216 lines
33 KiB
XML
216 lines
33 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.923.47104" ID-GBIF-Dataset="1e711d27-f2ea-4e62-9b52-19f33fa223b9" ID-PMC="PMC7142161" ID-Pensoft-Pub="1313-2970-923-65" ID-Pensoft-UUID="5A90240DCD655B319081ECB2F39B90B3" ID-PubMed="32292271" ID-ZooBank="B10E9E718A434560BBAC47F52E401045" ModsDocID="1313-2970-923-65" checkinTime="1585826883453" checkinUser="pensoft" docAuthor="Miller, Kelly B." docDate="2020" docId="9CD65445421C5F8CAE779D0522C38D60" docLanguage="en" docName="ZooKeys 923: 65-77" docOrigin="ZooKeys 923" docSource="http://dx.doi.org/10.3897/zookeys.923.47104" docTitle="Desmopachria manco Miller 2020, sp. nov." docType="treatment" docUuid="937F3475-5EDC-4269-B1FB-623F3DC78BC9" docUuidSource="ZooBank" docVersion="5" id="5A90240DCD655B319081ECB2F39B90B3" lastPageNumber="65" masterDocId="5A90240DCD655B319081ECB2F39B90B3" masterDocTitle="Two new species of Desmopachria Babington, 1841 in the D. convexa species group (Coleoptera, Adephaga, Dytiscidae, Hydroporinae, Hyphydrini)" masterLastPageNumber="77" masterPageNumber="65" pageNumber="65" updateTime="1701286543009" updateUser="admin">
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<mods:titleInfo id="9B9702AB4A7F5B61F361DD869582290A">
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<mods:title id="E363F48EF196E1F8A403D08639F2D02B">Two new species of Desmopachria Babington, 1841 in the D. convexa species group (Coleoptera, Adephaga, Dytiscidae, Hydroporinae, Hyphydrini)</mods:title>
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<mods:name id="A8B1C0699569C3CB96A21B93B1CDFF18" type="personal">
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<mods:roleTerm id="45370FB0E034C247C81327C71A679900">Author</mods:roleTerm>
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<mods:namePart id="1B27C89420529A47E8EE14371DC30623">Miller, Kelly B.</mods:namePart>
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<mods:affiliation id="842661B61AD242670ECAB2C576497D46">Department of Biology and Museum of Southwestern Biology, University of New Mexico, Albuquerque, NM 87131 - 0001 USA</mods:affiliation>
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<mods:nameIdentifier id="1C004EE9BED05AB5131053233A1914C1" type="email">kbmiller@unm.edu</mods:nameIdentifier>
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<mods:typeOfResource id="B1A7117E0DE9EF56126F4E969FB30B7F">text</mods:typeOfResource>
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<mods:title id="56FDD4FEAE78E80BE119F107E48D529E">ZooKeys</mods:title>
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<mods:part id="E97E1551208AEB4E6C8E2D8FD96EDB7F">
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<mods:date id="364E34ED4598B04097A8410A8FF83926">2020</mods:date>
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<mods:detail id="ADBC63F15BD7CCE64BB6F85D4F6469DB" type="volume">
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<mods:number id="CDCE8A8DFC70B1022812E6A40DE8A397">923</mods:number>
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<mods:start id="7321B85E387A7542CC22F326469F02C1">65</mods:start>
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<mods:end id="70AE24F3ED797EA9C27729D557E2A337">77</mods:end>
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<mods:location id="20E5EE27678BB136E3AF72E92AD07506">
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<mods:url id="CD740E5DEFAFBC54407912EFA7A0AFB3">http://dx.doi.org/10.3897/zookeys.923.47104</mods:url>
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<mods:classification id="B697103817C348796524628F5AE99803">journal article</mods:classification>
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<mods:identifier id="1AE2F4158E32F399587859E84709D46D" type="DOI">http://dx.doi.org/10.3897/zookeys.923.47104</mods:identifier>
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<treatment id="9CD65445421C5F8CAE779D0522C38D60" ID-GBIF-Taxon="163421240" LSID="urn:lsid:zoobank.org:act:937F3475-5EDC-4269-B1FB-623F3DC78BC9" httpUri="http://treatment.plazi.org/id/9CD65445421C5F8CAE779D0522C38D60" lastPageNumber="65" pageId="0" pageNumber="65">
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<subSubSection id="1A29AA4785BDBC645D7021ADB2845ADE" pageId="0" pageNumber="65" type="nomenclature">
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<paragraph id="1A64FB9C15126F9C86462C1E8EE40EE3" pageId="0" pageNumber="65">
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<taxonomicName id="E1A9E3A0140436C8A10D4EFD0E4B7382" ID-CoL="8LZ4N" LSID="9CD65445-421C-5F8C-AE77-9D0522C38D60" authority="Miller, 2020" authorityName="Miller" authorityYear="2020" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria manco" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="manco" status="sp. nov.">Desmopachria manco</taxonomicName>
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<taxonomicNameLabel id="AD020D5DF66AD7462090CF5B4870309B" pageId="0" pageNumber="65">sp. nov.</taxonomicNameLabel>
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<figureCitation id="DEEA37754EEAD3CDF2DFEAC32D0AFA80" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Desmopachria species. 1 - 5 D. manco 1 habitus 2 - 5 male genitalia 2 male median lobe, right lateral aspect 3 male median lobe, ventral aspect 4 male left lateral lobe, ventral aspect 5 male left lateral lobe, left lateral aspect 6 - 10 D. mortimer 6 habitus 7 - 10 male genitalia 7 male median lobe, right lateral aspect 8 male median lobe, ventral aspect 9 male left lateral lobe, ventral aspect 10 male left lateral lobe, left lateral aspect. Scale bars: 1.0 mm (1, 6)." figureDoi="10.3897/zookeys.923.47104.figures1-10" httpUri="https://binary.pensoft.net/fig/396455" pageId="0" pageNumber="65">Figures 1-5</figureCitation>
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<figureCitation id="C973283717794D5F43C26909C01EF3F7" captionStart="Figures 58, 59" captionStartId="F4" captionText="Figures 58, 59. Desmopachria new species, distribution. 58 D. manco (South America). 59 D. mortimer (Central America)." figureDoi="10.3897/zookeys.923.47104.figures58-59" httpUri="https://binary.pensoft.net/fig/396458" pageId="0" pageNumber="65">, 58</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="65A386086E471D8D46F4E9F57DDE3C88" pageId="0" pageNumber="65" type="type locality">
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<paragraph id="4BCF66B1AE2249D74D1C25999F0EC7AF" pageId="0" pageNumber="65">Type locality.</paragraph>
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<paragraph id="3366F9AC81F0EBA601E85FD9E063B58A" pageId="0" pageNumber="65">
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Guyana, Region IX, Parabara, trail to mines,
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<geoCoordinate id="80AFC01DCEB4A68221D348122D64557F" degrees="2" direction="north" minutes="05.095" orientation="latitude" precision="1" value="2.0849166">2°05.095'N</geoCoordinate>
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,
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<geoCoordinate id="BA99718749198395C451BD76DC7F4957" degrees="59" direction="west" minutes="14.174" orientation="longitude" precision="1" value="-59.236233">59°14.174'W</geoCoordinate>
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, 250m.
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</paragraph>
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</subSubSection>
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<subSubSection id="47B978C9E8B11839BF2B7232518E9FB3" pageId="0" pageNumber="65" type="type material">
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<paragraph id="61D3575D93F95474CAB72A68C22EB59F" pageId="0" pageNumber="65">Type material.</paragraph>
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<paragraph id="1B5DD23D7C461D6B3BF16E7062D85B41" pageId="0" pageNumber="65">
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Holotype in CSBD (currently in KUNHM, see above), male labeled, "Guyana: Region IX
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<geoCoordinate id="67C2F5DFFBE72D85E4EF4741A9570AB9" degrees="2" direction="north" minutes="05.095" orientation="latitude" precision="1" value="2.0849166">2°05.095'N</geoCoordinate>
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,
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<geoCoordinate id="05CA52C0E315ED34CCE87CFE4D25B609" degrees="59" direction="west" minutes="14.174" orientation="longitude" precision="1" value="-59.236233">59°14.174'W</geoCoordinate>
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, 250 m Parabara, Trail to mines detrital pools in forest leg. Short, Isaacs, Salisbury 2.xi.2013; GY13-1102-01A/ SEMC1271259 KUNHM-ENT/ Holotype
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<taxonomicName id="FA75CBBB2E46F4D7B48B881EDB26C40C" authorityName="Miller" authorityYear="2020" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria manco" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="manco">
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<emphasis id="4A32AD2F61C7A9C50C258252EB5DB442" italics="true" pageId="0" pageNumber="65">Desmopachria manco</emphasis>
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</taxonomicName>
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Miller, 2020 [red label with black line border]." Paratypes, 1 labeled, "Guyana: Region IX
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<geoCoordinate id="2687D0E4CD05C0CC9818D07DA3D3C640" degrees="2" direction="north" minutes="52.204" orientation="latitude" precision="1" value="2.8700666">2°52.204'N</geoCoordinate>
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,
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<geoCoordinate id="C508D0DF0456C22A2716035AB2649240" degrees="59" direction="west" minutes="55.003" orientation="longitude" precision="1" value="-59.916718">59°55.003'W</geoCoordinate>
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, 124 m nr. Kusad Mts., marshy area leg. Short, Isaacs, Salisbury 27x.2013; GY13-1027-01A/ SEMC1271270 KUNHM-ENT [barcode label]/ Paratype
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<taxonomicName id="1227D08153EF02495E79F480C2464EA1" authorityName="Miller" authorityYear="2020" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria manco" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="manco">
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<emphasis id="32B66AFD9372E9B3946E10C2C4D460AD" italics="true" pageId="0" pageNumber="65">Desmopachria manco</emphasis>
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</taxonomicName>
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Miller, 2020 [blue label with black line border]."
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</paragraph>
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<paragraph id="2B5BA27FB47E38050638F3F33C3FDE4D" pageId="0" pageNumber="65">The paratype specimen was not dissected but it has the same color pattern, size, and other features as the holotype. It is assigned to this species even though it is from some geographic distance away.</paragraph>
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</subSubSection>
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<subSubSection id="A63AD5EE183644902990A96E8E18BFAA" pageId="0" pageNumber="65" type="diagnosis">
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<paragraph id="E61EB701BFC48C19E5E990722E19BA4D" pageId="0" pageNumber="65">Diagnosis.</paragraph>
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<paragraph id="81C5121D4EE9B05996B5AC525F9CF026" pageId="0" pageNumber="65">
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This is an extremely small species among
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<taxonomicName id="FC261E0C108665736C1131D40A3BBAC3" family="Dytiscidae" lsidName="" pageId="0" pageNumber="65" rank="family">Dytiscidae</taxonomicName>
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and even among
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<taxonomicName id="C51AC6A6EA2968755DF502FDC4D2DADB" authorityName="Babington" authorityYear="1841" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="genus">
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<emphasis id="C2B19F9A3A29D4473DF104B01FE56C91" italics="true" pageId="0" pageNumber="65">Desmopachria</emphasis>
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</taxonomicName>
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(Fig.
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<figureCitation id="04B2E673D7436F4CF5EF69C7190C4B79" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Desmopachria species. 1 - 5 D. manco 1 habitus 2 - 5 male genitalia 2 male median lobe, right lateral aspect 3 male median lobe, ventral aspect 4 male left lateral lobe, ventral aspect 5 male left lateral lobe, left lateral aspect 6 - 10 D. mortimer 6 habitus 7 - 10 male genitalia 7 male median lobe, right lateral aspect 8 male median lobe, ventral aspect 9 male left lateral lobe, ventral aspect 10 male left lateral lobe, left lateral aspect. Scale bars: 1.0 mm (1, 6)." figureDoi="10.3897/zookeys.923.47104.figures1-10" httpUri="https://binary.pensoft.net/fig/396455" pageId="0" pageNumber="65">1</figureCitation>
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; TL = 1.2-1.3 mm). The dorsal diffuse maculae on the elytra are characteristic (Fig.
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<figureCitation id="C91198E2AC330CD6AFE9C2C869F35D08" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Desmopachria species. 1 - 5 D. manco 1 habitus 2 - 5 male genitalia 2 male median lobe, right lateral aspect 3 male median lobe, ventral aspect 4 male left lateral lobe, ventral aspect 5 male left lateral lobe, left lateral aspect 6 - 10 D. mortimer 6 habitus 7 - 10 male genitalia 7 male median lobe, right lateral aspect 8 male median lobe, ventral aspect 9 male left lateral lobe, ventral aspect 10 male left lateral lobe, left lateral aspect. Scale bars: 1.0 mm (1, 6)." figureDoi="10.3897/zookeys.923.47104.figures1-10" httpUri="https://binary.pensoft.net/fig/396455" pageId="0" pageNumber="65">1</figureCitation>
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). The male genitalia include a dispositively diagnostic set of features (Figs
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<figureCitation id="65C13672235BF4B7277CE20C7FC8AA02" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Desmopachria species. 1 - 5 D. manco 1 habitus 2 - 5 male genitalia 2 male median lobe, right lateral aspect 3 male median lobe, ventral aspect 4 male left lateral lobe, ventral aspect 5 male left lateral lobe, left lateral aspect 6 - 10 D. mortimer 6 habitus 7 - 10 male genitalia 7 male median lobe, right lateral aspect 8 male median lobe, ventral aspect 9 male left lateral lobe, ventral aspect 10 male left lateral lobe, left lateral aspect. Scale bars: 1.0 mm (1, 6)." figureDoi="10.3897/zookeys.923.47104.figures1-10" httpUri="https://binary.pensoft.net/fig/396455" pageId="0" pageNumber="65">2-5</figureCitation>
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) and place the species in the
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<taxonomicName id="F1426C27498F29E2D2DBA0CB70559640" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria convexa-signata" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="convexa-signata">
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<emphasis id="123B95625080489A2D1E2D41D0F525AE" italics="true" pageId="0" pageNumber="65">Desmopachria convexa-signata</emphasis>
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</taxonomicName>
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subgroup (see above). The median lobe is short (Figs
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<figureCitation id="F4AA518E06B8800F711F5DE1AB0542DC" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Desmopachria species. 1 - 5 D. manco 1 habitus 2 - 5 male genitalia 2 male median lobe, right lateral aspect 3 male median lobe, ventral aspect 4 male left lateral lobe, ventral aspect 5 male left lateral lobe, left lateral aspect 6 - 10 D. mortimer 6 habitus 7 - 10 male genitalia 7 male median lobe, right lateral aspect 8 male median lobe, ventral aspect 9 male left lateral lobe, ventral aspect 10 male left lateral lobe, left lateral aspect. Scale bars: 1.0 mm (1, 6)." figureDoi="10.3897/zookeys.923.47104.figures1-10" httpUri="https://binary.pensoft.net/fig/396455" pageId="0" pageNumber="65">2</figureCitation>
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,
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<figureCitation id="62A31E2018F81BD852EF39C3709A0ACF" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Desmopachria species. 1 - 5 D. manco 1 habitus 2 - 5 male genitalia 2 male median lobe, right lateral aspect 3 male median lobe, ventral aspect 4 male left lateral lobe, ventral aspect 5 male left lateral lobe, left lateral aspect 6 - 10 D. mortimer 6 habitus 7 - 10 male genitalia 7 male median lobe, right lateral aspect 8 male median lobe, ventral aspect 9 male left lateral lobe, ventral aspect 10 male left lateral lobe, left lateral aspect. Scale bars: 1.0 mm (1, 6)." figureDoi="10.3897/zookeys.923.47104.figures1-10" httpUri="https://binary.pensoft.net/fig/396455" pageId="0" pageNumber="65">3</figureCitation>
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). In lateral aspect it is irregular in shape, medially expanded ventrally and with the apical portion slender, short, slightly curved dorsad and apically narrowly rounded (Fig.
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<figureCitation id="A83535306B876AE362B0D58D4B4FE59C" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Desmopachria species. 1 - 5 D. manco 1 habitus 2 - 5 male genitalia 2 male median lobe, right lateral aspect 3 male median lobe, ventral aspect 4 male left lateral lobe, ventral aspect 5 male left lateral lobe, left lateral aspect 6 - 10 D. mortimer 6 habitus 7 - 10 male genitalia 7 male median lobe, right lateral aspect 8 male median lobe, ventral aspect 9 male left lateral lobe, ventral aspect 10 male left lateral lobe, left lateral aspect. Scale bars: 1.0 mm (1, 6)." figureDoi="10.3897/zookeys.923.47104.figures1-10" httpUri="https://binary.pensoft.net/fig/396455" pageId="0" pageNumber="65">2</figureCitation>
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). In ventral aspect it is very broad, basally deeply U-shaped, apically deeply bifid, each ramus broad basally, apically narrowed and slightly curved laterad (Fig.
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<figureCitation id="7AF02761968343FEC77FC124356911D6" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Desmopachria species. 1 - 5 D. manco 1 habitus 2 - 5 male genitalia 2 male median lobe, right lateral aspect 3 male median lobe, ventral aspect 4 male left lateral lobe, ventral aspect 5 male left lateral lobe, left lateral aspect 6 - 10 D. mortimer 6 habitus 7 - 10 male genitalia 7 male median lobe, right lateral aspect 8 male median lobe, ventral aspect 9 male left lateral lobe, ventral aspect 10 male left lateral lobe, left lateral aspect. Scale bars: 1.0 mm (1, 6)." figureDoi="10.3897/zookeys.923.47104.figures1-10" httpUri="https://binary.pensoft.net/fig/396455" pageId="0" pageNumber="65">3</figureCitation>
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). The lateral lobe in ventral aspect is elongate, broad, of subequal width throughout to a broadly truncate apex, with the subapical articulable lobe small and broad (Fig.
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<figureCitation id="D6EABE1EE27D01C95A71602AC08E8B48" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Desmopachria species. 1 - 5 D. manco 1 habitus 2 - 5 male genitalia 2 male median lobe, right lateral aspect 3 male median lobe, ventral aspect 4 male left lateral lobe, ventral aspect 5 male left lateral lobe, left lateral aspect 6 - 10 D. mortimer 6 habitus 7 - 10 male genitalia 7 male median lobe, right lateral aspect 8 male median lobe, ventral aspect 9 male left lateral lobe, ventral aspect 10 male left lateral lobe, left lateral aspect. Scale bars: 1.0 mm (1, 6)." figureDoi="10.3897/zookeys.923.47104.figures1-10" httpUri="https://binary.pensoft.net/fig/396455" pageId="0" pageNumber="65">4</figureCitation>
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). The lateral lobe in lateral aspect is very broad basally with the apex slender, subapically slightly expanded on the dorsal margin and apically sharply pointed with the subapical articulable process short and slender (Fig.
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<figureCitation id="A24FFED8C6C4FEE0D6B0D3BD3CC44746" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Desmopachria species. 1 - 5 D. manco 1 habitus 2 - 5 male genitalia 2 male median lobe, right lateral aspect 3 male median lobe, ventral aspect 4 male left lateral lobe, ventral aspect 5 male left lateral lobe, left lateral aspect 6 - 10 D. mortimer 6 habitus 7 - 10 male genitalia 7 male median lobe, right lateral aspect 8 male median lobe, ventral aspect 9 male left lateral lobe, ventral aspect 10 male left lateral lobe, left lateral aspect. Scale bars: 1.0 mm (1, 6)." figureDoi="10.3897/zookeys.923.47104.figures1-10" httpUri="https://binary.pensoft.net/fig/396455" pageId="0" pageNumber="65">5</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="5C3BAD6CC4194EC8891EAFE01AF0EF15" pageId="0" pageNumber="65" type="description">
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<paragraph id="1389119774B8E58A956B730FB21F4DD5" pageId="0" pageNumber="65">Description.</paragraph>
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<paragraph id="068BB266090800FB0EA4598A90DB1CAE" pageId="0" pageNumber="65">
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<emphasis id="B8E952034E683030118EF15B8E43B53C" bold="true" italics="true" pageId="0" pageNumber="65">Measurements.</emphasis>
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TL = 1.2-1.3 mm, GW = 0.8 mm, PW = 0.7 mm, HW = 0.4-0.5 mm, EW = 0.2 mm, TL/GW = 1.9, HW/EW = 2.0. Body round, subspherical, lateral margins continuous between pronotum and elytron (Fig.
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<figureCitation id="8A789A4C76E0E17BB7A69F749A725D5B" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Desmopachria species. 1 - 5 D. manco 1 habitus 2 - 5 male genitalia 2 male median lobe, right lateral aspect 3 male median lobe, ventral aspect 4 male left lateral lobe, ventral aspect 5 male left lateral lobe, left lateral aspect 6 - 10 D. mortimer 6 habitus 7 - 10 male genitalia 7 male median lobe, right lateral aspect 8 male median lobe, ventral aspect 9 male left lateral lobe, ventral aspect 10 male left lateral lobe, left lateral aspect. Scale bars: 1.0 mm (1, 6)." figureDoi="10.3897/zookeys.923.47104.figures1-10" httpUri="https://binary.pensoft.net/fig/396455" pageId="0" pageNumber="65">1</figureCitation>
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), dorsoventrally broad.
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</paragraph>
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<paragraph id="904306E6FFB4689A1888F26A0DCF4CB7" pageId="0" pageNumber="65">
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<emphasis id="1BACE21EBD449A23DC72CD03C6F032F7" bold="true" italics="true" pageId="0" pageNumber="65">Coloration</emphasis>
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(Fig.
|
||
<figureCitation id="41A37C830A0DD1E7E31EA03B75F2D30B" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Desmopachria species. 1 - 5 D. manco 1 habitus 2 - 5 male genitalia 2 male median lobe, right lateral aspect 3 male median lobe, ventral aspect 4 male left lateral lobe, ventral aspect 5 male left lateral lobe, left lateral aspect 6 - 10 D. mortimer 6 habitus 7 - 10 male genitalia 7 male median lobe, right lateral aspect 8 male median lobe, ventral aspect 9 male left lateral lobe, ventral aspect 10 male left lateral lobe, left lateral aspect. Scale bars: 1.0 mm (1, 6)." figureDoi="10.3897/zookeys.923.47104.figures1-10" httpUri="https://binary.pensoft.net/fig/396455" pageId="0" pageNumber="65">1</figureCitation>
|
||
). Head and pronotum yellow. Elytron orange, with distinct but weakly margined maculae at humeral angle, anterolaterally extending to near suture, lateromedially and apically; surface not iridescent. Ventral surfaces and appendages yellow.
|
||
</paragraph>
|
||
<paragraph id="3A1E08943E8808B8937C7421948511EA" pageId="0" pageNumber="65">
|
||
<emphasis id="FAE2D2F7D36972D0B9528A089BECE1A7" bold="true" italics="true" pageId="0" pageNumber="65">Sculpture and structure.</emphasis>
|
||
Head (Fig.
|
||
<figureCitation id="019352B6A9BF13CCA1F55BABD64B216E" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Desmopachria species. 1 - 5 D. manco 1 habitus 2 - 5 male genitalia 2 male median lobe, right lateral aspect 3 male median lobe, ventral aspect 4 male left lateral lobe, ventral aspect 5 male left lateral lobe, left lateral aspect 6 - 10 D. mortimer 6 habitus 7 - 10 male genitalia 7 male median lobe, right lateral aspect 8 male median lobe, ventral aspect 9 male left lateral lobe, ventral aspect 10 male left lateral lobe, left lateral aspect. Scale bars: 1.0 mm (1, 6)." figureDoi="10.3897/zookeys.923.47104.figures1-10" httpUri="https://binary.pensoft.net/fig/396455" pageId="0" pageNumber="65">1</figureCitation>
|
||
) broad, anteriorly produced in rounded margin; anterior margin of clypeus margined with conspicuous, continuous, flattened bead; surface of head shiny, very finely and sparsely punctate; eyes large (HW/EW = 1.2-1.4); antennae short, scape and pedicel relatively large and rounded, flagellomere III long and slender, apically expanded, antennomeres IV-X short and broad, lobe at anterodorsal angle, antennomere XI elongate, apically pointed. Pronotum short, lateral margins short, broadly curved with narrow, even bead; surface shiny, nearly impunctate medially, more but sparsely punctate along anterior and posterior margins, punctation variable, fine to course. Elytron moderately broad, laterally broadly curved; surface impunctate. Prosternum extremely short, longitudinally compressed, medially slightly carinate; prosternal process slender anteriorly, with distinctive, small medial tubercle, apically short and broad, concave, apically broadly pointed. Metaventrite broad and evenly smoothly convex medially, surface shiny, impunctate, anteromedially with curved transverse carina between posterior margins of mesocoxal cavities; metaventrite wings extremely slender. Metacoxa with medial portion short, about 1/3 length of metaventrite medially, metacoxal lines distinctly divergent anteriorly; lateral portion of metacoxa extremely large, anteriorly strongly expanded; surface shiny, impunctate, but slightly rugulose medially. Metatrochanter large, longer than ventral margin of metafemur anterior to metatrochanter apex; legs otherwise not noticeably modified. Abdomen with surfaces shiny and smooth, impunctate.
|
||
</paragraph>
|
||
<paragraph id="7EB259282BB078E067C023D8DEF671B0" pageId="0" pageNumber="65">
|
||
<emphasis id="B15181362D594FA187C242D2D986989D" bold="true" italics="true" pageId="0" pageNumber="65">Male genitalia.</emphasis>
|
||
Male median lobe in lateral aspect short, medially somewhat expanded on ventral margin, apically convergent to sharply angulate apex (Fig.
|
||
<figureCitation id="29696CDFFE2CF52B50963118D7A650AE" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Desmopachria species. 1 - 5 D. manco 1 habitus 2 - 5 male genitalia 2 male median lobe, right lateral aspect 3 male median lobe, ventral aspect 4 male left lateral lobe, ventral aspect 5 male left lateral lobe, left lateral aspect 6 - 10 D. mortimer 6 habitus 7 - 10 male genitalia 7 male median lobe, right lateral aspect 8 male median lobe, ventral aspect 9 male left lateral lobe, ventral aspect 10 male left lateral lobe, left lateral aspect. Scale bars: 1.0 mm (1, 6)." figureDoi="10.3897/zookeys.923.47104.figures1-10" httpUri="https://binary.pensoft.net/fig/396455" pageId="0" pageNumber="65">1</figureCitation>
|
||
). Median lobe in ventral aspect broad, base in broad
|
||
<normalizedToken id="1FBCC257817A22294C4E9BC96A5C51A7" originalValue="“U”">"U"</normalizedToken>
|
||
shape, apically deeply bifid, each ramus elongate, broad basally, and apically pointed (Fig.
|
||
<figureCitation id="0954BC6100072F6C17CB5A746E997343" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Desmopachria species. 1 - 5 D. manco 1 habitus 2 - 5 male genitalia 2 male median lobe, right lateral aspect 3 male median lobe, ventral aspect 4 male left lateral lobe, ventral aspect 5 male left lateral lobe, left lateral aspect 6 - 10 D. mortimer 6 habitus 7 - 10 male genitalia 7 male median lobe, right lateral aspect 8 male median lobe, ventral aspect 9 male left lateral lobe, ventral aspect 10 male left lateral lobe, left lateral aspect. Scale bars: 1.0 mm (1, 6)." figureDoi="10.3897/zookeys.923.47104.figures1-10" httpUri="https://binary.pensoft.net/fig/396455" pageId="0" pageNumber="65">2</figureCitation>
|
||
). Lateral lobe in ventral aspect moderately broad, apically gently curved laterad, with apex subtruncate, subapical articulable process oblique and flattened (Fig.
|
||
<figureCitation id="D4B1EEEA76CC0DF4EA897409C01B7D8A" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Desmopachria species. 1 - 5 D. manco 1 habitus 2 - 5 male genitalia 2 male median lobe, right lateral aspect 3 male median lobe, ventral aspect 4 male left lateral lobe, ventral aspect 5 male left lateral lobe, left lateral aspect 6 - 10 D. mortimer 6 habitus 7 - 10 male genitalia 7 male median lobe, right lateral aspect 8 male median lobe, ventral aspect 9 male left lateral lobe, ventral aspect 10 male left lateral lobe, left lateral aspect. Scale bars: 1.0 mm (1, 6)." figureDoi="10.3897/zookeys.923.47104.figures1-10" httpUri="https://binary.pensoft.net/fig/396455" pageId="0" pageNumber="65">3</figureCitation>
|
||
). Lateral lobe in lateral aspect very broad basally, constricted medially, apically slender, slightly and broadly curved dorsad, apex pointed, subapically articulable lobe slender, directed ventrad (Fig.
|
||
<figureCitation id="C5800444561FF84739B76F5C0277535B" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Desmopachria species. 1 - 5 D. manco 1 habitus 2 - 5 male genitalia 2 male median lobe, right lateral aspect 3 male median lobe, ventral aspect 4 male left lateral lobe, ventral aspect 5 male left lateral lobe, left lateral aspect 6 - 10 D. mortimer 6 habitus 7 - 10 male genitalia 7 male median lobe, right lateral aspect 8 male median lobe, ventral aspect 9 male left lateral lobe, ventral aspect 10 male left lateral lobe, left lateral aspect. Scale bars: 1.0 mm (1, 6)." figureDoi="10.3897/zookeys.923.47104.figures1-10" httpUri="https://binary.pensoft.net/fig/396455" pageId="0" pageNumber="65">4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="BFCF7382437CF8210BC6828A7A0B1950" pageId="0" pageNumber="65" type="etymology">
|
||
<paragraph id="3341525714934DE69E98D984F00117B4" pageId="0" pageNumber="65">Etymology.</paragraph>
|
||
<paragraph id="671A3D0E266D10AFDE2C44C079F641B4" pageId="0" pageNumber="65">
|
||
This species is named
|
||
<taxonomicName id="E4EB57B67A10AB040FB25B6853C4E3ED" authorityName="Miller" authorityYear="2020" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="manco" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="manco">
|
||
<emphasis id="D741499E461FD0930B860E8BD55AE719" italics="true" pageId="0" pageNumber="65">manco</emphasis>
|
||
</taxonomicName>
|
||
, after Manco, the younger bounty hunter in the Sergio Leone film "For a Few Dollars More".
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A97CB21742A052DC00892B9F2A378435" pageId="0" pageNumber="65" type="distribution">
|
||
<paragraph id="EF21B332FAA550FBC734C91CC8F790F2" pageId="0" pageNumber="65">Distribution.</paragraph>
|
||
<paragraph id="254E1031DFC1957B080AB4241CBE2DEA" pageId="0" pageNumber="65">
|
||
This species is known from two localities in Guyana, Region IX (Fig.
|
||
<figureCitation id="25D5BD31B3555DB323ED6AFD76E52152" captionStart="Figures 58, 59" captionStartId="F4" captionText="Figures 58, 59. Desmopachria new species, distribution. 58 D. manco (South America). 59 D. mortimer (Central America)." figureDoi="10.3897/zookeys.923.47104.figures58-59" httpUri="https://binary.pensoft.net/fig/396458" pageId="0" pageNumber="65">58</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0F0ACD7FAF46CF440962710A14B0D931" pageId="0" pageNumber="65" type="habitat">
|
||
<paragraph id="DE7C713FF4547FCE6C42B9105EA7D732" pageId="0" pageNumber="65">Habitat.</paragraph>
|
||
<paragraph id="2DECA3503222BA96D109D2E092EAD338" pageId="0" pageNumber="65">The type and paratype were collected in "detrital pools" and a "marshy area."</paragraph>
|
||
<caption id="D1B0E84A8476BD07F3F30E4E5ECBF60F" doi="10.3897/zookeys.923.47104.figures1-10" httpUri="https://binary.pensoft.net/fig/396455" pageId="0" pageNumber="65" start="Figures 1–10" startId="F1">
|
||
<paragraph id="7B63A280080EFF1931EC5E02CBDB677A" pageId="0" pageNumber="65">
|
||
<emphasis id="5AAE8EB79620C3219AB682A3A07BBD43" bold="true" pageId="0" pageNumber="65">Figures 1-10.</emphasis>
|
||
<taxonomicName id="8B6F4AF7AC05F446693DAF8A47D12566" authorityName="Babington" authorityYear="1841" class="Insecta" family="Dytiscidae" genus="Desmopachria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmopachria" order="Coleoptera" pageId="0" pageNumber="65" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6106FE2439C52FEC298D492037A0BD09" italics="true" pageId="0" pageNumber="65">Desmopachria</emphasis>
|
||
</taxonomicName>
|
||
species.
|
||
<emphasis id="2EBE06117B564BA7EDEEABB69559C224" bold="true" pageId="0" pageNumber="65">1-5</emphasis>
|
||
<taxonomicName id="8DCFF5A87EC26EDF79497D1F507973DF" lsidName="D. manco" pageId="0" pageNumber="65" rank="species" species="manco">
|
||
<emphasis id="CC2518771E7949B2D6290E6A66047589" italics="true" pageId="0" pageNumber="65">D. manco</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="FF03E5DB048FDBD7B53337FA4262209D" bold="true" pageId="0" pageNumber="65">1</emphasis>
|
||
habitus
|
||
<emphasis id="6C6D8CB6EE06EED524E84B6F39709E59" bold="true" pageId="0" pageNumber="65">2-5</emphasis>
|
||
male genitalia
|
||
<emphasis id="F99605C991BE238687468A49B756B1ED" bold="true" pageId="0" pageNumber="65">2</emphasis>
|
||
male median lobe, right lateral aspect
|
||
<emphasis id="6835D18139E81866B0BA1392EBE46820" bold="true" pageId="0" pageNumber="65">3</emphasis>
|
||
male median lobe, ventral aspect
|
||
<emphasis id="438393B1731A101C43D9CD620294599B" bold="true" pageId="0" pageNumber="65">4</emphasis>
|
||
male left lateral lobe, ventral aspect
|
||
<emphasis id="4784DA9C224E01B897B8063776E13DF1" bold="true" pageId="0" pageNumber="65">5</emphasis>
|
||
male left lateral lobe, left lateral aspect
|
||
<emphasis id="7DD46237E35695EC7562F0DC6CB2C1CD" bold="true" pageId="0" pageNumber="65">6-10</emphasis>
|
||
<taxonomicName id="B6A294350334513114353A3E0F657600" lsidName="D. mortimer" pageId="0" pageNumber="65" rank="species" species="mortimer">
|
||
<emphasis id="1D59F924A79EDAD5DF1EAF0E7ECD2970" italics="true" pageId="0" pageNumber="65">D. mortimer</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="F43C0656CC74679C9C877D439E8229F1" bold="true" pageId="0" pageNumber="65">6</emphasis>
|
||
habitus
|
||
<emphasis id="C267F685D83ACACAB45ADCD77EEDAD30" bold="true" pageId="0" pageNumber="65">7-10</emphasis>
|
||
male genitalia
|
||
<emphasis id="23A3E1688B9149CA841F1DDFE16C0A05" bold="true" pageId="0" pageNumber="65">7</emphasis>
|
||
male median lobe, right lateral aspect
|
||
<emphasis id="2D2F8694B7336C2C7B50115327056727" bold="true" pageId="0" pageNumber="65">8</emphasis>
|
||
male median lobe, ventral aspect
|
||
<emphasis id="F0BE7FE88285A658D255A280DB26AAB4" bold="true" pageId="0" pageNumber="65">9</emphasis>
|
||
male left lateral lobe, ventral aspect
|
||
<emphasis id="78D78BB06116FEBF866931CF67FBA52D" bold="true" pageId="0" pageNumber="65">10</emphasis>
|
||
male left lateral lobe, left lateral aspect. Scale bars: 1.0 mm (
|
||
<emphasis id="285966A605C07E6B4EA8AAB6FF5225C5" bold="true" pageId="0" pageNumber="65">1, 6</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |