245 lines
34 KiB
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245 lines
34 KiB
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<mods:title id="780EEFC111849F9D834C774A0F8692BF">The Luciolinae of S. E. Asia and the Australopacific region: a revisionary checklist (Coleoptera: Lampyridae) including description of three new genera and 13 new species</mods:title>
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<taxonomicName id="2394638951151878FF0EFDF8ED251EC6" ID-CoL="34RG" authority="Ballantyne 2009" authorityName="Ballantyne in Ballantyne & Lambkin" authorityYear="2009" box="[151,541,615,641]" class="Insecta" family="Lampyridae" genus="Asymmetricata" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="genus">
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<emphasis id="D6E0C41851151878FF0EFDF8ED251EC6" bold="true" box="[151,541,615,641]" pageId="46" pageNumber="47">
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<emphasis id="D6E0C41851151878FF0EFDF8EE741EC6" bold="true" box="[151,332,615,641]" italics="true" pageId="46" pageNumber="47">Asymmetricata</emphasis>
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<bibRefCitation id="800565FB51151878FECAFDF8ED251EC6" author="Ballantyne" box="[339,541,615,641]" firstAuthor="Ballantyne" pageId="46" pageNumber="47" pagination="1 - 188" refId="ref114542" refString="Ballantyne, L. A. & Lambkin, C. L. (2009) Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae). Zootaxa, 1997, 1 - 188." type="journal article" year="2009">Ballantyne 2009</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="AC8E4B8151151878FF0EFD14EE641EE2" box="[151,348,651,677]" pageId="46" pageNumber="47" type="description">
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<paragraph id="E42B180A51151878FF0EFD14EE641EE2" blockId="46.[151,541,615,677]" box="[151,348,651,677]" pageId="46" pageNumber="47">
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<figureCitation id="7CAF048F51151878FF0EFD14EFD41EE2" box="[151,236,651,677]" captionStart="FIGURES 9−16" captionStartId="12.[151,264,1864,1889]" captionTargetBox="[223,1364,166,1838]" captionTargetId="figure@12.[223,1364,166,1844]" captionTargetPageId="12" captionText="FIGURES 9−16. Key to genera characters 2. 9. Missimia flavida type male; 10 Luciola brahmina type male (MNHN); 11, 12 Luciola seriata type male (CMG); 13 Asymmetricata circumdata male; 14, 15 Kuantana menayah gen. et sp. nov.; 16 Curtos costipenis male. 9 anterior head (arrow indicates clypeus); 10, 11, 16 dorsal (arrows in 16 indicate humeral carina); 12 lateral tergite 8 aedeagal sheath and sclerites); 13 dorsal tergites 7 and 8; 14, 15, dorsal (14) and ventral tergite 8. Figure legends: AS aedeagal sheath; T7, 8 tergites 7, 8." figureDoi="http://doi.org/10.5281/zenodo.3508104" httpUri="https://zenodo.org/record/3508104/files/figure.png" pageId="46" pageNumber="47">Figs 13</figureCitation>
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,
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<figureCitation id="7CAF048F51151878FF61FD14EE641EE2" box="[248,348,651,677]" captionStart-0="FIGURES 139–147" captionStart-1="FIGURES 148–153" captionStart-2="FIGURES 154–164" captionStart-3="FIGURES 165−174" captionStart-4="FIGURES 175−179" captionStartId-0="49.[151,264,1924,1949]" captionStartId-1="50.[151,264,1359,1384]" captionStartId-2="52.[151,264,1906,1931]" captionStartId-3="53.[151,264,1924,1949]" captionStartId-4="55.[151,264,1713,1738]" captionTargetBox-0="[203,1385,166,1904]" captionTargetBox-1="[151,1435,172,1332]" captionTargetBox-2="[174,1435,166,1871]" captionTargetBox-3="[151,1322,166,1904]" captionTargetBox-4="[151,1435,166,1684]" captionTargetId-0="figure@49.[203,1385,166,1905]" captionTargetId-1="figure@50.[151,1435,166,1339]" captionTargetId-2="figure@52.[151,1435,166,1886]" captionTargetId-3="figure@53.[151,1333,166,1905]" captionTargetId-4="figure@55.[151,1435,166,1693]" captionTargetPageId-0="49" captionTargetPageId-1="50" captionTargetPageId-2="52" captionTargetPageId-3="55" captionTargetPageId-4="55" captionText-0="FIGURES 139–147. Asymmetricata spp. 139, 140 Luciola bicoloripes type male; 141–146 As. circumdata 141, 145–147 Tenasserim (Prague); 142, 143 Ranong (ANIC); 144 Thailand Mae Hong son (Prague); 139, 141, 143 ventral habitus; 140 type labels; 142, 144 dorsal habitus; 145–147 detail ventrites 6, 7." captionText-1="FIGURES 148–153. Asymmetricata circumdata female (Thailand, Chiang Mai, QSBG). 148 abdomen ventral; 149, 150 ovipositor, to left, and internal reproductive system (149 anterior end to left of page); 151 ovipositor ventral; 152, 153 median oviduct plate from above (152) and right (153). Figure legend: SDG spermatophore digesting gland. Scale lines are 1 mm except for 153 which is 0.5 mm." captionText-2="FIGURES 154–164. Asymmetricata spp. 154, 155, 158–164 Colophotia humeralis syntype male (NHML); 156, 157 male Nilgiri Hills labelled ‘doriae notatipennis’ (NHML). 154, 156, 157 dorsal habitus; 155 ventral habitus; 158, 159 abdomen only: 158 ventral; 159 dorsal; 160 type label; 161, 162 aedeagal sheath: 161 ventral; 162 dorsal; 164, 164 aedeagus: 163 dorsal; 164 left lateral. Scale lines are 1 mm." captionText-3="FIGURES 165−174. Luciola impressa holotype male (165−167, 171−174), cotype female (168−170). 165, 167, 168, 170 dorsal habitus (167 angled to show pronotal colour); 166, 169 ventral habitus; 171, 174 aedeagus: 171 ventral; 174 dorsal; 172 aedeagal sheath ventral; 173 type label." captionText-4="FIGURES 175−179. Asymmetricata ovalis female (Chiang Mai QSBG). 175 abdominal segments 7, 8, ventral; 176, 178, 179 reproductive system: 176 entire system SDG to right of picture, ovipositor to left; 177 female genitalia styli to foot of page 178 ventral outline of median oviduct plate with valvifers of ovipositor to right; 179 left lateral with median oviduct plate from side and ovipositor at bottom left.. Scale lines are 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3508166" figureDoi-1="http://doi.org/10.5281/zenodo.3508168" figureDoi-2="http://doi.org/10.5281/zenodo.3508170" figureDoi-3="http://doi.org/10.5281/zenodo.3508174" figureDoi-4="http://doi.org/10.5281/zenodo.3508176" httpUri-0="https://zenodo.org/record/3508166/files/figure.png" httpUri-1="https://zenodo.org/record/3508168/files/figure.png" httpUri-2="https://zenodo.org/record/3508170/files/figure.png" httpUri-3="https://zenodo.org/record/3508174/files/figure.png" httpUri-4="https://zenodo.org/record/3508176/files/figure.png" pageId="46" pageNumber="47">139−179</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="AC8E4B8151151878FF0EFD50EA521EAF" box="[151,1386,719,744]" pageId="46" pageNumber="47" type="reference_group">
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<paragraph id="E42B180A51151878FF0EFD50EA521EAF" blockId="46.[151,1386,719,744]" box="[151,1386,719,744]" pageId="46" pageNumber="47">
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<taxonomicName id="2394638951151878FF0EFD50EC291EA0" ID-CoL="34RG" authority="Ballantyne in Ballantyne & Lambkin 2009: 30" authorityName="Ballantyne in Ballantyne & Lambkin" authorityPageNumber="30" authorityYear="2009" box="[151,785,719,744]" class="Insecta" family="Lampyridae" genus="Asymmetricata" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="genus">
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<emphasis id="D6E0C41851151878FF0EFD50EE0B1EA0" bold="true" box="[151,307,719,743]" italics="true" pageId="46" pageNumber="47">Asymmetricata</emphasis>
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Ballantyne in
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<treatmentCitation id="65353E1B51151878FE51FD50EC291EA0" ID-DOI="http://doi.org/10.5281/zenodo.5324230" ID-Zenodo-Dep="5324230" author="Ballantyne, L. A. & Lambkin, C. L." box="[456,785,719,744]" httpUri="http://treatment.plazi.org/id/0394D665BE31FFA3FF3C510626BDEB22" page="30" pageId="46" pageNumber="47" year="2009">
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<bibRefCitation id="800565FB51151878FE51FD50EC291EA0" author="Ballantyne & Lambkin" box="[456,785,719,744]" firstAuthor="Ballantyne" pageId="46" pageNumber="47" pagination="1 - 188" refId="ref114542" refString="Ballantyne, L. A. & Lambkin, C. L. (2009) Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae). Zootaxa, 1997, 1 - 188." type="journal article" year="2009">Ballantyne & Lambkin 2009: 30</bibRefCitation>
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||
(figs 5–8, 108–114).
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<treatmentCitation id="65353E1B51151878FC72FD50EB401EA0" author="Yiu, V." box="[1003,1144,719,744]" page="88" pageId="46" pageNumber="47" year="2012">
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<bibRefCitation id="800565FB51151878FC72FD50EB401EA0" author="Yiu" box="[1003,1144,719,744]" firstAuthor="Yiu" pageId="46" pageNumber="47" refId="ref122578" refString="Yiu, V. (2012) Fireflies of Hong Kong. Popular Entomology Book Series No. 7. Hong Kong Entomological Society, Hong Kong, 96 pp." type="book" year="2012">Yiu, 2012: 88</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="65353E1B51151878FB1AFD50EBE71EA0" author="Yiu, V." box="[1155,1247,719,743]" page="88" pageId="46" pageNumber="47" year="2017">
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<bibRefCitation id="800565FB51151878FB1AFD50EBE71EA0" author="Yiu" box="[1155,1247,719,743]" firstAuthor="Yiu" pageId="46" pageNumber="47" pagination="67 - 119" refId="ref122609" refString="Yiu, V. (2017) A Study of Rhagophthalmidae and Lampyridae in Hong Kong with descriptions of new species (Coleoptera): Part 2. Lampyrid, 4, 67 - 119." type="journal article" year="2017">2017: 88</bibRefCitation>
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</treatmentCitation>
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.
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||
<treatmentCitation id="65353E1B51151878FB70FD4FEA5E1EAF" author="Fu, X. H." box="[1257,1382,719,744]" page="54" pageId="46" pageNumber="47" year="2014">
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<bibRefCitation id="800565FB51151878FB70FD4FEA5E1EAF" author="Fu" box="[1257,1382,719,744]" firstAuthor="Fu" pageId="46" pageNumber="47" refId="ref116462" refString="Fu, X. H. (2014) An illustrated handbook of Chinese fireflies. The Commercial Press, Beijing, 167 pp. [in Chinese]" type="book" year="2014">Fu 2014: 54</bibRefCitation>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="AC8E4B8151151878FF0EFC89EA241F77" box="[151,1308,789,816]" pageId="46" pageNumber="47" type="type_taxon">
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<paragraph id="E42B180A51151878FF0EFC89EA241F77" blockId="46.[151,1437,789,1536]" box="[151,1308,789,816]" pageId="46" pageNumber="47">
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<emphasis id="D6E0C41851151878FF0EFC89EE0C1F77" bold="true" box="[151,308,789,816]" pageId="46" pageNumber="47">
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<typeStatus id="3B2FA6A851151878FF0EFC89EFEA1F77" box="[151,210,790,816]" pageId="46" pageNumber="47">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName id="2394638951151878FEA5FC89EC381F77" ID-CoL="3WCPN" authority="(Motschulsky 1854)" baseAuthorityName="Motschulsky" baseAuthorityYear="1854" box="[316,768,789,816]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="circumdata">
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<emphasis id="D6E0C41851151878FEA5FC89ED2E1F77" box="[316,534,790,816]" italics="true" pageId="46" pageNumber="47">Luciola circumdata</emphasis>
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(
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<bibRefCitation id="800565FB51151878FDBCFC89EDC01F77" author="Motschulsky" box="[549,760,789,816]" firstAuthor="Motschulsky" pageId="46" pageNumber="47" pagination="47 - 62" refId="ref119634" refString="Motschulsky, V. de (1854) Lampyrides. Etudes Entomologiques, III, 47 - 62." type="journal article" year="1854">Motschulsky 1854</bibRefCitation>
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)
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</taxonomicName>
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designated by
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<bibRefCitation id="800565FB51151878FC30FC89EA201F77" author="Ballantyne & Lambkin" box="[937,1304,790,816]" firstAuthor="Ballantyne" pageId="46" pageNumber="47" pagination="1 - 188" refId="ref114542" refString="Ballantyne, L. A. & Lambkin, C. L. (2009) Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae). Zootaxa, 1997, 1 - 188." type="journal article" year="2009">Ballantyne & Lambkin 2009: 30</bibRefCitation>
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.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="AC8E4B8151151878FF5EFCA6EA75184F" pageId="46" pageNumber="47" type="diagnosis">
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<paragraph id="E42B180A51151878FF5EFCA6EA75184F" blockId="46.[151,1437,789,1536]" pageId="46" pageNumber="47">
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<emphasis id="D6E0C41851151878FF5EFCA6EE7A1F13" bold="true" box="[199,322,825,852]" pageId="46" pageNumber="47">Diagnosis.</emphasis>
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A genus with relatively large species (usually
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||
<quantity id="236CB5EF51151878FCFDFCA5EC8F1F13" box="[868,951,826,852]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="46" pageNumber="47" unit="mm" value="10.0">10 mm</quantity>
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or longer) having a broad pronotum with laterally divergent margins, and convex sided elytra which are often brightly coloured, many with dark brown to black elytra having pale margins. Known larvae are terrestrial. Males distinguished from all other
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<taxonomicName id="2394638951151878FB43FC1DEA6A1FDB" authorityName="Lacordaire" authorityYear="1857" box="[1242,1362,898,924]" class="Insecta" family="Lampyridae" genus="Luciolinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="166" phylum="Arthropoda" rank="genus">Luciolinae</taxonomicName>
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by the asymmetrical T8 which is emarginate on its left side (
|
||
<bibRefCitation id="800565FB51151878FD6BFC39EB0E1F87" author="Ballantyne & Lambkin" box="[754,1078,934,960]" firstAuthor="Ballantyne" pageId="46" pageNumber="47" pagination="1 - 188" refId="ref114542" refString="Ballantyne, L. A. & Lambkin, C. L. (2009) Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae). Zootaxa, 1997, 1 - 188." type="journal article" year="2009">Ballantyne & Lambkin 2009</bibRefCitation>
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||
fig. 7); all but one species have bipartite LO in
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<date id="902A3ECA51151878FEDDFC55EE561FA3" box="[324,366,970,996]" pageId="46" pageNumber="47">V7.</date>
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Females are macropterous and those examined have no bursa plates; in two species the median oviduct plate is subtriangular in outline; larvae have laterally explanate tergal margins (
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||
<bibRefCitation id="800565FB51151878FBF1FC71EA79184F" author="Fu" box="[1128,1345,1006,1032]" etAl="et al." firstAuthor="Fu" pageId="46" pageNumber="47" pagination="1 - 53" refId="ref116755" refString="Fu, X. H., Ballantyne, L. & Lambkin, C. (2012 a) Emeia gen. nov., a new genus of Luciolinae fireflies from China (Coleoptera: Lampyridae) with an unusual trilobite-like larva, and a redescription of the genus Curtos Motschulsky. Zootaxa, 3403 (1), 1 - 53. https: // doi. org / 10.11646 / zootaxa. 3403.1.1" type="journal article" year="2012" yearSuffix="a">
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||
Fu
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<emphasis id="D6E0C41851151878FB17FC70EBFF184F" box="[1166,1223,1006,1032]" italics="true" pageId="46" pageNumber="47">et al.</emphasis>
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||
2012a: 22
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||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="AC8E4B8151151878FF5EFB8DED4E1A47" pageId="46" pageNumber="47" type="discussion">
|
||
<paragraph id="E42B180A51151878FF5EFB8DEC601937" blockId="46.[151,1437,789,1536]" pageId="46" pageNumber="47">
|
||
<emphasis id="D6E0C41851151878FF5EFB8DEE0C186B" bold="true" box="[199,308,1042,1068]" pageId="46" pageNumber="47">Remarks</emphasis>
|
||
.
|
||
<taxonomicName id="2394638951151878FEDBFB8DEED2186B" authorityName="Ballantyne in Ballantyne & Lambkin" authorityYear="2009" box="[322,490,1042,1068]" class="Insecta" family="Lampyridae" genus="Asymmetricata" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="D6E0C41851151878FEDBFB8DEED2186B" box="[322,490,1042,1068]" italics="true" pageId="46" pageNumber="47">Asymmetricata</emphasis>
|
||
</taxonomicName>
|
||
was erected for two species transferred from
|
||
<taxonomicName id="2394638951151878FC75FB8DEB7A186B" box="[1004,1090,1042,1068]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="D6E0C41851151878FC75FB8DEB7A186B" box="[1004,1090,1042,1068]" italics="true" pageId="46" pageNumber="47">Luciola</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="2394638951151878FBCBFB8DEAA4186B" authority="Motsch. 1854" authorityName="Motschulsky" authorityYear="1854" box="[1106,1436,1041,1068]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="circumdata">
|
||
<emphasis id="D6E0C41851151878FBCBFB8DEBC3186B" box="[1106,1275,1042,1068]" italics="true" pageId="46" pageNumber="47">As. circumdata</emphasis>
|
||
Motsch. 1854
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="9C83589A51151878FF4FFBA9EE021817" box="[214,314,1078,1104]" name="Thailand" pageId="46" pageNumber="47">Thailand</collectingCountry>
|
||
and
|
||
<collectingCountry id="9C83589A51151878FEECFBA9EED31817" box="[373,491,1078,1104]" name="Cambodia" pageId="46" pageNumber="47">Cambodia</collectingCountry>
|
||
, and
|
||
<taxonomicName id="2394638951151878FDB3FBA9EDA11817" authorityName="Elytra" baseAuthorityName="Hope in Gray" baseAuthorityYear="1831" box="[554,665,1078,1104]" class="Insecta" family="Lampyridae" genus="Asymmetricata" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="ovalis">
|
||
<emphasis id="D6E0C41851151878FDB3FBA9ED7E1817" box="[554,582,1078,1104]" italics="true" pageId="46" pageNumber="47">As</emphasis>
|
||
.
|
||
<emphasis id="D6E0C41851151878FDCFFBA9EDA11817" box="[598,665,1078,1104]" italics="true" pageId="46" pageNumber="47">ovalis</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<bibRefCitation id="800565FB51151878FD31FBA9EC131817" author="Hope" box="[680,811,1077,1104]" firstAuthor="Hope" pageId="46" pageNumber="47" pagination="21 - 26" refId="ref117983" refString="Hope, F. W. (1831) Synopsis of the new species of Nepaul (sic) Insects in the collection of Major General Hardwicke. In: Gray, J. E. (Ed.), The Zoological Miscellany. Vol. 1. Cheapside, Stirling, pp. 21 - 26." type="book chapter" year="1831">Hope 1831</bibRefCitation>
|
||
] from
|
||
<collectingCountry id="9C83589A51151878FCE2FBA9EC8E1816" box="[891,950,1078,1105]" name="India" pageId="46" pageNumber="47">India</collectingCountry>
|
||
) and characterised primarily by the asymmetrical male abdominal T8 (
|
||
<bibRefCitation id="800565FB51151878FE7EFBC5EC651833" author="Ballantyne & Lambkin" box="[487,861,1114,1140]" firstAuthor="Ballantyne" pageId="46" pageNumber="47" pagination="1 - 188" refId="ref114542" refString="Ballantyne, L. A. & Lambkin, C. L. (2009) Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae). Zootaxa, 1997, 1 - 188." type="journal article" year="2009">Ballantyne & Lambkin 2009: 30</bibRefCitation>
|
||
). The species were keyed and distinguished by the nature of the light organs in
|
||
<date id="902A3ECA51151878FE42FBE1EEC518DF" box="[475,509,1150,1176]" pageId="46" pageNumber="47">V7</date>
|
||
(either bipartite or entire). Two further species with a similar configuration of T8 and bipartite LOs in
|
||
<date id="902A3ECA51151878FEE2FB3DEEA518FB" box="[379,413,1186,1212]" pageId="46" pageNumber="47">V7</date>
|
||
viz.
|
||
<taxonomicName id="2394638951151878FE4BFB3DEC2B18FB" authority="(Walker 1858)" authorityName="Ho" authorityYear="2019" baseAuthorityName="Walker" baseAuthorityYear="1858" box="[466,787,1185,1212]" class="Insecta" family="Lampyridae" genus="Asymmetricata" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="humeralis" status="comb. nov.">
|
||
<emphasis id="D6E0C41851151878FE4BFB3DEED618FB" box="[466,494,1186,1212]" italics="true" pageId="46" pageNumber="47">As</emphasis>
|
||
.
|
||
<emphasis id="D6E0C41851151878FE65FB3DED5418FB" box="[508,620,1186,1212]" italics="true" pageId="46" pageNumber="47">humeralis</emphasis>
|
||
(
|
||
<bibRefCitation id="800565FB51151878FDE0FB3DEC3218FB" author="Walker" box="[633,778,1185,1212]" firstAuthor="Walker" pageId="46" pageNumber="47" pagination="280 - 286" refId="ref122273" refString="Walker, F. (1858) XXX. Characters of some apparently undescribed Ceylon Insects. Annals and Magazine of natural history, Series 3, 2 (No. 10), 280 - 286. https: // doi. org / 10.1080 / 00222935808697026" type="journal article" year="1858">Walker 1858</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<emphasis id="D6E0C41851151878FC80FB3DECA018FB" bold="true" box="[793,920,1186,1212]" pageId="46" pageNumber="47">
|
||
<taxonomicNameLabel id="CDD3796351151878FC80FB3DECA018FB" box="[793,920,1186,1212]" pageId="46" pageNumber="47" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="2394638951151878FC54FB3DEBD618FB" authority="(Pic 1927)" authorityName="Ho" authorityYear="2019" baseAuthorityName="Pic" baseAuthorityYear="1927" box="[973,1262,1186,1212]" class="Insecta" family="Lampyridae" genus="Asymmetricata" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="bicoloripes" status="comb. nov.">
|
||
<emphasis id="D6E0C41851151878FC54FB3DECD118FB" box="[973,1001,1186,1212]" italics="true" pageId="46" pageNumber="47">As</emphasis>
|
||
.
|
||
<emphasis id="D6E0C41851151878FC6FFB3DEB4C18FB" box="[1014,1140,1186,1212]" italics="true" pageId="46" pageNumber="47">bicoloripes</emphasis>
|
||
(
|
||
<bibRefCitation id="800565FB51151878FB1BFB3DEBDE18FB" author="Pic" box="[1154,1254,1186,1212]" firstAuthor="Pic" pageId="46" pageNumber="47" pagination="37 - 52" refId="ref121296" refString="Pic, M. (1927) Malacodermes exotiques. L'Echange, Revue Linneenne, XLIII, 37 - 52." type="book chapter" year="1927">Pic 1927</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<emphasis id="D6E0C41851151878FB6DFB3DEA4C18FB" bold="true" box="[1268,1396,1186,1212]" pageId="46" pageNumber="47">
|
||
<taxonomicNameLabel id="CDD3796351151878FB6DFB3DEA4C18FB" box="[1268,1396,1186,1212]" pageId="46" pageNumber="47" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
are transferred from
|
||
<taxonomicName id="2394638951151878FECAFB59EE9118A7" box="[339,425,1222,1248]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="D6E0C41851151878FECAFB59EE9118A7" box="[339,425,1222,1248]" italics="true" pageId="46" pageNumber="47">Luciola</emphasis>
|
||
</taxonomicName>
|
||
. Three species (
|
||
<taxonomicName id="2394638951151878FDC5FB59ECAE18A7" authority="Olivier 1885" authorityName="Olivier" authorityYear="1885" box="[604,918,1221,1248]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="doriae">
|
||
<emphasis id="D6E0C41851151878FDC5FB59EC3818A7" box="[604,768,1222,1248]" italics="true" pageId="46" pageNumber="47">Luciola doriae</emphasis>
|
||
<bibRefCitation id="800565FB51151878FC9FFB59ECAE18A7" author="Olivier" box="[774,918,1221,1248]" firstAuthor="Olivier" pageId="46" pageNumber="47" pagination="333 - 374" refId="ref120140" refString="Olivier, E. (1885) Catalogue des Lampyrides faisant partie des collections du Musee Civique de Genes. Annali del Museo Civico di Storia Naturale di Genova, 2 (2 a), 333 - 374." type="journal article" year="1885">Olivier 1885</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="2394638951151878FC39FB58EBF918A7" authority="Olivier 1910 b" authorityName="Olivier" authorityYear="1910" box="[928,1217,1222,1248]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="impressa">
|
||
<emphasis id="D6E0C41851151878FC39FB58EC8818A7" box="[928,944,1223,1248]" italics="true" pageId="46" pageNumber="47">L</emphasis>
|
||
.
|
||
<emphasis id="D6E0C41851151878FC25FB58EB1818A7" box="[956,1056,1223,1248]" italics="true" pageId="46" pageNumber="47">impressa</emphasis>
|
||
<bibRefCitation id="800565FB51151878FBBFFB59EBF918A7" author="Olivier" box="[1062,1217,1222,1248]" firstAuthor="Olivier" pageId="46" pageNumber="47" pagination="532 - 533" refId="ref120801" refString="Olivier, E. (1910 b) XX. Descriptions of two new species of Luciola in the collection of Mr H. E. Andrewes. Transactions of the Entomological Society of London, 1910, 532 - 533. https: // doi. org / 10.1111 / j. 1365 - 2311.1910. tb 01183. x" type="journal article" year="1910" yearSuffix="b">Olivier 1910b</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="2394638951151878FB6CFB58EE0D1943" authority="Olivier 1909 a" authorityName="Olivier" authorityYear="1909" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="notatipennis">
|
||
<emphasis id="D6E0C41851151878FB6CFB58EAA418A7" box="[1269,1436,1223,1248]" italics="true" pageId="46" pageNumber="47">L. notatipennis</emphasis>
|
||
<bibRefCitation id="800565FB51151878FF0EFB75EE0D1943" author="Olivier" box="[151,309,1258,1284]" firstAuthor="Olivier" pageId="46" pageNumber="47" pagination="316 - 319" refId="ref120656" refString="Olivier, E. (1909 a) Descriptions de Lampyrides nouveaux du Musee Civique de Genes. Annali del Museo Civico di Storia Naturale di Genova, IV (3 a), 316 - 319." type="book chapter" year="1909" yearSuffix="a">Olivier 1909a</bibRefCitation>
|
||
</taxonomicName>
|
||
) are synonymised with
|
||
<taxonomicName id="2394638951151878FDDDFB75EDE71943" authorityName="Ho" authorityYear="2019" baseAuthorityName="Walker" baseAuthorityYear="1858" box="[580,735,1258,1284]" class="Insecta" family="Lampyridae" genus="Asymmetricata" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="humeralis" status="comb. nov.">
|
||
<emphasis id="D6E0C41851151878FDDDFB75ED581943" box="[580,608,1258,1284]" italics="true" pageId="46" pageNumber="47">As</emphasis>
|
||
.
|
||
<emphasis id="D6E0C41851151878FDF6FB75EDE71943" box="[623,735,1258,1284]" italics="true" pageId="46" pageNumber="47">humeralis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="D6E0C41851151878FD7EFB75EC511943" bold="true" box="[743,873,1258,1284]" pageId="46" pageNumber="47">
|
||
<taxonomicNameLabel id="CDD3796351151878FD7EFB75EC511943" box="[743,873,1258,1284]" pageId="46" pageNumber="47" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
However, the variability of the dorsal colour patterns indicated below could indicate further variability
|
||
<taxonomicName id="2394638951151878FC9CFA91EC98196F" authorityName="Ho" authorityYear="2019" baseAuthorityName="Walker" baseAuthorityYear="1858" box="[773,928,1294,1320]" class="Insecta" family="Lampyridae" genus="Asymmetricata" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="humeralis" status="comb. nov.">
|
||
<emphasis id="D6E0C41851151878FC9CFA91EC19196F" box="[773,801,1294,1320]" italics="true" pageId="46" pageNumber="47">As</emphasis>
|
||
.
|
||
<emphasis id="D6E0C41851151878FCA9FA91EC98196F" box="[816,928,1294,1320]" italics="true" pageId="46" pageNumber="47">humeralis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="D6E0C41851151878FC31FA91EB11196F" bold="true" box="[936,1065,1294,1320]" pageId="46" pageNumber="47">
|
||
<taxonomicNameLabel id="CDD3796351151878FC31FA91EB11196F" box="[936,1065,1294,1320]" pageId="46" pageNumber="47" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(interpretations were made from dead museum specimens where it may be difficult to interpret the original colour pattern). Assessing this variability requires examination of aedeagal and aedeagal sheath patterns.
|
||
</paragraph>
|
||
<paragraph id="E42B180A51151878FF5EFAE5ED4E1A47" blockId="46.[151,1437,789,1536]" pageId="46" pageNumber="47">
|
||
<bibRefCitation id="800565FB51151878FF5EFAE5ED2019D3" author="Ballantyne & Lambkin" box="[199,536,1402,1428]" firstAuthor="Ballantyne" pageId="46" pageNumber="47" pagination="1 - 188" refId="ref114542" refString="Ballantyne, L. A. & Lambkin, C. L. (2009) Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae). Zootaxa, 1997, 1 - 188." type="journal article" year="2009">Ballantyne & Lambkin (2009)</bibRefCitation>
|
||
suggested the possibility of a gradation from entire to bipartite LO in
|
||
<date id="902A3ECA51151878FA82FAE5EA0519D3" box="[1307,1341,1402,1428]" pageId="46" pageNumber="47">V7</date>
|
||
because of the anterior emargination of the LO in
|
||
<date id="902A3ECA51151878FDFAFA01EDBD19FF" box="[611,645,1438,1464]" pageId="46" pageNumber="47">
|
||
V
|
||
<quantity id="236CB5EF51151878FDE1FA01EDBD19FF" box="[632,645,1438,1464]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="46" pageNumber="47" unit="in" value="7.0">7</quantity>
|
||
</date>
|
||
in
|
||
<taxonomicName id="2394638951151878FD31FA01EC6819FF" authorityName="Motschulsky" authorityYear="1854" box="[680,848,1438,1464]" class="Insecta" family="Lampyridae" genus="Luciola" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="circumdata">
|
||
<emphasis id="D6E0C41851151878FD31FA01EC6819FF" box="[680,848,1438,1464]" italics="true" pageId="46" pageNumber="47">As. circumdata</emphasis>
|
||
</taxonomicName>
|
||
. On the basis of specimens examined here it appears that the LO in
|
||
<date id="902A3ECA51151878FEA0FA5DEE63199B" box="[313,347,1474,1500]" pageId="46" pageNumber="47">V7</date>
|
||
exists in two distinct forms either bipartite or entire (see below) but that there is no evidence of a gradation between the two different forms.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |