. Clark" authorityYear="1916" class="Ophiuroidea" family="Gorgonocephalidae" genus="Astrodendrum" kingdom="Animalia" order="Euryalida" pageId="20" pageNumber="309" phylum="Echinodermata" rank="species" species="galapagense">
<emphasis id="9FFBC737FF8C4163DCD1EDDBFECED2F7" italics="true" pageId="20" pageNumber="309">A. galapagense</emphasis>
</taxonomicName>
are plate-shaped and/or granular.
<taxonomicName id="6A8F60A6FF8C4163DBF4EDF7FCD7D2F7" box="[673,829,439,464]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Astrodendrum" kingdom="Animalia" order="Euryalida" pageId="20" pageNumber="309" phylum="Echinodermata" rank="species" species="laevigatum">
<emphasis id="9FFBC737FF8C4163DBF4EDF7FCD7D2F7" box="[673,829,439,464]" italics="true" pageId="20" pageNumber="309">A. laevigatum</emphasis>
</taxonomicName>
lacks external ossicles on the disc.
</paragraph>
<paragraph id="AD301B25FF8C4163D992ED93FECDD147" blockId="20.[151,1436,151,608]" pageId="20" pageNumber="309">
<taxonomicName id="6A8F60A6FF8C4163D992ED93FE86D2D3" authorityName="Doderlein" authorityYear="1911" box="[199,364,476,500]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Astrodendrum" kingdom="Animalia" order="Euryalida" pageId="20" pageNumber="309" phylum="Echinodermata" rank="species" species="sagaminum">
<emphasis id="9FFBC737FF8C4163D992ED93FE86D2D3" box="[199,364,476,500]" italics="true" pageId="20" pageNumber="309">A. sagaminum</emphasis>
</taxonomicName>
and
<taxonomicName id="6A8F60A6FF8C4163D8F9ED93FDBAD2D2" authority="Okanishi &amp; Fujita, 2018" authorityName="Okanishi &amp; Fujita" authorityYear="2018" box="[428,592,476,501]" class="Ophiuroidea" family="Gorgonocephalidae" genus="Astrodendrum" kingdom="Animalia" order="Euryalida" pageId="20" pageNumber="309" phylum="Echinodermata" rank="species" species="spinulosum" status="sp. nov.">
<emphasis id="9FFBC737FF8C4163D8F9ED93FDBAD2D2" box="[428,592,476,501]" italics="true" pageId="20" pageNumber="309">A. spinulosum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84C87A4CFF8C4163DB0EED93FD50D2D2" box="[603,698,476,501]" pageId="20" pageNumber="309" rank="species">
<emphasis id="9FFBC737FF8C4163DB0EED93FD50D2D2" bold="true" box="[603,698,476,501]" pageId="20" pageNumber="309">sp. nov.</emphasis>
</taxonomicNameLabel>
can be distinguished by absence/presence of granular external ossicles on disc, absence/presence of bulges of lateral edges on the proximal portion of the arms, and the maximum number of secondary teeth on hook-shaped arm spines in distal portion of the arms (Table 1, see also remarks for
<taxonomicName id="6A8F60A6FF8C4163DCD1EE6BFEFDD147" authorityName="Okanishi &amp; Fujita" authorityYear="2018" class="Ophiuroidea" family="Gorgonocephalidae" genus="Astrodendrum" kingdom="Animalia" order="Euryalida" pageId="20" pageNumber="309" phylum="Echinodermata" rank="species" species="spinulosum">
<emphasis id="9FFBC737FF8C4163DCD1EE6BFEFDD147" italics="true" pageId="20" pageNumber="309">A. spinulosum</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>