treatments-xml/data/42/8B/4D/428B4D9F4F50241B888F6F7A9334A700.xml
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<mods:title id="CD2EB9A5DE6DB0FFA3406D248E9AB856">Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)</mods:title>
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<mods:namePart id="725B3C9F8C66BC8B3ADA56A3E8984827">Bistroem, Olof</mods:namePart>
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<mods:namePart id="175F27FC09AB9DF20AA4F3CF5FC45FA2">Nilsson, Anders N.</mods:namePart>
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<mods:namePart id="7E63C877DB7BF2107D1D967E9CDB9C59">Bergsten, Johannes</mods:namePart>
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<treatment id="428B4D9F4F50241B888F6F7A9334A700" ID-GBIF-Taxon="127862777" LSID="urn:lsid:plazi:treatment:428B4D9F4F50241B888F6F7A9334A700" httpUri="http://treatment.plazi.org/id/428B4D9F4F50241B888F6F7A9334A700" lastPageId="228" lastPageNumber="229" pageId="225" pageNumber="226">
<subSubSection id="E1D487F76E7C968FBEC0A49D66B91CAB" pageId="225" pageNumber="226" type="multiple">
<paragraph id="17A1F1F48A741F4BEB53D7B189116E14" pageId="225" pageNumber="226">Taxon classification Animalia Coleoptera Dytiscidae</paragraph>
</subSubSection>
<subSubSection id="820A75F5960357B3A0E31FF415A37E0C" pageId="225" pageNumber="226" type="nomenclature">
<paragraph id="F84DEF82C350F3479C6BB463999D0533" pageId="225" pageNumber="226">
<taxonomicName id="7D443761309F8C07D74E0F052372B8EE" ID-CoL="3RM82" authority="Zimmermann, 1926" authorityName="Zimmermann" authorityYear="1926" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus inornatus" order="Coleoptera" pageId="225" pageNumber="226" phylum="Arthropoda" rank="species" species="inornatus">Laccophilus inornatus Zimmermann, 1926</taxonomicName>
Figs 182-183, 361, 510-511, 574
</paragraph>
</subSubSection>
<subSubSection id="79A0C060FD1E3D31D323D1DEB740A8C2" lastPageId="226" lastPageNumber="227" pageId="225" pageNumber="226" type="reference_group">
<paragraph id="9BFC9F094385154DCA3D61BF58DFE625" lastPageId="226" lastPageNumber="227" pageId="225" pageNumber="226">
<taxonomicName id="E8586A779662E08574B13465066693FF" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus inornatus" order="Coleoptera" pageId="225" pageNumber="226" phylum="Arthropoda" rank="species" species="inornatus">Laccophilus inornatus</taxonomicName>
<bibRefCitation id="D2424EBDF247229C7397FB39E08C7A36" author="Zimmermann, A" journalOrPublisher="Entomologische Blaetter" pageId="263" pageNumber="264" pagination="18 - 31" title="Schwimmkaefer-Material aus Deutsch-Ost-Afrika, gesammelt von Geh. Regierungsrat Methner in den Jahren 1902 - 1916." volume="22" year="1926">Zimmermann 1926</bibRefCitation>
: 24 (original description, faunistics);
<bibRefCitation id="6C5DF9D2E8DA19319D709F109F56869A" author="Gschwendtner, L" journalOrPublisher="Revue de Zoologie et Botanique Africaines" pageId="256" pageNumber="257" pagination="372 - 375" title="Weitere neue Schwimmkaefer aus Belgisch Kongo." volume="27" year="1935 b">Gschwendtner 1935b</bibRefCitation>
: 373 (faunistics);
<bibRefCitation id="E03E98579FC2CD4849C28B0B29BD6957" author="Guignot, F" journalOrPublisher="Revue Francaise d'Entomologie" pageId="256" pageNumber="257" pagination="81 - 103" title="Dytiscidae et Gyrinidae de la Cote d'Ivoire et du Soudan francais." volume="9" year="1943">Guignot 1943</bibRefCitation>
: 99 (faunistics);
<bibRefCitation id="2669F165F2A8AB8B9EA7CC421E86148C" author="Guignot, F" journalOrPublisher="Memoires du Museum National d'Histoire Naturelle Paris" pageId="256" pageNumber="257" pagination="215 - 322" title="Mission scientifique de l'Omo 6 (58). ColeopteraDytiscidae (2 e partie)." volume="19" year="1946 c">Guignot 1946c</bibRefCitation>
: 284, 312, 315 (discussion);
<bibRefCitation id="7130A0776275BD26150CBD754CA49BF1" author="Guignot, F" journalOrPublisher="Revue de Zoologie et Botanique Africaines" pageId="257" pageNumber="258" pagination="233 - 237" title="Hydrocanthares captures a la lumiere dans le Haut-Katanga." volume="47" year="1953 b">Guignot 1953b</bibRefCitation>
: 237 (discussion);
<bibRefCitation id="E308FA448EC95C129B31ED24F18583FE" author="Omer-Cooper, Joyce" journalOrPublisher="Proceedings of the Royal Entomological Society of London" pageId="260" pageNumber="261" pagination="23 - 31" title="Some new species of Dytiscidae (Coleoptera) from the Cape Province of South Africa." url="10.1111/j.1365-3113.1953.tb00040.x" volume="25" year="1953">Omer-Cooper 1953</bibRefCitation>
: 24 (discussion);
<bibRefCitation id="B8682155484EDE5E8930F5DC56A87291" author="Guignot, F" journalOrPublisher="Bulletin de l'Institut Francais d'Afrique Noire" pageId="257" pageNumber="258" pagination="1095 - 1101" title="Dytiscides et gyrinides de la Guinee francaise recoltes par M. A. Villiers." volume="17" year="1955 b">Guignot 1955b</bibRefCitation>
: 1096 (faunistics);
<bibRefCitation id="F2459F4418DFDD6514A53DAE4B8B85FC" author="Guignot, F" journalOrPublisher="Memoires de l'Institut Francais d'Afrique Noire" pageId="258" pageNumber="259" pagination="211 - 223" title="Coleopteres hydrocanthares. Le Parc National du Niokolo-Koba. 13." volume="48" year="1956 b">Guignot 1956b</bibRefCitation>
: 220, 221 (description, discussion);
<bibRefCitation id="CE1A17B45E7EFDFF3061D1E838F3D790" author="Guignot, F" journalOrPublisher="Annales du Musee Royal du Congo Belge (Serie 8 vo) Sciences Zoologiques" pageId="258" pageNumber="259" pagination="323 - 648" title="Revision des hydrocanthares d'Afrique (ColeopteraDytiscoidea). 2." volume="78" year="1959 a">Guignot 1959a</bibRefCitation>
: 579, 581 (description, faunistics);
<bibRefCitation id="988BEA0256215E9F0533D3FACA7DEE95" author="Guignot, F" journalOrPublisher="Resultats Scientifiques de l'Exploration Hydrobiologique des Lacs Kivu, Edouard et Albert (1952 - 1954)" pageId="258" pageNumber="259" pagination="133 - 169" title="Haliplidae et Dytiscidae (Coleoptera Hydrocanthares)." volume="3" year="1959 d">Guignot 1959d</bibRefCitation>
: 163 (faunistics, discussion);
<bibRefCitation id="A798D6A2B354106C70C82C8C6B8D28BC" author="Guignot, F" journalOrPublisher="Memoires de l'Institut Francais d'Afrique Noire" pageId="258" pageNumber="259" pagination="231 - 239" title="XXIV. ColeopteraDytiscoidea et Gyrinoidea (Supplement). Le Parc National du Niokolo Koba. 2." volume="62" year="1961 b">Guignot 1961b</bibRefCitation>
: 238 (faunistics, discussion);
<bibRefCitation id="8962FD33A1AF4046379A79D1148A5D01" author="Bruneau de Mire, P" journalOrPublisher="Bulletin de l'Institut Francais d'Afrique Noire" pageId="255" pageNumber="256" pagination="838 - 894" title="Les coleopteres hydrocanthares du Tibesti." volume="25" year="1963">
Bruneau de
<normalizedToken id="67DE9100109D1029DF400EF988BCF78D" originalValue="Miré">Mire</normalizedToken>
and Legros 1963
</bibRefCitation>
: 874, 888 (faunistics);
<bibRefCitation id="B48AC8494D1AF4C5D6F96E0A25743E02" author="Omer-Cooper, Joyce" journalOrPublisher="Journal of the Entomological Society of Southern Africa" lastPageId="226" lastPageNumber="227" pageId="261" pageNumber="262" pagination="48 - 63" title="Dytiscidae (Coleoptera) of Nyasaland and Southern Rhodesia. X. Cybisterini and addenda." volume="29" year="1967">
Omer-Cooper
<pageBreakToken id="47229BE84DEF862D242413384AC840CC" pageId="226" pageNumber="227" start="start">1967</pageBreakToken>
</bibRefCitation>
: 60 (discussion,
<taxonomicName id="A12B5BB695EA354F2573A908A9B90D19" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus livingstoni" order="Coleoptera" pageId="226" pageNumber="227" phylum="Arthropoda" rank="species" species="livingstoni">Laccophilus livingstoni</taxonomicName>
O-C. =
<taxonomicName id="BE4850BAA19B5529415B7616E931E254" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus inornatus" order="Coleoptera" pageId="226" pageNumber="227" phylum="Arthropoda" rank="species" species="inornatus">Laccophilus inornatus</taxonomicName>
Zimm.);
<bibRefCitation id="E6ADEE022DDD02CB150FB6ACE776C61A" author="Bilardo, A" journalOrPublisher="Memorie della Societa Entomologica Italiana" pageId="254" pageNumber="255" pagination="93 - 130" title="Recoltes de coleopteres aquatiques Haliplidae et Dytiscidae dans le Gabon et la Cote d'Ivoire." volume="56" year="1978">Bilardo and Pederzani 1978</bibRefCitation>
: 119 (faunistics, description);
<bibRefCitation id="CA021063B422185D3620149AB6063D61" author="Medler, JT" journalOrPublisher="Memoirs of the American Entomological Institute" pageId="259" pageNumber="260" pagination="1 - 919" title="Insects of Nigeria - Check list and bibliography." volume="30" year="1980">Medler 1980</bibRefCitation>
: 155 (faunistics, list);
<bibRefCitation id="6D941B677B25205C095BB1B947B12188" author="Forge, P" journalOrPublisher="- Cahiers de l'ORSTOM (Office de la Recherche Scientifique et Technique Outre-Mer) - Collection-Initiations-Documentations-Techniques" pageId="255" pageNumber="256" pagination="487 - 516" title="Flore et faune aquatiques de l'Afrique Sahelo-Soudanienne. Tome 2. - Coleopteres." volume="45" year="1981">Forge 1981</bibRefCitation>
: 500, 501 (description, faunistics);
<bibRefCitation id="557D67DC9B010D336E09D17C72A7A044" author="Bilardo, A" journalOrPublisher="Atti della Societa Italiana di Scienze Naturali del Museo Civivico di Storia Naturale di Milano" pageId="254" pageNumber="255" pagination="441 - 461" title="Contributo alla conoscenza dei Dytiscidae del Cameroun (Coleoptera)." volume="123" year="1982 a">Bilardo 1982a</bibRefCitation>
: 447 (description, faunistics, given as
<taxonomicName id="69BE2FC22024D140D6A50554BF26105D" genus="Laccophylus" lsidName="Laccophylus" pageId="226" pageNumber="227" rank="genus">Laccophylus</taxonomicName>
);
<bibRefCitation id="F98E11084EC4C357DFDDFC25C46CFFFA" author="Bilardo, A" journalOrPublisher="Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano" pageId="254" pageNumber="255" pagination="157 - 196" title="Haliplidae e Dytiscidae (Coleoptera) del Gabon con note sistematiche sulle specie di confronto. (Parte prima: Haliplidae, Methlinae, Hydroporinae, Noterinae, Laccophilinae)." volume="131" year="1990">Bilardo and Rocchi 1990</bibRefCitation>
: 162, 177 (faunistics, biology);
<bibRefCitation id="9E575828746061FAA325355DE338CEC3" author="Franciscolo, ME" journalOrPublisher="Accademia Nazionale dei Lincei" pageId="256" pageNumber="257" pagination="133 - 153" title="Four new species and new records of Hydradephaga from Sierra Leone (Coleoptera: Haliplidae, Gyrinidae, Dytiscidae). Ricerche biologiche in Sierra Leone. 3." volume="265" year="1990">Franciscolo and Sanfilippo 1990</bibRefCitation>
: 146 (faunistics, description);
<bibRefCitation id="D38F16955BBC3E8DB62EC27A41E5212E" author="Nilsson, AN" journalOrPublisher="Entomologica Fennica" pageId="260" pageNumber="261" pagination="57 - 94" title="Taxonomy, distribution and habitats of the Dytiscidae (Coleoptera) of Ethiopia." volume="4" year="1993">Nilsson and Persson 1993</bibRefCitation>
: 81, 94 (faunistics, biology);
<bibRefCitation id="E95B8B261B4171DFF68F2C396F2D14E7" author="Nilsson, AN" journalOrPublisher="Journal African Zoology" pageId="260" pageNumber="261" pagination="489 - 514" title="The diving beetles (Coleoptera, Dytiscidae) of Guinea Bissau in West Africa." volume="109" year="1995">Nilsson et al. 1995</bibRefCitation>
: 505 (faunistics);
<bibRefCitation id="30AF5EF9D264CD47507CEB4C8ACFF3DF" author="Rocchi, S" journalOrPublisher="Roma" pageId="262" pageNumber="263" pagination="11 - 33" title="Contributo alla conoscenza dei ditiscidi di Guinea con descrizione di duo nuove specie (Coleoptera, Dytiscidae)." volumeTitle="Fragmenta Entomologica" year="2000">Rocchi 2000</bibRefCitation>
: 24 (faunistics);
<bibRefCitation id="EFA84B9439AC1CB5D842F0B3E35F023A" author="Nilsson, AN" journalOrPublisher="World catalogue of Insects" pageId="260" pageNumber="261" pagination="1 - 395" title="Dytiscidae." volume="3" year="2001">Nilsson 2001</bibRefCitation>
: 245 (catalogue, faunistics);
<bibRefCitation id="5B76D2C27D8E61C887DE424FE567EFBA" author="Bilardo, A" journalOrPublisher="Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano" pageId="254" pageNumber="255" pagination="147 - 182" title="Haliplidae e Dytiscidae (Coleoptera) del Gabon. (Parte quarta)." volume="143" year="2002">Bilardo and Rocchi 2002</bibRefCitation>
: 164 (discussion, faunistics);
<bibRefCitation id="24DADD1BC1162CC9CDA920AD58E0B199" author="Reintjes, N" journalOrPublisher="Koleopterologische Rundschau" pageId="262" pageNumber="263" pagination="45 - 74" title="Noteridae and Dytiscidae of the Comoe National Park, Ivory Coast, West Africa (Coleoptera: Noteridae, Dytiscidae)." volume="74" year="2004">Reintjes 2004</bibRefCitation>
: 67 (faunistics); van
<bibRefCitation id="64405446E56778A159156AF829FCC9D7" author="Vondel, BJ van" journalOrPublisher="DEINSEA (Annual of the Natural History Museum Rotterdam)" pageId="263" pageNumber="264" pagination="119 - 138" title="Water beetles from Benin (Coleoptera: Haliplidae, Dytiscidae, Noteridae, Hydraenidae, Hydrochidae, Hydrophilidae, Gyrinidae, Elmidae)." volume="11" year="2005">Vondel 2005</bibRefCitation>
: 131 (faunistics, biology);
<bibRefCitation id="F25F0DEE7315AEC6D27B36303A454101" author="Nilsson, AN" journalOrPublisher="Apollo books, Stenstrup" pageId="260" pageNumber="261" title="A World Catalogue of the Family Dytiscidae, or the Diving Beetles (Coleoptera, Adephaga). Version 1. I. 2015" url="http://www2.emg.umu.se/projects/biginst/andersn/" year="2015">Nilsson 2015</bibRefCitation>
: 212 (catalogue, faunistics). [Comment: Many of the listed references are to be considered uncertain due to extensive species-confusion.]
</paragraph>
</subSubSection>
<subSubSection id="7A7A9C6BC24AB8146B08CC87F70FD765" pageId="226" pageNumber="227" type="type locality">
<paragraph id="D330DA1ABEC13FA371B089F8B23F32BE" pageId="226" pageNumber="227">Type locality.</paragraph>
<paragraph id="6ADB4473E49C8C855A3F4B12FA680D55" pageId="226" pageNumber="227">Tanzania: Usagara.</paragraph>
</subSubSection>
<subSubSection id="D48D7BC953049789E37C62F86F36AB91" pageId="226" pageNumber="227" type="type material">
<paragraph id="C29B7CC511A741DE495B4D90CF331D03" pageId="226" pageNumber="227">Type material studied</paragraph>
<paragraph id="FBB96892E5FFD1BFB04B92EC3BDF0DDC" pageId="226" pageNumber="227">
(3 exs.). Lectotype (by present designation): male: &quot;Usagara / Typus /
<taxonomicName id="17B1F0BA557619FE369290E2F1182DE0" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus inornatus" order="Coleoptera" pageId="226" pageNumber="227" phylum="Arthropoda" rank="species" species="inornatus">Laccophilus inornatus</taxonomicName>
sp. n. Type det. A. Zimmermann /
<taxonomicName id="4DD98D9923A9244B82937AAC23527A78" genus="Algae" lsidName="Algae inornatus" pageId="226" pageNumber="227" rank="species" species="inornatus">inornatus</taxonomicName>
Zimm. sp. n.&quot; (ZMHB). - Paralectotypes: &quot;Daressal. / Type / Samml. A. Zimmermann / Paratypus&quot; (1 ex. ZMSC); &quot;D.O. Afrika / Type / Samml. A. Zimmermann / Paratypus&quot; (1 ex. ZMSC; habitus in Fig. 510).
</paragraph>
</subSubSection>
<subSubSection id="74DBB7B246B63FFE2ABEC29F64850829" lastPageId="227" lastPageNumber="228" pageId="226" pageNumber="227" type="materials_examined">
<paragraph id="5E763362C9DEF1994D8F05185E76442D" pageId="226" pageNumber="227">Additional material studied</paragraph>
<paragraph id="C838862D5AEDFF905C4995E80670475F" lastPageId="227" lastPageNumber="228" pageId="226" pageNumber="227">
(283 exs.). Gambia: &quot;Abuko Nat. Res., at light at the bamboo pool 18.30-20.30 18.11. 1977 UTM 28PCK2181 / Cederholm &amp; al leg. /
<taxonomicName id="2B40DE9730C786F2107A1522E7B2D819" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus inornatus" order="Coleoptera" pageId="226" pageNumber="227" phylum="Arthropoda" rank="species" species="inornatus">Laccophilus inornatus</taxonomicName>
Zimm. Brancucci det.&quot; (1 ex. MZLU); &quot;Central R. Div. Lower Fulladu, Sapo Agric. St. 27.8. 1997, MV light, Woodcock&quot; (1 ex. NHMB); &quot;Bathurst jan. 1968 Palm /
<taxonomicName id="A004207380966030BB1F9021368FEFDD" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus inornatus" order="Coleoptera" pageId="226" pageNumber="227" phylum="Arthropoda" rank="species" species="inornatus">Laccophilus inornatus</taxonomicName>
Zimm. Persson det.&quot; (5 exs. MZLU); &quot;Tanji R, 3 km SW Brufut, at light 19-21.00, 28.2. 1977 Cederholm &amp; al.&quot; (1 ex. NHMB). - Guinea:
<normalizedToken id="8736B51D4C105CCA3A75309331FDE893" originalValue="“Sérédou">&quot;Seredou</normalizedToken>
, lux 4.4. 1975 Zott&quot; (26 exs. ZMHB, 5 exs. MZH; habitus in Fig. 511); same data but &quot;4.5. 1975&quot; (16 exs. ZMHB); same data but
<normalizedToken id="700CAF10C90C7F81C786878BF6366AD5" originalValue="“5.4.1975”">&quot;5.4.1975&quot;</normalizedToken>
(1 ex. MZH); same data but &quot;7-8.4. 1975&quot; (13 exs. ZMHB, 1 ex. MZH). - Liberia: &quot;Suakoko 10.3. 1953 light trap Blickenstaff&quot; (1 ex. USNM); &quot;Suakoko 14.3. 1952 / 6-9 pm light trap Blickenstaff&quot; (1 ex. USNM). - Ivory Coast:
<normalizedToken id="9F621B7083CEC1D439814A9B43F72FD0" originalValue="“Comoé">&quot;Comoe</normalizedToken>
Nat. P.,
<geoCoordinate id="CB643E625E00551E3763664FDAA5BF36" direction="north" orientation="latitude" precision="5555" value="8.5">N8,5°</geoCoordinate>
,
<geoCoordinate id="F086669A4404D25233A7750B62360F4E" direction="west" orientation="longitude" precision="5555" value="-3.5">W3,5°</geoCoordinate>
Reintjes leg. / 11.1. 1999&quot; (3 exs. NHMW); same but &quot;15.4. 1999&quot; (1 ex. NHMW); &quot;Bingerville 8. 1963 J. Decelle&quot; (1 ex. MRAC, 1 ex. MZH); same but &quot;12. 1961&quot; (1 ex. MRAC); same but &quot;9. 1963&quot; (1 ex. MRAC);
<normalizedToken id="109CC030D83505E789131839F07F582B" originalValue="“Danané">&quot;Danane</normalizedToken>
26.7. 1966 Thys &amp; Verheyen&quot; (1 ex. MRAC); &quot;Divo 28.11. 1963 J. Decelle&quot; (3 exs. MRAC, 1 ex. MZH); same but &quot;4.12. 1962&quot; (1 ex. MRAC); &quot;Toumodi / 12. 1930-IV. 1931 Alluaud &amp; Chappuis /
<taxonomicName id="682E54F7F646646CF5F1ECE53BC7F104" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus inornatus" order="Coleoptera" pageId="226" pageNumber="227" phylum="Arthropoda" rank="species" species="inornatus">Laccophilus inornatus</taxonomicName>
Zimm. det. Gschwendtner&quot; (2 exs. MNHN, 1 ex. OLML); &quot;Touba,
<normalizedToken id="DA023F0E60BD91BD2D0FA79E0F1424A2" originalValue="à">a</normalizedToken>
la
<normalizedToken id="7C49A7DE0DE363D0D7FFF4BE44838D78" originalValue="lumière">lumiere</normalizedToken>
4. 2002 Moretto /
<taxonomicName id="CE422F62BA62D235C269664174354E83" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus inornatus" order="Coleoptera" pageId="226" pageNumber="227" phylum="Arthropoda" rank="species" species="inornatus">Laccophilus inornatus</taxonomicName>
Zimm. det. Rocchi 2002&quot; (3 exs. CSR). - Ghana: &quot;Ashanti Reg. Kwadaso, agric. st.
<geoCoordinate id="9A9E09DE7F81F2176436C28E4CAE6E02" direction="north" orientation="latitude" precision="925" value="6.7">6°42'N</geoCoordinate>
,
<geoCoordinate id="988617F7ECEA282CFE5B8C216F95CEEB" direction="west" orientation="longitude" precision="925" value="-1.65">1°39'W</geoCoordinate>
/ light trap 26.2. 1969
<normalizedToken id="A05114BF52D1DECFB4C3A81C5E921A09" originalValue="Endrödy-Younga”">Endroedy-Younga&quot;</normalizedToken>
(4 exs. TMSA, 1 ex. MZH); &quot;Northern Region Damongo Game Res.,
<geoCoordinate id="F3AC7330AF8D2058C054A24E28D88D7A" direction="north" orientation="latitude" precision="925" value="9.066667">9°04'N</geoCoordinate>
, 1°48' / 12.11. 1970&quot; (1 ex. TMSA); &quot;N.R., Damongo Mole game res. 220 m,
<geoCoordinate id="E6CFC609D5CCE88B5151EDFBA3152C63" direction="north" orientation="latitude" precision="925" value="9.066667">9°04'N</geoCoordinate>
,
<geoCoordinate id="3ABF74B72BCDA014389C7F37F3F265C0" direction="south" orientation="latitude" precision="925" value="-1.8">1°48' S</geoCoordinate>
.
<normalizedToken id="09E50268BD6F3EA9A8D05FEE67EC718F" originalValue="Endrödy-Younga">Endroedy-Younga</normalizedToken>
/ on light 12.8. 1971&quot; (1 ex. OLML); &quot;Ghana Ashanti Kumasi, 330 m
<geoCoordinate id="0AA78FACD844D9EE89F2DE87DB73D434" direction="north" orientation="latitude" precision="555" value="6.43">N 6.43</geoCoordinate>
-
<geoCoordinate id="D78668D1798EAF77EE549F21E7122F93" direction="west" orientation="longitude" precision="555" value="-1.36">W 1.36</geoCoordinate>
/ 15.9. 1967 at light,
<normalizedToken id="B8A604B0BCE69EE2457EB6C8FE145075" originalValue="Endrödy-Younga”">Endroedy-Younga&quot;</normalizedToken>
(1 ex. TMSA); &quot;Kumasi 12.5. 67
<normalizedToken id="712E701E4A29756AF2F2566ABF602F88" originalValue="Endrödy-Younga">Endroedy-Younga</normalizedToken>
/
<taxonomicName id="E1503425C41F1C453587C8C448279224" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus inornatus" order="Coleoptera" pageId="226" pageNumber="227" phylum="Arthropoda" rank="species" species="inornatus">Laccophilus inornatus</taxonomicName>
Zimm. det. Wewalka 76&quot; (18 exs. MHNG); same data but &quot;16.6. 67&quot; (10 exs. MHNG); same
<pageBreakToken id="B41992B10CBB08A095BDBED5F1E028A3" pageId="227" pageNumber="228" start="start">data</pageBreakToken>
but &quot;18.5. 67&quot; (13 exs. MHNG). - Benin: &quot;Dep. du Zou, commune de
<normalizedToken id="5956B1BE51AF510DACDCAAA8000CE2D3" originalValue="Zogbodomè">Zogbodome</normalizedToken>
29.1.2006 Goergen / Lokoli Forest
<geoCoordinate id="B96CFFB532336DB934ACED7AB6362FC6" direction="north" orientation="latitude" precision="925" value="7.05">07°03'N</geoCoordinate>
/
<geoCoordinate id="EB6EC50DB1FD725D5DE587FF1C167899" direction="east" orientation="longitude" precision="925" value="2.25">02°15'E</geoCoordinate>
, 17 m a.s.l. light trap&quot; (7 exs. MZH); &quot;Dep. du Zou, commune de
<normalizedToken id="520C1C67FA89EC6BFF097EA864E51B08" originalValue="Zogbodomè">Zogbodome</normalizedToken>
, Lokoli (forest) Hlanzoun riv. 6.2.2006, Goergen, Komarek &amp;
<normalizedToken id="EDE0CEAC560C96333EE7BE1E5DF958E9" originalValue="Hounguè">Houngue</normalizedToken>
/
<geoCoordinate id="732775E3FFEB678E231B91FAC8F54EF9" direction="north" orientation="latitude" precision="925" value="7.5">07°30'N</geoCoordinate>
,
<geoCoordinate id="F0CF6378E1F783D4A2DC35B8F7B55CA8" direction="east" orientation="longitude" precision="925" value="2.25">02°15'E</geoCoordinate>
muddy stream&quot; (1 ex. MZH); &quot;Dep. Atlantique, Allada,
<normalizedToken id="DE286652C87DDA41363C41ED5C1A9BBC" originalValue="Glotomè">Glotome</normalizedToken>
(village) 31.1. 2006 Goergen, Komarek &amp;
<normalizedToken id="BBB188856AF035F14C16864C62C0F891" originalValue="Hounguè">Houngue</normalizedToken>
/
<geoCoordinate id="E4EABCE2FFABB2DE227C4AD5DCB20762" direction="north" orientation="latitude" precision="15" value="6.685">06°41'06&quot;N</geoCoordinate>
,
<geoCoordinate id="F886AF521C93CC4B108475C3D300207C" direction="east" orientation="longitude" precision="15" value="2.0433333">02°02'36&quot;E</geoCoordinate>
, 17 m a.s.l., slowly running stream&quot; (1 ex. MZH); &quot;Dep. Mono,
<normalizedToken id="5BB5627249570A434E9D1C194C053A29" originalValue="Comé">Come</normalizedToken>
,
<normalizedToken id="E5A3DD117DE8EE3BAD99324A4F911D9B" originalValue="Sè">Se</normalizedToken>
(village) 2.2.2006, Goergen, Komarek &amp;
<normalizedToken id="AB5F654CAD82630454F4B89684404599" originalValue="Hounguè">Houngue</normalizedToken>
/
<geoCoordinate id="1510721707783D56B52A758984F2F1F1" direction="north" orientation="latitude" precision="1" value="6.504028">06°30'14,5&quot;N</geoCoordinate>
,
<geoCoordinate id="58B793551B79CA843DE494AB8CFAEDE5" direction="east" orientation="longitude" precision="1" value="1.8303332">01°49'49,2&quot;E</geoCoordinate>
, ca. 20 m a.s.l. pond with muddy ground&quot; (1 ex. MZH); &quot;Dep. Littoral, Toho near Pahou (village) 2.2. 2006, Goergen, Komarek &amp;
<normalizedToken id="10F33A20B7FE9E3D2E6D22E09EFE7939" originalValue="Hounguè">Houngue</normalizedToken>
/
<geoCoordinate id="34701A676FF54E44111372524BDC859E" direction="north" orientation="latitude" precision="1" value="6.386472">06°23'11,3&quot;N</geoCoordinate>
,
<geoCoordinate id="407F55031442997B96B4989EF132ECEF" direction="east" orientation="longitude" precision="1" value="2.208389">02°12'30,2&quot;E</geoCoordinate>
lake with rich riparian vegetation&quot; (2 exs. MZH); &quot;Dep. Littoral Cotonou City, pond, 8.2. 2006, Goergen, Komarek &amp;
<normalizedToken id="16D4EA0C12190E80C49310B9197A1CF4" originalValue="Hounguè”">Houngue&quot;</normalizedToken>
(1 ex. MZH); &quot;Dep. Atlantique Toffo, near Toffo (village) 1.2. 2006, Goergen, Komarek &amp;
<normalizedToken id="40AD85D64FAA4FD4F5260782D8587165" originalValue="Hounguè">Houngue</normalizedToken>
/
<geoCoordinate id="949D6404F444DB0AE3369533F34DCF15" direction="north" orientation="latitude" precision="1" value="6.814389">06°48'51,8&quot;N</geoCoordinate>
,
<geoCoordinate id="B0B2DCF042CC334DF15888F4735D3A0C" direction="east" orientation="longitude" precision="1" value="2.1273887">02°07'38,6&quot;E</geoCoordinate>
, 25 m a.s.l. muddy stream&quot; (1 ex. MZH). - Nigeria: &quot;NW St. Badeggi rice field 8-9.8. 1973 Linnavuori&quot; (54 exs. MZH); &quot;NW St. Yelwa 23.7. 1973 Linnavuori&quot; (1 ex. MZH); &quot;W St. Ife 7-8.7. 1973 Linnavuori&quot; (1 ex. MZH); &quot;EC St. Norcap nr Abakaliki 29.6. 1973 Linnavuori&quot; (2 exs. MZH); same, but &quot;ad lucem&quot; (1 ex. MZH); &quot;Ibadan ca. Jan. - Juni 1954 Stenholt Clausen&quot; (9 exs. ZMUC, 1 ex. MZH); &quot;Ibadan, light trap: campus, Dec. 2003 Goergen&quot; (1 ex. NHMW); &quot;Ibadan at light 26.9. 1956 / G.H. Caswell&quot; (1 ex. BMNH); same data but &quot;27.11. 1955&quot; (1 ex. BMNH); &quot;Kontagora stream 3.IV. 1963&quot; (4 exs. AMGS); &quot;River crossing rd to Erugo about 79 miles from Makurdi 24.4. 1963 JOC.&quot; (4 exs. AMGS); &quot;Marsh, Katsina-Daurra rd. 6.IV. 1963&quot; (1 ex. AMGS); &quot;Stream 64 mi. From Bida on Jebba rd. 12.IV. 1963 JOC.&quot; (1 ex. AMGS); &quot;Stream crossing Kaduna rd. Nr. Zaria 4.IV. 1963 JOC.&quot; (2 exs. AMGS); &quot;Zaria,
<normalizedToken id="DBE53A3726B1AF87A253AF8E4D875C1F" originalValue="à">a</normalizedToken>
la
<normalizedToken id="DE3301D56C987B6CF5AECB4C1A38EE6B" originalValue="lumière">lumiere</normalizedToken>
1969 Roberts H.&quot;(4 exs. MRAC, 2 exs. MZH); &quot;River, Kaduna rd. 13,5 mi. From Jos 13.IV. 1963&quot; (3 exs. AMGS); &quot;R. Kaduna 4,5 mi. from Jos 13. IV. 1963 JOC.&quot; (2 exs. AMGS); &quot;Pools in dry stream bed, Kontagora 5.IV. 1963&quot; (1 ex. AMGS); &quot;Stream, reservoir Yom, Jos Plateau 11.IV. 1963 JOC.&quot; (1 ex. AMGS). - Sudan: &quot;R. Malia
<geoCoordinate id="431C153386D8587F581E63FB4AF84558" direction="east" orientation="longitude" precision="555" value="30.57">30,57E</geoCoordinate>
<geoCoordinate id="5994EC08A8987126A9DC42BABFEF393E" direction="north" orientation="latitude" precision="555" value="40.39">40,39N</geoCoordinate>
29.1. 1954 JJOC.&quot; (1 ex. AMGS); &quot;Minkammon
<geoCoordinate id="DD96BDDC8122482BC1ED5A3CA1B7C641" direction="east" orientation="longitude" precision="555" value="31.31">31,31E</geoCoordinate>
<geoCoordinate id="9340938A6566C9AF6FE9A4D6A67B6F1F" direction="north" orientation="latitude" precision="5555" value="6.2">6,2N</geoCoordinate>
16-17.1. 1954 JJOC.&quot; (1 ex. AMGS); &quot;Equatoria Nimule 11-13.3. 1963 Linnavuori&quot; (1 ex. MZH). - Cameroon: &quot;Yaounde, Bor to Kosti by boat 3-14.3. 1978 Perkins&quot; (3 exs. USNM, 1 ex. MZH); &quot;Minkama 15.4. 1970&quot; (3 exs. NHMB). - Central African Republic: &quot;Bozo 21.5. 1981 / Degallier&quot; (10 exs. NHMB); same but &quot;11. 1981&quot; (2 exs. NHMB); same but &quot;12. 1981&quot; (2 exs. MZH). - Kenya: &quot;Shimba Hills 10.12. 1989
<normalizedToken id="137F7B922E8E8B596EAE4B2D4D6539CC" originalValue="Jäch">Jaech</normalizedToken>
leg.&quot; (1 ex. NHMW). - Tanzania: &quot;Dar es Salaam Tendangiri (?) Exp. /
<taxonomicName id="172C80A233C62E15F1A128EB804AD5A7" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus inornatus" order="Coleoptera" pageId="227" pageNumber="228" phylum="Arthropoda" rank="species" species="inornatus">Laccophilus inornatus</taxonomicName>
<normalizedToken id="2E36704F25542834CEA184D38DC7F05E" originalValue="Rég">Reg</normalizedToken>
. det. M. Brancucci 81&quot; (1 ex. ZMHB);
<normalizedToken id="70391EB84D4AAE75BC94D61963034B35" originalValue="“Daressal.”">&quot;Daressal.&quot;</normalizedToken>
(1 ex. ZMHB). - South Africa: &quot;E Transvaal Hazyview
<geoCoordinate id="F90997ECEB3AC36185D2898FBC79C1B7" direction="south" orientation="latitude" precision="555" value="-25.04">25.04S-</geoCoordinate>
<geoCoordinate id="76BB40C48C99572FFBDA7FF53BC0346D" direction="east" orientation="longitude" precision="555" value="31.07">31.07E</geoCoordinate>
/ 3.4. 1990 E-Y: 2778 UV light trap leg.
<normalizedToken id="7F0F5C7B2003AD785F7BF5DE3B6E9BB4" originalValue="Endrödy-Younga”">Endroedy-Younga&quot;</normalizedToken>
(2 exs. TMSA); &quot;Natal, Waterton Timber Co. 3. 1985 N-28.20.5,
<geoCoordinate id="0EDFC5F147633A9936662C1B9C857B4D" direction="east" orientation="longitude" precision="555" value="32.14">E32.14</geoCoordinate>
, at light Atkinson&quot; (1 ex. NHMB); &quot;ECPr., 6 km S of Port of St Johns, outside Silaka Nature Reserve, pond, S31°38.862, E29°30.551, alt.90 m, 26.1. 2005 Bergsten&quot; (1 ex. NHRS).
</paragraph>
</subSubSection>
<subSubSection id="24B48B71EFEC0E9E4ED5E6A21F1E00C4" pageId="227" pageNumber="228" type="specimen with unclear labelling">
<paragraph id="D063599E6B3097572D3D7BAD6840EEDC" pageId="227" pageNumber="228">Specimen with unclear labelling.</paragraph>
<paragraph id="AC29CF38D39CA30D5F70F9B8D4FF21BD" pageId="227" pageNumber="228">
Egypt: &quot;Kairo /
<normalizedToken id="48BE62D85F687665D40463AAC20E8A77" originalValue="Trägårdh">Traegardh</normalizedToken>
/
<taxonomicName id="0835BAC36F712751B8B481E7CFA8372F" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus restrictus" order="Coleoptera" pageId="227" pageNumber="228" phylum="Arthropoda" rank="species" species="restrictus">Laccophilus restrictus</taxonomicName>
var.?&quot; (1 ex. NHRS).
</paragraph>
</subSubSection>
<subSubSection id="78713022DA6D45475A20CF637E1658B3" pageId="228" pageNumber="229" type="comments on synonymy">
<paragraph id="F31A6C1CA063C317C0458A8EF00EF8C7" pageId="228" pageNumber="229">
<pageBreakToken id="CBFEAB100F55437549209C100E56EA44" pageId="228" pageNumber="229" start="start">Comments</pageBreakToken>
on synonymy.
</paragraph>
<paragraph id="85FA171B2DC6D6E18A650FC1F6F2810F" pageId="228" pageNumber="229">
Examination of type material reveals that
<taxonomicName id="21846A57B6FFED50E4D5C7E054F0B698" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus inornatus" order="Coleoptera" pageId="228" pageNumber="229" phylum="Arthropoda" rank="species" species="inornatus">Laccophilus inornatus</taxonomicName>
is not synonymous with
<taxonomicName id="16A9CDD29C5E2992337E2F84F968AFB9" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus livingstoni" order="Coleoptera" pageId="228" pageNumber="229" phylum="Arthropoda" rank="species" species="livingstoni">Laccophilus livingstoni</taxonomicName>
O-C., as stated by
<bibRefCitation id="39BA8669BF91015B95DB49002A547C5C" author="Omer-Cooper, Joyce" journalOrPublisher="Journal of the Entomological Society of Southern Africa" pageId="261" pageNumber="262" pagination="48 - 63" title="Dytiscidae (Coleoptera) of Nyasaland and Southern Rhodesia. X. Cybisterini and addenda." volume="29" year="1967">Omer-Cooper (1967)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D56A92FC3FDC4DC1EF41DADF038D1F5C" pageId="228" pageNumber="229" type="diagnosis">
<paragraph id="8836C86316A84D1AFAA21E62BD03CF84" pageId="228" pageNumber="229">Diagnosis.</paragraph>
<paragraph id="6BE1E91038B7E90CFBFD3862D28D50A0" pageId="228" pageNumber="229">
Externally
<taxonomicName id="A9041D89064AB71874BEB9F5532ED078" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus inornatus" order="Coleoptera" pageId="228" pageNumber="229" phylum="Arthropoda" rank="species" species="inornatus">Laccophilus inornatus</taxonomicName>
resembles much of
<taxonomicName id="2E7F4E7F4AE9EB98BB33DCD64119B84E" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus canthydroides" order="Coleoptera" pageId="228" pageNumber="229" phylum="Arthropoda" rank="species" species="canthydroides">Laccophilus canthydroides</taxonomicName>
. The two species can be separated by examination of anatomical details exhibited by penis apex; external corner of penis apex distinctly extended in
<taxonomicName id="838DAC360EE5225D227F6F300C338A00" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus inornatus" order="Coleoptera" pageId="228" pageNumber="229" phylum="Arthropoda" rank="species" species="inornatus">Laccophilus inornatus</taxonomicName>
while less so in
<taxonomicName id="85ADBC14DEECA9D378246D34C76C774B" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus canthydroides" order="Coleoptera" pageId="228" pageNumber="229" phylum="Arthropoda" rank="species" species="canthydroides">Laccophilus canthydroides</taxonomicName>
. In general,
<taxonomicName id="CCBAAAA88AB4B719E43B60380B900F57" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus canthydroides" order="Coleoptera" pageId="228" pageNumber="229" phylum="Arthropoda" rank="species" species="canthydroides">Laccophilus canthydroides</taxonomicName>
has small, pale spots on elytral base while
<taxonomicName id="E1FB474DEC40AE38AD5FCD6A7B445CDF" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus inornatus" order="Coleoptera" pageId="228" pageNumber="229" phylum="Arthropoda" rank="species" species="inornatus">Laccophilus inornatus</taxonomicName>
lacks spots on elytral base or has in humeral region, a vague, pale spot.
</paragraph>
</subSubSection>
<subSubSection id="82A13A6B7DD53A3082851D9FB8397AA6" pageId="228" pageNumber="229" type="description">
<paragraph id="FD1FC36EE2AFB806551960DDC9AEDE9D" pageId="228" pageNumber="229">Description.</paragraph>
<paragraph id="6CEC22E0C5A7D09595C02A477274633D" pageId="228" pageNumber="229">Body Length 2.9-3.4 mm, width 1.5-1.9 mm. Dorsal colour pattern of body absent or vague; sometimes elytra with slightly paler base (Figs 505-506). From Ghana I have seen some totally black specimens - possible artefacts?</paragraph>
<paragraph id="FCB69D40BDA6CBD339D0052BA8931546" pageId="228" pageNumber="229">Head: Pale ferrugineous; posteriorly often, close to eyes slightly darker. Finely microsculptured. Reticulation almost simple; difficult to separate two kinds of meshes. Reticulation in part slightly irregular; shape of meshes variable. At eyes and in a narrow medial area between eyes with fine, irregular punctures.</paragraph>
<paragraph id="11047ED11C343B944B253A451636B4AF" pageId="228" pageNumber="229">Pronotum: Pale ferrugineous, frontally and medially at base often with vague, a little darker markings. Finely microsculptured; in part double reticulation discernible. A part of large meshes do not contain small meshes. Difference between large and small meshes indistinct or almost absent. Punctation rather indistinct. Fine, irregular punctures discernible at least at base and laterally.</paragraph>
<paragraph id="C99C66F947D382159D3AEFD3F84D6187" pageId="228" pageNumber="229">Elytra: Ferrugineous to dark ferrugineous. Sometimes elytra slightly paler in anterior half than posteriorly but change of colour is gradual and vague (Figs 510-511). Rather shiny, although finely microsculptured. Reticulation double, but posteriorly in particular, size-classes indistinct and difficult to discern. Anteriorly two kinds of meshes in general present; larger meshes which contain two to six fine meshes. Number of fine meshes sometimes difficult to count. Fine, rather indistinct and scattered puncture discernible. An irregular row of punctures laterally discernible (in posterior half of elytron distinct).</paragraph>
<paragraph id="F5865A4A9E2B567C3355B2D82B869153" pageId="228" pageNumber="229">Ventral aspect: Pale ferrugineous, abdomen, particularly posteriorly somewhat darker, ferrugineous to dark ferrugineous. Rather shiny, finely microsculptured except on abdomen where microsculpture is indistinct. Abdomen, basally with sparse, curved striae. Metacoxal plates with a number of mostly transversely located shallow furrows. Almost impunctate. Apex of prosternal process slender and pointed. Apical ventrite with a small knob on one side (Fig. 182).</paragraph>
<paragraph id="469FB0447B0CAC4FC4C11E98A49251E0" pageId="228" pageNumber="229">Legs: Pro- and mesotarsus rather slender, provided with fine suckers.</paragraph>
<paragraph id="D1BF31C74141A2DB0C209747D0CC1E81" pageId="228" pageNumber="229">Male genitalia: External apex of penis extended and sharp (Fig. 361).</paragraph>
<paragraph id="287AFD3D82396194CED44CE99C24CCCC" pageId="228" pageNumber="229">Female: Apical ventrite lacks knob (Fig. 183). Pro- and mesotarsus slender.</paragraph>
</subSubSection>
<subSubSection id="99690577A1860667C975FB6B196C55BF" pageId="228" pageNumber="229" type="distribution">
<paragraph id="6413403C912B0AFBEE611CEA09F08065" pageId="228" pageNumber="229">Distribution.</paragraph>
<paragraph id="ED63D98E83C4C5EDA3608336C163B2E6" pageId="228" pageNumber="229">Gambia, Guinea, Liberia, Ivory Coast, Ghana, Benin, Nigeria, Sudan, Cameroon, Central African Republic, Kenya, Tanzania, South Africa (Fig. 574). Only verified records are mapped. One specimen labelled Kairo (Egypt) is considered uncertain.</paragraph>
</subSubSection>
<subSubSection id="41B4009FB179ECFFEB538D8023256F10" pageId="228" pageNumber="229" type="collecting circumstances">
<paragraph id="716F23858CD43D36A9056F41B5D93F51" pageId="228" pageNumber="229">Collecting circumstances.</paragraph>
<paragraph id="9542AE920458ADF168053D570A501929" pageId="228" pageNumber="229">
Label data contains some information of the ecology of
<taxonomicName id="86E7A46EADFCD3F25899B652F2C57A07" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus inornatus" order="Coleoptera" pageId="228" pageNumber="229" phylum="Arthropoda" rank="species" species="inornatus">Laccophilus inornatus</taxonomicName>
. Accordingly, it has been collected both in running waters (&quot;muddy stream, slow running stream&quot;) and standing waters (&quot;muddy pond, pools in dry stream bed, in lake with rich riparian vegetation&quot;). Often captured at light collection.
</paragraph>
</subSubSection>
</treatment>
</document>