treatments-xml/data/25/20/87/252087CA1F1E9566FDA0FE2ED89AF80E.xml
2024-06-21 12:31:41 +02:00

1289 lines
154 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="49AFB4F0B59B4BE2F57FD1EB02EC330F" ID-CLB-Dataset="9775" ID-DOI="10.5852/ejt.2020.719.1107" ID-GBIF-Dataset="7ca72f76-4fae-4305-8601-4662f4cd2b96" ID-Zenodo-Dep="4064324" ID-ZooBank="A4500016-C219-4353-B81C-5E0BB520547F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1601679165585" checkinUser="valdenar" docAuthor="Williams, Paul H., Altanchimeg, Dorjsuren, Byvaltsev, Alexandr, Jonghe, Roland De, Jaffar, Saleem, Japoshvili, George, Kahono, Sih, Liang, Huan, Mei, Maurizio, Monfared, Alireza, Nidup, Tshering, Raina, Rifat, Ren, Zongxin, Thanoosing, Chawatat, Zhao, Yanhui &amp; Orr, Michael C." docDate="2020" docId="252087CA1F1E9566FDA0FE2ED89AF80E" docLanguage="en" docName="ejt-719_williams_altanchimeg_byvaltsev.pdf.imf" docOrigin="European Journal of Taxonomy 719" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Bombus separandus Vogt 1909, stat. rev." docType="treatment" docVersion="16" lastPageNumber="96" masterDocId="D919FFB21F429539FF87FFA1DC7AFFEB" masterDocTitle="Widespread polytypic species or complexes of local species? Revising bumblebees of the subgenus Melanobombus world-wide (Hymenoptera, Apidae, Bombus)" masterLastPageNumber="120" masterPageNumber="1" pageNumber="93" updateTime="1698845993937" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="E4F7980B5E892BFD7B739D61BBC092E6" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A4A18AB9E4911405083356F16103E8EF">
<mods:title id="A8F1E0A5E2B16D953F666400281A46E1">Widespread polytypic species or complexes of local species? Revising bumblebees of the subgenus Melanobombus world-wide (Hymenoptera, Apidae, Bombus)</mods:title>
</mods:titleInfo>
<mods:name id="0478B809C0D2FF35F08E03FF9DF90C79" type="personal">
<mods:role id="8B36239D088C6A8572762915769BA465">
<mods:roleTerm id="500BD43C0A8C97F2483E91846163FD37">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="41A8A78BE019D6D721C5D11F69423DF4">Williams, Paul H.</mods:namePart>
<mods:affiliation id="1FA32A5C5D3A538F2EA843B3E4B79F00">Natural History Museum, Cromwell Road, London SW 7 5 BD, UK. Institute of General and Experimental Biology, Peace Avenue 54 b, Ulaanbaatar 13330, Mongolia. Novosibirsk State University, ul. Pirogova 2, Novosibirsk, 630090 Russia. Langstraat 105, B- 2260 Westerlo, Belgium. South China Agricultural University, Guangzhou 510642, China. Agricultural University of Georgia, 240 Agmashenebli Alley, Tbilisi, Georgia. Indonesian Institute of Sciences (LIPI), Jakarta, Indonesia. Kunming Institute of Botany (Chinese Academy of Sciences), 132 Lanhei Road, Kunming, Yunnan 650201, China. Università di Roma Sapienza, Piazzale Valerio Massimo 6, Roma 00162, Italy. Yasouj University, Zirtol, Yasouj, Iran. Sherubtse College, Royal University of Bhutan, Trashigang, Bhutan. Zoological Survey of India, Pali Road, Jodhpur 342005, Rajasthan, India. Institute of Zoology (Chinese Academy of Sciences), 1 Beichen West Road, Chaoyang, Beijing 100101, China.</mods:affiliation>
<mods:nameIdentifier id="C75E73CAC1616555B63D5B8F874AF458" type="email">paw@nhm.ac.uk</mods:nameIdentifier>
</mods:name>
<mods:name id="47DD9C5B15B3DDFF0A38417F1B1D623C" type="personal">
<mods:role id="42CE7AA5314AF5E134D145E7C69FE9D6">
<mods:roleTerm id="E04B3D314D05A67336BB34D78792F776">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0226EFAEE9C6468C44273942B9E5E062">Altanchimeg, Dorjsuren</mods:namePart>
<mods:nameIdentifier id="72C6F7A48D32033533A69DB716154A70" type="email">altanchimegd@mas.ac.mn</mods:nameIdentifier>
</mods:name>
<mods:name id="82DB1B378252607C3CBC63399014CBD0" type="personal">
<mods:role id="E87BB2B35CB045C637545925E4EEADBF">
<mods:roleTerm id="B96A3BEA75C33984C8801B3472A8A118">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="09A2A4A2FDCC58474C351D112B0B5FE8">Byvaltsev, Alexandr</mods:namePart>
<mods:nameIdentifier id="0E51478F82754397D6DBB5170F64AB8F" type="email">byvam@yandex.ru</mods:nameIdentifier>
</mods:name>
<mods:name id="A4F2C5AEFED8BD1C5CF5EA19A867D300" type="personal">
<mods:role id="C5359BEAA1BFE2E6EA31A25C5AADF130">
<mods:roleTerm id="B3C59363783209E0D499B42A4D935FD3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C344F0C713C503D703D14288D76F6EDF">Jonghe, Roland De</mods:namePart>
<mods:nameIdentifier id="0FEE9592151A368DB783C31307494D8B" type="email">roland.de.jonghe@telenet.be</mods:nameIdentifier>
</mods:name>
<mods:name id="E9C71C233C9B49B7B3620409F1A635B5" type="personal">
<mods:role id="904755F6446059F85A7E8BF0DB1256E4">
<mods:roleTerm id="79E579814FBB8F2B0EBAB6793869B255">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A0F2BB712DE7AC851CAA047DC20C15D4">Jaffar, Saleem</mods:namePart>
<mods:nameIdentifier id="36A4055706D0B170972D2F3F8498A738" type="email">saleemjaffar@stu.scau.edu.cn</mods:nameIdentifier>
</mods:name>
<mods:name id="AC4CF9F349F5EA25C96FB14CEEB4FECA" type="personal">
<mods:role id="3A0E7DAEE0D3DF5A5153A19D5914A308">
<mods:roleTerm id="E9958805B7616C98874FEFE46AD0D462">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B29C18D8A683E50F943E4B1B0993FBFE">Japoshvili, George</mods:namePart>
<mods:nameIdentifier id="664ECAF6AD8D96EE78C13DBE3A67BCA2" type="email">g.japoshvili@agruni.edu.ge</mods:nameIdentifier>
</mods:name>
<mods:name id="FDC7A35231262D995526555574FF993E" type="personal">
<mods:role id="5F476AAFA7366B5373F27D2570D260C1">
<mods:roleTerm id="D4D515C8B0A67325F4860E356071133A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3E677E27231B641A767D7903E4D9C066">Kahono, Sih</mods:namePart>
<mods:nameIdentifier id="80D5F43EEE01571BD1B3D78BBFBAF41F" type="email">sihkahono@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="D21F017C1776E90DBA1D288770EBA175" type="personal">
<mods:role id="4AD3075C470A5AEB29E06F057887213A">
<mods:roleTerm id="939C46B77546FA048F4973B6EC6BC55E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9D4040DB6081F2A5079B6F986D12B66B">Liang, Huan</mods:namePart>
<mods:nameIdentifier id="10A352255873E7BC62CE79DF4B074CEB" type="email">lianghuan@mail.kib.ac.cn</mods:nameIdentifier>
</mods:name>
<mods:name id="8B87DC1ECB7FAA5447BB2D3FDB69DC98" type="personal">
<mods:role id="CE275366C1BF33F093539AF18F31F397">
<mods:roleTerm id="B4E10BE61B30971CF8C74C265E4C6425">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3892AC10F794F1C2FBDDC4B57CA00B6A">Mei, Maurizio</mods:namePart>
</mods:name>
<mods:name id="B754C62F27EC8D1E99C9B4381C9C2C6F" type="personal">
<mods:role id="76C03503EA9C3F57583BC4F037EB6253">
<mods:roleTerm id="A7EA7F337C99D31CFD98C210F45EA14B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="34B0B3553B35B1C6E86DA16240377A09">Monfared, Alireza</mods:namePart>
</mods:name>
<mods:name id="451BE4892B694CE5D79FD64CAE55D78B" type="personal">
<mods:role id="BAE7ADEEBCB4CC3EF7FC36B16FFABB77">
<mods:roleTerm id="D86EE97286E75585370AA446872A1185">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E78C14BE937EDB7D2E4C9F33F1D08E9D">Nidup, Tshering</mods:namePart>
</mods:name>
<mods:name id="FC46B5417C5AC235756CE1379B31B49D" type="personal">
<mods:role id="DE306A9C4AF6CE83CCD8229B56628003">
<mods:roleTerm id="BAEE44EE8C7CC7125DE134AE01B8F8D8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="834F62C8803DE5874068AF8E386ADCAA">Raina, Rifat</mods:namePart>
</mods:name>
<mods:name id="BBCFD0C89F86B69A91978B7FA4F0DB6B" type="personal">
<mods:role id="18892314FEC75036AA3F201A1BBE6D80">
<mods:roleTerm id="73E2474F318CD4E76ED52E05236C1D58">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8E829BA96BC214354631CA10747A3B5A">Ren, Zongxin</mods:namePart>
</mods:name>
<mods:name id="8E4F6E2E9434B7B5FF713AD4482D5703" type="personal">
<mods:role id="6188A579DD56D4E4D2BB1C44033B8473">
<mods:roleTerm id="02447A5E42B8EC25B3BA66C4E64769D9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="15DAD84EFAB55566600549B0F3664C6C">Thanoosing, Chawatat</mods:namePart>
</mods:name>
<mods:name id="8B0230668545DB4BEEDB236F251AD8EC" type="personal">
<mods:role id="A8B4498381FECDEE2ADC437B68932FD1">
<mods:roleTerm id="7717DDC6E0A7267BA9AA8CFF8A1CD1A2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8B01A45D50F66D50689D82E467FC6425">Zhao, Yanhui</mods:namePart>
</mods:name>
<mods:name id="03D5BC830F4CF45BB0F2C0774D3AED5A" type="personal">
<mods:role id="FACC4D084D979FD7291C640F4B090D6E">
<mods:roleTerm id="B9F491D5809FE7B9442E7B48C4CBF520">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D0607F3ACB70F459B37312BD27EE5245">Orr, Michael C.</mods:namePart>
</mods:name>
<mods:typeOfResource id="806F98FE9BAAB92A19B72562F619305E">text</mods:typeOfResource>
<mods:relatedItem id="1418997A4AD8B89A02D049DFDD5ED770" type="host">
<mods:titleInfo id="5A920B404220AC620D6AA692977B0795">
<mods:title id="F1EB0608AEF9A5E5482F6711AE4E8BF6">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="063CD5F850A65A5CFEBCA85811080419">
<mods:date id="DFA9726D47AC36F8960D92FC9AD1EEC2">2020</mods:date>
<mods:detail id="EF544FE21980893DA347251148D675CD" type="pubDate">
<mods:number id="0AE846AA4C2980281B0EFF96E30436BF">2020-10-02</mods:number>
</mods:detail>
<mods:detail id="2D74FE7A16890B0052CA659247D747ED" type="volume">
<mods:number id="63AB8D12A38EA2CF90947D4A61120684">719</mods:number>
</mods:detail>
<mods:extent id="80CBBC824F82DBE7119BCE1E0440CDCE" unit="page">
<mods:start id="D8BBBBC3183434FBC24EE806A7499477">1</mods:start>
<mods:end id="8329D2B631B7F9A75E901BC2DE8D90B1">120</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="9AAF10C252E2366653BB2D240F427080">journal article</mods:classification>
<mods:identifier id="824AD393D5F1AC658956F3189E11678C" type="CLB-Dataset">9775</mods:identifier>
<mods:identifier id="7AB071BE3AC66BAA289AEB4427235036" type="DOI">10.5852/ejt.2020.719.1107</mods:identifier>
<mods:identifier id="B84D4B4FF1C73A709647E16DFA2361E1" type="GBIF-Dataset">7ca72f76-4fae-4305-8601-4662f4cd2b96</mods:identifier>
<mods:identifier id="B9EE0078975138C3155AECFC29EC8A99" type="Zenodo-Dep">4064324</mods:identifier>
<mods:identifier id="D76794F3F1393FBAF4A73146FCD44E97" type="ZooBank">A4500016-C219-4353-B81C-5E0BB520547F</mods:identifier>
</mods:mods>
<treatment id="252087CA1F1E9566FDA0FE2ED89AF80E" ID-DOI="http://doi.org/10.5281/zenodo.4335562" ID-GBIF-Taxon="168220864" ID-Zenodo-Dep="4335562" LSID="urn:lsid:plazi:treatment:252087CA1F1E9566FDA0FE2ED89AF80E" httpUri="http://treatment.plazi.org/id/252087CA1F1E9566FDA0FE2ED89AF80E" lastPageId="95" lastPageNumber="96" pageId="92" pageNumber="93">
<subSubSection id="E59365571F1E9565FDA0FE2ED877FE41" box="[551,1037,399,426]" pageId="92" pageNumber="93" type="nomenclature">
<paragraph id="AD3636DC1F1E9565FDA0FE2ED877FE41" blockId="92.[551,1037,399,460]" box="[551,1037,399,426]" pageId="92" pageNumber="93">
<heading id="F67E81B01F1E9565FDA0FE2ED877FE41" box="[551,1037,399,426]" centered="true" fontSize="11" level="2" pageId="92" pageNumber="93" reason="2">
<taxonomicName id="6A894D5F1F1E9565FDA0FE2EDFD9FE42" ID-CoL="8P66B" authority="Vogt, 1909" authorityName="Vogt" authorityYear="1909" box="[551,931,399,426]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="separandus" status="stat. nov.">
<emphasis id="9FFDEACE1F1E9565FDA0FE2EDF66FE41" bold="true" box="[551,796,399,426]" italics="true" pageId="92" pageNumber="93">Bombus separandus</emphasis>
<bibRefCitation id="C9184B2D1F1E9565FCA4FE31DFD9FE42" author="Vogt O." box="[803,931,399,426]" pageId="92" pageNumber="93" pagination="28 - 84" refId="ref71967" refString="Vogt O. 1909. Studien ʾ ber das Artproblem. 1. Mitteilung. Uber das Variieren der Hummeln. 1. Teil. Sitzungsberichte der Gesellschaft naturforschender Freunde zu Berlin 1909: 28 - 84." type="journal article" year="1909">Vogt, 1909</bibRefCitation>
</taxonomicName>
<taxonomicNameLabel id="84CE57B51F1E9565FC2DFE31D877FE41" box="[938,1037,400,426]" pageId="92" pageNumber="93">stat. rev.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E59365571F1E9565FDDCFE13DFA2FE27" box="[603,984,434,460]" pageId="92" pageNumber="93" type="description">
<paragraph id="AD3636DC1F1E9565FDDCFE13DFA2FE27" blockId="92.[551,1037,399,460]" box="[603,984,434,460]" pageId="92" pageNumber="93">
<figureCitation id="35B22A591F1E9565FDDCFE13DECFFE27" box="[603,693,434,460]" captionStart="Figs 1416" captionStartId="25.[189,241,1956,1982]" captionTargetBox="[288,1321,265,1901]" captionTargetId="figure-11@25.[262,1325,265,1917]" captionTargetPageId="25" captionText="Figs 1416. Maps of sequenced samples as in Figs 1213. 14. The tanguticus-group and the lapidarius- group. 15. The sichelii-group. 16. The keriensis-group." figureDoi="http://doi.org/10.5281/zenodo.4064340" httpUri="https://zenodo.org/record/4064340/files/figure.png" pageId="92" pageNumber="93">Figs 16</figureCitation>
,
<figureCitation id="35B22A591F1E9565FD46FE13DF51FE27" box="[705,811,434,460]" captionStart="Figs 139180" captionStartId="42.[189,241,1497,1523]" captionTargetBox="[369,1221,268,1431]" captionTargetId="figure-11@42.[366,1221,265,1467]" captionTargetPageId="42" captionText="Figs 139180. Simplified diagrams for the colour patterns of the hair on the dorsum for the species from the integrative analysis. The dorsum is divided into regions, each of which shows only the predominant or most apparent colour for that region using a simplified colour palette, with olive indicating a mixture of black and yellow hair, and grey indicating a mixture of black and white hair. The keriensis-group. 139. Queen, China-Sichuan. 140. Worker, China-Sichuan. 141. Queen, China-Sichuan. 142. Queen, China-Sichuan. 143. Worker, China-Xizang. 144. Queen, India-Kashmir. 145. Male, China-Qinghai. 146. Male, India-Kashmir. 147. Male, China-Gansu. 148. Queen, Armenia. 149. Queen, Iran. 150. Male, Georgia. 151. Worker, China-Qinghai. 152. Male, China-Qinghai. 153. Queen, Mongolia. 154. Worker, Mongolia. 155. Male, Mongolia. 156. Male, Mongolia. 157. Queen, China-Sichuan. 158. Worker, China-Qinghai. 159. Worker, China-Qinghai. 160. Male, China-Qinghai. 161. Male, China-Qinghai. 162. Male, China-Qinghai. 163. Queen, Pakistan. 164. Queen, Pakistan. 165. Queen, Pakistan. 166. Queen, Pakistan. 167. Queen, Afghanistan. 168. Queen, Pakistan. 169. Queen, Pakistan. 170. Queen, India-Kashmir. 171. Male, India-Kashmir. 172. Male, India-Kashmir. 173. Queen, China- Xinjiang. 174. Queen, Kyrgyzstan. 175. Queen, Kyrgyzstan. 176. Queen, Mongolia. 177. Queen, India- Kashmir. 178. Male, Kazakhstan. 179. Male, China-Xinjiang. 180. Male, Mongolia." figureDoi="http://doi.org/10.5281/zenodo.4064358" httpUri="https://zenodo.org/record/4064358/files/figure.png" pageId="92" pageNumber="93">173180</figureCitation>
,
<figureCitation id="35B22A591F1E9565FCB0FE13DF1CFE27" box="[823,870,434,460]" captionStart="Figs 199204" captionStartId="45.[189,241,1852,1878]" captionTargetBox="[230,1342,827,1806]" captionTargetId="figure-159@45.[194,1393,771,1810]" captionTargetPageId="45" captionText="Figs 199204. Morphology of the male genitalia for species of the subgenus Melanobombus von Dalla Torre, 1880 from the dorsal aspect, anterior at the bottom of the image, posterior at the top. 199. Bombus alagesianus Reinig, 1930 stat. rev., Georgia. 200. B. tibeticus sp. nov., China-Qinghai. 201. B. incertoides Vogt, 1911 stat. rev., Mongolia. 202. B. qilianensis sp. nov., China-Qinghai. 203. B. keriensis Morawitz, 1887, India-Kashmir. 204. B. separandus Vogt, 1909 stat. rev., Kyrgyzstan. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4064364" httpUri="https://zenodo.org/record/4064364/files/figure.png" pageId="92" pageNumber="93">204</figureCitation>
,
<figureCitation id="35B22A591F1E9565FCF6FE13DFDAFE27" box="[881,928,434,460]" captionStart="Figs 209210" captionStartId="93.[189,241,1884,1910]" captionTargetBox="[190,1374,318,1788]" captionTargetId="figure-11@93.[189,1398,318,1792]" captionTargetPageId="93" captionText="Figs 209210. Images of queens of two cryptic species from around the western Qinghai-Tibetan plateau. 209. Bombus keriensis Morawitz, 1887 from Mt Apharwat (4000 m a.s.l.) in the Pir Panjal mountains (ML405). 210. B. separandus Vogt, 1909 stat. rev. from Nimaling plain (4800 m a.s.l.) in the Zanskar mountains (ML311). Viewed from the left lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.4064370" httpUri="https://zenodo.org/record/4064370/files/figure.png" pageId="92" pageNumber="93">210</figureCitation>
,
<figureCitation id="35B22A591F1E9565FC2BFE13DFA2FE27" box="[940,984,434,460]" captionStart="Figs 211212" captionStartId="100.[189,241,1742,1768]" captionTargetBox="[319,1269,265,1703]" captionTargetId="figure-10@100.[319,1269,265,1703]" captionTargetPageId="100" captionText="Figs 211212. Maps of sequenced samples showing the most frequent colour patterns locally. 211. The sichelii-complex, consisting of the single polytypic species B. sichelii Radoszkowski, 1859 (inset: B. sichelii worker, China-Neimenggu, photo PW). 212. The keriensis-complex, consisting of B. alagesianus Reinig, 1930 stat. rev. (blue spots), B. tibeticus sp. nov. (dark green spots), B. incertoides Vogt, 1911 stat. rev. (light green spots), B. qilianensis sp. nov. (yellow spots), B. keriensis Morawitz, 1887 (orange spots), and B. separandus Vogt, 1909 stat. rev. (red spots) (inset: B.qilianensis sp. nov. queen, China-Sichuan, photo PW). Maps projected onto a sphere in ArcGIS using World_Shaded_Relief basemap © 2014 ESRI and showing boundaries between countries as recognised by the UN." figureDoi="http://doi.org/10.5281/zenodo.4064372" httpUri="https://zenodo.org/record/4064372/files/figure.png" pageId="92" pageNumber="93">212</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="E59365571F1E9565FF3AFE55D95EFBB4" pageId="92" pageNumber="93" type="reference_group">
<paragraph id="AD3636DC1F1E9565FF3AFE55DFEBFDE5" blockId="92.[189,1399,499,1019]" box="[189,913,499,526]" pageId="92" pageNumber="93">
<treatmentCitationGroup id="8D9911F21F1E9565FF3AFE55DFEBFDE5" box="[189,913,499,526]" pageId="92" pageNumber="93">
<taxonomicName id="6A894D5F1F1E9565FF3AFE55DDE6FDE5" ID-CoL="MFMT" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[189,412,500,526]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="lapidarius">
<emphasis id="9FFDEACE1F1E9565FF3AFE55DDE6FDE5" box="[189,412,500,526]" italics="true" pageId="92" pageNumber="93">Bombus lapidarius</emphasis>
</taxonomicName>
Form [subsp.]
<treatmentCitation id="2C2810CD1F1E9565FDC8FE55DFF6FDE5" author="Vogt O." box="[591,908,500,526]" page="61" pageId="92" pageNumber="93" year="1909">
<taxonomicName id="6A894D5F1F1E9565FDC8FE55DFF6FDE5" ID-CoL="8P66B" authority="Vogt, 1909: 61" authorityName="Vogt" authorityPageNumber="61" authorityYear="1909" box="[591,908,500,526]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="separandus">
<emphasis id="9FFDEACE1F1E9565FDC8FE55DEACFDE5" box="[591,726,500,526]" italics="true" pageId="92" pageNumber="93">separandus</emphasis>
<bibRefCitation id="C9184B2D1F1E9565FD5AFE55DFF6FDE5" author="Vogt O." box="[733,908,500,526]" pageId="92" pageNumber="93" pagination="28 - 84" refId="ref71967" refString="Vogt O. 1909. Studien ʾ ber das Artproblem. 1. Mitteilung. Uber das Variieren der Hummeln. 1. Teil. Sitzungsberichte der Gesellschaft naturforschender Freunde zu Berlin 1909: 28 - 84." type="journal article" year="1909">Vogt, 1909: 61</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="AD3636DC1F1E9565FF3AFDB6DD22FDBF" blockId="92.[189,1399,499,1019]" pageId="92" pageNumber="93">
<treatmentCitationGroup id="8D9911F21F1E9565FF3AFDB6DD22FDBF" pageId="92" pageNumber="93">
<taxonomicName id="6A894D5F1F1E9565FF3AFDB6DDE0FDDA" ID-CoL="MFMT" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[189,410,535,561]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="lapidarius">
<emphasis id="9FFDEACE1F1E9565FF3AFDB6DDE0FDDA" box="[189,410,535,561]" italics="true" pageId="92" pageNumber="93">Bombus lapidarius</emphasis>
</taxonomicName>
Form [subsp.]
<emphasis id="9FFDEACE1F1E9565FDCFFDB6DEF9FDDA" box="[584,643,535,561]" italics="true" pageId="92" pageNumber="93">kohli</emphasis>
<treatmentCitation id="2C2810CD1F1E9565FD0FFDB9DF48FDDA" author="Vogt O." box="[648,818,535,562]" page="61" pageId="92" pageNumber="93" year="1909">
<bibRefCitation id="C9184B2D1F1E9565FD0FFDB9DF48FDDA" author="Vogt O." box="[648,818,535,562]" pageId="92" pageNumber="93" pagination="28 - 84" refId="ref71967" refString="Vogt O. 1909. Studien ʾ ber das Artproblem. 1. Mitteilung. Uber das Variieren der Hummeln. 1. Teil. Sitzungsberichte der Gesellschaft naturforschender Freunde zu Berlin 1909: 28 - 84." type="journal article" year="1909">Vogt, 1909: 61</bibRefCitation>
</treatmentCitation>
(non
<treatmentCitation id="2C2810CD1F1E9565FCF3FDB6D822FDD9" author="Cockerell T. D. A." box="[884,1112,535,562]" page="75" pageId="92" pageNumber="93" year="1906">
<bibRefCitation id="C9184B2D1F1E9565FCF3FDB6D822FDD9" author="Cockerell T. D. A." box="[884,1112,535,562]" pageId="92" pageNumber="93" pagination="69 - 75" refId="ref64506" refString="Cockerell T. D. A. 1906. Descriptions and records of bees. - XII. Annals and Magazine of Natural History Series 7 18: 69 - 75. https: // doi. org / 10.1080 / 00222930608562581" type="journal article" year="1906">Cockerell, 1906: 75</bibRefCitation>
</treatmentCitation>
=
<taxonomicName id="6A894D5F1F1E9565FBF2FDB9D909FDD9" ID-CoL="MFQP" authority="(Swederus))" authorityName="" baseAuthorityName="Swederus" baseAuthorityYear="1787" box="[1141,1395,535,562]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="morio">
<emphasis id="9FFDEACE1F1E9565FBF2FDB9D8A0FDDA" box="[1141,1242,536,561]" italics="true" pageId="92" pageNumber="93">B. morio</emphasis>
(Swederus))
</taxonomicName>
.
<emphasis id="9FFDEACE1F1E9565FF6DFD9BDD22FDBF" bold="true" box="[234,344,570,596]" pageId="92" pageNumber="93">syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="AD3636DC1F1E9565FF3AFDFCD883FD9C" blockId="92.[189,1399,499,1019]" box="[189,1273,605,632]" pageId="92" pageNumber="93">
<treatmentCitationGroup id="8D9911F21F1E9565FF3AFDFCD883FD9C" box="[189,1273,605,632]" pageId="92" pageNumber="93">
<treatmentCitation id="2C2810CD1F1E9565FF3AFDFCDE06FD9C" author="Skorikov A. S." box="[189,636,605,631]" page="413" pageId="92" pageNumber="93" year="1910">
<taxonomicName id="6A894D5F1F1E9565FF3AFDFCDE06FD9C" ID-CoL="8P65W" authority="Skorikov, 1910 a: 413" authorityName="Skorikov" authorityPageNumber="413" authorityYear="1910" box="[189,636,605,631]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="kozlovi">
<emphasis id="9FFDEACE1F1E9565FF3AFDFCDD0CFD9C" box="[189,374,605,631]" italics="true" pageId="92" pageNumber="93">Bombus kozlovi</emphasis>
<bibRefCitation id="C9184B2D1F1E9565FEFAFDFCDE06FD9C" author="Skorikov A. S." box="[381,636,605,631]" pageId="92" pageNumber="93" pagination="409 - 413" refId="ref70619" refString="Skorikov A. S. 1910 a. [New forms of bumblebees] (Hymenoptera, Bombidae). ([Preliminary diagnoses]). III. Russkoe entomologicheskoe Obozrenie 9 (1909): 409 - 413. [In Russian.]" type="journal article" year="1910">Skorikov, 1910a: 413</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, replacement name for
<emphasis id="9FFDEACE1F1E9565FC09FDFCDFB3FD9C" box="[910,969,605,631]" italics="true" pageId="92" pageNumber="93">kohli</emphasis>
<treatmentCitation id="2C2810CD1F1E9565FC57FDFFD8FAFD9C" author="Vogt O." box="[976,1152,605,632]" page="61" pageId="92" pageNumber="93" year="1909">
<bibRefCitation id="C9184B2D1F1E9565FC57FDFFD8FAFD9C" author="Vogt O." box="[976,1152,605,632]" pageId="92" pageNumber="93" pagination="28 - 84" refId="ref71967" refString="Vogt O. 1909. Studien ʾ ber das Artproblem. 1. Mitteilung. Uber das Variieren der Hummeln. 1. Teil. Sitzungsberichte der Gesellschaft naturforschender Freunde zu Berlin 1909: 28 - 84." type="journal article" year="1909">Vogt, 1909: 61</bibRefCitation>
</treatmentCitation>
.
<emphasis id="9FFDEACE1F1E9565FB0BFDFCD889FD9C" bold="true" box="[1164,1267,605,631]" pageId="92" pageNumber="93">syn. nov</emphasis>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="AD3636DC1F1E9565FF3AFD21DFA3FD71" blockId="92.[189,1399,499,1019]" box="[189,985,640,667]" pageId="92" pageNumber="93">
<treatmentCitationGroup id="8D9911F21F1E9565FF3AFD21DFA3FD71" box="[189,985,640,667]" pageId="92" pageNumber="93">
<taxonomicName id="6A894D5F1F1E9565FF3AFD21DDE6FD71" ID-CoL="MFMT" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[189,412,640,666]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="lapidarius">
<emphasis id="9FFDEACE1F1E9565FF3AFD21DDE6FD71" box="[189,412,640,666]" italics="true" pageId="92" pageNumber="93">Bombus lapidarius</emphasis>
</taxonomicName>
var. [subsp.]
<treatmentCitation id="2C2810CD1F1E9565FDBBFD21DF1AFD71" author="Friese H." box="[572,864,640,667]" page="86" pageId="92" pageNumber="93" year="1913">
<taxonomicName id="6A894D5F1F1E9565FDBBFD21DF1AFD71" ID-CoL="8PF53" authority="Friese, 1913: 86" authorityName="Friese" authorityPageNumber="86" authorityYear="1913" box="[572,864,640,667]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="tenellus">
<emphasis id="9FFDEACE1F1E9565FDBBFD21DEEDFD71" box="[572,663,640,666]" italics="true" pageId="92" pageNumber="93">tenellus</emphasis>
<bibRefCitation id="C9184B2D1F1E9565FD19FD21DF1AFD71" author="Friese H." box="[670,864,640,667]" pageId="92" pageNumber="93" pagination="85 - 89" refId="ref65615" refString="Friese H. 1913. Uber einige neue Apiden (Hym.). Archiv fur Naturgeschichte 78: 85 - 89." type="journal article" year="1913">Friese, 1913: 86</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<emphasis id="9FFDEACE1F1E9565FCECFD20DFA8FD70" bold="true" box="[875,978,641,667]" pageId="92" pageNumber="93">syn. nov</emphasis>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="AD3636DC1F1E9565FF3AFD05DF5AFD0A" blockId="92.[189,1399,499,1019]" pageId="92" pageNumber="93">
<treatmentCitationGroup id="8D9911F21F1E9565FF3AFD05DF5AFD0A" pageId="92" pageNumber="93">
<taxonomicName id="6A894D5F1F1E9565FF3AFD05DD05FD55" ID-CoL="8PF53" authorityName="Friese" authorityYear="1913" box="[189,383,676,702]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="tenellus">
<emphasis id="9FFDEACE1F1E9565FF3AFD05DD05FD55" box="[189,383,676,702]" italics="true" pageId="92" pageNumber="93">Bombus tenellus</emphasis>
</taxonomicName>
var. [not subsp.]
<emphasis id="9FFDEACE1F1E9565FDD5FD05DEBEFD55" box="[594,708,676,702]" italics="true" pageId="92" pageNumber="93">alpivagus</emphasis>
<treatmentCitation id="2C2810CD1F1E9565FD4BFD05DFC5FD55" author="Richards O. W." box="[716,959,676,702]" page="639" pageId="92" pageNumber="93" year="1930">
<bibRefCitation id="C9184B2D1F1E9565FD4BFD05DFC5FD55" author="Richards O. W." box="[716,959,676,702]" pageId="92" pageNumber="93" pagination="633 - 658" refId="ref70041" refString="Richards O. W. 1930. The humble-bees captured on the expeditions to Mt. Everest (Hymenoptera, Bombidae). Annals and Magazine of Natural History Series 10 5: 633 - 658. https: // doi. org / 10.1080 / 00222933008673177" type="journal article" year="1930">Richards, 1930: 639</bibRefCitation>
</treatmentCitation>
, but not infrasubspecific after
<treatmentCitation id="2C2810CD1F1E9565FAAFFD05DD1AFD0A" author="Reinig W. F." page="333" pageId="92" pageNumber="93" year="1935">
<bibRefCitation id="C9184B2D1F1E9565FAAFFD05DD1AFD0A" author="Reinig W. F." pageId="92" pageNumber="93" pagination="321 - 356" refId="ref69854" refString="Reinig W. F. 1935. On the variation of Bombus lapidarius L. and its cuckoo, Psithyrus rupestris Fabr., with notes on mimetic similarity. Journal of Genetics 30: 321 - 356. https: // doi. org / 10.1007 / BF 02982243" type="journal article" year="1935">Reinig 1935: 333</bibRefCitation>
</treatmentCitation>
(
<bibRefCitation id="C9184B2D1F1E9565FEF7FD66DD86FD0A" author="ICZN" box="[368,508,711,737]" pageId="92" pageNumber="93" refId="ref66549" refString="ICZN 1999. International Code of Zoological Nomenclature 4 th Ed .. International Commission on Zoological Nomenclature, London. Available from https: // www. iczn. org / the-code / the-international-code-of-zoological-nomenclature / the-code-online / [accessed 14 Aug. 2020]." type="url" year="1999">ICZN 1999</bibRefCitation>
: Art. 45.6.4.1).
<emphasis id="9FFDEACE1F1E9565FD34FD66DF5AFD0A" bold="true" box="[691,800,711,737]" pageId="92" pageNumber="93">syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="AD3636DC1F1E9565FF3AFD4BD846FCC3" blockId="92.[189,1399,499,1019]" pageId="92" pageNumber="93">
<treatmentCitationGroup id="8D9911F21F1E9565FF3AFD4BD846FCC3" pageId="92" pageNumber="93">
<treatmentCitation id="2C2810CD1F1E9565FF3AFD4BDFBDFCEF" author="Skorikov A. S." box="[189,967,746,772]" page="226" pageId="92" pageNumber="93" year="1931">
<taxonomicName id="6A894D5F1F1E9565FF3AFD4BDFBDFCEF" ID-CoL="8QDW7" authority="Skorikov, 1931: 226" authorityName="Skorikov" authorityPageNumber="226" authorityYear="1931" box="[189,967,746,772]" class="Insecta" family="Apidae" genus="Lapidariobombus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="subSpecies" species="alagesianus" subSpecies="pamirus">
<emphasis id="9FFDEACE1F1E9565FF3AFD4BDE5AFCEF" box="[189,544,746,772]" italics="true" pageId="92" pageNumber="93">Lapidariobombus alagesianus</emphasis>
subsp.
<emphasis id="9FFDEACE1F1E9565FDF1FD4ADEACFCEF" box="[630,726,747,772]" italics="true" pageId="92" pageNumber="93">pamirus</emphasis>
<bibRefCitation id="C9184B2D1F1E9565FD5BFD4BDFBDFCEF" author="Skorikov A. S." box="[732,967,746,772]" pageId="92" pageNumber="93" pagination="175 - 247" refId="ref70808" refString="Skorikov A. S. 1931. Die Hummelfauna Turkestans und ihre Beziehungen zur zentralasiatischen Fauna (Hymenoptera, Bombidae). In: Lindholm V. A. (ed.) Abhandlungen der Pamir-Expedition 1928: 175 - 247. Academy of Sciences of the USSR, Leningrad." type="journal article" year="1931">Skorikov, 1931: 226</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(non
<treatmentCitation id="2C2810CD1F1E9565FB8EFD4BD88FFCEF" author="Skorikov A. S." box="[1033,1269,746,772]" page="232" pageId="92" pageNumber="93" year="1931">
<bibRefCitation id="C9184B2D1F1E9565FB8EFD4BD88FFCEF" author="Skorikov A. S." box="[1033,1269,746,772]" pageId="92" pageNumber="93" pagination="175 - 247" refId="ref70808" refString="Skorikov A. S. 1931. Die Hummelfauna Turkestans und ihre Beziehungen zur zentralasiatischen Fauna (Hymenoptera, Bombidae). In: Lindholm V. A. (ed.) Abhandlungen der Pamir-Expedition 1928: 175 - 247. Academy of Sciences of the USSR, Leningrad." type="journal article" year="1931">Skorikov, 1931: 232</bibRefCitation>
</treatmentCitation>
=
<taxonomicName id="6A894D5F1F1E9565FA96FD4ADD25FCCC" ID-CoL="MFRK" authority="Morawitz" authorityName="Morawitz" authorityYear="1883" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="oberti">
<emphasis id="9FFDEACE1F1E9565FA96FD4AD90CFCEF" box="[1297,1398,746,772]" italics="true" pageId="92" pageNumber="93">B. oberti</emphasis>
Morawitz
</taxonomicName>
, not
<taxonomicName id="6A894D5F1F1E9565FE1FFCAFDECCFCCC" ID-CoL="MFH3" authority="Skorikov" authorityName="Skorikov" authorityYear="1912" box="[408,694,781,807]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="difficillimus">
<emphasis id="9FFDEACE1F1E9565FE1FFCAFDE39FCCC" box="[408,579,781,807]" italics="true" pageId="92" pageNumber="93">B. difficillimus</emphasis>
Skorikov
</taxonomicName>
as given by Skorikov).
<emphasis id="9FFDEACE1F1E9565FC49FCACD846FCC3" bold="true" box="[974,1084,781,808]" pageId="92" pageNumber="93">syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="AD3636DC1F1E9565FF3AFC91DE3AFC85" blockId="92.[189,1399,499,1019]" pageId="92" pageNumber="93">
<treatmentCitationGroup id="8D9911F21F1E9565FF3AFC91DE3AFC85" pageId="92" pageNumber="93">
<treatmentCitation id="2C2810CD1F1E9565FF3AFC91D85AFCA1" author="Skorikov A. S." box="[189,1056,816,843]" page="226" pageId="92" pageNumber="93" year="1931">
<taxonomicName id="6A894D5F1F1E9565FF3AFC91D85AFCA1" ID-CoL="8QFN4" authority="Skorikov, 1931: 226" authorityName="Skorikov" authorityPageNumber="226" authorityYear="1931" box="[189,1056,816,843]" class="Insecta" family="Apidae" genus="Lapidariobombus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="subSpecies" species="alagesianus" subSpecies="mongolicus">
<emphasis id="9FFDEACE1F1E9565FF3AFC91DE52FCA1" box="[189,552,816,842]" italics="true" pageId="92" pageNumber="93">Lapidariobombus alagesianus</emphasis>
subsp.
<emphasis id="9FFDEACE1F1E9565FD09FC91DF6FFCA1" box="[654,789,816,842]" italics="true" pageId="92" pageNumber="93">mongolicus</emphasis>
<bibRefCitation id="C9184B2D1F1E9565FCA3FC91D85AFCA1" author="Skorikov A. S." box="[804,1056,816,843]" pageId="92" pageNumber="93" pagination="175 - 247" refId="ref70808" refString="Skorikov A. S. 1931. Die Hummelfauna Turkestans und ihre Beziehungen zur zentralasiatischen Fauna (Hymenoptera, Bombidae). In: Lindholm V. A. (ed.) Abhandlungen der Pamir-Expedition 1928: 175 - 247. Academy of Sciences of the USSR, Leningrad." type="journal article" year="1931">Skorikov, 1931: 226</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(non
<treatmentCitation id="2C2810CD1F1E9565FBF5FC91D931FCA0" author="Friese H." box="[1138,1355,816,843]" page="110" pageId="92" pageNumber="93" year="1916">
<bibRefCitation id="C9184B2D1F1E9565FBF5FC91D931FCA0" author="Friese H." box="[1138,1355,816,843]" pageId="92" pageNumber="93" pagination="489 - 520" refId="ref65657" refString="Friese H. 1916. Uber einige neue Hummelformen. Bombus 1916: 107 - 110. Friese H. 1918. Wissenschaftlishe Ergebnisse einer Forschungsreise nach Ostindien, ausgef ʾ hrt im Auftrage der Kgl. Preuss. Akademie der Wissenschaften zu Berlin von Herrn Prof. Dr. v. Buttel-Reepen. VII. Bienen aus Sumatra, Java, Malakka und Ceylon. Gesammelt von Herrn Prof. Dr. Buttel-Reepen in den Jahren 1911 - 1912. Zoologische Jahrbucher, Abteilung fur Systematik, Geographie und Biologie der Tiere 41: 489 - 520." type="journal article" year="1916">Friese, 1916: 110</bibRefCitation>
</treatmentCitation>
?=
<taxonomicName id="6A894D5F1F1E9565FF6DFCF4DDC5FC84" ID-CoL="MFH9" authority="Smith" authorityName="Smith" authorityYear="1869" box="[234,447,852,879]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="diversus">
<emphasis id="9FFDEACE1F1E9565FF6DFCF4DD17FC85" box="[234,365,852,878]" italics="true" pageId="92" pageNumber="93">B. diversus</emphasis>
Smith
</taxonomicName>
).
<emphasis id="9FFDEACE1F1E9565FE54FCF5DE3AFC85" bold="true" box="[467,576,852,878]" pageId="92" pageNumber="93">syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="AD3636DC1F1E9565FF3AFCD6DF84FC7A" blockId="92.[189,1399,499,1019]" box="[189,1022,886,914]" pageId="92" pageNumber="93">
<treatmentCitationGroup id="8D9911F21F1E9565FF3AFCD6DF84FC7A" box="[189,1022,886,914]" pageId="92" pageNumber="93">
<taxonomicName id="6A894D5F1F1E9565FF3AFCD6DDF1FC7A" ID-CoL="MFM2" authorityName="F. Morawitz" authorityYear="1887" box="[189,395,887,913]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="keriensis">
<emphasis id="9FFDEACE1F1E9565FF3AFCD6DDF1FC7A" box="[189,395,887,913]" italics="true" pageId="92" pageNumber="93">Bombus keriensis</emphasis>
</taxonomicName>
f.g. [subsp.]
<emphasis id="9FFDEACE1F1E9565FDA1FCD6DED2FC7A" box="[550,680,887,913]" italics="true" pageId="92" pageNumber="93">bucharicus</emphasis>
<treatmentCitation id="2C2810CD1F1E9565FD37FCD6DFFDFC7A" author="Reinig W. F." box="[688,903,887,914]" page="340" pageId="92" pageNumber="93" year="1935">
<bibRefCitation id="C9184B2D1F1E9565FD37FCD6DFFDFC7A" author="Reinig W. F." box="[688,903,887,914]" pageId="92" pageNumber="93" pagination="321 - 356" refId="ref69854" refString="Reinig W. F. 1935. On the variation of Bombus lapidarius L. and its cuckoo, Psithyrus rupestris Fabr., with notes on mimetic similarity. Journal of Genetics 30: 321 - 356. https: // doi. org / 10.1007 / BF 02982243" type="journal article" year="1935">Reinig, 1935: 340</bibRefCitation>
</treatmentCitation>
.
<emphasis id="9FFDEACE1F1E9565FC16FCD6DF84FC7A" bold="true" box="[913,1022,887,913]" pageId="92" pageNumber="93">syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="AD3636DC1F1E9565FF3AFC3BD90DFC5E" blockId="92.[189,1399,499,1019]" box="[189,1399,922,949]" pageId="92" pageNumber="93">
<treatmentCitationGroup id="8D9911F21F1E9565FF3AFC3BD90DFC5E" box="[189,1399,922,949]" pageId="92" pageNumber="93">
<taxonomicName id="6A894D5F1F1E9565FF3AFC3BDDF0FC5F" ID-CoL="MFM2" authorityName="F. Morawitz" authorityYear="1887" box="[189,394,922,948]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="keriensis">
<emphasis id="9FFDEACE1F1E9565FF3AFC3BDDF0FC5F" box="[189,394,922,948]" italics="true" pageId="92" pageNumber="93">Bombus keriensis</emphasis>
</taxonomicName>
f.g. [subsp.]
<emphasis id="9FFDEACE1F1E9565FDA4FC3BDEF7FC5F" box="[547,653,922,948]" italics="true" pageId="92" pageNumber="93">richardsi</emphasis>
<treatmentCitation id="2C2810CD1F1E9565FD13FC3BDF1CFC5F" author="Reinig W. F." box="[660,870,922,949]" page="341" pageId="92" pageNumber="93" year="1935">
<bibRefCitation id="C9184B2D1F1E9565FD13FC3BDF1CFC5F" author="Reinig W. F." box="[660,870,922,949]" pageId="92" pageNumber="93" pagination="321 - 356" refId="ref69854" refString="Reinig W. F. 1935. On the variation of Bombus lapidarius L. and its cuckoo, Psithyrus rupestris Fabr., with notes on mimetic similarity. Journal of Genetics 30: 321 - 356. https: // doi. org / 10.1007 / BF 02982243" type="journal article" year="1935">Reinig, 1935: 341</bibRefCitation>
</treatmentCitation>
(non
<treatmentCitation id="2C2810CD1F1E9565FC2EFC3BD821FC5F" author="Frison T. H." box="[937,1115,922,948]" page="6" pageId="92" pageNumber="93" year="1930">
<bibRefCitation id="C9184B2D1F1E9565FC2EFC3BD821FC5F" author="Frison T. H." box="[937,1115,922,948]" pageId="92" pageNumber="93" pagination="1 - 22" refId="ref65809" refString="Frison T. H. 1930. The bumblebees of Java, Sumatra and Borneo (Bremidae: Hymenoptera). Treubia 12: 1 - 22." type="journal article" year="1930">Frison, 1930: 6</bibRefCitation>
</treatmentCitation>
=
<taxonomicName id="6A894D5F1F1E9565FBFFFC3AD911FC5E" ID-CoL="MFTW" authority="Lepeletier" authorityName="Lepeletier de Saint-Fargeau" authorityYear="1835" box="[1144,1387,922,949]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="rufipes">
<emphasis id="9FFDEACE1F1E9565FBFFFC3AD892FC5F" box="[1144,1256,922,948]" italics="true" pageId="92" pageNumber="93">B. rufipes</emphasis>
Lepeletier
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="AD3636DC1F1E9565FF3AFC1CDDE9FC10" blockId="92.[189,1399,499,1019]" pageId="92" pageNumber="93">
<treatmentCitationGroup id="8D9911F21F1E9565FF3AFC1CDDE9FC10" pageId="92" pageNumber="93">
<taxonomicName id="6A894D5F1F1E9565FF3AFC1CDE3FFC3C" ID-CoL="8P66H" authorityName="Wang" authorityYear="1982" box="[189,581,957,983]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="subSpecies" species="tenellus" subSpecies="tibetensis">
<emphasis id="9FFDEACE1F1E9565FF3AFC1CDD04FC3C" box="[189,382,957,983]" italics="true" pageId="92" pageNumber="93">Bombus tenellus</emphasis>
subsp.
<emphasis id="9FFDEACE1F1E9565FE51FC1CDE3FFC3C" box="[470,581,957,983]" italics="true" pageId="92" pageNumber="93">tibetensis</emphasis>
</taxonomicName>
S.-F.
<treatmentCitation id="2C2810CD1F1E9565FD0EFC1FDF2EFC3C" author="Wang S. - F." box="[649,852,957,984]" page="439" pageId="92" pageNumber="93" year="1982">
<bibRefCitation id="C9184B2D1F1E9565FD0EFC1FDF2EFC3C" author="Wang S. - F." box="[649,852,957,984]" pageId="92" pageNumber="93" pagination="427 - 447" refId="ref72195" refString="Wang S. - F. 1982. Hymenoptera: Apidae - Bombus. In: Qinghai-Tibet Plateau Comprehensive Scientific Investigation Team of the Chinese Academy of Sciences (ed.) Insects of Xizang: 427 - 447. Science Press, Beijing." type="book chapter" year="1982">Wang, 1982: 439</bibRefCitation>
</treatmentCitation>
, replacement name for
<emphasis id="9FFDEACE1F1E9565FBE1FC1CD8AAFC3C" box="[1126,1232,957,983]" italics="true" pageId="92" pageNumber="93">richardsi</emphasis>
<treatmentCitation id="2C2810CD1F1E9565FB50FC1CDD63FC10" author="Reinig W. F." page="341" pageId="92" pageNumber="93" year="1935">
<bibRefCitation id="C9184B2D1F1E9565FB50FC1CDD63FC10" author="Reinig W. F." pageId="92" pageNumber="93" pagination="321 - 356" refId="ref69854" refString="Reinig W. F. 1935. On the variation of Bombus lapidarius L. and its cuckoo, Psithyrus rupestris Fabr., with notes on mimetic similarity. Journal of Genetics 30: 321 - 356. https: // doi. org / 10.1007 / BF 02982243" type="journal article" year="1935">Reinig, 1935: 341</bibRefCitation>
</treatmentCitation>
.
<emphasis id="9FFDEACE1F1E9565FEA2FC40DDE9FC10" bold="true" box="[293,403,993,1019]" pageId="92" pageNumber="93">syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="AD3636DC1F1E9565FF3AFB83DFF2FBD7" blockId="92.[189,1316,1058,1120]" box="[189,904,1058,1085]" pageId="92" pageNumber="93">
<treatmentCitationGroup id="8D9911F21F1E9565FF3AFB83DFF2FBD7" box="[189,904,1058,1085]" pageId="92" pageNumber="93">
<taxonomicName id="6A894D5F1F1E9565FF3AFB83DD07FBD7" ID-CoL="8P65X" authorityName="Skorikov" authorityYear="1910" box="[189,381,1058,1084]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="kozlowi">
<emphasis id="9FFDEACE1F1E9565FF3AFB83DD07FBD7" box="[189,381,1058,1084]" italics="true" pageId="92" pageNumber="93">Bombus kozlowi</emphasis>
</taxonomicName>
<treatmentCitation id="2C2810CD1F1E9565FE1DFB83DE12FBD7" author="Bischoff H." box="[410,616,1058,1084]" page="9" pageId="92" pageNumber="93" year="1936">
<bibRefCitation id="C9184B2D1F1E9565FE1DFB83DE12FBD7" author="Bischoff H." box="[410,616,1058,1084]" pageId="92" pageNumber="93" pagination="1 - 27" refId="ref63947" refString="Bischoff H. 1936. Schwedisch-chinesische wissenschaftliche Expedition nach den nordwestlichen Provinzen Chinas, unter Leitung von Dr. Sven Hedin und Prof. S ʾ Ping-chang. Insekten gesammelt vom schwedischen Arzt der Expedition Dr. David Hummel 1927 - 1930. 56. Hymenoptera. 10. Bombinae. Arkiv f ˆ r zoologi 27 A: 1 - 27." type="journal article" year="1936">Bischoff, 1936: 9</bibRefCitation>
</treatmentCitation>
, unjustified emendation.
</treatmentCitationGroup>
</paragraph>
<paragraph id="AD3636DC1F1E9565FF3AFBE4D95EFBB4" blockId="92.[189,1316,1058,1120]" box="[189,1316,1093,1120]" pageId="92" pageNumber="93">
<treatmentCitationGroup id="8D9911F21F1E9565FF3AFBE4D95EFBB4" box="[189,1316,1093,1120]" pageId="92" pageNumber="93">
<taxonomicName id="6A894D5F1F1E9565FF3AFBE4DDF1FBB4" ID-CoL="MFM2" authorityName="F. Morawitz" authorityYear="1887" box="[189,395,1093,1119]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="keriensis">
<emphasis id="9FFDEACE1F1E9565FF3AFBE4DDF1FBB4" box="[189,395,1093,1119]" italics="true" pageId="92" pageNumber="93">Bombus keriensis</emphasis>
</taxonomicName>
(part)
<treatmentCitation id="2C2810CD1F1E9565FE76FBE4DEBAFBB4" author="Reinig W. F." box="[497,704,1093,1120]" page="341" pageId="92" pageNumber="93" year="1935">
<bibRefCitation id="C9184B2D1F1E9565FE76FBE4DEBAFBB4" author="Reinig W. F." box="[497,704,1093,1120]" pageId="92" pageNumber="93" pagination="321 - 356" refId="ref69854" refString="Reinig W. F. 1935. On the variation of Bombus lapidarius L. and its cuckoo, Psithyrus rupestris Fabr., with notes on mimetic similarity. Journal of Genetics 30: 321 - 356. https: // doi. org / 10.1007 / BF 02982243" type="journal article" year="1935">Reinig 1935: 341</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="2C2810CD1F1E9565FD68FBE4DFADFBB4" author="Williams P. H." box="[751,983,1093,1119]" page="134" pageId="92" pageNumber="93" year="1998">
<bibRefCitation id="C9184B2D1F1E9565FD68FBE4DFADFBB4" author="Williams P. H." box="[751,983,1093,1119]" pageId="92" pageNumber="93" pagination="79 - 152" refId="ref72435" refString="Williams P. H. 1998. An annotated checklist of bumble bees with an analysis of patterns of description (Hymenoptera: Apidae, Bombini). Bulletin of The Natural History Museum (Entomology) 67: 79 - 152. Available and updated from www. nhm. ac. uk / bombus / [accessed 2019]." type="journal article" year="1998">Williams 1998: 134</bibRefCitation>
</treatmentCitation>
(non
<treatmentCitation id="2C2810CD1F1E9565FB9BFBE4D96DFBB4" author="Morawitz F. F." box="[1052,1303,1093,1119]" page="199" pageId="92" pageNumber="93" year="1887">
<bibRefCitation id="C9184B2D1F1E9565FB9BFBE4D96DFBB4" author="Morawitz F. F." box="[1052,1303,1093,1119]" pageId="92" pageNumber="93" pagination="195 - 229" refId="ref68420" refString="Morawitz F. F. 1887. Insecta in itinere cl. N. Przewalskii in Asia centrali novissime lecta. I. Apidae. Trudy Russkago entomologicheskago obshchestva 20 (1886): 195 - 229." type="journal article" year="1887">Morawitz, 1887: 199</bibRefCitation>
</treatmentCitation>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E59365571F1E9567FF3AFB26DEE9FB24" lastPageId="94" lastPageNumber="95" pageId="92" pageNumber="93" type="description">
<paragraph id="AD3636DC1F1E9565FF3AFB26DFA6FA51" blockId="92.[189,1399,1159,1467]" pageId="92" pageNumber="93">
<bibRefCitation id="C9184B2D1F1E9565FF3AFB26DD06FB49" author="Vogt O." box="[189,380,1159,1186]" pageId="92" pageNumber="93" pagination="28 - 84" refId="ref71967" refString="Vogt O. 1909. Studien ʾ ber das Artproblem. 1. Mitteilung. Uber das Variieren der Hummeln. 1. Teil. Sitzungsberichte der Gesellschaft naturforschender Freunde zu Berlin 1909: 28 - 84." type="journal article" year="1909">Vogt (1909: 46)</bibRefCitation>
stated that his “Formen” are subpopulations, which can be considered equivalent to subspecies.
<bibRefCitation id="C9184B2D1F1E9565FECBFB0BDE55FB2F" author="Reinig W. F." box="[332,559,1194,1221]" pageId="92" pageNumber="93" pagination="321 - 356" refId="ref69854" refString="Reinig W. F. 1935. On the variation of Bombus lapidarius L. and its cuckoo, Psithyrus rupestris Fabr., with notes on mimetic similarity. Journal of Genetics 30: 321 - 356. https: // doi. org / 10.1007 / BF 02982243" type="journal article" year="1935">Reinig (1935: 333)</bibRefCitation>
stated explicitly that his “f.g.” (
<emphasis id="9FFDEACE1F1E9565FC31FB0BD8E0FB2F" box="[950,1178,1194,1220]" italics="true" pageId="92" pageNumber="93">forma geographica</emphasis>
) are equivalent to subspecies.
<bibRefCitation id="C9184B2D1F1E9565FECFFB6FDE7BFB03" author="Vogt O." box="[328,513,1229,1256]" pageId="92" pageNumber="93" pagination="28 - 84" refId="ref71967" refString="Vogt O. 1909. Studien ʾ ber das Artproblem. 1. Mitteilung. Uber das Variieren der Hummeln. 1. Teil. Sitzungsberichte der Gesellschaft naturforschender Freunde zu Berlin 1909: 28 - 84." type="journal article" year="1909">Vogt (1909: 61)</bibRefCitation>
described his taxon
<taxonomicName id="6A894D5F1F1E9565FD77FB6CDF0DFB0C" authorityName="Vogt" authorityYear="1909" box="[752,887,1229,1255]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="separandus">
<emphasis id="9FFDEACE1F1E9565FD77FB6CDF0DFB0C" box="[752,887,1229,1255]" italics="true" pageId="92" pageNumber="93">separandus</emphasis>
</taxonomicName>
as having the hair of the face and top of the head (“Gesicht und Stirn”) black and the underside of the thorax dark haired. He described his taxon
<emphasis id="9FFDEACE1F1E9565FF3AFAB2DC82FAC6" box="[189,248,1299,1325]" italics="true" pageId="92" pageNumber="93">kohli</emphasis>
(later renamed
<taxonomicName id="6A894D5F1F1E9565FE37FAB2DE79FAC6" authorityName="Skorikov" authorityYear="1910" box="[432,515,1299,1325]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="kozlovi">
<emphasis id="9FFDEACE1F1E9565FE37FAB2DE79FAC6" box="[432,515,1299,1325]" italics="true" pageId="92" pageNumber="93">kozlovi</emphasis>
</taxonomicName>
) as having the face with a few yellow hairs. However,
<bibRefCitation id="C9184B2D1F1E9565FB02FAB5D947FAC5" author="Vogt O." box="[1157,1341,1299,1326]" pageId="92" pageNumber="93" pagination="31 - 74" refId="ref72005" refString="Vogt O. 1911. Studien ʾ ber das Artproblem. 2. Mitteilung. Uber das Variieren der Hummeln. 2. Teil. (Schluss). Sitzungsberichte der Gesellschaft naturforschender Freunde zu Berlin 1911: 31 - 74." type="journal article" year="1911">Vogt (1911: 58)</bibRefCitation>
then characterised the taxon
<taxonomicName id="6A894D5F1F1E9565FE50FA96DE24FABA" authorityName="Vogt" authorityYear="1909" box="[471,606,1335,1361]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="separandus">
<emphasis id="9FFDEACE1F1E9565FE50FA96DE24FABA" box="[471,606,1335,1361]" italics="true" pageId="92" pageNumber="93">separandus</emphasis>
</taxonomicName>
as differing from the taxon
<taxonomicName id="6A894D5F1F1E9565FC29FA96D86CFABA" authorityName="F. Morawitz" authorityYear="1887" box="[942,1046,1335,1361]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="keriensis">
<emphasis id="9FFDEACE1F1E9565FC29FA96D86CFABA" box="[942,1046,1335,1361]" italics="true" pageId="92" pageNumber="93">keriensis</emphasis>
</taxonomicName>
only in having an absence of pale hairs on the face (he did not include the taxon
<emphasis id="9FFDEACE1F1E9565FC89FAFBDF33FA9F" box="[782,841,1370,1396]" italics="true" pageId="92" pageNumber="93">kohli</emphasis>
/
<taxonomicName id="6A894D5F1F1E9565FCDFFAFBDFD1FA9F" authorityName="Skorikov" authorityYear="1910" box="[856,939,1370,1396]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="kozlovi">
<emphasis id="9FFDEACE1F1E9565FCDFFAFBDFD1FA9F" box="[856,939,1370,1396]" italics="true" pageId="92" pageNumber="93">kozlovi</emphasis>
</taxonomicName>
as part of the taxon
<taxonomicName id="6A894D5F1F1E9565FB10FAFBD964FA9F" authorityName="Vogt" authorityYear="1909" box="[1175,1310,1370,1396]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="separandus">
<emphasis id="9FFDEACE1F1E9565FB10FAFBD964FA9F" box="[1175,1310,1370,1396]" italics="true" pageId="92" pageNumber="93">separandus</emphasis>
</taxonomicName>
but did include the taxon
<taxonomicName id="6A894D5F1F1E9565FE08FADCDE6BFA7C" authorityName="Vogt" authorityYear="1911" baseAuthorityName="Vogt" baseAuthorityYear="1911" box="[399,529,1405,1431]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="incertoides">
<emphasis id="9FFDEACE1F1E9565FE08FADCDE6BFA7C" box="[399,529,1405,1431]" italics="true" pageId="92" pageNumber="93">incertoides</emphasis>
</taxonomicName>
as part of
<taxonomicName id="6A894D5F1F1E9565FD08FADCDF6CFA7C" authorityName="Vogt" authorityYear="1909" box="[655,790,1405,1431]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="separandus">
<emphasis id="9FFDEACE1F1E9565FD08FADCDF6CFA7C" box="[655,790,1405,1431]" italics="true" pageId="92" pageNumber="93">separandus</emphasis>
</taxonomicName>
). The taxon
<taxonomicName id="6A894D5F1F1E9565FC2AFADCD84EFA7C" authorityName="Vogt" authorityYear="1909" box="[941,1076,1405,1431]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="separandus">
<emphasis id="9FFDEACE1F1E9565FC2AFADCD84EFA7C" box="[941,1076,1405,1431]" italics="true" pageId="92" pageNumber="93">separandus</emphasis>
</taxonomicName>
was treated as a part of the species
<taxonomicName id="6A894D5F1F1E9565FE9EFA00DDD8FA51" authorityName="F. Morawitz" authorityYear="1887" box="[281,418,1440,1466]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" sensu="lato" species="keriensis">
<emphasis id="9FFDEACE1F1E9565FE9EFA00DDD8FA51" box="[281,418,1440,1466]" italics="true" pageId="92" pageNumber="93">B. keriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84CE57B51F1E9565FE2EFA00DD92FA51" box="[425,488,1440,1467]" pageId="92" pageNumber="93" sensu="lato">s. lat.</taxonomicNameLabel>
by
<bibRefCitation id="C9184B2D1F1E9565FD93FA01DEC2FA50" author="Reinig W. F." box="[532,696,1440,1467]" pageId="92" pageNumber="93" pagination="321 - 356" refId="ref69854" refString="Reinig W. F. 1935. On the variation of Bombus lapidarius L. and its cuckoo, Psithyrus rupestris Fabr., with notes on mimetic similarity. Journal of Genetics 30: 321 - 356. https: // doi. org / 10.1007 / BF 02982243" type="journal article" year="1935">Reinig (1935)</bibRefCitation>
and by
<bibRefCitation id="C9184B2D1F1E9565FC92FA01DFA2FA51" author="Williams P. H." box="[789,984,1440,1466]" pageId="92" pageNumber="93" pagination="79 - 152" refId="ref72435" refString="Williams P. H. 1998. An annotated checklist of bumble bees with an analysis of patterns of description (Hymenoptera: Apidae, Bombini). Bulletin of The Natural History Museum (Entomology) 67: 79 - 152. Available and updated from www. nhm. ac. uk / bombus / [accessed 2019]." type="journal article" year="1998">Williams (1998)</bibRefCitation>
.
</paragraph>
<paragraph id="AD3636DC1F1E9565FF3AFA43DE8DF98E" blockId="92.[189,1399,1506,1638]" pageId="92" pageNumber="93">
Our PTP analysis (
<figureCitation id="35B22A591F1E9565FE27FA43DD8DFA17" box="[416,503,1506,1532]" captionStart="Fig" captionStartId="21.[189,232,1953,1979]" captionTargetBox="[233,1316,282,1913]" captionTargetId="figure-12@21.[233,1355,265,1914]" captionTargetPageId="21" captionText="Fig. 10. October 2019 (Fig. 8) Bayesian PTP analysis of MrBayes tree of unique COI barcodes (UHF- PTP). Symbols as in Fig. 9." figureDoi="http://doi.org/10.5281/zenodo.4064332" httpUri="https://zenodo.org/record/4064332/files/figure.png" pageId="92" pageNumber="93">Fig. 10</figureCitation>
) of coalescents in the COI gene within the
<taxonomicName id="6A894D5F1F1E9565FB80FA43D815FA17" authorityName="F. Morawitz" authorityYear="1887" box="[1031,1135,1506,1532]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="keriensis">
<emphasis id="9FFDEACE1F1E9565FB80FA43D815FA17" box="[1031,1135,1506,1532]" italics="true" pageId="92" pageNumber="93">keriensis</emphasis>
</taxonomicName>
-complex supports six species including
<taxonomicName id="6A894D5F1F1E9565FE11F9A7DE3BF9F4" authorityName="Vogt" authorityYear="1909" box="[406,577,1541,1567]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="separandus">
<emphasis id="9FFDEACE1F1E9565FE11F9A7DE3BF9F4" box="[406,577,1541,1567]" italics="true" pageId="92" pageNumber="93">B. separandus</emphasis>
</taxonomicName>
, corroborated by differences in morphology. These species are also supported by the absence of a positive divergence-with-distance relationship among them (
<figureCitation id="35B22A591F1E9565FB64F989D942F9A9" box="[1251,1336,1576,1602]" captionStart="Figs 1720" captionStartId="26.[189,241,1773,1799]" captionTargetBox="[191,1390,661,1666]" captionTargetId="figure-140@26.[189,1398,653,1671]" captionTargetPageId="26" captionText="Figs 1720. Plots of (y-axis) pairwise proportion of genetic divergence between COI barcode region sequences against (x-axis) pairwise Great Circle geographical distance in km between sample sites with linear trend lines (black). 17. Genetic divergences within the candidate species for the sichelii-complex identified using Bayesian UF-PTP (Mantel r = 0.52). 18. Genetic divergences among the candidate species for the sichelii-complex identified using Bayesian UF-PTP (Mantel r = 0.21). 19. Genetic divergences within the candidate species for the keriensis-complex identified using Bayesian UF-PTP (Mantel r = 0.16). 20. Genetic divergences among the candidate species for the keriensis-complex identified using Bayesian UF-PTP (relationship not positive). For details of the measurements see the" figureDoi="http://doi.org/10.5281/zenodo.4064342" httpUri="https://zenodo.org/record/4064342/files/figure.png" pageId="92" pageNumber="93">Fig. 20</figureCitation>
) (see Divergence and geographical distance, page 12).
</paragraph>
<paragraph id="AD3636DC1F1E9567FF3AF92CDFFBFD8B" blockId="92.[189,1399,1677,2020]" lastBlockId="94.[189,1399,265,608]" lastPageId="94" lastPageNumber="95" pageId="92" pageNumber="93">
From morphology, after examining&gt;1000 specimens of the
<taxonomicName id="6A894D5F1F1E9565FC03F92CDF96F94C" authorityName="F. Morawitz" authorityYear="1887" box="[900,1004,1677,1703]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="keriensis">
<emphasis id="9FFDEACE1F1E9565FC03F92CDF96F94C" box="[900,1004,1677,1703]" italics="true" pageId="92" pageNumber="93">keriensis</emphasis>
</taxonomicName>
-group from across Asia and after comparing the distribution of states of several characters as part of this study (PW), Vogts character state of exclusively black hair on the face does appear to remain one of the most consistently diagnostic characters (in comparison with COI sequences) for
<taxonomicName id="6A894D5F1F1E9565FCB0F956DF99F8FB" authorityName="Vogt" authorityYear="1909" box="[823,995,1782,1808]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="separandus">
<emphasis id="9FFDEACE1F1E9565FCB0F956DF99F8FB" box="[823,995,1782,1808]" italics="true" pageId="92" pageNumber="93">B. separandus</emphasis>
</taxonomicName>
in the QTP and Central Asia, at least for queens (but see the comments on
<taxonomicName id="6A894D5F1F1E9565FD34F8BBDF47F8D8" authorityName="F. Morawitz" authorityYear="1887" box="[691,829,1817,1843]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="keriensis">
<emphasis id="9FFDEACE1F1E9565FD34F8BBDF47F8D8" box="[691,829,1817,1843]" italics="true" pageId="92" pageNumber="93">B. keriensis</emphasis>
</taxonomicName>
). Nonetheless, there are some individuals of the yellow-banded taxon
<taxonomicName id="6A894D5F1F1E9565FE3EF89CDE76F8BC" authorityName="Skorikov" authorityYear="1910" box="[441,524,1853,1879]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="kozlovi">
<emphasis id="9FFDEACE1F1E9565FE3EF89CDE76F8BC" box="[441,524,1853,1879]" italics="true" pageId="92" pageNumber="93">kozlovi</emphasis>
</taxonomicName>
from
<collectingCountry id="D59E764C1F1E9565FDD5F89CDEBEF8BC" box="[594,708,1853,1879]" name="Mongolia" pageId="92" pageNumber="93">Mongolia</collectingCountry>
(part of
<taxonomicName id="6A894D5F1F1E9565FCA0F89FDFB4F8BC" authorityName="Vogt" authorityYear="1909" box="[807,974,1853,1879]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" sensu="lato" species="separandus">
<emphasis id="9FFDEACE1F1E9565FCA0F89FDFB4F8BC" box="[807,974,1853,1879]" italics="true" pageId="92" pageNumber="93">B. separandus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84CE57B51F1E9565FC53F89CD868F8BC" box="[980,1042,1853,1879]" pageId="92" pageNumber="93" sensu="lato">s. lat.</taxonomicNameLabel>
according our COI-coalescent results,
<figureCitation id="35B22A591F1E9565FE9EF8C1DD14F891" box="[281,366,1888,1914]" captionStart="Fig" captionStartId="21.[189,232,1953,1979]" captionTargetBox="[233,1316,282,1913]" captionTargetId="figure-12@21.[233,1355,265,1914]" captionTargetPageId="21" captionText="Fig. 10. October 2019 (Fig. 8) Bayesian PTP analysis of MrBayes tree of unique COI barcodes (UHF- PTP). Symbols as in Fig. 9." figureDoi="http://doi.org/10.5281/zenodo.4064332" httpUri="https://zenodo.org/record/4064332/files/figure.png" pageId="92" pageNumber="93">Fig. 10</figureCitation>
) that do have a few yellow hairs on the face, so that exclusively black hair on the face is not diagnostic in all parts of the range. Several aspects of the colour pattern vary strongly within the species: (1)
<taxonomicName id="6A894D5F1F1E9565FECFF806DD94F82B" authorityName="Vogt" authorityYear="1909" box="[328,494,1958,1984]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="separandus">
<emphasis id="9FFDEACE1F1E9565FECFF806DD94F82B" box="[328,494,1958,1984]" italics="true" pageId="92" pageNumber="93">B. separandus</emphasis>
</taxonomicName>
includes both yellow-banded and white-banded colour patterns; (2) the extent of the black hair between the wing bases varies; (3) the extent of the pale hair on the side and ventral area (leg bases) of the thorax varies; (4) the extent of the pale hair posteriorly on T3 varies (the ciliation of authors); (5) the contrast of the pale posterior fringes on T45 varies. Many of these characters might appear to show a continuum of variation that is also continuous with the variation within
<taxonomicName id="6A894D5F1F1C9567FB6BFEF1D90CFE82" authorityName="F. Morawitz" authorityYear="1887" box="[1260,1398,335,361]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" sensu="stricto" species="keriensis">
<emphasis id="9FFDEACE1F1C9567FB6BFEF1D90CFE82" box="[1260,1398,335,361]" italics="true" pageId="94" pageNumber="95">B. keriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84CE57B51F1C9567FF3AFED2DC86FE66" box="[189,252,371,397]" pageId="94" pageNumber="95" sensu="stricto">s. str.</taxonomicNameLabel>
(
<bibRefCitation id="C9184B2D1F1C9567FE8AFED3DDBAFE66" author="Williams P. H." box="[269,448,370,397]" pageId="94" pageNumber="95" pagination="1 - 204" refId="ref72395" refString="Williams P. H. 1991. The bumble bees of the Kashmir Himalaya (Hymenoptera: Apidae, Bombini). Bulletin of the British Museum (Natural History) (Entomology) 60: 1 - 204." type="journal article" year="1991">Williams 1991</bibRefCitation>
), which might then appear consistent with the two taxa being parts of the same species (
<bibRefCitation id="C9184B2D1F1C9567FEA7FE37DDCCFE5A" author="Reinig W. F." box="[288,438,406,433]" pageId="94" pageNumber="95" pagination="321 - 356" refId="ref69854" refString="Reinig W. F. 1935. On the variation of Bombus lapidarius L. and its cuckoo, Psithyrus rupestris Fabr., with notes on mimetic similarity. Journal of Genetics 30: 321 - 356. https: // doi. org / 10.1007 / BF 02982243" type="journal article" year="1935">Reinig 1935</bibRefCitation>
). However, as noted above, evidence from COI barcodes shows that the two genecoalescent groups differ most consistently in the absence of pale hairs on the face for
<taxonomicName id="6A894D5F1F1C9567FB42FE1BD915FE38" authorityName="Vogt" authorityYear="1909" box="[1221,1391,441,467]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="separandus">
<emphasis id="9FFDEACE1F1C9567FB42FE1BD915FE38" box="[1221,1391,441,467]" italics="true" pageId="94" pageNumber="95">B. separandus</emphasis>
</taxonomicName>
, supporting Vogts characterisation of the species, providing integrated morphological support for its status as a separate species. The overlapping colour variation of
<taxonomicName id="6A894D5F1F1C9567FC43FDA1D814FDF2" authorityName="Vogt" authorityYear="1909" box="[964,1134,511,537]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="separandus">
<emphasis id="9FFDEACE1F1C9567FC43FDA1D814FDF2" box="[964,1134,511,537]" italics="true" pageId="94" pageNumber="95">B. separandus</emphasis>
</taxonomicName>
but only in
<collectingCountry id="D59E764C1F1C9567FA83FE5ED90CFDF2" box="[1284,1398,511,537]" name="Mongolia" pageId="94" pageNumber="95">Mongolia</collectingCountry>
(where
<taxonomicName id="6A894D5F1F1C9567FE91FD82DDE4FDD7" authorityName="F. Morawitz" authorityYear="1887" box="[278,414,546,572]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" sensu="stricto" species="keriensis">
<emphasis id="9FFDEACE1F1C9567FE91FD82DDE4FDD7" box="[278,414,546,572]" italics="true" pageId="94" pageNumber="95">B. keriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84CE57B51F1C9567FE22FD82DD99FDD6" box="[421,483,547,573]" pageId="94" pageNumber="95" sensu="stricto">s. str.</taxonomicNameLabel>
does not occur) does not negate the fact that
<taxonomicName id="6A894D5F1F1C9567FC73FD82D8E1FDD7" authorityName="Vogt" authorityYear="1909" box="[1012,1179,546,572]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="separandus">
<emphasis id="9FFDEACE1F1C9567FC73FD82D8E1FDD7" box="[1012,1179,546,572]" italics="true" pageId="94" pageNumber="95">B. separandus</emphasis>
</taxonomicName>
is recognisable by morphological characters throughout its geographical range.
</paragraph>
<caption id="F9F666541F1F9564FF3AF8FDDFC3F80B" ID-DOI="http://doi.org/10.5281/zenodo.4064370" ID-Zenodo-Dep="4064370" httpUri="https://zenodo.org/record/4064370/files/figure.png" pageId="93" pageNumber="94" startId="93.[189,241,1884,1910]" targetBox="[190,1374,318,1788]" targetPageId="93">
<paragraph id="AD3636DC1F1F9564FF3AF8FDDFC3F80B" blockId="93.[189,1399,1884,2016]" pageId="93" pageNumber="94">
<emphasis id="9FFDEACE1F1F9564FF3AF8FDDD16F89D" bold="true" box="[189,364,1884,1910]" pageId="93" pageNumber="94">Figs 209210.</emphasis>
Images of queens of two cryptic species from around the western Qinghai-Tibetan plateau.
<emphasis id="9FFDEACE1F1F9564FEA3F8DEDD2AF872" bold="true" box="[292,336,1919,1945]" pageId="93" pageNumber="94">209</emphasis>
.
<taxonomicName id="6A894D5F1F1F9564FEE5F8DEDE87F872" authority="Morawitz, 1887" authorityName="F. Morawitz" authorityYear="1887" box="[354,765,1919,1945]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="keriensis">
<emphasis id="9FFDEACE1F1F9564FEE5F8DEDE49F872" box="[354,563,1919,1945]" italics="true" pageId="93" pageNumber="94">Bombus keriensis</emphasis>
<bibRefCitation id="C9184B2D1F1F9564FDB9F8DEDE87F872" author="Morawitz F. F." box="[574,765,1919,1945]" pageId="93" pageNumber="94" pagination="195 - 229" refId="ref68420" refString="Morawitz F. F. 1887. Insecta in itinere cl. N. Przewalskii in Asia centrali novissime lecta. I. Apidae. Trudy Russkago entomologicheskago obshchestva 20 (1886): 195 - 229." type="journal article" year="1887">Morawitz, 1887</bibRefCitation>
</taxonomicName>
from Mt Apharwat (4000 m a.s.l.) in the Pir Panjal mountains (ML405).
<emphasis id="9FFDEACE1F1F9564FE3FF803DD9EF857" bold="true" box="[440,484,1954,1980]" pageId="93" pageNumber="94">210</emphasis>
.
<taxonomicName id="6A894D5F1F1F9564FE75F802DF65F857" authority="Vogt, 1909" authorityName="Vogt" authorityYear="1909" box="[498,799,1954,1981]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="separandus" status="stat. nov.">
<emphasis id="9FFDEACE1F1F9564FE75F802DEE3F857" box="[498,665,1954,1980]" italics="true" pageId="93" pageNumber="94">B. separandus</emphasis>
<bibRefCitation id="C9184B2D1F1F9564FD18F802DF65F857" author="Vogt O." box="[671,799,1954,1981]" pageId="93" pageNumber="94" pagination="28 - 84" refId="ref71967" refString="Vogt O. 1909. Studien ʾ ber das Artproblem. 1. Mitteilung. Uber das Variieren der Hummeln. 1. Teil. Sitzungsberichte der Gesellschaft naturforschender Freunde zu Berlin 1909: 28 - 84." type="journal article" year="1909">Vogt, 1909</bibRefCitation>
</taxonomicName>
<taxonomicNameLabel id="84CE57B51F1F9564FCA1F802DFF2F856" box="[806,904,1955,1981]" pageId="93" pageNumber="94">stat. rev.</taxonomicNameLabel>
from Nimaling plain (4800 m a.s.l.) in the Zanskar mountains (ML311). Viewed from the left lateral aspect.
</paragraph>
</caption>
<paragraph id="AD3636DC1F1C9567FF3AFD26DE6AFBE9" blockId="94.[189,1399,647,1026]" pageId="94" pageNumber="95">
There are some queens of the yellow-banded taxon
<emphasis id="9FFDEACE1F1C9567FC99FD26DFF2FD4A" box="[798,904,647,673]" italics="true" pageId="94" pageNumber="95">richardsi</emphasis>
(later renamed
<taxonomicName id="6A894D5F1F1C9567FBC5FD26D8CBFD4A" authorityName="Wang" authorityYear="1982" box="[1090,1201,647,673]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="tibetensis">
<emphasis id="9FFDEACE1F1C9567FBC5FD26D8CBFD4A" box="[1090,1201,647,673]" italics="true" pageId="94" pageNumber="95">tibetensis</emphasis>
</taxonomicName>
) from the northwestern QTP that have no pale hairs on the face, very few or no yellow hairs on the leg bases, and very few black hairs within the yellow bands of the thoracic dorsum. Many of the available specimens have failed to sequence, so the species diagnosis remains tentative. Particularly influential in the interpretation of these individuals is one yellow-banded queen with a reduced extent of pale hair on the side of the thorax from the Zanskar range (PW: ML311,
<figureCitation id="35B22A591F1C9567FDFEFC96DEA5FCBA" box="[633,735,823,849]" captionStart="Figs 139180" captionStartId="42.[189,241,1497,1523]" captionTargetBox="[369,1221,268,1431]" captionTargetId="figure-11@42.[366,1221,265,1467]" captionTargetPageId="42" captionText="Figs 139180. Simplified diagrams for the colour patterns of the hair on the dorsum for the species from the integrative analysis. The dorsum is divided into regions, each of which shows only the predominant or most apparent colour for that region using a simplified colour palette, with olive indicating a mixture of black and yellow hair, and grey indicating a mixture of black and white hair. The keriensis-group. 139. Queen, China-Sichuan. 140. Worker, China-Sichuan. 141. Queen, China-Sichuan. 142. Queen, China-Sichuan. 143. Worker, China-Xizang. 144. Queen, India-Kashmir. 145. Male, China-Qinghai. 146. Male, India-Kashmir. 147. Male, China-Gansu. 148. Queen, Armenia. 149. Queen, Iran. 150. Male, Georgia. 151. Worker, China-Qinghai. 152. Male, China-Qinghai. 153. Queen, Mongolia. 154. Worker, Mongolia. 155. Male, Mongolia. 156. Male, Mongolia. 157. Queen, China-Sichuan. 158. Worker, China-Qinghai. 159. Worker, China-Qinghai. 160. Male, China-Qinghai. 161. Male, China-Qinghai. 162. Male, China-Qinghai. 163. Queen, Pakistan. 164. Queen, Pakistan. 165. Queen, Pakistan. 166. Queen, Pakistan. 167. Queen, Afghanistan. 168. Queen, Pakistan. 169. Queen, Pakistan. 170. Queen, India-Kashmir. 171. Male, India-Kashmir. 172. Male, India-Kashmir. 173. Queen, China- Xinjiang. 174. Queen, Kyrgyzstan. 175. Queen, Kyrgyzstan. 176. Queen, Mongolia. 177. Queen, India- Kashmir. 178. Male, Kazakhstan. 179. Male, China-Xinjiang. 180. Male, Mongolia." figureDoi="http://doi.org/10.5281/zenodo.4064358" httpUri="https://zenodo.org/record/4064358/files/figure.png" pageId="94" pageNumber="95">Figs 177</figureCitation>
,
<figureCitation id="35B22A591F1C9567FD6CFC96DF61FCBA" box="[747,795,823,849]" captionStart="Figs 209210" captionStartId="93.[189,241,1884,1910]" captionTargetBox="[190,1374,318,1788]" captionTargetId="figure-11@93.[189,1398,318,1792]" captionTargetPageId="93" captionText="Figs 209210. Images of queens of two cryptic species from around the western Qinghai-Tibetan plateau. 209. Bombus keriensis Morawitz, 1887 from Mt Apharwat (4000 m a.s.l.) in the Pir Panjal mountains (ML405). 210. B. separandus Vogt, 1909 stat. rev. from Nimaling plain (4800 m a.s.l.) in the Zanskar mountains (ML311). Viewed from the left lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.4064370" httpUri="https://zenodo.org/record/4064370/files/figure.png" pageId="94" pageNumber="95">210</figureCitation>
), which has produced a short sequence (382 bp) that nonetheless shows bases diagnostic for
<taxonomicName id="6A894D5F1F1C9567FD12FCFADF47FC9F" authorityName="Vogt" authorityYear="1909" box="[661,829,858,884]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="separandus">
<emphasis id="9FFDEACE1F1C9567FD12FCFADF47FC9F" box="[661,829,858,884]" italics="true" pageId="94" pageNumber="95">B. separandus</emphasis>
</taxonomicName>
compared to
<taxonomicName id="6A894D5F1F1C9567FC65FCFAD816FC9F" authorityName="F. Morawitz" authorityYear="1887" box="[994,1132,858,884]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="keriensis">
<emphasis id="9FFDEACE1F1C9567FC65FCFAD816FC9F" box="[994,1132,858,884]" italics="true" pageId="94" pageNumber="95">B. keriensis</emphasis>
</taxonomicName>
(base positions 267A, 286C, 301C, 306T, 339T, 363C, 423T, 498C, 529G, 540T, 541C, 556C). This is one of the specimens that most closely resemble some yellow-banded individuals of the candidate species
<taxonomicName id="6A894D5F1F1C9567FB11FC00D884FC50" authorityName="F. Morawitz" authorityYear="1887" box="[1174,1278,929,955]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" sensu="stricto" species="keriensis">
<emphasis id="9FFDEACE1F1C9567FB11FC00D884FC50" box="[1174,1278,929,955]" italics="true" pageId="94" pageNumber="95">keriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84CE57B51F1C9567FA82FC00D93BFC50" box="[1285,1345,929,955]" pageId="94" pageNumber="95" sensu="stricto">s. str.</taxonomicNameLabel>
also from Kashmir (
<figureCitation id="35B22A591F1C9567FEF5FC65DDA7FC35" box="[370,477,964,990]" captionStart="Figs 139180" captionStartId="42.[189,241,1497,1523]" captionTargetBox="[369,1221,268,1431]" captionTargetId="figure-11@42.[366,1221,265,1467]" captionTargetPageId="42" captionText="Figs 139180. Simplified diagrams for the colour patterns of the hair on the dorsum for the species from the integrative analysis. The dorsum is divided into regions, each of which shows only the predominant or most apparent colour for that region using a simplified colour palette, with olive indicating a mixture of black and yellow hair, and grey indicating a mixture of black and white hair. The keriensis-group. 139. Queen, China-Sichuan. 140. Worker, China-Sichuan. 141. Queen, China-Sichuan. 142. Queen, China-Sichuan. 143. Worker, China-Xizang. 144. Queen, India-Kashmir. 145. Male, China-Qinghai. 146. Male, India-Kashmir. 147. Male, China-Gansu. 148. Queen, Armenia. 149. Queen, Iran. 150. Male, Georgia. 151. Worker, China-Qinghai. 152. Male, China-Qinghai. 153. Queen, Mongolia. 154. Worker, Mongolia. 155. Male, Mongolia. 156. Male, Mongolia. 157. Queen, China-Sichuan. 158. Worker, China-Qinghai. 159. Worker, China-Qinghai. 160. Male, China-Qinghai. 161. Male, China-Qinghai. 162. Male, China-Qinghai. 163. Queen, Pakistan. 164. Queen, Pakistan. 165. Queen, Pakistan. 166. Queen, Pakistan. 167. Queen, Afghanistan. 168. Queen, Pakistan. 169. Queen, Pakistan. 170. Queen, India-Kashmir. 171. Male, India-Kashmir. 172. Male, India-Kashmir. 173. Queen, China- Xinjiang. 174. Queen, Kyrgyzstan. 175. Queen, Kyrgyzstan. 176. Queen, Mongolia. 177. Queen, India- Kashmir. 178. Male, Kazakhstan. 179. Male, China-Xinjiang. 180. Male, Mongolia." figureDoi="http://doi.org/10.5281/zenodo.4064358" httpUri="https://zenodo.org/record/4064358/files/figure.png" pageId="94" pageNumber="95">Figs 170</figureCitation>
,
<figureCitation id="35B22A591F1C9567FE6FFC65DE6CFC35" box="[488,534,964,990]" captionStart="Figs 209210" captionStartId="93.[189,241,1884,1910]" captionTargetBox="[190,1374,318,1788]" captionTargetId="figure-11@93.[189,1398,318,1792]" captionTargetPageId="93" captionText="Figs 209210. Images of queens of two cryptic species from around the western Qinghai-Tibetan plateau. 209. Bombus keriensis Morawitz, 1887 from Mt Apharwat (4000 m a.s.l.) in the Pir Panjal mountains (ML405). 210. B. separandus Vogt, 1909 stat. rev. from Nimaling plain (4800 m a.s.l.) in the Zanskar mountains (ML311). Viewed from the left lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.4064370" httpUri="https://zenodo.org/record/4064370/files/figure.png" pageId="94" pageNumber="95">209</figureCitation>
) (the difference in the breadth of the black band between the wings is not diagnostic for these species).
</paragraph>
<paragraph id="AD3636DC1F1C9567FF3AFB88DEE9FB24" blockId="94.[189,1399,1065,1232]" pageId="94" pageNumber="95">
<taxonomicName id="6A894D5F1F1C9567FF3AFB88DDD2FBA8" authorityName="Vogt" authorityYear="1909" box="[189,424,1065,1091]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="separandus">
<emphasis id="9FFDEACE1F1C9567FF3AFB88DDD2FBA8" box="[189,424,1065,1091]" italics="true" pageId="94" pageNumber="95">Bombus separandus</emphasis>
</taxonomicName>
co-occurs locally with the similar
<taxonomicName id="6A894D5F1F1C9567FCB0FB88DFA2FBA8" authorityName="Vogt" authorityYear="1911" baseAuthorityName="Vogt" baseAuthorityYear="1911" box="[823,984,1065,1091]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="incertoides">
<emphasis id="9FFDEACE1F1C9567FCB0FB88DFA2FBA8" box="[823,984,1065,1091]" italics="true" pageId="94" pageNumber="95">B. incertoides</emphasis>
</taxonomicName>
in the Altai and Khangai mountains of
<collectingCountry id="D59E764C1F1C9567FF5CFBEDDD2BFB8D" box="[219,337,1100,1126]" name="Mongolia" pageId="94" pageNumber="95">Mongolia</collectingCountry>
.
<taxonomicName id="6A894D5F1F1C9567FEDDFBEDDE3FFB8D" authorityName="Vogt" authorityYear="1909" box="[346,581,1100,1126]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="separandus">
<emphasis id="9FFDEACE1F1C9567FEDDFBEDDE3FFB8D" box="[346,581,1100,1126]" italics="true" pageId="94" pageNumber="95">Bombus separandus</emphasis>
</taxonomicName>
also co-occurs locally with the similar
<taxonomicName id="6A894D5F1F1C9567FB82FBECD8F6FB8D" authorityName="F. Morawitz" authorityYear="1887" box="[1029,1164,1100,1126]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" sensu="stricto" species="keriensis">
<emphasis id="9FFDEACE1F1C9567FB82FBECD8F6FB8D" box="[1029,1164,1100,1126]" italics="true" pageId="94" pageNumber="95">B. keriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84CE57B51F1C9567FB15FBEDD8B7FB8D" box="[1170,1229,1100,1126]" pageId="94" pageNumber="95" sensu="stricto">s. str.</taxonomicNameLabel>
in the Zanskar mountains of Kashmir. Examination of specimens in the IOZ collection labelled
<taxonomicName id="6A894D5F1F1C9567FBE0FBD1D89CFB62" authorityName="Morawitz" authorityYear="1881" box="[1127,1254,1136,1161]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="incertus">
<emphasis id="9FFDEACE1F1C9567FBE0FBD1D89CFB62" box="[1127,1254,1136,1161]" italics="true" pageId="94" pageNumber="95">B. incertus</emphasis>
</taxonomicName>
(S.-F.
<bibRefCitation id="C9184B2D1F1C9567FAB4FBD1DC81FB46" author="Wang S. - F." pageId="94" pageNumber="95" pagination="160 - 165" refId="ref72240" refString="Wang S. - F. 1985. Apidae - Bombus. In: [Organisms of the Tumuefeng region of Tianshan]: 160 - 165." type="book chapter" year="1985">Wang 1985</bibRefCitation>
; S.-F.
<bibRefCitation id="C9184B2D1F1C9567FEC3FB32DE67FB47" author="Wang S. - F. &amp; Yao J." box="[324,541,1170,1197]" pageId="94" pageNumber="95" pagination="303 - 309" refId="ref72268" refString="Wang S. - F. &amp; Yao J. 1996. Hymenoptera: Apodae [sic] - Bombini. In: Huang F. - S. (ed.) Insects of the Karakorum-Kunlun Mountains: 303 - 309. Science Press, Beijing." type="book chapter" year="1996">Wang &amp; Yao 1996</bibRefCitation>
;
<bibRefCitation id="C9184B2D1F1C9567FDAEFB33DEA3FB47" author="Niu Z. - Q. &amp; Yuan F. &amp; Zhu C. - D." box="[553,729,1170,1196]" pageId="94" pageNumber="95" refId="ref68540" refString="Niu Z. - Q., Yuan F. &amp; Zhu C. - D. 2018. Species Catalogue of China. Volume 2 Animals. Insecta (IV). Apoidea (Apidae, Melittidae, Halictidae). Science Press, Beijing." type="book" year="2018">
Niu
<emphasis id="9FFDEACE1F1C9567FDDBFB32DEECFB47" box="[604,662,1170,1196]" italics="true" pageId="94" pageNumber="95">et al.</emphasis>
2018
</bibRefCitation>
) shows that these are mostly misidentified
<taxonomicName id="6A894D5F1F1C9567FB48FB32D90CFB47" authorityName="Vogt" authorityYear="1909" box="[1231,1398,1170,1196]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="separandus">
<emphasis id="9FFDEACE1F1C9567FB48FB32D90CFB47" box="[1231,1398,1170,1196]" italics="true" pageId="94" pageNumber="95">B. separandus</emphasis>
</taxonomicName>
(with a few
<taxonomicName id="6A894D5F1F1C9567FECDFB17DDA8FB24" authorityName="F. Morawitz" authorityYear="1887" box="[330,466,1205,1231]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="keriensis">
<emphasis id="9FFDEACE1F1C9567FECDFB17DDA8FB24" box="[330,466,1205,1231]" italics="true" pageId="94" pageNumber="95">B. keriensis</emphasis>
</taxonomicName>
) from Xinjiang.
</paragraph>
</subSubSection>
<subSubSection id="E59365571F1C9567FF3AFB56D88AFA26" pageId="94" pageNumber="95" type="diagnosis">
<paragraph id="AD3636DC1F1C9567FF3AFB56DD4CFAFA" blockId="94.[189,310,1271,1297]" box="[189,310,1271,1297]" pageId="94" pageNumber="95">
<heading id="F67E81B01F1C9567FF3AFB56DD4CFAFA" bold="true" box="[189,310,1271,1297]" fontSize="11" level="3" pageId="94" pageNumber="95" reason="3">
<emphasis id="9FFDEACE1F1C9567FF3AFB56DD4CFAFA" bold="true" box="[189,310,1271,1297]" pageId="94" pageNumber="95">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="AD3636DC1F1C9567FF6DFA87DD2BFAAB" blockId="94.[189,1399,1318,1485]" box="[234,337,1318,1344]" pageId="94" pageNumber="95">
<emphasis id="9FFDEACE1F1C9567FF6DFA87DD2BFAAB" bold="true" box="[234,337,1318,1344]" pageId="94" pageNumber="95">Females</emphasis>
</paragraph>
<paragraph id="AD3636DC1F1C9567FF3AFAE8D88AFA26" blockId="94.[189,1399,1318,1485]" pageId="94" pageNumber="95">
Queens medium-sized body length
<quantity id="6A719B391F1C9567FDDCFAE8DEA7FA88" box="[603,733,1353,1379]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="2.0" metricValueMin="1.7" pageId="94" pageNumber="95" unit="mm" value="18.5" valueMax="20.0" valueMin="17.0">1720 mm</quantity>
, workers
<quantity id="6A719B391F1C9567FCCAFAE8DFBAFA88" box="[845,960,1353,1379]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.4" metricValueMin="0.9" pageId="94" pageNumber="95" unit="mm" value="11.5" valueMax="14.0" valueMin="9.0">914 mm</quantity>
. Can be distinguished in Central Asia and
<collectingCountry id="D59E764C1F1C9567FF76FACDDD19FA6D" box="[241,355,1388,1414]" name="Mongolia" pageId="94" pageNumber="95">Mongolia</collectingCountry>
by the combination of hair of the face usually without pale hairs (except in
<collectingCountry id="D59E764C1F1C9567FA83FACDD90CFA6D" box="[1284,1398,1388,1414]" name="Mongolia" pageId="94" pageNumber="95">Mongolia</collectingCountry>
where
<taxonomicName id="6A894D5F1F1C9567FE8BFA31DDEEFA42" authorityName="F. Morawitz" authorityYear="1887" box="[268,404,1423,1449]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="keriensis">
<emphasis id="9FFDEACE1F1C9567FE8BFA31DDEEFA42" box="[268,404,1423,1449]" italics="true" pageId="94" pageNumber="95">B. keriensis</emphasis>
</taxonomicName>
does not occur), the leg bases usually without pale hairs, T2 posteriorly entirely pale without black hairs, and T3 laterally usually without pale hairs (cf.
<taxonomicName id="6A894D5F1F1C9567FC55FA12D83DFA27" authorityName="Radoszkowski" authorityYear="1860" box="[978,1095,1458,1484]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="sichelii">
<emphasis id="9FFDEACE1F1C9567FC55FA12D83DFA27" box="[978,1095,1458,1484]" italics="true" pageId="94" pageNumber="95">B. sichelii</emphasis>
</taxonomicName>
,
<taxonomicName id="6A894D5F1F1C9567FBD1FA12D8A4FA27" authorityName="F. Morawitz" authorityYear="1887" box="[1110,1246,1458,1484]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="keriensis">
<emphasis id="9FFDEACE1F1C9567FBD1FA12D8A4FA27" box="[1110,1246,1458,1484]" italics="true" pageId="94" pageNumber="95">B. keriensis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="E59365571F1C9567FF6DFA55DF06F90A" pageId="94" pageNumber="95" type="description">
<paragraph id="AD3636DC1F1C9567FF6DFA55DD4FF9E5" blockId="94.[189,1399,1524,1761]" box="[234,309,1524,1550]" pageId="94" pageNumber="95">
<emphasis id="9FFDEACE1F1C9567FF6DFA55DD4FF9E5" bold="true" box="[234,309,1524,1550]" pageId="94" pageNumber="95">Males</emphasis>
</paragraph>
<paragraph id="AD3636DC1F1C9567FF3AF9B6DF06F90A" blockId="94.[189,1399,1524,1761]" pageId="94" pageNumber="95">
Body length
<quantity id="6A719B391F1C9567FED2F9B6DDADF9DA" box="[341,471,1559,1586]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="94" pageNumber="95" unit="mm" value="12.5" valueMax="15.0" valueMin="10.0">1015 mm</quantity>
. Can be distinguished reliably at present only by their COI sequence, although in the western Himalaya they may be distinguished by entirely black hair between the wing bases and hair of T3 entirely black. Genitalia (
<figureCitation id="35B22A591F1C9567FDF3F9FCDEA1F99C" box="[628,731,1629,1655]" captionStart="Figs 199204" captionStartId="45.[189,241,1852,1878]" captionTargetBox="[230,1342,827,1806]" captionTargetId="figure-159@45.[194,1393,771,1810]" captionTargetPageId="45" captionText="Figs 199204. Morphology of the male genitalia for species of the subgenus Melanobombus von Dalla Torre, 1880 from the dorsal aspect, anterior at the bottom of the image, posterior at the top. 199. Bombus alagesianus Reinig, 1930 stat. rev., Georgia. 200. B. tibeticus sp. nov., China-Qinghai. 201. B. incertoides Vogt, 1911 stat. rev., Mongolia. 202. B. qilianensis sp. nov., China-Qinghai. 203. B. keriensis Morawitz, 1887, India-Kashmir. 204. B. separandus Vogt, 1909 stat. rev., Kyrgyzstan. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4064364" httpUri="https://zenodo.org/record/4064364/files/figure.png" pageId="94" pageNumber="95">Fig. 204</figureCitation>
) with the gonostylus shorter than broad, its inner basal projection reduced to a short stub (cf.
<emphasis id="9FFDEACE1F1C9567FDFAF920DEADF970" box="[637,727,1665,1691]" italics="true" pageId="94" pageNumber="95">rufipes-</emphasis>
group,
<emphasis id="9FFDEACE1F1C9567FCADF920DFF6F970" box="[810,908,1665,1691]" italics="true" pageId="94" pageNumber="95">festivus-</emphasis>
group,
<taxonomicName id="6A894D5F1F1C9567FC58F920D802F970" authorityName="Smith" authorityYear="1852" box="[991,1144,1665,1691]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="rufofasciatus">
<emphasis id="9FFDEACE1F1C9567FC58F920D802F970" box="[991,1144,1665,1691]" italics="true" pageId="94" pageNumber="95">rufofasciatus</emphasis>
</taxonomicName>
-group); volsella with the inner distal corner broadly produced but without a narrow hook (cf.
<emphasis id="9FFDEACE1F1C9567FB9CF905D80FF955" box="[1051,1141,1700,1726]" italics="true" pageId="94" pageNumber="95">rufipes-</emphasis>
group,
<emphasis id="9FFDEACE1F1C9567FB4EF905D951F955" box="[1225,1323,1700,1726]" italics="true" pageId="94" pageNumber="95">festivus-</emphasis>
group,
<taxonomicName id="6A894D5F1F1C9567FF3AF966DD2CF90A" authorityName="Smith" authorityYear="1852" box="[189,342,1735,1761]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="rufofasciatus">
<emphasis id="9FFDEACE1F1C9567FF3AF966DD2CF90A" box="[189,342,1735,1761]" italics="true" pageId="94" pageNumber="95">rufofasciatus</emphasis>
</taxonomicName>
-group); eye unenlarged relative to female eye.
</paragraph>
</subSubSection>
<subSubSection id="E59365571F1C9566FF3AF8A9DF52F9BF" lastPageId="95" lastPageNumber="96" pageId="94" pageNumber="95" type="materials_examined">
<paragraph id="AD3636DC1F1C9567FF3AF8A9DDD0F8C9" blockId="94.[189,426,1800,1826]" box="[189,426,1800,1826]" pageId="94" pageNumber="95">
<heading id="F67E81B01F1C9567FF3AF8A9DDD0F8C9" bold="true" box="[189,426,1800,1826]" fontSize="11" level="3" pageId="94" pageNumber="95" reason="3">
<emphasis id="9FFDEACE1F1C9567FF3AF8A9DDD0F8C9" bold="true" box="[189,426,1800,1826]" pageId="94" pageNumber="95">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="AD3636DC1F1C9567FF6DF896DD1CF8BA" blockId="94.[189,1398,1847,1944]" box="[234,358,1847,1873]" pageId="94" pageNumber="95">
<emphasis id="9FFDEACE1F1C9567FF6DF896DD1CF8BA" bold="true" box="[234,358,1847,1873]" pageId="94" pageNumber="95">
<typeStatus id="7232887E1F1C9567FF6DF896DD1CF8BA" box="[234,358,1847,1873]" pageId="94" pageNumber="95" type="lectotype">Lectotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="AD3636DC1F1C9567FF3AF8FBD8FCF873" blockId="94.[189,1398,1847,1944]" pageId="94" pageNumber="95">
<collectingCountry id="D59E764C1F1C9567FF3AF8FBDD62F89F" box="[189,280,1882,1908]" name="China" pageId="94" pageNumber="95">CHINA</collectingCountry>
<specimenCount id="BB8FFD551F1C9567FEB1F8FADD36F89E" box="[310,332,1883,1909]" pageId="94" pageNumber="95" type="female"></specimenCount>
(queen),
<typeStatus id="7232887E1F1C9567FE47F8FBDE56F89F" box="[448,556,1882,1908]" pageId="94" pageNumber="95" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="6A894D5F1F1C9567FDD0F8FBDF42F89F" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[599,824,1882,1908]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="lapidarius">
<emphasis id="9FFDEACE1F1C9567FDD0F8FBDF42F89F" box="[599,824,1882,1908]" italics="true" pageId="94" pageNumber="95">Bombus lapidarius</emphasis>
</taxonomicName>
Form
<taxonomicName id="6A894D5F1F1C9567FC0DF8FBD8E7F89F" authority="Vogt, 1909" authorityName="Vogt" authorityYear="1909" box="[906,1181,1882,1909]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="separandus">
<emphasis id="9FFDEACE1F1C9567FC0DF8FBD86BF89F" box="[906,1041,1882,1908]" italics="true" pageId="94" pageNumber="95">separandus</emphasis>
<bibRefCitation id="C9184B2D1F1C9567FB9DF8FAD8E7F89F" author="Vogt O." box="[1050,1181,1882,1909]" pageId="94" pageNumber="95" pagination="28 - 84" refId="ref71967" refString="Vogt O. 1909. Studien ʾ ber das Artproblem. 1. Mitteilung. Uber das Variieren der Hummeln. 1. Teil. Sitzungsberichte der Gesellschaft naturforschender Freunde zu Berlin 1909: 28 - 84." type="journal article" year="1909">Vogt, 1909</bibRefCitation>
</taxonomicName>
by designation of
<bibRefCitation id="C9184B2D1F1C9567FF3AF8DCDDD7F87C" author="Williams P. H." box="[189,429,1917,1943]" pageId="94" pageNumber="95" pagination="1 - 204" refId="ref72395" refString="Williams P. H. 1991. The bumble bees of the Kashmir Himalaya (Hymenoptera: Apidae, Bombini). Bulletin of the British Museum (Natural History) (Entomology) 60: 1 - 204." type="journal article" year="1991">Williams (1991: 97)</bibRefCitation>
;
<collectingRegion id="6F4DF83E1F1C9567FE3EF8DCDE59F87C" box="[441,547,1917,1943]" country="China" name="Xinjiang Uygur" pageId="94" pageNumber="95">Xinjiang</collectingRegion>
, Borohoro Shan, Tisilikau;
<collectionCode id="CB98AE191F1C9567FCE1F8DFDFC4F873" box="[870,958,1918,1944]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="94" pageNumber="95" type="Museum">RMNH</collectionCode>
(examined
<collectionCode id="CB98AE191F1C9567FBCFF8DFD803F873" box="[1096,1145,1918,1944]" country="Thailand" name="Paleontological Collections" pageId="94" pageNumber="95">PW</collectionCode>
).
</paragraph>
<paragraph id="AD3636DC1F1D9566FF3AFEA8DE16FECF" blockId="95.[189,620,265,292]" box="[189,620,265,292]" pageId="95" pageNumber="96">
<emphasis id="9FFDEACE1F1D9566FF3AFEA8DDC8FEC8" bold="true" box="[189,434,265,291]" pageId="95" pageNumber="96">Material sequenced</emphasis>
(
<specimenCount id="BB8FFD551F1D9566FE45FEA8DE1EFECF" box="[450,612,265,292]" pageId="95" pageNumber="96" type="generic">25 specimens</specimenCount>
)
</paragraph>
<paragraph id="AD3636DC1F1D9566FF3AFE99DE13FDCE" blockId="95.[189,1399,311,549]" pageId="95" pageNumber="96">
<materialsCitation id="1DE13C811F1D9566FF3AFE99DD34FE9E" ID-GBIF-Occurrence="2867606355" collectingDate="2013-06-22" collectionCode="NHMUK, PW" country="Mongolia" latitude="47.4257" location="Bulgan" longLatPrecision="6" longitude="103.694" pageId="95" pageNumber="96" specimenCode="ML147" specimenCount="1" specimenCount-female="1" stateProvince="Bulgan">
<collectingCountry id="D59E764C1F1D9566FF3AFE99DD21FEB9" box="[189,347,312,338]" name="Mongolia" pageId="95" pageNumber="96">MONGOLIA</collectingCountry>
<specimenCount id="BB8FFD551F1D9566FEF4FE99DDE4FEB9" box="[371,414,312,338]" pageId="95" pageNumber="96" type="female">1 ♀</specimenCount>
(queen);
<collectingRegion id="6F4DF83E1F1D9566FD8AFE96DE1FFEBA" box="[525,613,311,337]" country="Mongolia" name="Bulgan" pageId="95" pageNumber="96">Bulgan</collectingRegion>
;
<geoCoordinate id="C8BD501B1F1D9566FDF6FE96DE81FEB9" box="[625,763,311,338]" degrees="47.4257" direction="north" orientation="latitude" pageId="95" pageNumber="96" precision="5" value="47.4257">47.4257° N</geoCoordinate>
,
<geoCoordinate id="C8BD501B1F1D9566FC81FE96DFE1FEB9" box="[774,923,311,338]" degrees="103.6940" direction="east" orientation="longitude" pageId="95" pageNumber="96" precision="5" value="103.694">103.6940° E</geoCoordinate>
;
<date id="D937101C1F1D9566FC20FE99D83AFEB9" box="[935,1088,312,338]" pageId="95" pageNumber="96" value="2013-06-22">
<collectingDate id="C973E9F41F1D9566FC20FE99D83AFEB9" box="[935,1088,312,338]" pageId="95" pageNumber="96" value="2013-06-22">22 Jun. 2013</collectingDate>
</date>
;
<collectionCode id="CB98AE191F1D9566FBCBFE99D8C1FEB9" box="[1100,1211,312,338]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="95" pageNumber="96" type="Museum">NHMUK</collectionCode>
seq: NHMSR1;
<collectionCode id="CB98AE191F1D9566FF3AFEFADC97FE9E" box="[189,237,347,373]" country="Thailand" name="Paleontological Collections" pageId="95" pageNumber="96">PW</collectionCode>
:
<specimenCode id="FD2F9EA71F1D9566FF71FEFADD34FE9E" box="[246,334,347,373]" collectionCode="PW" country="Thailand" name="Paleontological Collections" pageId="95" pageNumber="96">ML147</specimenCode>
</materialsCitation>
<materialsCitation id="1DE13C811F1D9566FEE4FEFADE29FE72" ID-GBIF-Occurrence="2867606361" collectingDate="1992-07-26" collectionCode="PW" collectorName="Culverwell" country="Mongolia" latitude="46.9333" location="Bayanolgiy" longLatPrecision="6" longitude="91.1667" pageId="95" pageNumber="96" specimenCode="ML151" specimenCount="1" specimenCount-female="1" stateProvince="Bulgan">
<specimenCount id="BB8FFD551F1D9566FEE4FEFADDF4FE9E" box="[355,398,347,373]" pageId="95" pageNumber="96" type="female">1 ♀</specimenCount>
(worker);
<location id="A85660071F1D9566FD8FFEFADEEAFE9E" LSID="urn:lsid:plazi:treatment:252087CA1F1E9566FDA0FE2ED89AF80E:A85660071F1D9566FD8FFEFADEEAFE9E" box="[520,656,347,373]" country="Mongolia" latitude="46.9333" longLatPrecision="6" longitude="91.1667" name="Bayanolgiy" pageId="95" pageNumber="96" stateProvince="Bulgan">Bayanolgiy</location>
,
<collectingRegion id="6F4DF83E1F1D9566FD1DFEFADE88FE9E" box="[666,754,347,373]" country="Mongolia" name="Bulgan" pageId="95" pageNumber="96">Bulgan</collectingRegion>
;
<geoCoordinate id="C8BD501B1F1D9566FD7AFEFADFFCFE9E" box="[765,902,347,373]" degrees="46.9333" direction="north" orientation="latitude" pageId="95" pageNumber="96" precision="5" value="46.9333">46.9333° N</geoCoordinate>
,
<geoCoordinate id="C8BD501B1F1D9566FC17FEFAD86CFE9E" box="[912,1046,347,373]" degrees="91.1667" direction="east" orientation="longitude" pageId="95" pageNumber="96" precision="5" value="91.1667">91.1667° E</geoCoordinate>
;
<date id="D937101C1F1D9566FBA7FEFAD8CBFE9E" box="[1056,1201,347,373]" pageId="95" pageNumber="96" value="1992-07-26">
<collectingDate id="C973E9F41F1D9566FBA7FEFAD8CBFE9E" box="[1056,1201,347,373]" pageId="95" pageNumber="96" value="1992-07-26">26 Jul. 1992</collectingDate>
</date>
;
<collectorName id="007C530A1F1D9566FB3CFEFAD947FE9E" box="[1211,1341,347,373]" pageId="95" pageNumber="96">Culverwell</collectorName>
leg.; BOLD seq: 6877B11;
<collectionCode id="CB98AE191F1D9566FE46FEDFDD8BFE73" box="[449,497,382,408]" country="Thailand" name="Paleontological Collections" pageId="95" pageNumber="96">PW</collectionCode>
:
<specimenCode id="FD2F9EA71F1D9566FE7CFEDFDE29FE72" box="[507,595,382,409]" collectionCode="PW" country="Thailand" name="Paleontological Collections" pageId="95" pageNumber="96">ML151</specimenCode>
</materialsCitation>
<materialsCitation id="1DE13C811F1D9566FDEEFEDFDFDCFE50" ID-GBIF-Occurrence="2867606519" collectingDate="2015-06-16" collectorName="R. DeJonghe" country="Mongolia" latitude="47.9905" location="Terelj" longLatPrecision="6" longitude="107.3719" municipality="Khenteyn Nuruu" pageId="95" pageNumber="96" specimenCount="1" specimenCount-female="1">
<specimenCount id="BB8FFD551F1D9566FDEEFEDFDEEEFE73" box="[617,660,382,408]" pageId="95" pageNumber="96" type="female">1 ♀</specimenCount>
(queen);
<collectingMunicipality id="4D52ACA61F1D9566FC85FEDFDFB3FE73" box="[770,969,382,408]" pageId="95" pageNumber="96">Khenteyn Nuruu</collectingMunicipality>
,
<location id="A85660071F1D9566FC54FEDFD861FE73" LSID="urn:lsid:plazi:treatment:252087CA1F1E9566FDA0FE2ED89AF80E:A85660071F1D9566FC54FEDFD861FE73" box="[979,1051,382,408]" country="Mongolia" latitude="47.9905" longLatPrecision="6" longitude="107.3719" municipality="Khenteyn Nuruu" name="Terelj" pageId="95" pageNumber="96">Terelj</location>
;
<geoCoordinate id="C8BD501B1F1D9566FBA2FEDFD8D4FE73" box="[1061,1198,382,409]" degrees="47.9905" direction="north" orientation="latitude" pageId="95" pageNumber="96" precision="5" value="47.9905">47.9905° N</geoCoordinate>
,
<geoCoordinate id="C8BD501B1F1D9566FB3EFEDFD934FE73" box="[1209,1358,382,408]" degrees="107.3719" direction="east" orientation="longitude" pageId="95" pageNumber="96" precision="5" value="107.3719">107.3719° E</geoCoordinate>
;
<date id="D937101C1F1D9566FADEFEDFDD48FE57" pageId="95" pageNumber="96" value="2015-06-16">
<collectingDate id="C973E9F41F1D9566FADEFEDFDD48FE57" pageId="95" pageNumber="96" value="2015-06-16">16 Jun. 2015</collectingDate>
</date>
;
<collectorName id="007C530A1F1D9566FEB9FE03DDAFFE50" box="[318,469,417,444]" pageId="95" pageNumber="96">R. DeJonghe</collectorName>
leg.; BOLD seq: 6877A08; RJ: ML333
</materialsCitation>
<materialsCitation id="1DE13C811F1D9566FC3AFE00D954FE35" ID-GBIF-Occurrence="2867606576" collectingDate="2015-06-17" collectionCode="RDJ" collectorName="R. DeJonghe" country="Mongolia" latitude="47.9956" location="Terelj" longLatPrecision="6" longitude="107.3303" municipality="Khenteyn Nuruu" pageId="95" pageNumber="96" specimenCode="ML489" specimenCount="1" specimenCount-female="1">
<specimenCount id="BB8FFD551F1D9566FC3AFE00DF93FE57" box="[957,1001,417,444]" pageId="95" pageNumber="96" type="female">1 ♀</specimenCount>
(queen);
<collectingMunicipality id="4D52ACA61F1D9566FBDEFE00D95AFE57" box="[1113,1312,417,444]" pageId="95" pageNumber="96">Khenteyn Nuruu</collectingMunicipality>
,
<location id="A85660071F1D9566FAADFE00D908FE50" LSID="urn:lsid:plazi:treatment:252087CA1F1E9566FDA0FE2ED89AF80E:A85660071F1D9566FAADFE00D908FE50" box="[1322,1394,417,443]" country="Mongolia" latitude="47.9956" longLatPrecision="6" longitude="107.3303" municipality="Khenteyn Nuruu" name="Terelj" pageId="95" pageNumber="96">Terelj</location>
;
<geoCoordinate id="C8BD501B1F1D9566FF3AFE65DD33FE34" box="[189,329,452,479]" degrees="47.9956" direction="north" orientation="latitude" pageId="95" pageNumber="96" precision="5" value="47.9956">47.9956° N</geoCoordinate>
,
<geoCoordinate id="C8BD501B1F1D9566FED1FE65DD97FE34" box="[342,493,452,479]" degrees="107.3303" direction="east" orientation="longitude" pageId="95" pageNumber="96" precision="5" value="107.3303">107.3303° E</geoCoordinate>
;
<date id="D937101C1F1D9566FE7CFE65DEE2FE34" box="[507,664,452,479]" pageId="95" pageNumber="96" value="2015-06-17">
<collectingDate id="C973E9F41F1D9566FE7CFE65DEE2FE34" box="[507,664,452,479]" pageId="95" pageNumber="96" value="2015-06-17">17 Jun. 2015</collectingDate>
</date>
;
<collectorName id="007C530A1F1D9566FD21FE64DF44FE35" box="[678,830,452,479]" pageId="95" pageNumber="96">R. DeJonghe</collectorName>
leg.; BOLD seq: 6880B05;
<collectionCode id="CB98AE191F1D9566FB17FE64D8B3FE34" box="[1168,1225,453,479]" pageId="95" pageNumber="96">RDJ</collectionCode>
:
<specimenCode id="FD2F9EA71F1D9566FB52FE65D954FE35" box="[1237,1326,452,478]" collectionCode="RDJ" pageId="95" pageNumber="96">ML489</specimenCode>
</materialsCitation>
<materialsCitation id="1DE13C811F1D9566FACEFE65DE1FFDCE" ID-GBIF-Occurrence="2867606403" collectingDate="2018-08-11" collectionCode="RDJ" collectorName="R. DeJonghe" country="Mongolia" latitude="47.575" location="Chuluut Khujirt Mts" longLatPrecision="6" longitude="101.1092" pageId="95" pageNumber="96" specimenCode="ML497" specimenCount="1" specimenCount-female="1" stateProvince="Arhangay">
<specimenCount id="BB8FFD551F1D9566FACEFE65D90CFE34" box="[1353,1398,452,479]" pageId="95" pageNumber="96" type="female">1 ♀</specimenCount>
(worker);
<collectingRegion id="6F4DF83E1F1D9566FEB5FE46DDCBFDEA" box="[306,433,487,513]" country="Mongolia" name="Arhangay" pageId="95" pageNumber="96">Arkhangai</collectingRegion>
,
<location id="A85660071F1D9566FE3BFE46DECAFDE9" LSID="urn:lsid:plazi:treatment:252087CA1F1E9566FDA0FE2ED89AF80E:A85660071F1D9566FE3BFE46DECAFDE9" box="[444,688,487,514]" country="Mongolia" latitude="47.575" longLatPrecision="6" longitude="101.1092" name="Chuluut Khujirt Mts" pageId="95" pageNumber="96" stateProvince="Arhangay">Chuluut Khujirt Mts</location>
;
<geoCoordinate id="C8BD501B1F1D9566FD3BFE46DF3DFDE9" box="[700,839,487,514]" degrees="47.5750" direction="north" orientation="latitude" pageId="95" pageNumber="96" precision="5" value="47.575">47.5750° N</geoCoordinate>
,
<geoCoordinate id="C8BD501B1F1D9566FCD4FE49DF93FDE9" box="[851,1001,488,514]" degrees="101.1092" direction="east" orientation="longitude" pageId="95" pageNumber="96" precision="5" value="101.1092">101.1092° E</geoCoordinate>
;
<date id="D937101C1F1D9566FC71FE49D8EDFDE9" box="[1014,1175,488,514]" pageId="95" pageNumber="96" value="2018-08-11">
<collectingDate id="C973E9F41F1D9566FC71FE49D8EDFDE9" box="[1014,1175,488,514]" pageId="95" pageNumber="96" value="2018-08-11">11 Aug. 2018</collectingDate>
</date>
;
<collectorName id="007C530A1F1D9566FB23FE49D941FDEA" box="[1188,1339,487,514]" pageId="95" pageNumber="96">R. DeJonghe</collectorName>
leg.; BOLD seq: 6880B12;
<collectionCode id="CB98AE191F1D9566FE41FDAADD85FDCE" box="[454,511,523,549]" pageId="95" pageNumber="96">RDJ</collectionCode>
:
<specimenCode id="FD2F9EA71F1D9566FD8EFDAADE1FFDCE" box="[521,613,523,549]" collectionCode="RDJ" pageId="95" pageNumber="96">ML497</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="AD3636DC1F1D9566FF3AFDEDD91FFCB6" blockId="95.[189,1399,588,861]" pageId="95" pageNumber="96">
<materialsCitation id="1DE13C811F1D9566FF3AFDEDDF4DFD62" ID-GBIF-Occurrence="2867606468" collectingDate="2017-07-06" collectingDateMax="2017-07-07" collectingDateMin="2017-07-06" collectionCode="PW" collectorName="S. Knyazev" country="Russia" latitude="50.3158" location="Ulagansky" longLatPrecision="6" longitude="87.7036" pageId="95" pageNumber="96" specimenCode="ML264" specimenCount="1" specimenCount-female="1" stateProvince="Altay">
<collectingCountry id="D59E764C1F1D9566FF3AFDEDDD5EFD8D" box="[189,292,588,614]" name="Russia" pageId="95" pageNumber="96">RUSSIA</collectingCountry>
<specimenCount id="BB8FFD551F1D9566FEBBFDEDDD12FD8C" box="[316,360,588,615]" pageId="95" pageNumber="96" type="female">1 ♀</specimenCount>
(worker);
<collectingRegion id="6F4DF83E1F1D9566FE64FDEDDE5BFD8D" box="[483,545,588,614]" country="Russia" name="Altay" pageId="95" pageNumber="96">Altai</collectingRegion>
,
<location id="A85660071F1D9566FDABFDEDDED1FD8D" LSID="urn:lsid:plazi:treatment:252087CA1F1E9566FDA0FE2ED89AF80E:A85660071F1D9566FDABFDEDDED1FD8D" box="[556,683,588,614]" country="Russia" latitude="50.3158" longLatPrecision="6" longitude="87.7036" name="Ulagansky" pageId="95" pageNumber="96" stateProvince="Altay">Ulagansky</location>
,
<locationDeviation id="BC842FFE1F1D9566FD31FDEDDFE6FD8D" box="[694,924,588,615]" location="Aktash" pageId="95" pageNumber="96">
<quantity id="6A719B391F1D9566FD31FDEDDE8BFD8D" box="[694,753,588,614]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="95" pageNumber="96" unit="km" value="6.0">6 km</quantity>
NE of Aktash
</locationDeviation>
;
<geoCoordinate id="C8BD501B1F1D9566FC20FDEDD84BFD8C" box="[935,1073,588,615]" degrees="50.3158" direction="north" orientation="latitude" pageId="95" pageNumber="96" precision="5" value="50.3158">50.3158° N</geoCoordinate>
,
<geoCoordinate id="C8BD501B1F1D9566FBBAFDEDD8BEFD8C" box="[1085,1220,588,615]" degrees="87.7036" direction="east" orientation="longitude" pageId="95" pageNumber="96" precision="5" value="87.7036">87.7036° E</geoCoordinate>
;
<date id="D937101C1F1D9566FB57FDEDD908FD8D" box="[1232,1394,588,614]" pageId="95" pageNumber="96" value="2017-07-06" valueMax="2017-07-07" valueMin="2017-07-06">
<collectingDate id="C973E9F41F1D9566FB57FDEDD908FD8D" box="[1232,1394,588,614]" pageId="95" pageNumber="96" value="2017-07-06" valueMax="2017-07-07" valueMin="2017-07-06">67 Jul. 2017</collectingDate>
</date>
;
<collectorName id="007C530A1F1D9566FF3AFDCEDD3DFD61" box="[189,327,623,650]" pageId="95" pageNumber="96">S. Knyazev</collectorName>
leg.; BOLD seq: 1555F07;
<collectionCode id="CB98AE191F1D9566FD27FDD1DEAAFD61" box="[672,720,624,650]" country="Thailand" name="Paleontological Collections" pageId="95" pageNumber="96">PW</collectionCode>
:
<specimenCode id="FD2F9EA71F1D9566FD59FDCEDF4DFD62" box="[734,823,623,649]" collectionCode="PW" country="Thailand" name="Paleontological Collections" pageId="95" pageNumber="96">ML264</specimenCode>
</materialsCitation>
<materialsCitation id="1DE13C811F1D9566FCD1FDCED91EFD46" ID-GBIF-Occurrence="2867606381" collectingDate="2017-07-05" collectingDateMax="2017-07-06" collectingDateMin="2017-07-05" collectionCode="PW" collectorName="S. Knyazev" country="Russia" latitude="50.316" location="Ulagansky" longLatPrecision="6" longitude="87.7253" pageId="95" pageNumber="96" specimenCode="ML265" specimenCount="1" specimenCount-female="1" stateProvince="Altay">
<specimenCount id="BB8FFD551F1D9566FCD1FDCEDFFFFD61" box="[854,901,623,650]" pageId="95" pageNumber="96" type="female">1 ♀</specimenCount>
(worker);
<collectingRegion id="6F4DF83E1F1D9566FB8FFDCED83CFD62" box="[1032,1094,623,649]" country="Russia" name="Altay" pageId="95" pageNumber="96">Altai</collectingRegion>
,
<location id="A85660071F1D9566FBD2FDCED8AEFD62" LSID="urn:lsid:plazi:treatment:252087CA1F1E9566FDA0FE2ED89AF80E:A85660071F1D9566FBD2FDCED8AEFD62" box="[1109,1236,623,649]" country="Russia" latitude="50.316" longLatPrecision="6" longitude="87.7253" name="Ulagansky" pageId="95" pageNumber="96" stateProvince="Altay">Ulagansky</location>
,
<locationDeviation id="BC842FFE1F1D9566FB65FDCEDD6EFD47" location="Aktash" pageId="95" pageNumber="96">
<quantity id="6A719B391F1D9566FB65FDCED95BFD62" box="[1250,1313,623,649]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="95" pageNumber="96" unit="km" value="8.0">8 km</quantity>
NE of Aktash
</locationDeviation>
;
<geoCoordinate id="C8BD501B1F1D9566FEA7FD33DDD1FD46" box="[288,427,658,685]" degrees="50.3160" direction="north" orientation="latitude" pageId="95" pageNumber="96" precision="5" value="50.316">50.3160° N</geoCoordinate>
,
<geoCoordinate id="C8BD501B1F1D9566FE3FFD33DE3AFD46" box="[440,576,658,685]" degrees="87.7253" direction="east" orientation="longitude" pageId="95" pageNumber="96" precision="5" value="87.7253">87.7253° E</geoCoordinate>
;
<date id="D937101C1F1D9566FDCAFD33DE8BFD46" box="[589,753,658,685]" pageId="95" pageNumber="96" value="2017-07-05" valueMax="2017-07-06" valueMin="2017-07-05">
<collectingDate id="C973E9F41F1D9566FDCAFD33DE8BFD46" box="[589,753,658,685]" pageId="95" pageNumber="96" value="2017-07-05" valueMax="2017-07-06" valueMin="2017-07-05">56 Jul. 2017</collectingDate>
</date>
;
<collectorName id="007C530A1F1D9566FD79FD32DFFCFD46" box="[766,902,659,685]" pageId="95" pageNumber="96">S. Knyazev</collectorName>
leg.; BOLD seq: 1555F08;
<collectionCode id="CB98AE191F1D9566FB56FD32D97BFD46" box="[1233,1281,659,685]" country="Thailand" name="Paleontological Collections" pageId="95" pageNumber="96">PW</collectionCode>
:
<specimenCode id="FD2F9EA71F1D9566FA8BFD33D91EFD46" box="[1292,1380,658,685]" collectionCode="PW" country="Thailand" name="Paleontological Collections" pageId="95" pageNumber="96">ML265</specimenCode>
</materialsCitation>
<materialsCitation id="1DE13C811F1D9566FF3AFD17DE72FD18" ID-GBIF-Occurrence="2867606383" collectingDate="2014-07-11" collectingDateMax="2014-07-12" collectingDateMin="2014-07-11" collectionCode="PW" collectorName="A. Lelej" country="Russia" latitude="50.0049" location="27 km S of Erzin" longLatPrecision="6" longitude="95.1719" pageId="95" pageNumber="96" specimenCode="ML266" specimenCount="1" specimenCount-female="1" stateProvince="Tyva">
<specimenCount id="BB8FFD551F1D9566FF3AFD17DC93FD3B" box="[189,233,694,720]" pageId="95" pageNumber="96" type="female">1 ♀</specimenCount>
(worker);
<collectingRegion id="6F4DF83E1F1D9566FEE0FD17DDDFFD3B" box="[359,421,694,720]" country="Russia" name="Tyva" pageId="95" pageNumber="96">Tuva</collectingRegion>
,
<location id="A85660071F1D9566FE35FD17DEFAFD3B" LSID="urn:lsid:plazi:treatment:252087CA1F1E9566FDA0FE2ED89AF80E:A85660071F1D9566FE35FD17DEFAFD3B" box="[434,640,694,720]" country="Russia" latitude="50.0049" longLatPrecision="6" longitude="95.1719" name="27 km S of Erzin" pageId="95" pageNumber="96" stateProvince="Tyva">
<quantity id="6A719B391F1D9566FE35FD17DD86FD3B" box="[434,508,694,720]" metricMagnitude="4" metricUnit="m" metricValue="2.7" pageId="95" pageNumber="96" unit="km" value="27.0">27 km</quantity>
S of Erzin
</location>
;
<geoCoordinate id="C8BD501B1F1D9566FD0BFD17DF6DFD3B" box="[652,791,694,721]" degrees="50.0049" direction="north" orientation="latitude" pageId="95" pageNumber="96" precision="5" value="50.0049">50.0049° N</geoCoordinate>
,
<geoCoordinate id="C8BD501B1F1D9566FCA3FD17DFD1FD3B" box="[804,939,694,721]" degrees="95.1719" direction="east" orientation="longitude" pageId="95" pageNumber="96" precision="5" value="95.1719">95.1719° E</geoCoordinate>
;
<date id="D937101C1F1D9566FC3FFD17D802FD3B" box="[952,1144,694,720]" pageId="95" pageNumber="96" value="2014-07-11" valueMax="2014-07-12" valueMin="2014-07-11">
<collectingDate id="C973E9F41F1D9566FC3FFD17D802FD3B" box="[952,1144,694,720]" pageId="95" pageNumber="96" value="2014-07-11" valueMax="2014-07-12" valueMin="2014-07-11">1112 Jul. 2014</collectingDate>
</date>
;
<collectorName id="007C530A1F1D9566FB04FD17D899FD3B" box="[1155,1251,694,720]" pageId="95" pageNumber="96">A. Lelej</collectorName>
leg.; BOLD seq: 1555F09;
<collectionCode id="CB98AE191F1D9566FEF5FD78DDD8FD18" box="[370,418,729,755]" country="Thailand" name="Paleontological Collections" pageId="95" pageNumber="96">PW</collectionCode>
:
<specimenCode id="FD2F9EA71F1D9566FE37FD78DE72FD18" box="[432,520,729,755]" collectionCode="PW" country="Thailand" name="Paleontological Collections" pageId="95" pageNumber="96">ML266</specimenCode>
</materialsCitation>
<materialsCitation id="1DE13C811F1D9566FDAFFD78DFEBFCFD" ID-GBIF-Occurrence="2867606515" collectingDate="2014-07-14" collectingDateMax="2014-07-15" collectingDateMin="2014-07-14" collectionCode="PW" collectorName="A. Lelej" country="Russia" latitude="50.1611" location="Erzin" longLatPrecision="6" longitude="95.4778" pageId="95" pageNumber="96" specimenCode="ML267" specimenCount="1" specimenCount-female="1" stateProvince="Tyva">
<specimenCount id="BB8FFD551F1D9566FDAFFD78DE2DFD18" box="[552,599,729,755]" pageId="95" pageNumber="96" type="female">1 ♀</specimenCount>
(worker);
<collectingRegion id="6F4DF83E1F1D9566FD5CFD78DF63FD18" box="[731,793,729,755]" country="Russia" name="Tyva" pageId="95" pageNumber="96">Tuva</collectingRegion>
,
<quantity id="6A719B391F1D9566FCAFFD78DF0FFD18" box="[808,885,729,756]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="95" pageNumber="96" unit="km" value="25.0">25 km</quantity>
of
<location id="A85660071F1D9566FC23FD78DF9DFD18" LSID="urn:lsid:plazi:treatment:252087CA1F1E9566FDA0FE2ED89AF80E:A85660071F1D9566FC23FD78DF9DFD18" box="[932,999,729,755]" country="Russia" latitude="50.1611" longLatPrecision="6" longitude="95.4778" name="Erzin" pageId="95" pageNumber="96" stateProvince="Tyva">Erzin</location>
;
<geoCoordinate id="C8BD501B1F1D9566FC71FD78D8F9FD18" box="[1014,1155,729,756]" degrees="50.1611" direction="north" orientation="latitude" pageId="95" pageNumber="96" precision="5" value="50.1611">50.1611° N</geoCoordinate>
,
<geoCoordinate id="C8BD501B1F1D9566FB15FD78D967FD18" box="[1170,1309,729,756]" degrees="95.4778" direction="east" orientation="longitude" pageId="95" pageNumber="96" precision="5" value="95.4778">95.4778° E</geoCoordinate>
;
<date id="D937101C1F1D9566FAAAFD78DD4AFCFD" pageId="95" pageNumber="96" value="2014-07-14" valueMax="2014-07-15" valueMin="2014-07-14">
<collectingDate id="C973E9F41F1D9566FAAAFD78DD4AFCFD" pageId="95" pageNumber="96" value="2014-07-14" valueMax="2014-07-15" valueMin="2014-07-14">1415 Jul. 2014</collectingDate>
</date>
;
<collectorName id="007C530A1F1D9566FEB9FD5DDDDBFCFD" box="[318,417,764,790]" pageId="95" pageNumber="96">A. Lelej</collectorName>
leg.; BOLD seq: 1555F10;
<collectionCode id="CB98AE191F1D9566FD7CFD5CDF51FCFC" box="[763,811,765,791]" country="Thailand" name="Paleontological Collections" pageId="95" pageNumber="96">PW</collectionCode>
:
<specimenCode id="FD2F9EA71F1D9566FCBEFD5DDFEBFCFD" box="[825,913,764,790]" collectionCode="PW" country="Thailand" name="Paleontological Collections" pageId="95" pageNumber="96">ML267</specimenCode>
</materialsCitation>
<materialsCitation id="1DE13C811F1D9566FC36FD5DD8C6FCD2" ID-GBIF-Occurrence="2867606581" collectingDate="2014-07-18" collectionCode="PW" collectorName="A. Lelej" country="Russia" latitude="50.0366" location="Erzin" longLatPrecision="6" longitude="95.4348" pageId="95" pageNumber="96" specimenCode="ML269" specimenCount="1" specimenCount-female="1" stateProvince="Tyva">
<specimenCount id="BB8FFD551F1D9566FC36FD5DDF9BFCFC" box="[945,993,764,791]" pageId="95" pageNumber="96" type="female">1 ♀</specimenCount>
(worker);
<collectingRegion id="6F4DF83E1F1D9566FBE2FD5CD8D9FCFC" box="[1125,1187,765,791]" country="Russia" name="Tyva" pageId="95" pageNumber="96">Tuva</collectingRegion>
,
<quantity id="6A719B391F1D9566FB34FD5DD97AFCFD" box="[1203,1280,764,790]" metricMagnitude="4" metricUnit="m" metricValue="3.1" pageId="95" pageNumber="96" unit="km" value="31.0">31 km</quantity>
of
<location id="A85660071F1D9566FAA8FD5DD908FCFD" LSID="urn:lsid:plazi:treatment:252087CA1F1E9566FDA0FE2ED89AF80E:A85660071F1D9566FAA8FD5DD908FCFD" box="[1327,1394,764,790]" country="Russia" latitude="50.0366" longLatPrecision="6" longitude="95.4348" name="Erzin" pageId="95" pageNumber="96" stateProvince="Tyva">Erzin</location>
;
<geoCoordinate id="C8BD501B1F1D9566FF3AFCBEDD3DFCD1" box="[189,327,799,826]" degrees="50.0366" direction="north" orientation="latitude" pageId="95" pageNumber="96" precision="5" value="50.0366">50.0366° N</geoCoordinate>
,
<geoCoordinate id="C8BD501B1F1D9566FED4FCBEDDA0FCD1" box="[339,474,799,826]" degrees="95.4348" direction="east" orientation="longitude" pageId="95" pageNumber="96" precision="5" value="95.4348">95.4348° E</geoCoordinate>
;
<date id="D937101C1F1D9566FE61FCBEDE03FCD2" box="[486,633,799,825]" pageId="95" pageNumber="96" value="2014-07-18">
<collectingDate id="C973E9F41F1D9566FE61FCBEDE03FCD2" box="[486,633,799,825]" pageId="95" pageNumber="96" value="2014-07-18">18 Jul. 2014</collectingDate>
</date>
;
<collectorName id="007C530A1F1D9566FD03FC81DE99FCD2" box="[644,739,799,826]" pageId="95" pageNumber="96">A. Lelej</collectorName>
leg.; BOLD seq: 1555F12;
<collectionCode id="CB98AE191F1D9566FBADFC81D820FCD1" box="[1066,1114,800,826]" country="Thailand" name="Paleontological Collections" pageId="95" pageNumber="96">PW</collectionCode>
:
<specimenCode id="FD2F9EA71F1D9566FBE3FCBED8C6FCD2" box="[1124,1212,799,825]" collectionCode="PW" country="Thailand" name="Paleontological Collections" pageId="95" pageNumber="96">ML269</specimenCode>
</materialsCitation>
<materialsCitation id="1DE13C811F1D9566FB52FCBED91BFCB6" ID-GBIF-Occurrence="2867606401" collectingDate="2014-07-16" collectionCode="PW" collectorName="A. Lelej" country="Russia" latitude="50.0366" location="Erzin" longLatPrecision="6" longitude="95.4348" pageId="95" pageNumber="96" specimenCode="ML270" specimenCount="1" specimenCount-female="1" stateProvince="Tyva">
<specimenCount id="BB8FFD551F1D9566FB52FCBED97BFCD1" box="[1237,1281,799,826]" pageId="95" pageNumber="96" type="female">1 ♀</specimenCount>
(worker); same collection data as for preceding;
<date id="D937101C1F1D9566FD07FCE3DF6EFCB6" box="[640,788,834,861]" pageId="95" pageNumber="96" value="2014-07-16">
<collectingDate id="C973E9F41F1D9566FD07FCE3DF6EFCB6" box="[640,788,834,861]" pageId="95" pageNumber="96" value="2014-07-16">16 Jul. 2014</collectingDate>
</date>
;
<collectorName id="007C530A1F1D9566FC99FCE2DF04FCB7" box="[798,894,834,861]" pageId="95" pageNumber="96">A. Lelej</collectorName>
leg.; BOLD seq: 1555G01;
<collectionCode id="CB98AE191F1D9566FB4CFCE2D881FCB6" box="[1227,1275,835,861]" country="Thailand" name="Paleontological Collections" pageId="95" pageNumber="96">PW</collectionCode>
:
<specimenCode id="FD2F9EA71F1D9566FA82FCE2D91BFCB6" box="[1285,1377,835,861]" collectionCode="PW" country="Thailand" name="Paleontological Collections" pageId="95" pageNumber="96">ML270</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="AD3636DC1F1D9566FF3AFC25DEADFA67" blockId="95.[189,1399,900,1420]" pageId="95" pageNumber="96">
<materialsCitation id="1DE13C811F1D9566FF3AFC25DEE8FC29" ID-GBIF-Occurrence="2867606567" collectingDate="2009-05-27" collectionCode="PW" collectorName="L. Best" country="Kyrgyzstan" latitude="42.6522" location="Jalalabad" longLatPrecision="6" longitude="74.4812" municipality="Jylamysh" pageId="95" pageNumber="96" specimenCode="ML156" specimenCount="1" specimenCount-female="1">
<collectingCountry id="D59E764C1F1D9566FF3AFC25DDFBFC75" box="[189,385,900,926]" name="Kyrgyzstan" pageId="95" pageNumber="96">KYRGYZSTAN</collectingCountry>
<specimenCount id="BB8FFD551F1D9566FE19FC25DDB6FC74" box="[414,460,900,927]" pageId="95" pageNumber="96" type="female">1 ♀</specimenCount>
(queen);
<location id="A85660071F1D9566FDC7FC25DECAFC75" LSID="urn:lsid:plazi:treatment:252087CA1F1E9566FDA0FE2ED89AF80E:A85660071F1D9566FDC7FC25DECAFC75" box="[576,688,900,926]" country="Kyrgyzstan" latitude="42.6522" longLatPrecision="6" longitude="74.4812" municipality="Jylamysh" name="Jalalabad" pageId="95" pageNumber="96">Jalalabad</location>
,
<collectingMunicipality id="4D52ACA61F1D9566FD3AFC25DF4AFC75" box="[701,816,900,926]" pageId="95" pageNumber="96">Jylamysh</collectingMunicipality>
;
<geoCoordinate id="C8BD501B1F1D9566FCBAFC25DFB0FC74" box="[829,970,900,927]" degrees="42.6522" direction="north" orientation="latitude" pageId="95" pageNumber="96" precision="5" value="42.6522">42.6522° N</geoCoordinate>
,
<geoCoordinate id="C8BD501B1F1D9566FC50FC25D81AFC74" box="[983,1120,900,927]" degrees="74.4812" direction="east" orientation="longitude" pageId="95" pageNumber="96" precision="5" value="74.4812">74.4812° E</geoCoordinate>
;
<date id="D937101C1F1D9566FBE8FC25D968FC75" box="[1135,1298,900,927]" pageId="95" pageNumber="96" value="2009-05-27">
<collectingDate id="C973E9F41F1D9566FBE8FC25D968FC75" box="[1135,1298,900,927]" pageId="95" pageNumber="96" value="2009-05-27">27 May 2009</collectingDate>
</date>
;
<collectorName id="007C530A1F1D9566FAA7FC24D90CFC74" box="[1312,1398,901,927]" pageId="95" pageNumber="96">L. Best</collectorName>
leg.; BOLD seq: 1552H03;
<collectionCode id="CB98AE191F1D9566FD87FC09DE4AFC29" box="[512,560,936,962]" country="Thailand" name="Paleontological Collections" pageId="95" pageNumber="96">PW</collectionCode>
:
<specimenCode id="FD2F9EA71F1D9566FDBDFC06DEE8FC29" box="[570,658,935,962]" collectionCode="PW" country="Thailand" name="Paleontological Collections" pageId="95" pageNumber="96">ML156</specimenCode>
</materialsCitation>
<materialsCitation id="1DE13C811F1D9566FD2DFC06DF85FC0F" ID-GBIF-Occurrence="2867606317" collectingDate="1998-07-18" collectionCode="OLML" collectorName="H. Rausch" country="Kyrgyzstan" latitude="39.4761" location="Oshakaya" longLatPrecision="6" longitude="71.8933" municipality="Kyzyl-Suu" pageId="95" pageNumber="96" specimenCode="ML191" specimenCount="1" specimenCount-female="1">
<specimenCount id="BB8FFD551F1D9566FD2DFC06DEACFC29" box="[682,726,935,962]" pageId="95" pageNumber="96" type="female">1 ♀</specimenCount>
(queen);
<location id="A85660071F1D9566FCC2FC06DFC6FC2A" LSID="urn:lsid:plazi:treatment:252087CA1F1E9566FDA0FE2ED89AF80E:A85660071F1D9566FCC2FC06DFC6FC2A" box="[837,956,935,961]" country="Kyrgyzstan" latitude="39.4761" longLatPrecision="6" longitude="71.8933" municipality="Kyzyl-Suu" name="Oshakaya" pageId="95" pageNumber="96">Oshakaya</location>
,
<collectingMunicipality id="4D52ACA61F1D9566FC40FC06D830FC2A" box="[967,1098,935,961]" pageId="95" pageNumber="96">Kyzyl-Suu</collectingMunicipality>
;
<geoCoordinate id="C8BD501B1F1D9566FBD2FC06D8A5FC29" box="[1109,1247,935,962]" degrees="39.4761" direction="north" orientation="latitude" pageId="95" pageNumber="96" precision="5" value="39.4761">39.4761° N</geoCoordinate>
,
<geoCoordinate id="C8BD501B1F1D9566FB6DFC06D90BFC29" box="[1258,1393,935,962]" degrees="71.8933" direction="east" orientation="longitude" pageId="95" pageNumber="96" precision="5" value="71.8933">71.8933° E</geoCoordinate>
;
<date id="D937101C1F1D9566FF3AFC6BDD2DFC0F" box="[189,343,970,996]" pageId="95" pageNumber="96" value="1998-07-18">
<collectingDate id="C973E9F41F1D9566FF3AFC6BDD2DFC0F" box="[189,343,970,996]" pageId="95" pageNumber="96" value="1998-07-18">18 Jul. 1998</collectingDate>
</date>
;
<collectorName id="007C530A1F1D9566FEE0FC6ADD9EFC0F" box="[359,484,970,997]" pageId="95" pageNumber="96">H. Rausch</collectorName>
leg.; BOLD seq: 1552A08;
<collectionCode id="CB98AE191F1D9566FCC6FC6BDFE3FC0F" box="[833,921,970,996]" country="Austria" httpUri="http://grbio.org/cool/pp0t-48s3" name="Oberösterreichisches Landesmuseum" pageId="95" pageNumber="96" type="Museum">OLML</collectionCode>
:
<specimenCode id="FD2F9EA71F1D9566FC20FC6BDF85FC0F" box="[935,1023,970,996]" collectionCode="OLML" country="Austria" httpUri="http://grbio.org/cool/pp0t-48s3" name="Oberösterreichisches Landesmuseum" pageId="95" pageNumber="96" type="Museum">ML191</specimenCode>
</materialsCitation>
<materialsCitation id="1DE13C811F1D9566FB98FC6BDD3EFBC0" ID-GBIF-Occurrence="2867606627" collectingDate="2016-06-26" collectorName="R. DeJonghe" country="Kyrgyzstan" latitude="42.8386" location="Chong Aksuu Valley" longLatPrecision="6" longitude="77.3889" municipality="Karakol lake" pageId="95" pageNumber="96" specimenCount="1" specimenCount-female="1">
<specimenCount id="BB8FFD551F1D9566FB98FC6BD834FC0E" box="[1055,1102,970,997]" pageId="95" pageNumber="96" type="female">1 ♀</specimenCount>
(worker);
<location id="A85660071F1D9566FB54FC6BDD72FBEC" LSID="urn:lsid:plazi:treatment:252087CA1F1E9566FDA0FE2ED89AF80E:A85660071F1D9566FB54FC6BDD72FBEC" country="Kyrgyzstan" latitude="42.8386" longLatPrecision="6" longitude="77.3889" municipality="Karakol lake" name="Chong Aksuu Valley" pageId="95" pageNumber="96">Chong Aksuu Valley</location>
nr
<collectingMunicipality id="4D52ACA61F1D9566FEA9FC4CDDB3FBEC" box="[302,457,1005,1031]" pageId="95" pageNumber="96">Karakol lake</collectingMunicipality>
;
<geoCoordinate id="C8BD501B1F1D9566FE54FC4CDE27FBE3" box="[467,605,1005,1032]" degrees="42.8386" direction="north" orientation="latitude" pageId="95" pageNumber="96" precision="5" value="42.8386">42.8386° N</geoCoordinate>
,
<geoCoordinate id="C8BD501B1F1D9566FDEEFC4FDE8AFBE3" box="[617,752,1006,1032]" degrees="77.3889" direction="east" orientation="longitude" pageId="95" pageNumber="96" precision="5" value="77.3889">77.3889° E</geoCoordinate>
;
<date id="D937101C1F1D9566FD7BFC4CDFEFFBEC" box="[764,917,1005,1032]" pageId="95" pageNumber="96" value="2016-06-26">
<collectingDate id="C973E9F41F1D9566FD7BFC4CDFEFFBEC" box="[764,917,1005,1032]" pageId="95" pageNumber="96" value="2016-06-26">26 Jun. 2016</collectingDate>
</date>
;
<collectorName id="007C530A1F1D9566FC26FC4FD842FBEC" box="[929,1080,1005,1032]" pageId="95" pageNumber="96">R. DeJonghe</collectorName>
leg.; BOLD seq: 1555F04; RJ: ML261
</materialsCitation>
<materialsCitation id="1DE13C811F1D9566FED8FBB0D91EFBC0" ID-GBIF-Occurrence="2867606424" box="[351,1380,1041,1068]" collectingDate="2016-06-26" collectorName="R. DeJonghe" country="Kyrgyzstan" latitude="42.8386" location="Chong Aksuu Valley" longLatPrecision="6" longitude="77.3889" municipality="Karakol lake" pageId="95" pageNumber="96" specimenCount="1" specimenCount-female="1">
<specimenCount id="BB8FFD551F1D9566FED8FBB0DDF6FBC0" box="[351,396,1041,1067]" pageId="95" pageNumber="96" type="female">1 ♀</specimenCount>
(worker); same collection data as for preceding; BOLD seq: 1555F05; RJ: ML262
</materialsCitation>
<materialsCitation id="1DE13C811F1D9566FF3AFB95D8BEFBA5" ID-GBIF-Occurrence="2867606473" box="[189,1220,1076,1103]" collectingDate="2016-06-26" collectorName="R. DeJonghe" country="Kyrgyzstan" latitude="42.8386" location="Chong Aksuu Valley" longLatPrecision="6" longitude="77.3889" municipality="Karakol lake" pageId="95" pageNumber="96" specimenCount="1" specimenCount-female="1">
<specimenCount id="BB8FFD551F1D9566FF3AFB95DC90FBA4" box="[189,234,1076,1103]" pageId="95" pageNumber="96" type="female">1 ♀</specimenCount>
(worker); same collection data as for preceding; BOLD seq: 1555F06; RJ: ML263
</materialsCitation>
<materialsCitation id="1DE13C811F1D9566FB58FB95DE2FFB7F" ID-GBIF-Occurrence="2867606615" collectingDate="2016-06-25" collectorName="R. DeJonghe" country="Kyrgyzstan" county="Karakol lake" latitude="42.8397" location="Kungey Ala-Too" longLatPrecision="407" longitude="77.37" municipality="Chong Aksuu" pageId="95" pageNumber="96" specimenCount="1" specimenCount-female="1">
<specimenCount id="BB8FFD551F1D9566FB58FB95D976FBA4" box="[1247,1292,1076,1103]" pageId="95" pageNumber="96" type="female">1 ♀</specimenCount>
(queen);
<location id="A85660071F1D9566FF3AFBF9DDF9FB9A" LSID="urn:lsid:plazi:treatment:252087CA1F1E9566FDA0FE2ED89AF80E:A85660071F1D9566FF3AFBF9DDF9FB9A" box="[189,387,1111,1138]" country="Kyrgyzstan" county="Karakol lake" latitude="42.8397" longLatPrecision="407" longitude="77.37" municipality="Chong Aksuu" name="Kungey Ala-Too" pageId="95" pageNumber="96">Kungey Ala-Too</location>
,
<collectingMunicipality id="4D52ACA61F1D9566FE09FBF6DE54FB9A" box="[398,558,1111,1137]" pageId="95" pageNumber="96">Chong Aksuu</collectingMunicipality>
nr
<collectingCounty id="44574E501F1D9566FDD4FBF6DE97FB9A" box="[595,749,1111,1137]" pageId="95" pageNumber="96">Karakol lake</collectingCounty>
;
<geoCoordinate id="C8BD501B1F1D9566FD7FFBF6DFF8FB99" box="[760,898,1111,1138]" degrees="42.8397" direction="north" orientation="latitude" pageId="95" pageNumber="96" precision="5" value="42.8397">42.8397° N</geoCoordinate>
,
<geoCoordinate id="C8BD501B1F1D9566FC0AFBF6DF8CFB99" box="[909,1014,1111,1138]" degrees="77.37" direction="east" orientation="longitude" pageId="95" pageNumber="96" precision="555" value="77.37">77.37° E</geoCoordinate>
;
<date id="D937101C1F1D9566FB85FBF6D8E0FB9A" box="[1026,1178,1111,1138]" pageId="95" pageNumber="96" value="2016-06-25">
<collectingDate id="C973E9F41F1D9566FB85FBF6D8E0FB9A" box="[1026,1178,1111,1138]" pageId="95" pageNumber="96" value="2016-06-25">25 Jun. 2016</collectingDate>
</date>
;
<collectorName id="007C530A1F1D9566FB21FBF9D946FB9A" box="[1190,1340,1111,1138]" pageId="95" pageNumber="96">R. DeJonghe</collectorName>
leg.; BOLD seq: 6877A09; RJ: ML336
</materialsCitation>
<materialsCitation id="1DE13C811F1D9566FDF5FBDBD80EFB53" ID-GBIF-Occurrence="2867606574" collectingDate="2016-07-02" collectorName="R. DeJonghe" country="Kyrgyzstan" latitude="42.4183" location="Terskey Ala-Too" longLatPrecision="6" longitude="78.9467" municipality="Turgon-Aksuu river" pageId="95" pageNumber="96" specimenCount="1" specimenCount-female="1">
<specimenCount id="BB8FFD551F1D9566FDF5FBDBDEDAFB7E" box="[626,672,1146,1173]" pageId="95" pageNumber="96" type="female">1 ♀</specimenCount>
(queen);
<location id="A85660071F1D9566FC94FBDBDFA1FB7F" LSID="urn:lsid:plazi:treatment:252087CA1F1E9566FDA0FE2ED89AF80E:A85660071F1D9566FC94FBDBDFA1FB7F" box="[787,987,1146,1172]" country="Kyrgyzstan" latitude="42.4183" longLatPrecision="6" longitude="78.9467" municipality="Turgon-Aksuu river" name="Terskey Ala-Too" pageId="95" pageNumber="96">Terskey Ala-Too</location>
,
<collectingMunicipality id="4D52ACA61F1D9566FC6EFBDBD8A2FB7F" box="[1001,1240,1146,1172]" pageId="95" pageNumber="96">Turgon-Aksuu river</collectingMunicipality>
;
<geoCoordinate id="C8BD501B1F1D9566FB61FBDBD908FB7E" box="[1254,1394,1146,1173]" degrees="42.4183" direction="north" orientation="latitude" pageId="95" pageNumber="96" precision="5" value="42.4183">42.4183° N</geoCoordinate>
,
<geoCoordinate id="C8BD501B1F1D9566FF3AFB3CDD32FB53" box="[189,328,1181,1208]" degrees="78.9467" direction="east" orientation="longitude" pageId="95" pageNumber="96" precision="5" value="78.9467">78.9467° E</geoCoordinate>
;
<date id="D937101C1F1D9566FEDFFB3FDD99FB5C" box="[344,483,1181,1208]" pageId="95" pageNumber="96" value="2016-07-02">
<collectingDate id="C973E9F41F1D9566FEDFFB3FDD99FB5C" box="[344,483,1181,1208]" pageId="95" pageNumber="96" value="2016-07-02">2 Jul. 2016</collectingDate>
</date>
;
<collectorName id="007C530A1F1D9566FE74FB3FDEF7FB5C" box="[499,653,1181,1208]" pageId="95" pageNumber="96">R. DeJonghe</collectorName>
leg.; BOLD seq: 6877A11; RJ: ML341
</materialsCitation>
<materialsCitation id="1DE13C811F1D9566FB13FB3FDF9CFB30" ID-GBIF-Occurrence="2867606368" collectingDate="2016-07-02" collectorName="R. DeJonghe" country="Kyrgyzstan" latitude="42.4183" location="Terskey Ala-Too" longLatPrecision="6" longitude="78.9467" municipality="Turgon-Aksuu river" pageId="95" pageNumber="96" specimenCount="1" specimenCount-female="1">
<specimenCount id="BB8FFD551F1D9566FB13FB3FD8B9FB53" box="[1172,1219,1182,1208]" pageId="95" pageNumber="96" type="female">1 ♀</specimenCount>
(queen); same collection data as for preceding; BOLD seq: 6877A10; RJ: ML342
</materialsCitation>
<materialsCitation id="1DE13C811F1D9566FB82FB60DD6FFACA" ID-GBIF-Occurrence="2867606412" collectingDate="2016-07-03" collectorName="R. DeJonghe" country="Kyrgyzstan" latitude="42.4019" location="Terskey Ala-Too" longLatPrecision="6" longitude="79.0486" municipality="Turgon-Aksuu river" pageId="95" pageNumber="96" specimenCount="1" specimenCount-female="1">
<specimenCount id="BB8FFD551F1D9566FB82FB60D849FB30" box="[1029,1075,1217,1243]" pageId="95" pageNumber="96" type="female">1 ♀</specimenCount>
(queen);
<location id="A85660071F1D9566FB2EFB60D908FB30" LSID="urn:lsid:plazi:treatment:252087CA1F1E9566FDA0FE2ED89AF80E:A85660071F1D9566FB2EFB60D908FB30" box="[1193,1394,1217,1243]" country="Kyrgyzstan" latitude="42.4019" longLatPrecision="6" longitude="79.0486" municipality="Turgon-Aksuu river" name="Terskey Ala-Too" pageId="95" pageNumber="96">Terskey Ala-Too</location>
,
<collectingMunicipality id="4D52ACA61F1D9566FF3AFB45DDD0FB15" box="[189,426,1252,1278]" pageId="95" pageNumber="96">Turgon-Aksuu river</collectingMunicipality>
;
<geoCoordinate id="C8BD501B1F1D9566FE31FB45DE3AFB14" box="[438,576,1252,1279]" degrees="42.4019" direction="north" orientation="latitude" pageId="95" pageNumber="96" precision="5" value="42.4019">42.4019° N</geoCoordinate>
,
<geoCoordinate id="C8BD501B1F1D9566FDCCFB45DEA8FB14" box="[587,722,1252,1279]" degrees="79.0486" direction="east" orientation="longitude" pageId="95" pageNumber="96" precision="5" value="79.0486">79.0486° E</geoCoordinate>
;
<date id="D937101C1F1D9566FD59FB45DF18FB15" box="[734,866,1252,1278]" pageId="95" pageNumber="96" value="2016-07-03">
<collectingDate id="C973E9F41F1D9566FD59FB45DF18FB15" box="[734,866,1252,1278]" pageId="95" pageNumber="96" value="2016-07-03">3 Jul. 2016</collectingDate>
</date>
;
<collectorName id="007C530A1F1D9566FCE9FB44D87FFB15" box="[878,1029,1252,1279]" pageId="95" pageNumber="96">R. DeJonghe</collectorName>
leg.; BOLD seq: 6877A12; RJ: ML343
</materialsCitation>
<materialsCitation id="1DE13C811F1D9566FEB7FAA6DF71FAAF" ID-GBIF-Occurrence="2867606607" collectingDate="2016-07-06" collectorName="R. DeJonghe" country="Kyrgyzstan" county="Juktuu river" latitude="42.0947" location="Terskey Ala-Too" longLatPrecision="6" longitude="77.9617" municipality="Saruu" pageId="95" pageNumber="96" specimenCount="1" specimenCount-female="1">
<specimenCount id="BB8FFD551F1D9566FEB7FAA6DD24FAC9" box="[304,350,1287,1314]" pageId="95" pageNumber="96" type="female">1 ♀</specimenCount>
(queen);
<location id="A85660071F1D9566FE57FAA6DEEDFACA" LSID="urn:lsid:plazi:treatment:252087CA1F1E9566FDA0FE2ED89AF80E:A85660071F1D9566FE57FAA6DEEDFACA" box="[464,663,1287,1313]" country="Kyrgyzstan" county="Juktuu river" latitude="42.0947" longLatPrecision="6" longitude="77.9617" municipality="Saruu" name="Terskey Ala-Too" pageId="95" pageNumber="96">Terskey Ala-Too</location>
,
<collectingMunicipality id="4D52ACA61F1D9566FD22FAA6DE96FACA" box="[677,748,1287,1313]" pageId="95" pageNumber="96">Saruu</collectingMunicipality>
,
<collectingCounty id="44574E501F1D9566FD7EFAA6DFF1FACA" box="[761,907,1287,1313]" pageId="95" pageNumber="96">Juktuu river</collectingCounty>
;
<geoCoordinate id="C8BD501B1F1D9566FC1FFAA6D859FAC9" box="[920,1059,1287,1314]" degrees="42.0947" direction="north" orientation="latitude" pageId="95" pageNumber="96" precision="5" value="42.0947">42.0947° N</geoCoordinate>
,
<geoCoordinate id="C8BD501B1F1D9566FBB7FAA6D8C3FAC9" box="[1072,1209,1287,1314]" degrees="77.9617" direction="east" orientation="longitude" pageId="95" pageNumber="96" precision="5" value="77.9617">77.9617° E</geoCoordinate>
;
<date id="D937101C1F1D9566FB40FAA6D934FACA" box="[1223,1358,1287,1313]" pageId="95" pageNumber="96" value="2016-07-06">
<collectingDate id="C973E9F41F1D9566FB40FAA6D934FACA" box="[1223,1358,1287,1313]" pageId="95" pageNumber="96" value="2016-07-06">6 Jul. 2016</collectingDate>
</date>
;
<collectorName id="007C530A1F1D9566FADBFAA9DD48FAAF" pageId="95" pageNumber="96">R. DeJonghe</collectorName>
leg.; BOLD seq: 6877B01; RJ: ML348
</materialsCitation>
<materialsCitation id="1DE13C811F1D9566FCA0FA8BDF01FA83" ID-GBIF-Occurrence="2867606384" collectingDate="2016-06-29" collectorName="R. DeJonghe" country="Kyrgyzstan" latitude="42.7389" location="Santash" longLatPrecision="40803" longitude="79.0" pageId="95" pageNumber="96" specimenCount="1" specimenCount-female="1">
<specimenCount id="BB8FFD551F1D9566FCA0FA8BDF2EFAAE" box="[807,852,1322,1349]" pageId="95" pageNumber="96" type="female">1 ♀</specimenCount>
(queen);
<location id="A85660071F1D9566FC40FA8BD85CFAAF" LSID="urn:lsid:plazi:treatment:252087CA1F1E9566FDA0FE2ED89AF80E:A85660071F1D9566FC40FA8BD85CFAAF" box="[967,1062,1322,1348]" country="Kyrgyzstan" latitude="42.7389" longLatPrecision="40803" longitude="79.0" name="Santash" pageId="95" pageNumber="96">Santash</location>
;
<geoCoordinate id="C8BD501B1F1D9566FBB4FA8BD8C5FAAE" box="[1075,1215,1322,1349]" degrees="42.7389" direction="north" orientation="latitude" pageId="95" pageNumber="96" precision="5" value="42.7389">42.7389° N</geoCoordinate>
,
<geoCoordinate id="C8BD501B1F1D9566FB4BFA8BD969FAAE" box="[1228,1299,1322,1349]" degrees="79" direction="east" orientation="longitude" pageId="95" pageNumber="96" precision="55555" value="79.0">79° E</geoCoordinate>
;
<date id="D937101C1F1D9566FAA7FA8BDC81FA8C" pageId="95" pageNumber="96" value="2016-06-29">
<collectingDate id="C973E9F41F1D9566FAA7FA8BDC81FA8C" pageId="95" pageNumber="96" value="2016-06-29">29 Jun. 2016</collectingDate>
</date>
;
<collectorName id="007C530A1F1D9566FE8EFAEFDDDBFA8C" box="[265,417,1357,1384]" pageId="95" pageNumber="96">R. DeJonghe</collectorName>
leg.; BOLD seq: 6877B02; RJ: ML356
</materialsCitation>
<materialsCitation id="1DE13C811F1D9566FC10FAEFDEA9FA67" ID-GBIF-Occurrence="2867606594" collectingDate="2016-06-29" collectorName="R. DeJonghe" country="Kyrgyzstan" latitude="42.7389" location="Santash" longLatPrecision="40803" longitude="79.0" pageId="95" pageNumber="96" specimenCount="1" specimenCount-female="1">
<specimenCount id="BB8FFD551F1D9566FC10FAEFDFBFFA83" box="[919,965,1358,1384]" pageId="95" pageNumber="96" type="female">1 ♀</specimenCount>
(queen); same collection data as for preceding; BOLD seq: 6877B03; RJ: ML357
</materialsCitation>
.
</paragraph>
<paragraph id="AD3636DC1F1D9566FF3AFA13DFACFA1B" blockId="95.[189,1399,1458,1520]" pageId="95" pageNumber="96">
<materialsCitation id="1DE13C811F1D9566FF3AFA13DFA9FA1B" ID-GBIF-Occurrence="2867606305" collectingDate="1984-08-08" collectionCode="PW" collectorName="W. Budenburg" country="Pakistan" latitude="36.315" location="Tirich Valley" longLatPrecision="6" longitude="71.9824" municipality="Chitral" pageId="95" pageNumber="96" specimenCode="ML383" specimenCount="1" specimenCount-female="1" stateProvince="Khyber Pakhtunkhwa">
<collectingCountry id="D59E764C1F1D9566FF3AFA13DD33FA27" box="[189,329,1458,1484]" name="Pakistan" pageId="95" pageNumber="96">PAKISTAN</collectingCountry>
<specimenCount id="BB8FFD551F1D9566FEE1FA13DDEEFA26" box="[358,404,1458,1485]" pageId="95" pageNumber="96" type="female">1 ♀</specimenCount>
(worker);
<collectingMunicipality id="4D52ACA61F1D9566FD91FA13DE10FA27" box="[534,618,1458,1484]" pageId="95" pageNumber="96">Chitral</collectingMunicipality>
,
<collectingRegion id="6F4DF83E1F1D9566FDFFFA13DF06FA27" box="[632,892,1458,1484]" country="Pakistan" name="Khyber Pakhtunkhwa" pageId="95" pageNumber="96">Khyber-Pakhtunkhwa</collectingRegion>
,
<location id="A85660071F1D9566FC0EFA13D852FA27" LSID="urn:lsid:plazi:treatment:252087CA1F1E9566FDA0FE2ED89AF80E:A85660071F1D9566FC0EFA13D852FA27" box="[905,1064,1458,1484]" country="Pakistan" latitude="36.315" longLatPrecision="6" longitude="71.9824" municipality="Chitral" name="Tirich Valley" pageId="95" pageNumber="96" stateProvince="Khyber Pakhtunkhwa">Tirich Valley</location>
;
<geoCoordinate id="C8BD501B1F1D9566FBB2FA13D8B8FA26" box="[1077,1218,1458,1485]" degrees="36.3150" direction="north" orientation="latitude" pageId="95" pageNumber="96" precision="5" value="36.315">36.3150° N</geoCoordinate>
,
<geoCoordinate id="C8BD501B1F1D9566FB57FA13D923FA26" box="[1232,1369,1458,1485]" degrees="71.9824" direction="east" orientation="longitude" pageId="95" pageNumber="96" precision="5" value="71.9824">71.9824° E</geoCoordinate>
;
<date id="D937101C1F1D9566FAEFFA13DD46FA1B" pageId="95" pageNumber="96" value="1984-08-08">
<collectingDate id="C973E9F41F1D9566FAEFFA13DD46FA1B" pageId="95" pageNumber="96" value="1984-08-08">8 Aug. 1984</collectingDate>
</date>
;
<collectorName id="007C530A1F1D9566FECFFA77DD8BFA04" box="[328,497,1493,1520]" pageId="95" pageNumber="96">W. Budenburg</collectorName>
leg.; BOLD seq: 6877C02;
<collectionCode id="CB98AE191F1D9566FCBBFA77DF16FA1B" box="[828,876,1494,1520]" country="Thailand" name="Paleontological Collections" pageId="95" pageNumber="96">PW</collectionCode>
:
<specimenCode id="FD2F9EA71F1D9566FCF1FA77DFA9FA1B" box="[886,979,1494,1520]" collectionCode="PW" country="Thailand" name="Paleontological Collections" pageId="95" pageNumber="96">ML383</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="AD3636DC1F1D9566FF3AF9B6DF52F9BF" blockId="95.[189,1399,1559,1621]" pageId="95" pageNumber="96">
<materialsCitation id="1DE13C811F1D9566FF3AF9B6DF5EF9BF" ID-GBIF-Occurrence="2867606466" collectingDate="1980-07-15" collectionCode="PW" collectorName="P. Williams" country="India" county="Nimaling Plain" latitude="33.7927" location="Kashmir" longLatPrecision="6" longitude="77.5881" municipality="Zanskar" pageId="95" pageNumber="96" specimenCode="ML311" specimenCount="1" specimenCount-female="1">
<collectingCountry id="D59E764C1F1D9566FF3AF9B6DD74F9DA" box="[189,270,1559,1585]" name="India" pageId="95" pageNumber="96">INDIA</collectingCountry>
<specimenCount id="BB8FFD551F1D9566FEB5F9B6DD19F9DA" box="[306,355,1559,1585]" pageId="95" pageNumber="96" type="female">1 ♀</specimenCount>
(queen);
<location id="A85660071F1D9566FE59F9B6DE3FF9DA" LSID="urn:lsid:plazi:treatment:252087CA1F1E9566FDA0FE2ED89AF80E:A85660071F1D9566FE59F9B6DE3FF9DA" box="[478,581,1559,1585]" country="India" county="Nimaling Plain" latitude="33.7927" longLatPrecision="6" longitude="77.5881" municipality="Zanskar" name="Kashmir" pageId="95" pageNumber="96">Kashmir</location>
,
<collectingMunicipality id="4D52ACA61F1D9566FDD1F9B6DECCF9DA" box="[598,694,1559,1585]" pageId="95" pageNumber="96">Zanskar</collectingMunicipality>
,
<collectingCounty id="44574E501F1D9566FD4FF9B6DFF9F9DA" box="[712,899,1559,1585]" pageId="95" pageNumber="96">Nimaling Plain</collectingCounty>
;
<geoCoordinate id="C8BD501B1F1D9566FC13F9B6D85EF9DA" box="[916,1060,1559,1585]" degrees="33.7927" direction="north" orientation="latitude" pageId="95" pageNumber="96" precision="5" value="33.7927">33.7927° N</geoCoordinate>
,
<geoCoordinate id="C8BD501B1F1D9566FBB2F9B6D8B8F9DA" box="[1077,1218,1559,1586]" degrees="77.5881" direction="east" orientation="longitude" pageId="95" pageNumber="96" precision="5" value="77.5881">77.5881° E</geoCoordinate>
;
<date id="D937101C1F1D9566FB54F9B6D908F9DA" box="[1235,1394,1559,1586]" pageId="95" pageNumber="96" value="1980-07-15">
<collectingDate id="C973E9F41F1D9566FB54F9B6D908F9DA" box="[1235,1394,1559,1586]" pageId="95" pageNumber="96" value="1980-07-15">15 Jul. 1980</collectingDate>
</date>
;
<collectorName id="007C530A1F1D9566FF3AF99ADD38F9BF" box="[189,322,1594,1621]" pageId="95" pageNumber="96">P. Williams</collectorName>
leg.; BOLD seq: 1555H10;
<collectionCode id="CB98AE191F1D9566FD08F99ADEC5F9BE" box="[655,703,1595,1621]" country="Thailand" name="Paleontological Collections" pageId="95" pageNumber="96">PW</collectionCode>
:
<specimenCode id="FD2F9EA71F1D9566FD4EF99BDF5EF9BF" box="[713,804,1594,1620]" collectionCode="PW" country="Thailand" name="Paleontological Collections" pageId="95" pageNumber="96">ML311</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E59365571F1D9566FF3AF9DADE57F8BA" pageId="95" pageNumber="96" type="distribution">
<paragraph id="AD3636DC1F1D9566FF3AF9DADDD7F97E" blockId="95.[189,429,1659,1685]" box="[189,429,1659,1685]" pageId="95" pageNumber="96">
<heading id="F67E81B01F1D9566FF3AF9DADDD7F97E" bold="true" box="[189,429,1659,1685]" fontSize="11" level="3" pageId="95" pageNumber="96" reason="3">
<emphasis id="9FFDEACE1F1D9566FF3AF9DADDD7F97E" bold="true" box="[189,429,1659,1685]" pageId="95" pageNumber="96">Global distribution</emphasis>
</heading>
</paragraph>
<paragraph id="AD3636DC1F1D9566FF3AF90BDE57F8BA" blockId="95.[189,1399,1706,1873]" pageId="95" pageNumber="96">
(Central Asian and Mongolian mountain species extending into the northwestern corner of the QTP)
<emphasis id="9FFDEACE1F1D9566FF3AF96CDD37F90C" bold="true" box="[189,333,1741,1767]" pageId="95" pageNumber="96">North Asia</emphasis>
:
<collectingCountry id="D59E764C1F1D9566FEE1F96CDE73F90C" box="[358,521,1741,1767]" name="Mongolia" pageId="95" pageNumber="96">MONGOLIA</collectingCountry>
,
<collectingCountry id="D59E764C1F1D9566FD9AF96CDEF0F90C" box="[541,650,1741,1767]" name="Russia" pageId="95" pageNumber="96">RUSSIA</collectingCountry>
:
<collectingRegion id="6F4DF83E1F1D9566FD1BF96FDEA0F903" box="[668,730,1742,1768]" country="Russia" name="Tyva" pageId="95" pageNumber="96">Tuva</collectingRegion>
,
<collectingRegion id="6F4DF83E1F1D9566FD6AF96CDF52F90C" box="[749,808,1741,1767]" country="Russia" name="Altay" pageId="95" pageNumber="96">Altai</collectingRegion>
Mountains.
<emphasis id="9FFDEACE1F1D9566FC6CF96CD8EBF90C" bold="true" box="[1003,1169,1741,1767]" pageId="95" pageNumber="96">Central Asia</emphasis>
:
<collectingCountry id="D59E764C1F1D9566FB2EF96CD908F90C" box="[1193,1394,1741,1767]" name="Kazakhstan" pageId="95" pageNumber="96">KAZAKHSTAN</collectingCountry>
,
<collectingCountry id="D59E764C1F1D9566FF3AF950DDFFF8E0" box="[189,389,1777,1803]" name="Kyrgyzstan" pageId="95" pageNumber="96">KYRGYZSTAN</collectingCountry>
,
<collectingCountry id="D59E764C1F1D9566FE14F950DE40F8E0" box="[403,570,1777,1803]" name="Tajikistan" pageId="95" pageNumber="96">TAJIKISTAN</collectingCountry>
.
<emphasis id="9FFDEACE1F1D9566FDE5F950DEA0F8E1" bold="true" box="[610,730,1776,1803]" pageId="95" pageNumber="96">East Asia</emphasis>
:
<collectingCountry id="D59E764C1F1D9566FD6AF950DF35F8E0" box="[749,847,1777,1803]" name="China" pageId="95" pageNumber="96">CHINA</collectingCountry>
:
<collectingRegion id="6F4DF83E1F1D9566FCDAF951DFB2F8E1" box="[861,968,1776,1802]" country="China" name="Xinjiang Uygur" pageId="95" pageNumber="96">Xinjiang</collectingRegion>
.
<emphasis id="9FFDEACE1F1D9566FC77F951D810F8E1" bold="true" box="[1008,1130,1776,1802]" pageId="95" pageNumber="96">Himalaya</emphasis>
:
<collectingCountry id="D59E764C1F1D9566FBFAF950D976F8E0" box="[1149,1292,1777,1803]" name="Pakistan" pageId="95" pageNumber="96">PAKISTAN</collectingCountry>
,
<collectingCountry id="D59E764C1F1D9566FA9CF950D909F8E0" box="[1307,1395,1777,1803]" name="India" pageId="95" pageNumber="96">INDIA</collectingCountry>
: Kashmir. (IAR, IOZ, NHMUK, OLML, PW, RDJ, RMNH, ZIN, ZMHB.). This species is widespread and can be common (
<figureCitation id="35B22A591F1D9566FE3FF896DE5AF8BA" box="[440,544,1847,1873]" captionStart="Figs 211212" captionStartId="100.[189,241,1742,1768]" captionTargetBox="[319,1269,265,1703]" captionTargetId="figure-10@100.[319,1269,265,1703]" captionTargetPageId="100" captionText="Figs 211212. Maps of sequenced samples showing the most frequent colour patterns locally. 211. The sichelii-complex, consisting of the single polytypic species B. sichelii Radoszkowski, 1859 (inset: B. sichelii worker, China-Neimenggu, photo PW). 212. The keriensis-complex, consisting of B. alagesianus Reinig, 1930 stat. rev. (blue spots), B. tibeticus sp. nov. (dark green spots), B. incertoides Vogt, 1911 stat. rev. (light green spots), B. qilianensis sp. nov. (yellow spots), B. keriensis Morawitz, 1887 (orange spots), and B. separandus Vogt, 1909 stat. rev. (red spots) (inset: B.qilianensis sp. nov. queen, China-Sichuan, photo PW). Maps projected onto a sphere in ArcGIS using World_Shaded_Relief basemap © 2014 ESRI and showing boundaries between countries as recognised by the UN." figureDoi="http://doi.org/10.5281/zenodo.4064372" httpUri="https://zenodo.org/record/4064372/files/figure.png" pageId="95" pageNumber="96">Fig. 212</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E59365571F1D9566FF3AF8D9D89AF80E" pageId="95" pageNumber="96" type="biology_ecology">
<paragraph id="AD3636DC1F1D9566FF3AF8D9DD45F879" blockId="95.[189,319,1912,1938]" box="[189,319,1912,1938]" pageId="95" pageNumber="96">
<heading id="F67E81B01F1D9566FF3AF8D9DD45F879" bold="true" box="[189,319,1912,1938]" fontSize="11" level="3" pageId="95" pageNumber="96" reason="3">
<emphasis id="9FFDEACE1F1D9566FF3AF8D9DD45F879" bold="true" box="[189,319,1912,1938]" pageId="95" pageNumber="96">Behaviour</emphasis>
</heading>
</paragraph>
<paragraph id="AD3636DC1F1D9566FF3AF806D89AF80E" blockId="95.[189,1399,1959,2021]" pageId="95" pageNumber="96">
Very few food-plant records but expected to be generalists (
<taxonomicName id="6A894D5F1F1D9566FCF7F806D94CF82A" authority="in Williams 1991" authorityName="in Williams" authorityYear="1991" box="[880,1334,1959,1985]" class="Magnoliopsida" family="Fabaceae" genus="Caragana" kingdom="Plantae" order="Fabales" pageId="95" pageNumber="96" phylum="Tracheophyta" rank="species" species="versicolor">
<emphasis id="9FFDEACE1F1D9566FCF7F806D81EF82A" box="[880,1124,1959,1985]" italics="true" pageId="95" pageNumber="96">Caragana versicolor</emphasis>
in
<bibRefCitation id="C9184B2D1F1D9566FB00F806D94CF82A" author="Williams P. H." box="[1159,1334,1959,1985]" pageId="95" pageNumber="96" pagination="1 - 204" refId="ref72395" refString="Williams P. H. 1991. The bumble bees of the Kashmir Himalaya (Hymenoptera: Apidae, Bombini). Bulletin of the British Museum (Natural History) (Entomology) 60: 1 - 204." type="journal article" year="1991">Williams 1991</bibRefCitation>
</taxonomicName>
). The male mate-searching behaviour is expected to resemble the patrolling of
<taxonomicName id="6A894D5F1F1D9566FB94F86AD8E1F80F" authorityName="F. Morawitz" authorityYear="1887" box="[1043,1179,1994,2020]" class="Insecta" family="Apidae" genus="Bombus" kingdom="Animalia" order="Hymenoptera" pageId="95" pageNumber="96" phylum="Arthropoda" rank="species" sensu="stricto" species="keriensis">
<emphasis id="9FFDEACE1F1D9566FB94F86AD8E1F80F" box="[1043,1179,1994,2020]" italics="true" pageId="95" pageNumber="96">B. keriensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84CE57B51F1D9566FB24F86AD89AF80E" box="[1187,1248,1995,2021]" pageId="95" pageNumber="96" sensu="stricto">s. str.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>