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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.827.31961" ID-GBIF-Dataset="7b9a822b-a3de-4ee7-b1ec-3bf85820332e" ID-PMC="PMC6472300" ID-Pensoft-Pub="1313-2970-827-139" ID-PubMed="31114427" ID-ZBK="ADF1A9E3027745DC9E3ED3D453FC1430" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1313-2970-827-139" ModsDocOrigin="ZooKeys 827" ModsDocTitle="New dustywings (Neuroptera, Coniopterygidae) from mid-Cretaceous amber of Myanmar reveal spectacular diversity" checkinTime="1551875345984" checkinUser="pensoft" docAuthor="Ruzickova, Dominika, Nel, Andre &amp; Prokop, Jakub" docDate="2019" docId="50B09083F0A817773340322042FE20B7" docLanguage="en" docName="ZooKeys 827: 139-152" docOrigin="ZooKeys 827" docSource="http://dx.doi.org/10.3897/zookeys.827.31961" docTitle="Palaeoconis azari gen. et., sp. n." docType="treatment" docUuid="D2FC29B5-FFBE-42C7-AD0B-FD637B5011E7" docUuidSource="ZooBank" docVersion="5" lastPageNumber="146" masterDocId="FFC0FF8F1E5A8E786508FFAA0574FFDB" masterDocTitle="New dustywings (Neuroptera, Coniopterygidae) from mid-Cretaceous amber of Myanmar reveal spectacular diversity" masterLastPageNumber="152" masterPageNumber="139" pageNumber="143" updateTime="1668166975016" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>New dustywings (Neuroptera, Coniopterygidae) from mid-Cretaceous amber of Myanmar reveal spectacular diversity</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ruzickova, Dominika</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nel, Andre</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Prokop, Jakub</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>827</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>139</mods:start>
<mods:end>152</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.827.31961</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.827.31961</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-827-139</mods:identifier>
<mods:identifier type="ZBK">ADF1A9E3027745DC9E3ED3D453FC1430</mods:identifier>
<mods:identifier type="ZooBank">ADF1A9E3027745DC9E3ED3D453FC1430</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="154175314" LSID="urn:lsid:zoobank.org:act:D2FC29B5-FFBE-42C7-AD0B-FD637B5011E7" httpUri="http://treatment.plazi.org/id/50B09083F0A817773340322042FE20B7" lastPageId="8" lastPageNumber="146" pageId="4" pageNumber="143">
<subSubSection pageId="4" pageNumber="143" type="nomenclature">
<paragraph pageId="4" pageNumber="143">
<taxonomicName LSID="http://zoobank.org/D2FC29B5-FFBE-42C7-AD0B-FD637B5011E7" authority="gen. et." class="Insecta" family="Coniopterygidae" genus="Palaeoconis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Palaeoconis azari" order="Neuroptera" pageId="4" pageNumber="143" phylum="Arthropoda" rank="species" species="azari">Palaeoconis azari gen. et.</taxonomicName>
<taxonomicNameLabel pageId="4" pageNumber="143">sp. n.</taxonomicNameLabel>
Figures 3, 4
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="144" type="holotype">
<paragraph pageId="5" pageNumber="144">
<pageBreakToken pageId="5" pageNumber="144" start="start">Holotype</pageBreakToken>
.
</paragraph>
<paragraph pageId="5" pageNumber="144">
BUB 2914; lowermost Cenomanian amber (
<bibRefCitation author="Shi, G" journalOrPublisher="Cretaceous Research" pageId="11" pageNumber="150" pagination="155 - 163" title="Age constraint on Burmese amber based on U-Pb dating of zircons." url="https://doi.org/10.1016/j.cretres.2012.03.014" volume="37" year="2012">Shi et al. 2012</bibRefCitation>
); Myanmar, Kachin, Hukawng Valley; preserved in a polished, transparent yellow piece of amber (9.18
<normalizedToken originalValue="×">x</normalizedToken>
6.77
<normalizedToken originalValue="×">x</normalizedToken>
2.18 mm), deposited in the Museum
<normalizedToken originalValue="für">fuer</normalizedToken>
Naturkunde, Berlin (ex. coll. Patrick
<normalizedToken originalValue="Müller">Mueller</normalizedToken>
). The amber piece contains also one syninclusion of an imago of
<taxonomicName lsidName="Auchenorrhyncha" order="Diptera" pageId="5" pageNumber="144" rank="order">Diptera</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="144" type="etymology">
<paragraph pageId="5" pageNumber="144">Etymology.</paragraph>
<paragraph pageId="5" pageNumber="144">The specific epithet honors Prof. Dany Azar (Lebanese University, Fanar, Lebanon), friend and colleague of AN and JP and worldwide known palaeoentomologist.</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="144" type="diagnosis">
<paragraph pageId="5" pageNumber="144">Diagnosis.</paragraph>
<paragraph pageId="5" pageNumber="144">As for the genus (vide supra).</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="146" pageId="5" pageNumber="144" type="description">
<paragraph pageId="5" pageNumber="144">Description.</paragraph>
<paragraph lastPageId="6" lastPageNumber="145" pageId="5" pageNumber="144">
Male. Body length ca. 1.93 mm (measured from tip of head to tip of genitalia). Head hypognathous, ca. 0.34 mm. Compound eyes well developed, 0.22 mm x 0.14 mm (from lateral view). Antennae 21-segmented (with 19 flagellomeres),
<pageBreakToken pageId="6" pageNumber="145" start="start">scape</pageBreakToken>
and pedicel stouter, longer and broader than flagellomeres, first flagellomere longer and wider than remaining flagellomeres, flagellomeres subquadrate, nearly as long as wide, terminal flagellomere conical. Maxillary palps five-segmented, fifth segment distinctly larger than other palpomeres, length of fifth segment ca. 0.12 mm. Labial palps three-segmented, third segment larger than remaining. Thorax well developed. Prothorax narrower and overall smaller than meso- and metathorax.
</paragraph>
<paragraph lastPageId="7" lastPageNumber="146" pageId="6" pageNumber="145">
Forewing ca. 2.19 mm long, ca. 0.93 mm wide; two distinct apical crossveins in costal area; division of ScP1 and ScP2 0.31 mm from wing apex; R branching into RA
<pageBreakToken pageId="7" pageNumber="146" start="start">and</pageBreakToken>
RP 0.48 mm from wing base; RA simple, distally ending connecting with branch of ScP2; RP branched into RP1 and RP2; ra-rp1 present; first spot on basal abscissa of RP2 near RP1 (near place where is RP forked); crossvein rp-m oblique crossing dark spot about mid-length; M with three branches (MA, MP1 and MP2), M with one stiff seta near crossvein rp-m (Figs 3A, 4C); crossvein rp2-ma relatively long, approximately as long as basal abscissa of RP2; crossvein 1
<normalizedToken originalValue="mcua">m-cua</normalizedToken>
near rp-m, but not aligned; crossvein 2mp2-cua near to wing apex, with third spot over it; crossveins cua-cup and a1 present. Hindwing ca. 2.12 mm long, ca. 0.92 mm wide; venation pattern very similar to forewing with exception of only single crossvein between C and ScP, wing membrane in hindwing hyaline without any pigmented spots; crossveins ra-rp1 and rp2-ma present; basal part of hindwing without clearly recognizable venation.
</paragraph>
<caption pageId="7" pageNumber="146">
<paragraph pageId="7" pageNumber="146">
Figure 3.
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Palaeoconis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Palaeoconis azari" order="Neuroptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="species" species="azari">Palaeoconis azari</taxonomicName>
gen. et sp. n., holotype BUB 2914. A Drawing of right fore wing B drawing of left hindwing. Scale bars: 100
<normalizedToken originalValue="µm">µm</normalizedToken>
.
</paragraph>
</caption>
<caption pageId="7" pageNumber="146">
<paragraph pageId="7" pageNumber="146">
Figure 4.
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Palaeoconis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Palaeoconis azari" order="Neuroptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="species" species="azari">Palaeoconis azari</taxonomicName>
gen. et sp. n., holotype BUB 2914. A Habitus lateral view B wing venation and spots C detail of forewing venation with stiff seta on vein M D head from dorso-lateral view showing antennae, terminal segment of maxillary palp and eye. Scale bars: 500µm (A, B); not in scale: (C); 50
<normalizedToken originalValue="µm">µm</normalizedToken>
(D).
</paragraph>
</caption>
<paragraph pageId="7" pageNumber="146">Legs long and slender; femora covered with fine and sparse setae; tibiae covered with stiff and dense setae; tarsi five-segmented, first tarsomere distinctly longer than remaining tarsomeres, terminal tarsomere elongated ending with two claws, tarsi covered with fine dense setae. Abdomen large and broad, tapered to the end (visible from lateral view only); plicatures present on sternites II-IV; length of abdomen including genitalia 1.22 mm, width 0.54 mm. Structures of external genitalia hardly discernible with exception of domed and well-sclerotized ectoproct in lateral view and the caudal projection of gonarcus below.</paragraph>
</subSubSection>
<subSubSection lastPageId="8" lastPageNumber="147" pageId="7" pageNumber="146" type="discussion">
<paragraph pageId="7" pageNumber="146">Discussion.</paragraph>
<paragraph pageId="7" pageNumber="146">
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Palaeoconis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Palaeoconis" order="Neuroptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="genus">Palaeoconis</taxonomicName>
gen. n. can be attributed to the
<taxonomicName lsidName="" pageId="7" pageNumber="146" rank="subfamily" subfamily="Aleuropteryginae">Aleuropteryginae</taxonomicName>
based on the presence of two crossveins between RP and M and also the presence of plicatures on abdominal sternites II-IV. Nevertheless,
<bibRefCitation author="Zimmermann, D" journalOrPublisher="European Journal of Entomology" pageId="11" pageNumber="150" pagination="651 - 662" title="The first holistic SEM study of Coniopterygidae (Neuroptera) - structural evidence and phylogenetic implications." url="https://doi.org/10.14411/eje.2009.081" volume="106" year="2009">Zimmermann et al. (2009)</bibRefCitation>
considered this last character as a symplesiomorphy of the
<taxonomicName family="Coniopterygidae" lsidName="" pageId="7" pageNumber="146" rank="family">Coniopterygidae</taxonomicName>
. However,
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Palaeoconis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Palaeoconis" order="Neuroptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="genus">Palaeoconis</taxonomicName>
does not possess in the forewings two long stiff setae on M, which is the diagnostic character for
<taxonomicName lsidName="" pageId="7" pageNumber="146" rank="subfamily" subfamily="Aleuropteryginae">Aleuropteryginae</taxonomicName>
. In contrast to this, the absence of these setae on the media and the presence of three-branched media are considered as plesiomorphic characters typical for Mesozoic
<taxonomicName family="Coniopterygidae" lsidName="" pageId="7" pageNumber="146" rank="family">Coniopterygidae</taxonomicName>
(Engel 2002). In addition,
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Palaeoconis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Palaeoconis" order="Neuroptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="genus">Palaeoconis</taxonomicName>
possesses in forewing crossvein a1 connecting A1 to posterior wing margin. According to
<bibRefCitation author="Engel, MS" journalOrPublisher="Novitates Paleoentomologicae" pageId="10" pageNumber="149" pagination="1 - 16" title="Two new genera of Cretaceous dustywings in amber from Myanmar (Neuroptera: Coniopterygidae)." url="https://doi.org/10.17161/np.v0i17.6465" volume="17" year="2016">Engel (2016)</bibRefCitation>
the subfamily
<taxonomicName lsidName="" pageId="7" pageNumber="146" rank="subfamily" subfamily="Aleuropteryginae">Aleuropteryginae</taxonomicName>
contains 14 genera (Mesozoic and Cenozoic), i.e.,
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Achlyoconis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Achlyoconis" order="Neuroptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="genus">Achlyoconis</taxonomicName>
Engel, 2016;
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Alboconis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alboconis" order="Neuroptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="genus">Alboconis</taxonomicName>
Nel et al., 2005;
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Apoglaesoconis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Apoglaesoconis" order="Neuroptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="genus">Apoglaesoconis</taxonomicName>
Grimaldi, 2000;
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Archiconiocompsa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Archiconiocompsa" order="Neuroptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="genus">Archiconiocompsa</taxonomicName>
Enderlein, 1910;
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Archiconis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Archiconis" order="Neuroptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="genus">Archiconis</taxonomicName>
Enderlein, 1930;
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Garnaconis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Garnaconis" order="Neuroptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="genus">Garnaconis</taxonomicName>
Perrichot and Nel in
<bibRefCitation pageId="7" pageNumber="146">Perrichot et al., 2014</bibRefCitation>
;
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Geroconiocompsa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geroconiocompsa" order="Neuroptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="genus">Geroconiocompsa</taxonomicName>
Engel, 2010;
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Glaesoconis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glaesoconis" order="Neuroptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="genus">Glaesoconis</taxonomicName>
Meinander, 1975;
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Hemisemidalis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemisemidalis" order="Neuroptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="genus">Hemisemidalis</taxonomicName>
Meinander, 1972;
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Juraconiopteryx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Juraconiopteryx" order="Neuroptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="genus">Juraconiopteryx</taxonomicName>
Meinander, 1975;
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Libanoconis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Libanoconis" order="Neuroptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="genus">Libanoconis</taxonomicName>
Engel, 2002;
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Neoconis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neoconis" order="Neuroptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="genus">Neoconis</taxonomicName>
Enderlein, 1930;
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Pararchiconis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pararchiconis" order="Neuroptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="genus">Pararchiconis</taxonomicName>
Nel, 1991, and
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Spiloconis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Spiloconis" order="Neuroptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="genus">Spiloconis</taxonomicName>
Enderlein, 1907.
</paragraph>
<paragraph lastPageId="8" lastPageNumber="147" pageId="7" pageNumber="146">
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Achlyoconis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Achlyoconis" order="Neuroptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="genus">Achlyoconis</taxonomicName>
,
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Alboconis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alboconis" order="Neuroptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="genus">Alboconis</taxonomicName>
,
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Apoglaesoconis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Apoglaesoconis" order="Neuroptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="genus">Apoglaesoconis</taxonomicName>
,
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Glaesoconis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glaesoconis" order="Neuroptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="genus">Glaesoconis</taxonomicName>
, and
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Libanoconis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Libanoconis" order="Neuroptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="genus">Libanoconis</taxonomicName>
share the presence of two distinct basal crossveins in the costal area, and in addition
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Achlyoconis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Achlyoconis" order="Neuroptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="genus">Achlyoconis</taxonomicName>
,
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Alboconis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alboconis" order="Neuroptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="genus">Alboconis</taxonomicName>
and
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Libanoconis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Libanoconis" order="Neuroptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="genus">Libanoconis</taxonomicName>
share the presence of crossvein a1-a2, vein A2 and crossvein a2. These traits separate these genera from
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Palaeoconis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Palaeoconis" order="Neuroptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="genus">Palaeoconis</taxonomicName>
gen. n. In addition,
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Glaesoconis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glaesoconis" order="Neuroptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="genus">Glaesoconis</taxonomicName>
described from the Lower Cretaceous Myanmar amber differs from
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Palaeoconis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Palaeoconis" order="Neuroptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="genus">Palaeoconis</taxonomicName>
gen. n. by a distinctly shorter crossvein rp2-ma, single crossvein
<normalizedToken originalValue="mcua">m-cua</normalizedToken>
, presence of A2 and four pigmented spots on wing membrane instead of three in
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Palaeoconis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Palaeoconis" order="Neuroptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="genus">Palaeoconis</taxonomicName>
(
<bibRefCitation author="Engel, MS" journalOrPublisher="Journal of Systematic Palaeontology" pageId="10" pageNumber="149" pagination="133 - 136" title="The dustywings in Cretaceous Burmese amber (Insecta: NeuropteraConiopterygidae)." url="https://doi.org/10.1017/S1477201904001191" volume="2" year="2004">Engel 2004</bibRefCitation>
).
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Libanoconis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Libanoconis" order="Neuroptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="genus">Libanoconis</taxonomicName>
, known from the Lower Cretaceous amber of Lebanon, differs from
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Palaeoconis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Palaeoconis" order="Neuroptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="genus">Palaeoconis</taxonomicName>
gen. n. in the positions of crossveins between M and CuA (1
<normalizedToken originalValue="mcua">m-cua</normalizedToken>
is near to wing base; 2
<normalizedToken originalValue="mcua">m-cua</normalizedToken>
approximately at mid-length of wing), and lack pigmented spots
<pageBreakToken pageId="8" pageNumber="147" start="start">on</pageBreakToken>
membrane (Engel 2002;
<bibRefCitation author="Nel, A" journalOrPublisher="Annales Zoologici" pageId="11" pageNumber="150" pagination="1 - 7" title="New and poorly known fossil Coniopterygidae in Cretaceous and Cenozoic ambers (Insecta: Neuroptera)." volume="55" year="2004">Nel et al. 2004</bibRefCitation>
).
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Achlyoconis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Achlyoconis" order="Neuroptera" pageId="8" pageNumber="147" phylum="Arthropoda" rank="genus">Achlyoconis</taxonomicName>
described from the Upper Cretaceous amber of northern Myanmar differs from
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Palaeoconis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Palaeoconis" order="Neuroptera" pageId="8" pageNumber="147" phylum="Arthropoda" rank="genus">Palaeoconis</taxonomicName>
gen. n. in the presence of seven thickenings with specialized setae on M, positions of crossveins 1m-cua and 2m-cua, presence of a single crossvein between CuP and A1, and four pigmented spots (
<bibRefCitation author="Engel, MS" journalOrPublisher="Novitates Paleoentomologicae" pageId="10" pageNumber="149" pagination="1 - 16" title="Two new genera of Cretaceous dustywings in amber from Myanmar (Neuroptera: Coniopterygidae)." url="https://doi.org/10.17161/np.v0i17.6465" volume="17" year="2016">Engel 2016</bibRefCitation>
).
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Alboconis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alboconis" order="Neuroptera" pageId="8" pageNumber="147" phylum="Arthropoda" rank="genus">Alboconis</taxonomicName>
, described from the Lower Cretaceous French amber, differs from
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Palaeoconis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Palaeoconis" order="Neuroptera" pageId="8" pageNumber="147" phylum="Arthropoda" rank="genus">Palaeoconis</taxonomicName>
gen. n. by a two branched media bearing two long setae, crossvein between CuA and R+M, and two crossveins between CuP and A1 (
<bibRefCitation author="Nel, A" journalOrPublisher="Annales Zoologici" pageId="11" pageNumber="150" pagination="1 - 7" title="New and poorly known fossil Coniopterygidae in Cretaceous and Cenozoic ambers (Insecta: Neuroptera)." volume="55" year="2004">Nel et al. 2004</bibRefCitation>
).
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Apoglaesoconis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Apoglaesoconis" order="Neuroptera" pageId="8" pageNumber="147" phylum="Arthropoda" rank="genus">Apoglaesoconis</taxonomicName>
and
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Garnaconis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Garnaconis" order="Neuroptera" pageId="8" pageNumber="147" phylum="Arthropoda" rank="genus">Garnaconis</taxonomicName>
share the following features such as M with two long setae, presence of crossveins
<normalizedToken originalValue="cupa">cup-a</normalizedToken>
1; a1-a2, vein A2 and absence of pigmented spots on wing membrane.
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Apoglaesoconis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Apoglaesoconis" order="Neuroptera" pageId="8" pageNumber="147" phylum="Arthropoda" rank="genus">Apoglaesoconis</taxonomicName>
, from the Late Cretaceous amber of New Jersey, differs from
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Palaeoconis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Palaeoconis" order="Neuroptera" pageId="8" pageNumber="147" phylum="Arthropoda" rank="genus">Palaeoconis</taxonomicName>
gen. n. by one crossvein between RP and M, branches RP2 and MA forming a cross, and by a single crossvein between M and CuA (Engel 2002).
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Garnaconis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Garnaconis" order="Neuroptera" pageId="8" pageNumber="147" phylum="Arthropoda" rank="genus">Garnaconis</taxonomicName>
, from the Late Cretaceous amber of
<normalizedToken originalValue="Vendée">Vendee</normalizedToken>
in northwestern France, differs from
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Palaeoconis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Palaeoconis" order="Neuroptera" pageId="8" pageNumber="147" phylum="Arthropoda" rank="genus">Palaeoconis</taxonomicName>
gen. n. in the remote positions of separated transverse ScP2 and crossvein between RA+ScP2 and RP1 (see Fig. 3), unlike both being aligned in
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Garnaconis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Garnaconis" order="Neuroptera" pageId="8" pageNumber="147" phylum="Arthropoda" rank="genus">Garnaconis</taxonomicName>
.
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Palaeoconis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Palaeoconis" order="Neuroptera" pageId="8" pageNumber="147" phylum="Arthropoda" rank="genus">Palaeoconis</taxonomicName>
gen. n. bears two crossveins beween RP and M, a two-branched media with setae instead of only one crossvein between RP2 and MA and a three-branched media without setae in
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Garnaconis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Garnaconis" order="Neuroptera" pageId="8" pageNumber="147" phylum="Arthropoda" rank="genus">Garnaconis</taxonomicName>
(
<bibRefCitation pageId="8" pageNumber="147">Perrichot et al. 2014</bibRefCitation>
).
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Juraconiopteryx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Juraconiopteryx" order="Neuroptera" pageId="8" pageNumber="147" phylum="Arthropoda" rank="genus">Juraconiopteryx</taxonomicName>
, Upper Jurassic of Kazakhstan, represents the oldest record of
<taxonomicName family="Coniopterygidae" lsidName="" pageId="8" pageNumber="147" rank="family">Coniopterygidae</taxonomicName>
. Wing venation of
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Juraconiopteryx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Juraconiopteryx" order="Neuroptera" pageId="8" pageNumber="147" phylum="Arthropoda" rank="genus">Juraconiopteryx</taxonomicName>
is very poorly preserved (
<bibRefCitation author="Meinander, M" journalOrPublisher="Notulae Entomologicae" pageId="10" pageNumber="149" pagination="53 - 57" title="Fossil Coniopterygidae (Neuroptera)." volume="55" year="1975">Meinander 1975</bibRefCitation>
), therefore we are unable to make a reliable comparison with
<taxonomicName class="Insecta" family="Coniopterygidae" genus="Palaeoconis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Palaeoconis" order="Neuroptera" pageId="8" pageNumber="147" phylum="Arthropoda" rank="genus">Palaeoconis</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>