231 lines
18 KiB
XML
231 lines
18 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.64.464" ID-GBIF-Dataset="6b201dfe-5a6c-4f3d-a08b-d149b0595eaf" ID-PMC="PMC3088402" ID-Pensoft-Pub="1313-2970-64-33" ID-PubMed="21594024" ModsDocAuthor="" ModsDocDate="2010" ModsDocID="1313-2970-64-33" ModsDocOrigin="ZooKeys 64" ModsDocTitle="New species and new records of Mydidae from the Afrotropical and Oriental regions (Insecta, Diptera, Asiloidea)" checkinTime="1451250921296" checkinUser="pensoft" docAuthor="Dikow, Torsten" docDate="2010" docId="15C93CF3D6F07077DF29E6901470A158" docLanguage="en" docName="ZooKeys 64: 33-75" docOrigin="ZooKeys 64" docSource="http://dx.doi.org/10.3897/zookeys.64.464" docTitle="Leptomydas rapti Dikow, 2010, sp. n." docType="treatment" docVersion="7" lastPageNumber="40" masterDocId="4628053AE3231F40CA513918057EFFF9" masterDocTitle="New species and new records of Mydidae from the Afrotropical and Oriental regions (Insecta, Diptera, Asiloidea)" masterLastPageNumber="75" masterPageNumber="33" pageNumber="38" updateTime="1668163853975" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>New species and new records of Mydidae from the Afrotropical and Oriental regions (Insecta, Diptera, Asiloidea)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Dikow, Torsten</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2010</mods:date>
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<mods:detail type="volume">
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<mods:number>64</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>33</mods:start>
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<mods:end>75</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.64.464</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.64.464</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-64-33</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="159360439" LSID="urn:lsid:zoobank.org:act:E4EEC8E2-C144-4884-89EF-CC4C369CAA2A" httpUri="http://treatment.plazi.org/id/15C93CF3D6F07077DF29E6901470A158" lastPageId="7" lastPageNumber="40" pageId="5" pageNumber="38">
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<subSubSection pageId="5" pageNumber="38" type="nomenclature">
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<paragraph pageId="5" pageNumber="38">
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<taxonomicName LSID="urn:lsid:zoobank.org:act:E4EEC8E2-C144-4884-89EF-CC4C369CAA2A" class="Insecta" family="Mydidae" genus="Leptomydas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptomydas rapti" order="Diptera" pageId="5" pageNumber="38" phylum="Arthropoda" rank="species" species="rapti">Leptomydas rapti</taxonomicName>
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<taxonomicNameLabel pageId="5" pageNumber="38">sp. n.</taxonomicNameLabel>
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Figs 4-63146
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="38" type="etymology">
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<paragraph pageId="5" pageNumber="38">Etymology:</paragraph>
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<paragraph pageId="5" pageNumber="38">
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The specific epithet refers to the Rapti Valley, Nepal in which the
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<typeStatus>type</typeStatus>
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locality is situated and is a noun in apposition.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="38" type="diagnosis">
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<paragraph pageId="5" pageNumber="38">Diagnosis:</paragraph>
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<paragraph pageId="5" pageNumber="38">The species is distinguished from congeners by the entirely light brown stained wings, the long brown setae on the posterior abdominal sternites, the fusion of abdominal sternite 8 with tergite 8, and its apparent distribution in south-central Nepal.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="6" lastPageNumber="39" pageId="5" pageNumber="38" type="description male">
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<paragraph pageId="5" pageNumber="38">Description Male:</paragraph>
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<paragraph pageId="5" pageNumber="38">
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Head: brown, in general grey pruinose; width distinctly greater than thorax, interocular distance on vertex larger than at ventral eye margin, vertex between compound eyes
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<normalizedToken originalValue="±">+/-</normalizedToken>
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horizontally straight, medially only slightly below dorsal eye margin, parafacial area less than
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<normalizedToken originalValue="½">1/2</normalizedToken>
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the width of central facial gibbosity; facial gibbosity distinct, well-developed and discernible in lateral view; mystax white, covering only lateral facial gibbosity (asetose medially); frons medially apruinose, laterally grey pruinose, vertex predominantly apruinose, only lateral margins grey pruinose, postgenae apruinose; setation: vertex white, frons white, ocp setae white, pocl setae white; ocellar triangle apruinose; proboscis brown, long, projecting beyond fronto-clypeal suture; labella large, much wider than prementum, as long as prementum, unsclerotised laterally; maxillary palpi cylindrical, brown, shorter than length of pedicel.
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</paragraph>
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<paragraph pageId="5" pageNumber="38">
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Antenna: brown, scape and pedicel white setose dorsally and ventrally; postpedicel cylindrical in proximal
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<normalizedToken originalValue="½">1/2</normalizedToken>
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, symmetrically bulbous in distal
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<normalizedToken originalValue="½">1/2</normalizedToken>
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, &ge; 6.0 times as long as combined length of scape and pedicel; apical
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<normalizedToken originalValue="‚seta-like‘">'seta-like'</normalizedToken>
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sensory element situated apically in cavity on postpedicel.
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</paragraph>
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<paragraph lastPageId="6" lastPageNumber="39" pageId="5" pageNumber="38">
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Thorax: brown, scutum predominantly grey pruinose, pleura predominantly apruinose; scutum medially bluish-black, laterally brown, surface entirely smooth, predominantly grey pruinose, only broad sublateral stripes and narrow paramedial stripes (not reaching posterior margin) apruinose, scutal setation comprised of distinct rows of long dorsocentral setae and lateral scutal setae; dc setae pre- and postsuturally white, acr setae absent, lateral scutal setae white, npl, spal, and pal setae absent; postpronotal lobe light brown, grey pruinose; proepisternum, lateral postpronotum, and postpronotal lobes long brown setose or long white setose; scutellum grey pruinose proximally, apruinose distally, asetose, apical scutellar setae absent; mesopostnotum, anatergite, and katatergite grey pruinose, mesopostnotum asetose, anatergite asetose, katatergite long white setose; katatergite
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<normalizedToken originalValue="±">+/-</normalizedToken>
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flat; anterior anepisternum asetose, supero-posterior
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<pageBreakToken pageId="6" pageNumber="39" start="start">anepisternum</pageBreakToken>
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asetose; posterior anepimeron long white setose, katepimeron asetose; metepimeron
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<normalizedToken originalValue="±">+/-</normalizedToken>
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flat, same colour as T1, grey pruinose, long white setose; metepisternum grey pruinose, asetose.
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</paragraph>
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<paragraph pageId="6" pageNumber="39">
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Leg: brown, setation predominantly white; all coxae apruinose, long white setose; met trochanter setose medially; femora brown, met femora evenly clubbed in distal
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<normalizedToken originalValue="¾">3/4</normalizedToken>
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, in distal
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<normalizedToken originalValue="½">1/2</normalizedToken>
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macrosetose, 1 antero-ventral and 1 postero-ventral row of macrosetae; all tibiae laterally arched, met tibia cylindrical, ventral keel absent; pro and mes tarsomere 1 about as long as individual tarsomeres 2, 3, or 4, met tarsomere 1 as long as combined length of tarsomeres 2-3; pulvilli well-developed, as long as well-developed claws, and as wide as base of claws; empodium absent.
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</paragraph>
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<paragraph pageId="6" pageNumber="39">
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Wing: length = 9.0-
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<quantity metricMagnitude="-2" metricUnit="m" metricValue="9.3" unit="mm" value="93.0">9.3 mm</quantity>
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; slightly brown stained throughout, veins light brown, microtrichia absent; cells r1, r4, r5, m3, + cup closed except r5 open; C terminates at junction with M1 (or
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<collectionCode>M</collectionCode>
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1+
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<collectionCode>M</collectionCode>
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2); R4 terminates in R1; R5 terminates in R1; stump vein (
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<collectionCode>R</collectionCode>
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3) at base of R4 present, short not reaching R2; R4 and R5 widest apart medially; r-m distinct, R4+5 and M1 apart, connected by crossvein or indistinct, R4+5 and M1 fused; M1 curves anteriorly at r-m, M1 (or
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<collectionCode>M</collectionCode>
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1+
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<collectionCode>M</collectionCode>
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2) terminates in C; CuA1 and CuA2 split proximally to m-cu (cell m3 narrow proximally); M3+CuA1 do not terminate together in C; A1 undulating, cell a1 wide, A1 and wing margin further apart proximally than distally, alula well-developed; halter brown.
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</paragraph>
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<paragraph pageId="6" pageNumber="39">
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Abdomen: brown to bluish-black; setation comprised of scattered white setae, surface entirely smooth; T1-7 brown, yellow posterior margins; T1 and anterior
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<normalizedToken originalValue="½">1/2</normalizedToken>
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of T2 long white setose, remaining T2 and T3 short white setose; T predominantly apruinose; S1-7 brown, yellow posterior margins; S1 asetose, S2 long white setose, S3 short white setose; S predominantly apruinose; T2-4 parallel-sided and not constricted waist-like; bullae on T2 brown, transversely elongate, surface entirely smooth, T2 surface anterior to bullae smooth.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="39" type="male terminalia">
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<paragraph pageId="6" pageNumber="39">Male terminalia:</paragraph>
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<paragraph pageId="6" pageNumber="39">
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T1-7 well-developed, entirely sclerotised, T8 postero-medially weakly sclerotised, with anterior transverse sclerotised bridge connecting lateral sclerites; T7-8 anteriorly with 2 lateral apodemes; S6 regular, without any special setation postero-medially, S8 medially divided into two halves, fused to T8 dorso-laterally, divided entirely ventro-medially; epandrium formed by single sclerite (fused medially
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<normalizedToken originalValue="±">+/-</normalizedToken>
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entirely), blunt, evenly rounded; subepandrial sclerite without lateral or median protuberances; hypandrium slightly concave, partially divided ventro-medially, anteriorly with heavily sclerotised transverse bridge, entirely fused with gonocoxite, forming a gonocoxite-hypandrial complex; gonocoxites dorso-ventrally flattened (same height throughout, expanded laterally and medially), with median protuberance, gonocoxal apodeme absent; 1 functional aedeagal prong; aedeagal epimere absent; lateral ejaculatory processes absent; ejaculatory apodeme formed by single dorso-ventrally oriented plate; ventro-median margin of dorsal aedeagal sheath heavily sclerotised (appearing entirely closed); dorsal aedeagal sheath short, sperm sac entirely free; sperm sac appearing
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<normalizedToken originalValue="±">+/-</normalizedToken>
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heavily sclerotised.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="39" type="description female">
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<paragraph pageId="6" pageNumber="39">Description Female:</paragraph>
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<paragraph pageId="6" pageNumber="39">unknown.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="7" lastPageNumber="40" pageId="6" pageNumber="39" type="material examined">
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<paragraph pageId="6" pageNumber="39">Material examined:</paragraph>
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<paragraph lastPageId="7" lastPageNumber="40" pageId="6" pageNumber="39">
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<materialsCitation ID-GBIF-Occurrence="2818988337" collectingDate="1962-03-29" collectingDateMax="1962-04-04" collectingDateMin="1962-03-29" collectionCode="ZSMC" collectorName="G. Ebert & H. Falkner" country="Nepal" elevation="0" latitude="27.583334" location="Megouli (= Megauli), Rapti Valley" longLatPrecision="20" longitude="84.23333" specimenCode="AAM- 000113" specimenCount="1" specimenCount-male="1" stateProvince="Bagmati" typeStatus="paratype">
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<collectingCountry name="Nepal" value="Nepal">Nepal</collectingCountry>
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:
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<collectingRegion>Bagmati</collectingRegion>
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:
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<specimenCount type="male">1 ♂</specimenCount>
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<location LSID="urn:lsid:plazi:treatment:15C93CF3D6F07077DF29E6901470A158:9275077FE66C73EB095564AB63A24B7A" country="Nepal" latitude="27.583334" longLatPrecision="20" longitude="84.23333" name="Megouli (= Megauli), Rapti Valley" stateProvince="Bagmati">Megouli (= Megauli), Rapti Valley</location>
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,
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<geoCoordinate degrees="27" direction="north" minutes="35" orientation="latitude" precision="15" seconds="00" value="27.583334">27°35'00"N</geoCoordinate>
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;
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<geoCoordinate degrees="084" direction="east" minutes="14" orientation="longitude" precision="15" seconds="00" value="84.23333">084°14'00"E</geoCoordinate>
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,
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<date value="1962-03-29" valueMax="1962-04-04" valueMin="1962-03-29">
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<collectingDate value="1962-03-29" valueMax="1962-04-04" valueMin="1962-03-29">
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29.iii.-4.iv.
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<quantity metricMagnitude="3" metricUnit="m" metricValue="1.9622999999999997" unit="m" value="1962.3">
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1962,
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<elevation>300 m</elevation>
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</quantity>
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</collectingDate>
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</date>
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,
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<collectorName>G. Ebert</collectorName>
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<collectorName>H. Falkner</collectorName>
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(
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<specimenCode>
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AAM-
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<pageBreakToken pageId="7" pageNumber="40" start="start">000113</pageBreakToken>
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</specimenCode>
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<typeStatus>paratype</typeStatus>
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,
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<collectionCode collectionName="Germany, Muenchen [= Munich], Zoologische Staatssammlung">ZSMC</collectionCode>
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)
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</materialsCitation>
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;
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<materialsCitation ID-GBIF-Occurrence="2818988312" collectingDate="1967-04-16" collectionCode="ZSMC" collectorName="Dierl & Forster & Schacht" country="Nepal" elevation="200" latitude="27.583334" location="Jhawani, Rapti Valley" longLatPrecision="20" longitude="84.51667" specimenCode="AAM-000115, AAM-000114" specimenCount="2" specimenCount-male="2" stateProvince="Bagmati" typeStatus="holotype">
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<specimenCount type="male">2 ♂</specimenCount>
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<location LSID="urn:lsid:plazi:treatment:15C93CF3D6F07077DF29E6901470A158:313B67688E9B6C158FB27989447BA746" country="Nepal" latitude="27.583334" longLatPrecision="20" longitude="84.51667" name="Jhawani, Rapti Valley" stateProvince="Bagmati">Jhawani, Rapti Valley</location>
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,
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<geoCoordinate degrees="27" direction="north" minutes="35" orientation="latitude" precision="15" seconds="00" value="27.583334">27°35'00"N</geoCoordinate>
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;
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<geoCoordinate degrees="084" direction="east" minutes="31" orientation="longitude" precision="15" seconds="00" value="84.51667">084°31'00"E</geoCoordinate>
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,
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<quantity metricMagnitude="2" metricUnit="m" metricValue="2.0" unit="m" value="200.0">
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<elevation metricMagnitude="2" metricUnit="m" metricValue="2.0" unit="m" value="200.0">200 m</elevation>
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</quantity>
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,
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<date value="1967-04-16">
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<collectingDate value="1967-04-16">16.iv.1967</collectingDate>
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</date>
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,
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<collectorName>Dierl</collectorName>
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<collectorName>Forster</collectorName>
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<collectorName>Schacht</collectorName>
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(
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<specimenCode>AAM-000115</specimenCode>
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1 ♂
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<typeStatus>holotype</typeStatus>
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,
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<specimenCode>AAM-000114</specimenCode>
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<typeStatus>paratype</typeStatus>
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,
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<collectionCode collectionName="Germany, Muenchen [= Munich], Zoologische Staatssammlung">ZSMC</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="40" type="type locality and distribution">
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<paragraph pageId="7" pageNumber="40">
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<typeStatus>Type</typeStatus>
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locality and distribution:
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</paragraph>
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<paragraph pageId="7" pageNumber="40">
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Jhawani (
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<geoCoordinate degrees="27" direction="north" minutes="35" orientation="latitude" precision="15" seconds="00" value="27.583334">
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27°35'00"
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<collectionCode>N</collectionCode>
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</geoCoordinate>
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;
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<geoCoordinate degrees="084" direction="east" minutes="31" orientation="longitude" precision="15" seconds="00" value="84.51667">
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084°31'00"
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<collectionCode>E</collectionCode>
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</geoCoordinate>
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), Nepal (Fig. 46). Biodiversity hotspot/high-biodiversity wilderness area: Himalaya/-.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="40" type="remarks">
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<paragraph pageId="7" pageNumber="40">Remarks:</paragraph>
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<paragraph pageId="7" pageNumber="40">
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The particular shape of the tip of the aedeagus, which is not tapered distally (Fig. 4) as in other members of
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<taxonomicName class="Insecta" family="Mydidae" genus="Leptomydas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptomydas" order="Diptera" pageId="7" pageNumber="40" phylum="Arthropoda" rank="genus">Leptomydas</taxonomicName>
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, is generally indicative of a species of
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<taxonomicName class="Insecta" family="Mydidae" genus="Eremomidas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eremomidas" order="Diptera" pageId="7" pageNumber="40" phylum="Arthropoda" rank="genus">Eremomidas</taxonomicName>
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Semenov, 1896 as suggested by Richter and Ovtshinnikova (1996) and
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<bibRefCitation author="Richter, VA" editor="Papp, L" journalOrPublisher="Science Herald, Budapest" pageId="32" pageNumber="65" pagination="539 - 547" title="Family Mydidae" volumeTitle="Contribution to a Manual of Palaearctic Diptera (with special reference to flies of economic importance). Volume 2, Nematocera and Lower Brachycera" year="1997">Richter (1997)</bibRefCitation>
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. However, the proboscis of this species is projecting well-beyond the fronto-clypeal suture, a feature considered to be specific to
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<taxonomicName class="Insecta" family="Mydidae" genus="Leptomydas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptomydas" order="Diptera" pageId="7" pageNumber="40" phylum="Arthropoda" rank="genus">Leptomydas</taxonomicName>
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. The correct generic placement of this species needs to be established with a phylogenetic analysis of
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<taxonomicName family="Mydidae" lsidName="" pageId="7" pageNumber="40" rank="family">Mydidae</taxonomicName>
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genera that is currently in preparation by the author.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |