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<document id="23A9EBCEC2B97B022046FB13639781C6" ID-CLB-Dataset="198864" ID-DOI="10.11646/zootaxa.5214.2.2" ID-GBIF-Dataset="7f28c5c2-9723-4be8-ba5c-0261df939f3c" ID-ISSN="1175-5326" ID-Zenodo-Dep="7384822" ID-ZooBank="9B1E49B6-1057-4994-8BC4-8C11ED69C4E1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1669872087637" checkinUser="plazi" docAuthor="Sáfián, Szabolcs &amp; Collins, Steve C." docDate="2022" docId="33367D24FF85FFDD39AFFF0CA2D042AD" docLanguage="en" docName="zootaxa.5214.2.2.pdf" docOrigin="Zootaxa 5214 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Iolaus brazza Sáfián &amp; Collins 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="185" masterDocId="CF0F055CFF82FFD43938FF9BA16B4300" masterDocTitle="Revisional notes on Iolaus aequatorialis Stempffer &amp; Bennett, 1958 and related species in the subgenus Philiolaus Stempffer &amp; Bennett, 1958 (Lepidoptera, Lycaenidae, Theclinae), with description of two new species" masterLastPageNumber="188" masterPageNumber="176" pageNumber="183" updateTime="1699412660392" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="553A0A74096DA7FC88D40920E2A3173D">Revisional notes on Iolaus aequatorialis Stempffer &amp; Bennett, 1958 and related species in the subgenus Philiolaus Stempffer &amp; Bennett, 1958 (Lepidoptera, Lycaenidae, Theclinae), with description of two new species</mods:title>
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<mods:name id="A8CE589703F331DFAD624301BAD23B40" type="personal">
<mods:role id="57F085186F591DD95735BC4F98B2F4FB">
<mods:roleTerm id="96B8380B4C353AB0F034C250830AF1C9">Author</mods:roleTerm>
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<mods:namePart id="6974745105900479445D9AAD9DFDE18B">Sáfián, Szabolcs</mods:namePart>
<mods:affiliation id="48FA2D196CF748BF5BE66300E73A4942">African Butterfly Research Institute, P. O. Box 14308, 0800 Nairobi, Kenya.</mods:affiliation>
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<mods:role id="6D03C7209CB8AA345876049110441A9D">
<mods:roleTerm id="234270E2E7027930E15F31EDF3E58364">Author</mods:roleTerm>
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<mods:namePart id="F2494A02F0B0532DECC47020EC971F08">Collins, Steve C.</mods:namePart>
<mods:nameIdentifier id="9F47E2CF2CE737C3B765184BFA766767" type="ORCID">0000-0001-8018-5997</mods:nameIdentifier>
<mods:affiliation id="388B473C9741A10E1C57783D971558BB">African Butterfly Research Institute, P. O. Box 14308, 0800 Nairobi, Kenya. &amp; collinsabri @ gmail. com; https: // orcid. org / 0000 - 0001 - 8018 - 5997</mods:affiliation>
<mods:nameIdentifier id="DA459BEF338817EDE45CCFBCD43CA695" type="email">collinsabri@gmail.com</mods:nameIdentifier>
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<mods:typeOfResource id="80A9EED5CFA8CBBE8E14F9D28B8594FE">text</mods:typeOfResource>
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<mods:title id="2C399066D3A58B7C625F98D73DB70407">Zootaxa</mods:title>
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<mods:part id="BF602A4D4A52ACF232B8E5DE86EFE645">
<mods:date id="EAC1984470671B9E46A8E9B4C94E81FE">2022</mods:date>
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<mods:number id="613AE475C5B1824D27583996D0148F09">2022-12-01</mods:number>
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<mods:detail id="53897FB0E465C72762F8D15E8A793B3F" type="volume">
<mods:number id="3D5264ED0E435B4D1006A34837EE31CE">5214</mods:number>
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<mods:classification id="C13CE7E04F8310BF9DD2F2D107BA9DB6">journal article</mods:classification>
<mods:identifier id="2FE31F3060C643BDA55298A7B1DD4E58" type="CLB-Dataset">198864</mods:identifier>
<mods:identifier id="A68E420E2CEA65206EF4D9DB7B3781D8" type="DOI">10.11646/zootaxa.5214.2.2</mods:identifier>
<mods:identifier id="5AECB79BAFD1270483AAD04FB92CEF6F" type="GBIF-Dataset">7f28c5c2-9723-4be8-ba5c-0261df939f3c</mods:identifier>
<mods:identifier id="54A0AEBBCABD6D80B13711974367A799" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="00136CC9E6A2CF51A6C2F20DA0AF945A" type="Zenodo-Dep">7384822</mods:identifier>
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<treatment id="33367D24FF85FFDD39AFFF0CA2D042AD" ID-DOI="http://doi.org/10.5281/zenodo.7386152" ID-GBIF-Taxon="205341206" ID-Zenodo-Dep="7386152" LSID="urn:lsid:plazi:treatment:33367D24FF85FFDD39AFFF0CA2D042AD" httpUri="http://treatment.plazi.org/id/33367D24FF85FFDD39AFFF0CA2D042AD" lastPageId="9" lastPageNumber="185" pageId="7" pageNumber="183">
<subSubSection id="F3859FB9FF85FFD339AFFF0CA0F543B1" box="[151,414,151,178]" pageId="7" pageNumber="183" type="nomenclature">
<paragraph id="BB20CC32FF85FFD339AFFF0CA0F543B1" blockId="7.[151,609,151,214]" box="[151,414,151,178]" pageId="7" pageNumber="183">
<heading id="E0687B5EFF85FFD339AFFF0CA0F543B1" bold="true" box="[151,414,151,178]" fontSize="11" level="1" pageId="7" pageNumber="183" reason="1">
<emphasis id="89EB1020FF85FFD339AFFF0CA0F543B1" bold="true" box="[151,414,151,178]" pageId="7" pageNumber="183">
<taxonomicName id="7C9FB7B1FF85FFD339AFFF0CA05243B2" authority="Sáfián &amp; Collins, 2022" authorityName="Sáfián &amp; Collins" authorityYear="2022" box="[151,313,151,178]" class="Insecta" family="Lycaenidae" genus="Iolaus" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="183" phylum="Arthropoda" rank="species" species="brazza" status="sp. nov.">
<emphasis id="89EB1020FF85FFD339AFFF0CA05243B2" bold="true" box="[151,313,151,178]" italics="true" pageId="7" pageNumber="183">Iolaus brazza</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92D8AD5BFF85FFD33879FF0CA0F543B1" box="[321,414,151,177]" pageId="7" pageNumber="183" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F3859FB9FF85FFD339AFFF20A30A43D5" box="[151,609,187,214]" pageId="7" pageNumber="183" type="description">
<paragraph id="BB20CC32FF85FFD339AFFF20A30A43D5" blockId="7.[151,609,151,214]" box="[151,609,187,214]" pageId="7" pageNumber="183">
<figureCitation id="23A4D0B7FF85FFD339AFFF20A07243D5" box="[151,281,187,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[179,1406,191,1907]" captionTargetId="figure-27@3.[164,1424,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Iolaus males: I. aequatorialis (Uganda, Itwara Forest) upperside—A, underside—D; I. aequatorialis (DRC, Kivu) upperside—B. underside—E; I. mane (Liberia, Nimba Mountains) upperside—C, underside—F; I. ivani (Zambia, Mundwiji Plains) upperside—G, underside—I; I. brazza (Congo, Southwest Brazzaville) upperside—H, underside—J." figureDoi="http://doi.org/10.5281/zenodo.7384828" httpUri="https://zenodo.org/record/7384828/files/figure.png" pageId="7" pageNumber="183">FIGS. 2H,J</figureCitation>
;
<figureCitation id="23A4D0B7FF85FFD3381DFF20A03743D5" box="[293,348,187,213]" captionStart="FIGURE 3" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[165,1416,760,1874]" captionTargetId="figure-292@5.[151,1436,741,1885]" captionTargetPageId="5" captionText="FIGURE 3. Iolaus male genitalia: I. aequatorialis (Uganda, Itwara, Gen. prep.: SAFI00184) lateral view—A, aedeagus—D; I. aequatorialis (DRC, Kivu, Gen. prep.: SAFI00187) lateral view—B, aedeagus—E; I. mane (Liberia, Nimba Mountains, Gen. prep.: SAFI00101) lateral view—C, aedeagus—F; I. ivani (holotype, SAFI00180) lateral view—G, aedeagus—I (vesica everted); I. brazza (holotype, Gen. prep.: SAFI00182) lateral view—H, aedeagus—J." figureDoi="http://doi.org/10.5281/zenodo.7384830" httpUri="https://zenodo.org/record/7384830/files/figure.png" pageId="7" pageNumber="183">3H,J</figureCitation>
;
<figureCitation id="23A4D0B7FF85FFD33850FF20A0E143D5" box="[360,394,187,213]" captionStart="FIGURE 4" captionStartId="6.[151,250,1865,1889]" captionTargetBox="[182,1411,1009,1836]" captionTargetId="figure-445@6.[151,1436,993,1842]" captionTargetPageId="6" captionText="FIGURE 4. Iolaus male genitalia (dorsal view): I. aequatorialis (Uganda, Itwara, Gen. prep.: SAFI00184)—A; I. aequatorialis (DRC, Kivu, Gen. prep.: SAFI00187)—B; I. mane (Liberia, Nimba Mountains, Gen. prep.: SAFI00101)—C; I. ivani (holotype, Gen. prep.: SAFI00180)—D; I. brazza (holotype, Gen. prep.: SAFI00182)—E." figureDoi="http://doi.org/10.5281/zenodo.7384832" httpUri="https://zenodo.org/record/7384832/files/figure.png" pageId="7" pageNumber="183">4E</figureCitation>
;
<figureCitation id="23A4D0B7FF85FFD338AEFF20A0DC43D6" box="[406,439,187,214]" captionStart="FIGURE5" captionStartId="7.[151,250,1941,1965]" captionTargetBox="[186,1389,385,1911]" captionTargetId="figure-71@7.[151,1436,345,1917]" captionTargetPageId="7" captionText="FIGURE5. Iolaus male genitalia (posteriorview):I.aequatorialis (Uganda,Itwara,Gen.prep.:SAFI00184)—A; I.aequatorialis (DRC, Kivu, Gen. prep.: SAFI00187)—B; I. mane (Liberia, Nimba Mountains, Gen. prep.: SAFI00101)—C; I. ivani (holotype, Gen. prep.: SAFI00180)—D; I. brazza (holotype, Gen. prep.: SAFI00182)—E." figureDoi="http://doi.org/10.5281/zenodo.7384834" httpUri="https://zenodo.org/record/7384834/files/figure.png" pageId="7" pageNumber="183">5E</figureCitation>
;
<figureCitation id="23A4D0B7FF85FFD338FBFF20A36E43D6" box="[451,517,187,214]" captionStart="FIGURE 6" captionStartId="8.[151,250,1670,1694]" captionTargetBox="[160,1426,966,1639]" captionTargetId="figure-356@8.[151,1436,957,1647]" captionTargetPageId="8" captionText="FIGURE 6. Iolaus females: I. aequatorialis (Uganda, Itwara Forest) upperside—A, underside—E; I. mane (Guinea, Fouta Djallon) upperside—B, underside—F; I. ivani (paratype, Zambia, Mundwiji Plain) upperside—C, underside—G; I. brazza (paratype, Congo, Southwest Brazzaville) upperside—D, underside—H." figureDoi="http://doi.org/10.5281/zenodo.7384840" httpUri="https://zenodo.org/record/7384840/files/figure.png" pageId="7" pageNumber="183">6C, F</figureCitation>
;
<figureCitation id="23A4D0B7FF85FFD33B28FF20A35F43D5" box="[528,564,187,213]" captionStart="FIGURE 7" captionStartId="9.[151,250,1720,1744]" captionTargetBox="[166,1417,486,1688]" captionTargetId="figure-155@9.[151,1436,453,1697]" captionTargetPageId="9" captionText="FIGURE 7. Female genitalia (lateral view): Iolaus aequatorialis (Uganda, Itwara Forest, Gen. prep.: SAFI00185)—A; I. ivani (paratype, Zambia, Mundwiji Plain, Gen. prep.: SAFI00181)—B; I. brazza (paratype, Congo, southwest Brazzaville, Gen. prep.: SAFI00183)—C." figureDoi="http://doi.org/10.5281/zenodo.7384842" httpUri="https://zenodo.org/record/7384842/files/figure.png" pageId="7" pageNumber="183">7C</figureCitation>
;
<figureCitation id="23A4D0B7FF85FFD33B78FF20A30A43D5" box="[576,609,187,213]" captionStart="FIGURE 8" captionStartId="10.[151,250,1122,1146]" captionTargetBox="[169,1415,200,1090]" captionTargetId="figure-19@10.[151,1436,181,1098]" captionTargetPageId="10" captionText="FIGURE 8. Female genitalia, papillae anales and lamellae (ventral view): Iolaus aequatorialis (Uganda, Itwara Forest, Gen. prep.: SAFI00185)—A; I. ivani (paratype, Zambia, Mundwiji Plain, Gen. prep.: SAFI00181)—B; I. brazza (paratype, Congo, southwest Brazzaville, Gen. prep.: SAFI00183)—C." figureDoi="http://doi.org/10.5281/zenodo.7384844" httpUri="https://zenodo.org/record/7384844/files/figure.png" pageId="7" pageNumber="183">8C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="F3859FB9FF85FFDC39AFFE98A3B343F9" lastPageId="8" lastPageNumber="184" pageId="7" pageNumber="183" type="materials_examined">
<paragraph id="BB20CC32FF85FFD339AFFE98A47B421D" blockId="7.[151,1417,259,322]" box="[151,1296,259,286]" pageId="7" pageNumber="183">
<materialsCitation id="0BF7C66FFF85FFD339AFFE98A466421D" ID-GBIF-Occurrence="3979380305" box="[151,1293,259,286]" collectionCode="R, ABRI" collectorName="Km SW Brazzaville I &amp; S. Collins. Deposited" country="Democratic Republic of the Congo" location="Foulakari" pageId="7" pageNumber="183" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="89EB1020FF85FFD339AFFE98A066421D" bold="true" box="[151,269,259,285]" pageId="7" pageNumber="183">
<typeStatus id="64247290FF85FFD339AFFE98A063421D" box="[151,264,259,285]" pageId="7" pageNumber="183" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="AD9907BBFF85FFD3382CFE9FA042421E" box="[276,297,260,286]" pageId="7" pageNumber="183" type="male">♂</specimenCount>
<collectingCountry id="C3888CA2FF85FFD33808FE98A0FD421D" box="[304,406,259,285]" name="Democratic Republic of the Congo" pageId="7" pageNumber="183">CONGO</collectingCountry>
,
<location id="BE409AE9FF85FFD3389AFE98A366421D" LSID="urn:lsid:plazi:treatment:33367D24FF85FFDD39AFFF0CA2D042AD:BE409AE9FF85FFD3389AFE98A366421D" box="[418,525,259,285]" country="Democratic Republic of the Congo" name="Foulakari" pageId="7" pageNumber="183">Foulakari</location>
<collectionCode id="DD8E54F7FF85FFD33B2CFE9FA341421E" box="[532,554,260,286]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="183">R</collectionCode>
. 85
<collectorName id="166AA9E4FF85FFD33B60FE9FA221421D" box="[600,842,259,286]" pageId="7" pageNumber="183">Km SW Brazzaville I</collectorName>
/93. Leg.:
<collectorName id="166AA9E4FF85FFD33A84FE98A5CF421D" box="[956,1188,259,285]" pageId="7" pageNumber="183">S.Collins. Deposited</collectorName>
in
<collectionCode id="DD8E54F7FF85FFD33DFEFE98A466421D" box="[1222,1293,259,285]" pageId="7" pageNumber="183">ABRI</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="BB20CC32FF85FFD339FFFEBCA4E24241" blockId="7.[151,1417,259,322]" box="[199,1417,295,322]" pageId="7" pageNumber="183">
<materialsCitation id="0BF7C66FFF85FFD339FFFEBCA3314242" ID-GBIF-Occurrence="3979380303" box="[199,602,295,322]" collectionCode="R" country="Democratic Republic of the Congo" location="Foulakari" pageId="7" pageNumber="183" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<emphasis id="89EB1020FF85FFD339FFFEBCA0564241" bold="true" box="[199,317,295,321]" pageId="7" pageNumber="183">
<typeStatus id="64247290FF85FFD339FFFEBCA0534241" box="[199,312,295,321]" pageId="7" pageNumber="183" type="paratype">Paratype</typeStatus>
:
</emphasis>
<specimenCount id="AD9907BBFF85FFD3387CFEB3A0324242" box="[324,345,296,322]" pageId="7" pageNumber="183" type="female">♀</specimenCount>
<collectingCountry id="C3888CA2FF85FFD33858FEBCA0AD4241" box="[352,454,295,321]" name="Democratic Republic of the Congo" pageId="7" pageNumber="183">CONGO</collectingCountry>
,
<location id="BE409AE9FF85FFD338EAFEBCA3564241" LSID="urn:lsid:plazi:treatment:33367D24FF85FFDD39AFFF0CA2D042AD:BE409AE9FF85FFD338EAFEBCA3564241" box="[466,573,295,321]" country="Democratic Republic of the Congo" name="Foulakari" pageId="7" pageNumber="183">Foulakari</location>
<collectionCode id="DD8E54F7FF85FFD33B7CFEB3A3314242" box="[580,602,296,322]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="183">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="0BF7C66FFF85FFD33B5DFEBCA4ED4241" ID-GBIF-Occurrence="3979380306" box="[613,1414,295,322]" collectionCode="ABRI" collectorName="Km SW Brazzaville &amp; S. Collins. Deposited" country="Democratic Republic of the Congo" pageId="7" pageNumber="183" specimenCount="1" typeStatus="paratype">
85
<collectorName id="166AA9E4FF85FFD33BB0FEB3A2024241" box="[648,873,295,322]" pageId="7" pageNumber="183">Km SW Brazzaville</collectorName>
7/92 (bred) Leg.:
<collectorName id="166AA9E4FF85FFD33D0DFEBCA4764241" box="[1077,1309,295,321]" pageId="7" pageNumber="183">S.Collins. Deposited</collectorName>
in
<collectionCode id="DD8E54F7FF85FFD33C07FEBCA4ED4241" box="[1343,1414,295,321]" pageId="7" pageNumber="183">ABRI</collectionCode>
</materialsCitation>
.
</paragraph>
<caption id="EFE09CBAFF85FFD339AFF80EA2D144F5" ID-DOI="http://doi.org/10.5281/zenodo.7384834" ID-Zenodo-Dep="7384834" httpUri="https://zenodo.org/record/7384834/files/figure.png" pageId="7" pageNumber="183" startId="7.[151,250,1941,1965]" targetBox="[186,1389,385,1911]" targetPageId="7">
<paragraph id="BB20CC32FF85FFD339AFF80EA2D144F5" blockId="7.[151,1436,1941,2037]" pageId="7" pageNumber="183">
<emphasis id="89EB1020FF85FFD339AFF80EA07B44AD" bold="true" box="[151,272,1941,1965]" pageId="7" pageNumber="183">FIGURE 5.</emphasis>
<taxonomicName id="7C9FB7B1FF85FFD3382CF80EA03844AD" authorityName="Hubner" authorityYear="1819" box="[276,339,1941,1965]" class="Insecta" family="Lycaenidae" genus="Iolaus" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="183" phylum="Arthropoda" rank="genus">
<emphasis id="89EB1020FF85FFD3382CF80EA03844AD" box="[276,339,1941,1965]" italics="true" pageId="7" pageNumber="183">Iolaus</emphasis>
</taxonomicName>
male genitalia (posterior view):
<taxonomicName id="7C9FB7B1FF85FFD33BAAF80DA24044AD" authorityName="Stempffer &amp; Bennett" authorityYear="1958" box="[658,811,1941,1965]" class="Insecta" family="Lycaenidae" genus="Iolaus" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="183" phylum="Arthropoda" rank="species" species="aequatorialis">
<emphasis id="89EB1020FF85FFD33BAAF80DA24044AD" box="[658,811,1941,1965]" italics="true" pageId="7" pageNumber="183">I. aequatorialis</emphasis>
</taxonomicName>
(Uganda, Itwara, Gen.prep.: SAFI00184)—A;
<taxonomicName id="7C9FB7B1FF85FFD33C3BF80DA4F744AD" authorityName="Stempffer &amp; Bennett" authorityYear="1958" box="[1283,1436,1941,1965]" class="Insecta" family="Lycaenidae" genus="Iolaus" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="183" phylum="Arthropoda" rank="species" species="aequatorialis">
<emphasis id="89EB1020FF85FFD33C3BF80DA4F744AD" box="[1283,1436,1941,1965]" italics="true" pageId="7" pageNumber="183">I. aequatorialis</emphasis>
</taxonomicName>
(DRC, Kivu, Gen. prep.: SAFI00187)—B;
<taxonomicName id="7C9FB7B1FF85FFD33B72F821A3FD44D1" authorityName="Collins &amp; Larsen" authorityYear="2003" box="[586,662,1978,2001]" class="Insecta" family="Lycaenidae" genus="Iolaus" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="183" phylum="Arthropoda" rank="species" species="mane">
<emphasis id="89EB1020FF85FFD33B72F821A3FD44D1" box="[586,662,1978,2001]" italics="true" pageId="7" pageNumber="183">I. mane</emphasis>
</taxonomicName>
(Liberia, Nimba Mountains, Gen. prep.: SAFI00101)—C;
<taxonomicName id="7C9FB7B1FF85FFD33DD0F821A44444D1" authorityName="Sáfián &amp; Collins" authorityYear="2022" box="[1256,1327,1978,2001]" class="Insecta" family="Lycaenidae" genus="Iolaus" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="183" phylum="Arthropoda" rank="species" species="ivani">
<emphasis id="89EB1020FF85FFD33DD0F821A44444D1" box="[1256,1327,1978,2001]" italics="true" pageId="7" pageNumber="183">I. ivani</emphasis>
</taxonomicName>
(holotype, Gen. prep.: SAFI00180)—D;
<taxonomicName id="7C9FB7B1FF85FFD338FEF845A34B44F5" authorityName="Sáfián &amp; Collins" authorityYear="2022" box="[454,544,2013,2037]" class="Insecta" family="Lycaenidae" genus="Iolaus" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="183" phylum="Arthropoda" rank="species" species="brazza">
<emphasis id="89EB1020FF85FFD338FEF845A34B44F5" box="[454,544,2013,2037]" italics="true" pageId="7" pageNumber="183">I. brazza</emphasis>
</taxonomicName>
(holotype, Gen. prep.: SAFI00182)—E.
</paragraph>
</caption>
<paragraph id="BB20CC32FF8AFFDC39FFFF0CA3B343F9" blockId="8.[151,1437,151,933]" pageId="8" pageNumber="184">
<emphasis id="89EB1020FF8AFFDC39FFFF0CA06343B1" bold="true" box="[199,264,151,177]" pageId="8" pageNumber="184">Note:</emphasis>
The male specimen illustrated by
<bibRefCitation id="DF0EB1C3FF8AFFDC3B46FF0CA25843B1" author="d'Abrera, B." box="[638,819,151,177]" pageId="8" pageNumber="184" pagination="531 - 880" refId="ref5349" refString="d'Abrera, B. (2009) s. n. In: Butterflies of the Afrotropical Region. Part III. Lycaenidae, Riodinidae. New and Revised Edition. Hill House Publishers, Melbourne and London, pp. i - xxvi + 531 - 880." type="book chapter" year="2009">d’Abrera (2009)</bibRefCitation>
as
<taxonomicName id="7C9FB7B1FF8AFFDC3A6DFF0CA59043B1" box="[853,1275,151,177]" class="Insecta" family="Lycaenidae" genus="Iolaphilus" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="184" phylum="Arthropoda" rank="subSpecies" species="aequatorialis" subSpecies="aequatorialis">
<emphasis id="89EB1020FF8AFFDC3A6DFF0CA59043B1" box="[853,1275,151,177]" italics="true" pageId="8" pageNumber="184">Iolaphilus aequatorialis aequatorialis</emphasis>
</taxonomicName>
may represent this taxon, as its upperside colour and other features are identical with that of the
<typeStatus id="64247290FF8AFFDC3D2BFF20A51E43D5" box="[1043,1141,187,213]" pageId="8" pageNumber="184" type="holotype">holotype</typeStatus>
(although the blue colours of
<taxonomicName id="7C9FB7B1FF8AFFDC398EFF44A19743F9" authorityName="Hubner" authorityYear="1819" box="[182,252,223,249]" class="Insecta" family="Lycaenidae" genus="Iolaus" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="184" phylum="Arthropoda" rank="genus">
<emphasis id="89EB1020FF8AFFDC398EFF44A19743F9" box="[182,252,223,249]" italics="true" pageId="8" pageNumber="184">Iolaus</emphasis>
</taxonomicName>
are difficult to reproduce in high fidelity).
</paragraph>
</subSubSection>
<subSubSection id="F3859FB9FF8AFFDC39FFFE99A26B40A5" pageId="8" pageNumber="184" type="description">
<paragraph id="BB20CC32FF8AFFDC39FFFE99A3894265" blockId="8.[151,1437,151,933]" pageId="8" pageNumber="184">
<emphasis id="89EB1020FF8AFFDC39FFFE99A0DA421D" bold="true" box="[199,433,258,285]" pageId="8" pageNumber="184">Description of male.</emphasis>
Forewing length:
<quantity id="7C6761D7FF8AFFDC3B4EFE98A3AE421D" box="[630,709,259,285]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="8" pageNumber="184" unit="mm" value="19.0">19 mm</quantity>
.Wingspan:
<quantity id="7C6761D7FF8AFFDC3A7FFE99A2C0421D" box="[839,939,258,285]" metricMagnitude="-2" metricUnit="m" metricValue="3.75" pageId="8" pageNumber="184" unit="mm" value="37.5">37.5 mm</quantity>
. General appearance and main morphological features as for
<taxonomicName id="7C9FB7B1FF8AFFDC3806FEBCA0E64240" authorityName="Sáfián &amp; Collins" authorityYear="2022" box="[318,397,295,320]" class="Insecta" family="Lycaenidae" genus="Iolaus" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="184" phylum="Arthropoda" rank="species" species="ivani">
<emphasis id="89EB1020FF8AFFDC3806FEBCA0E64240" box="[318,397,295,320]" italics="true" pageId="8" pageNumber="184">I. ivani</emphasis>
</taxonomicName>
, but blue colour on upperside royal blue, lacking the turquoise or slight greenish tinge. Submarginal line on forewing underside fine, very faint.
</paragraph>
<paragraph id="BB20CC32FF8AFFDC39FFFEF4A2184119" blockId="8.[151,1437,151,933]" pageId="8" pageNumber="184">
<emphasis id="89EB1020FF8AFFDC39FFFEF4A01D4288" bold="true" box="[199,374,366,393]" pageId="8" pageNumber="184">Male genitalia.</emphasis>
General morphology as for other species in the group as described above in detail. Dorsoventrally the posterior one-third of the right valva narrows down sharply towards the inward turning tip. The left valva remains broad almost all the way to its unusually broad and blunt tip. The ventral edge of the valvae is almost straight in lateral view, slightly bent inwardly. The two strong cornuti on the tip of the aedeagus and on the membrane of the vesica are shorter than those of
<taxonomicName id="7C9FB7B1FF8AFFDC38C2FE64A3CD4119" baseAuthorityName="Stempffer &amp; Bennett" baseAuthorityYear="1958" box="[506,678,511,537]" class="Insecta" family="Lycaenidae" genus="Iolaphilus" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="184" phylum="Arthropoda" rank="species" species="aequatorialis">
<emphasis id="89EB1020FF8AFFDC38C2FE64A3CD4119" box="[506,678,511,537]" italics="true" pageId="8" pageNumber="184">I. aequatorialis</emphasis>
</taxonomicName>
and
<taxonomicName id="7C9FB7B1FF8AFFDC3BE5FE64A2474118" authority="Sáfián &amp; Collins, 2022" authorityName="Sáfián &amp; Collins" authorityYear="2022" box="[733,812,511,536]" class="Insecta" family="Lycaenidae" genus="Iolaus" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="184" phylum="Arthropoda" rank="species" species="ivani" status="sp. nov.">
<emphasis id="89EB1020FF8AFFDC3BE5FE64A2474118" box="[733,812,511,536]" italics="true" pageId="8" pageNumber="184">I. ivani</emphasis>
</taxonomicName>
<emphasis id="89EB1020FF8AFFDC3A0BFE64A2184119" bold="true" box="[819,883,511,537]" pageId="8" pageNumber="184">
<taxonomicNameLabel id="92D8AD5BFF8AFFDC3A0BFE64A2184119" box="[819,883,511,537]" pageId="8" pageNumber="184" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="BB20CC32FF8AFFDC39FFFDB9A02041A9" blockId="8.[151,1437,151,933]" pageId="8" pageNumber="184">
<emphasis id="89EB1020FF8AFFDC39FFFDB9A0B6413C" bold="true" box="[199,477,546,572]" pageId="8" pageNumber="184">Description of female.</emphasis>
Forewing length:
<quantity id="7C6761D7FF8AFFDC3BFBFDB9A259413D" box="[707,818,546,573]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" pageId="8" pageNumber="184" unit="mm" value="18.5">18.5 mm</quantity>
. Wingspan:
<quantity id="7C6761D7FF8AFFDC3AF4FDB8A54D413D" box="[972,1062,547,573]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="8" pageNumber="184" unit="mm" value="36.0">36 mm</quantity>
. General appearance and main morphological features as for
<taxonomicName id="7C9FB7B1FF8AFFDC38D2FDDCA3514160" authority="Sáfián &amp; Collins, 2022" authorityName="Sáfián &amp; Collins" authorityYear="2022" box="[490,570,583,608]" class="Insecta" family="Lycaenidae" genus="Iolaus" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="184" phylum="Arthropoda" rank="species" species="ivani" status="sp. nov.">
<emphasis id="89EB1020FF8AFFDC38D2FDDCA3514160" box="[490,570,583,608]" italics="true" pageId="8" pageNumber="184">I. ivani</emphasis>
</taxonomicName>
<emphasis id="89EB1020FF8AFFDC3B7AFDDCA3E94161" bold="true" box="[578,642,583,609]" pageId="8" pageNumber="184">
<taxonomicNameLabel id="92D8AD5BFF8AFFDC3B7AFDDCA3E94161" box="[578,642,583,609]" pageId="8" pageNumber="184" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, but outer edge of blue area on forewing upperside more scalloped by black indentations along veins 2, 3 and 4. Whitish area completely missing. Underside orange line on forewing brighter orange.
</paragraph>
<paragraph id="BB20CC32FF8AFFDC39FFFD28A26B40A5" blockId="8.[151,1437,151,933]" pageId="8" pageNumber="184">
<emphasis id="89EB1020FF8AFFDC39FFFD28A0F941CC" bold="true" box="[199,402,690,717]" pageId="8" pageNumber="184">Female genitalia.</emphasis>
Papillae anales rather small (&lt;
<quantity id="7C6761D7FF8AFFDC3BD5FD28A24141CD" box="[749,810,691,717]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="184" unit="mm" value="1.0">1 mm</quantity>
dorsoventrally) with almost no hairs. Posterior edge flat with rounded dorsal and ventral edges. Apophyses posterior straight, longer than dorsoventral length of papillae. Only middle section of ductus bursae sclerotised and creased longitudinally; antrum swollen, membranous, including its anterior section before mouth of corpus. Corpus bursae oval, completely membranous with no signum; length slightly longer than half the length of ductus. The lamellae ante and postvaginalis are almost fused, shell-shaped in lateral view; lamella antevaginalis shield-shaped in ventral view, its width approximately
<quantity id="7C6761D7FF8AFFDC3DA8FCFCA5B94081" box="[1168,1234,871,897]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="184" unit="mm" value="2.0">2 mm</quantity>
at its widest with two ear-like lobes laterally, and straight posterior edge.
</paragraph>
</subSubSection>
<caption id="EFE09CBAFF8AFFDC39AFF91DA21D45E7" ID-DOI="http://doi.org/10.5281/zenodo.7384840" ID-Zenodo-Dep="7384840" httpUri="https://zenodo.org/record/7384840/files/figure.png" pageId="8" pageNumber="184" startId="8.[151,250,1670,1694]" targetBox="[160,1426,966,1639]" targetPageId="8">
<paragraph id="BB20CC32FF8AFFDC39AFF91DA21D45E7" blockId="8.[151,1437,1670,1767]" pageId="8" pageNumber="184">
<emphasis id="89EB1020FF8AFFDC39AFF91DA07D459F" bold="true" box="[151,278,1670,1695]" pageId="8" pageNumber="184">FIGURE 6.</emphasis>
<taxonomicName id="7C9FB7B1FF8AFFDC3827F91CA035459F" authorityName="Hubner" authorityYear="1819" box="[287,350,1671,1695]" class="Insecta" family="Lycaenidae" genus="Iolaus" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="184" phylum="Arthropoda" rank="genus">
<emphasis id="89EB1020FF8AFFDC3827F91CA035459F" box="[287,350,1671,1695]" italics="true" pageId="8" pageNumber="184">Iolaus</emphasis>
</taxonomicName>
females:
<taxonomicName id="7C9FB7B1FF8AFFDC38FFF91CA30E459F" baseAuthorityName="Stempffer &amp; Bennett" baseAuthorityYear="1958" box="[455,613,1671,1695]" class="Insecta" family="Lycaenidae" genus="Iolaphilus" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="184" phylum="Arthropoda" rank="species" species="aequatorialis">
<emphasis id="89EB1020FF8AFFDC38FFF91CA30E459F" box="[455,613,1671,1695]" italics="true" pageId="8" pageNumber="184">I. aequatorialis</emphasis>
</taxonomicName>
(Uganda, Itwara Forest) upperside—A, underside—E;
<taxonomicName id="7C9FB7B1FF8AFFDC3D91F91CA593459E" authorityName="Collins &amp; Larsen" authorityYear="2003" box="[1193,1272,1671,1694]" class="Insecta" family="Lycaenidae" genus="Iolaus" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="184" phylum="Arthropoda" rank="species" species="mane">
<emphasis id="89EB1020FF8AFFDC3D91F91CA593459E" box="[1193,1272,1671,1694]" italics="true" pageId="8" pageNumber="184">I. mane</emphasis>
</taxonomicName>
(Guinea, Fouta Djallon) upperside—B, underside—F;
<taxonomicName id="7C9FB7B1FF8AFFDC3B0CF930A31445C2" authorityName="Sáfián &amp; Collins" authorityYear="2022" box="[564,639,1707,1730]" class="Insecta" family="Lycaenidae" genus="Iolaus" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="184" phylum="Arthropoda" rank="species" species="ivani">
<emphasis id="89EB1020FF8AFFDC3B0CF930A31445C2" box="[564,639,1707,1730]" italics="true" pageId="8" pageNumber="184">I. ivani</emphasis>
</taxonomicName>
(paratype, Zambia, Mundwiji Plain) upperside—C, underside—G;
<taxonomicName id="7C9FB7B1FF8AFFDC3C79F930A4F645C3" authorityName="Sáfián &amp; Collins" authorityYear="2022" box="[1345,1437,1707,1731]" class="Insecta" family="Lycaenidae" genus="Iolaus" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="184" phylum="Arthropoda" rank="species" species="brazza">
<emphasis id="89EB1020FF8AFFDC3C79F930A4F645C3" box="[1345,1437,1707,1731]" italics="true" pageId="8" pageNumber="184">I. brazza</emphasis>
</taxonomicName>
(paratype, Congo, Southwest Brazzaville) upperside—D, underside—H.
</paragraph>
</caption>
<subSubSection id="F3859FB9FF8AFFDD39FFF88EA54543F9" lastPageId="9" lastPageNumber="185" pageId="8" pageNumber="184" type="diagnosis">
<paragraph id="BB20CC32FF8AFFDD39FFF88EA54543F9" blockId="8.[151,1436,1813,2019]" lastBlockId="9.[151,1437,151,429]" lastPageId="9" lastPageNumber="185" pageId="8" pageNumber="184">
<emphasis id="89EB1020FF8AFFDC39FFF88EA029442F" bold="true" box="[199,322,1813,1839]" pageId="8" pageNumber="184">Diagnosis.</emphasis>
The male of
<taxonomicName id="7C9FB7B1FF8AFFDC38EFF88DA352442F" authority="Sáfián &amp; Collins, 2022" authorityName="Sáfián &amp; Collins" authorityYear="2022" box="[471,569,1813,1839]" class="Insecta" family="Lycaenidae" genus="Iolaus" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="184" phylum="Arthropoda" rank="species" species="brazza" status="sp. nov.">
<emphasis id="89EB1020FF8AFFDC38EFF88DA352442F" box="[471,569,1813,1839]" italics="true" pageId="8" pageNumber="184">I. brazza</emphasis>
</taxonomicName>
<emphasis id="89EB1020FF8AFFDC3B78F88EA3EB442F" bold="true" box="[576,640,1813,1839]" pageId="8" pageNumber="184">
<taxonomicNameLabel id="92D8AD5BFF8AFFDC3B78F88EA3EB442F" box="[576,640,1813,1839]" pageId="8" pageNumber="184" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
has a distinctive tone of royal blue on the upperside, with no turquoise or greenish tinge (
<figureCitation id="23A4D0B7FF8AFFDC3858F8A2A0B14453" box="[352,474,1849,1875]" captionStart="FIGURE 2" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[179,1406,191,1907]" captionTargetId="figure-27@3.[164,1424,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Iolaus males: I. aequatorialis (Uganda, Itwara Forest) upperside—A, underside—D; I. aequatorialis (DRC, Kivu) upperside—B. underside—E; I. mane (Liberia, Nimba Mountains) upperside—C, underside—F; I. ivani (Zambia, Mundwiji Plains) upperside—G, underside—I; I. brazza (Congo, Southwest Brazzaville) upperside—H, underside—J." figureDoi="http://doi.org/10.5281/zenodo.7384828" httpUri="https://zenodo.org/record/7384828/files/figure.png" pageId="8" pageNumber="184">Figs. 2H, J</figureCitation>
). The female lacks the whitish-blue area (
<figureCitation id="23A4D0B7FF8AFFDC3A90F8A2A54E4453" box="[936,1061,1849,1875]" captionStart="FIGURE 6" captionStartId="8.[151,250,1670,1694]" captionTargetBox="[160,1426,966,1639]" captionTargetId="figure-356@8.[151,1436,957,1647]" captionTargetPageId="8" captionText="FIGURE 6. Iolaus females: I. aequatorialis (Uganda, Itwara Forest) upperside—A, underside—E; I. mane (Guinea, Fouta Djallon) upperside—B, underside—F; I. ivani (paratype, Zambia, Mundwiji Plain) upperside—C, underside—G; I. brazza (paratype, Congo, Southwest Brazzaville) upperside—D, underside—H." figureDoi="http://doi.org/10.5281/zenodo.7384840" httpUri="https://zenodo.org/record/7384840/files/figure.png" pageId="8" pageNumber="184">Figs. 6C, F</figureCitation>
) present in the three other species in the group. In the male genitalia
<taxonomicName id="7C9FB7B1FF8AFFDC3B2DF8C5A31C4477" authority="Sáfián &amp; Collins, 2022" authorityName="Sáfián &amp; Collins" authorityYear="2022" box="[533,631,1885,1911]" class="Insecta" family="Lycaenidae" genus="Iolaus" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="184" phylum="Arthropoda" rank="species" species="brazza" status="sp. nov.">
<emphasis id="89EB1020FF8AFFDC3B2DF8C5A31C4477" box="[533,631,1885,1911]" italics="true" pageId="8" pageNumber="184">I. brazza</emphasis>
</taxonomicName>
<emphasis id="89EB1020FF8AFFDC3B45F8C6A3D74477" bold="true" box="[637,700,1885,1911]" pageId="8" pageNumber="184">
<taxonomicNameLabel id="92D8AD5BFF8AFFDC3B45F8C6A3D74477" box="[637,700,1885,1911]" pageId="8" pageNumber="184" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="7C9FB7B1FF8AFFDC3A76F8C5A2914477" baseAuthorityName="Stempffer &amp; Bennett" baseAuthorityYear="1958" box="[846,1018,1885,1911]" class="Insecta" family="Lycaenidae" genus="Iolaphilus" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="184" phylum="Arthropoda" rank="species" species="aequatorialis">
<emphasis id="89EB1020FF8AFFDC3A76F8C5A2914477" box="[846,1018,1885,1911]" italics="true" pageId="8" pageNumber="184">I. aequatorialis</emphasis>
</taxonomicName>
and
<taxonomicName id="7C9FB7B1FF8AFFDC3D08F8C5A5154477" authorityName="Sáfián &amp; Collins" authorityYear="2022" box="[1072,1150,1886,1911]" class="Insecta" family="Lycaenidae" genus="Iolaus" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="184" phylum="Arthropoda" rank="species" species="ivani">
<emphasis id="89EB1020FF8AFFDC3D08F8C5A5154477" box="[1072,1150,1886,1911]" italics="true" pageId="8" pageNumber="184">I. ivani</emphasis>
</taxonomicName>
in the shorter and blunter tip and the straighter ventral edge of the valvae (
<figureCitation id="23A4D0B7FF8AFFDC3B97F81AA278449B" box="[687,787,1921,1947]" captionStart="FIGURE 3" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[165,1416,760,1874]" captionTargetId="figure-292@5.[151,1436,741,1885]" captionTargetPageId="5" captionText="FIGURE 3. Iolaus male genitalia: I. aequatorialis (Uganda, Itwara, Gen. prep.: SAFI00184) lateral view—A, aedeagus—D; I. aequatorialis (DRC, Kivu, Gen. prep.: SAFI00187) lateral view—B, aedeagus—E; I. mane (Liberia, Nimba Mountains, Gen. prep.: SAFI00101) lateral view—C, aedeagus—F; I. ivani (holotype, SAFI00180) lateral view—G, aedeagus—I (vesica everted); I. brazza (holotype, Gen. prep.: SAFI00182) lateral view—H, aedeagus—J." figureDoi="http://doi.org/10.5281/zenodo.7384830" httpUri="https://zenodo.org/record/7384830/files/figure.png" pageId="8" pageNumber="184">Figs. 3H</figureCitation>
;
<figureCitation id="23A4D0B7FF8AFFDC3A26F81AA22B449B" box="[798,832,1921,1947]" captionStart="FIGURE 4" captionStartId="6.[151,250,1865,1889]" captionTargetBox="[182,1411,1009,1836]" captionTargetId="figure-445@6.[151,1436,993,1842]" captionTargetPageId="6" captionText="FIGURE 4. Iolaus male genitalia (dorsal view): I. aequatorialis (Uganda, Itwara, Gen. prep.: SAFI00184)—A; I. aequatorialis (DRC, Kivu, Gen. prep.: SAFI00187)—B; I. mane (Liberia, Nimba Mountains, Gen. prep.: SAFI00101)—C; I. ivani (holotype, Gen. prep.: SAFI00180)—D; I. brazza (holotype, Gen. prep.: SAFI00182)—E." figureDoi="http://doi.org/10.5281/zenodo.7384832" httpUri="https://zenodo.org/record/7384832/files/figure.png" pageId="8" pageNumber="184">4E</figureCitation>
;
<figureCitation id="23A4D0B7FF8AFFDC3A73F81AA206449C" box="[843,877,1921,1948]" captionStart="FIGURE5" captionStartId="7.[151,250,1941,1965]" captionTargetBox="[186,1389,385,1911]" captionTargetId="figure-71@7.[151,1436,345,1917]" captionTargetPageId="7" captionText="FIGURE5. Iolaus male genitalia (posteriorview):I.aequatorialis (Uganda,Itwara,Gen.prep.:SAFI00184)—A; I.aequatorialis (DRC, Kivu, Gen. prep.: SAFI00187)—B; I. mane (Liberia, Nimba Mountains, Gen. prep.: SAFI00101)—C; I. ivani (holotype, Gen. prep.: SAFI00180)—D; I. brazza (holotype, Gen. prep.: SAFI00182)—E." figureDoi="http://doi.org/10.5281/zenodo.7384834" httpUri="https://zenodo.org/record/7384834/files/figure.png" pageId="8" pageNumber="184">5E</figureCitation>
). The cornuti on the tip of the aedeagus and on the membrane of the vesica are very short (more similar to
<taxonomicName id="7C9FB7B1FF8AFFDC3BC4F83DA23B44BF" authorityName="Collins &amp; Larsen" authorityYear="2003" box="[764,848,1958,1983]" class="Insecta" family="Lycaenidae" genus="Iolaus" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="184" phylum="Arthropoda" rank="species" species="mane">
<emphasis id="89EB1020FF8AFFDC3BC4F83DA23B44BF" box="[764,848,1958,1983]" italics="true" pageId="8" pageNumber="184">I. mane</emphasis>
</taxonomicName>
) (
<figureCitation id="23A4D0B7FF8AFFDC3A5FF83EA2B044BF" box="[871,987,1957,1983]" captionStart="FIGURE 3" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[165,1416,760,1874]" captionTargetId="figure-292@5.[151,1436,741,1885]" captionTargetPageId="5" captionText="FIGURE 3. Iolaus male genitalia: I. aequatorialis (Uganda, Itwara, Gen. prep.: SAFI00184) lateral view—A, aedeagus—D; I. aequatorialis (DRC, Kivu, Gen. prep.: SAFI00187) lateral view—B, aedeagus—E; I. mane (Liberia, Nimba Mountains, Gen. prep.: SAFI00101) lateral view—C, aedeagus—F; I. ivani (holotype, SAFI00180) lateral view—G, aedeagus—I (vesica everted); I. brazza (holotype, Gen. prep.: SAFI00182) lateral view—H, aedeagus—J." figureDoi="http://doi.org/10.5281/zenodo.7384830" httpUri="https://zenodo.org/record/7384830/files/figure.png" pageId="8" pageNumber="184">Figs. 3F, J</figureCitation>
). In the female genitalia only the middle section of the ductus bursae is sclerotised (the antrum and the anterior section are membranous), while almost the entire bursa is sclerotised in
<taxonomicName id="7C9FB7B1FF8BFFDD38E7FF0CA3E643B1" baseAuthorityName="Stempffer &amp; Bennett" baseAuthorityYear="1958" box="[479,653,151,177]" class="Insecta" family="Lycaenidae" genus="Iolaphilus" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="185" phylum="Arthropoda" rank="species" species="aequatorialis">
<emphasis id="89EB1020FF8BFFDD38E7FF0CA3E643B1" box="[479,653,151,177]" italics="true" pageId="9" pageNumber="185">I. aequatorialis</emphasis>
</taxonomicName>
and
<taxonomicName id="7C9FB7B1FF8BFFDD3BFFFF0CA27343B0" authorityName="Sáfián &amp; Collins" authorityYear="2022" box="[711,792,151,176]" class="Insecta" family="Lycaenidae" genus="Iolaus" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="185" phylum="Arthropoda" rank="species" species="ivani">
<emphasis id="89EB1020FF8BFFDD3BFFFF0CA27343B0" box="[711,792,151,176]" italics="true" pageId="9" pageNumber="185">I. ivani</emphasis>
</taxonomicName>
(
<figureCitation id="23A4D0B7FF8BFFDD3A11FF0CA20443B1" box="[809,879,151,177]" captionStart="FIGURE 7" captionStartId="9.[151,250,1720,1744]" captionTargetBox="[166,1417,486,1688]" captionTargetId="figure-155@9.[151,1436,453,1697]" captionTargetPageId="9" captionText="FIGURE 7. Female genitalia (lateral view): Iolaus aequatorialis (Uganda, Itwara Forest, Gen. prep.: SAFI00185)—A; I. ivani (paratype, Zambia, Mundwiji Plain, Gen. prep.: SAFI00181)—B; I. brazza (paratype, Congo, southwest Brazzaville, Gen. prep.: SAFI00183)—C." figureDoi="http://doi.org/10.5281/zenodo.7384842" httpUri="https://zenodo.org/record/7384842/files/figure.png" pageId="9" pageNumber="185">Fig. 7</figureCitation>
). The lamella antevaginalis of
<taxonomicName id="7C9FB7B1FF8BFFDD3DE9FF0CA45E43B1" authority="Sáfián &amp; Collins, 2022" authorityName="Sáfián &amp; Collins" authorityYear="2022" box="[1233,1333,151,177]" class="Insecta" family="Lycaenidae" genus="Iolaus" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="185" phylum="Arthropoda" rank="species" species="brazza" status="sp. nov.">
<emphasis id="89EB1020FF8BFFDD3DE9FF0CA45E43B1" box="[1233,1333,151,177]" italics="true" pageId="9" pageNumber="185">I. brazza</emphasis>
</taxonomicName>
<emphasis id="89EB1020FF8BFFDD3C05FF0CA41543B1" bold="true" box="[1341,1406,151,177]" pageId="9" pageNumber="185">
<taxonomicNameLabel id="92D8AD5BFF8BFFDD3C05FF0CA41543B1" box="[1341,1406,151,177]" pageId="9" pageNumber="185" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
in ventral view is shield-shape with two ear-like lobes ventrally, similar to that of
<taxonomicName id="7C9FB7B1FF8BFFDD3D1CFF20A5BA43D5" baseAuthorityName="Stempffer &amp; Bennett" baseAuthorityYear="1958" box="[1060,1233,187,213]" class="Insecta" family="Lycaenidae" genus="Iolaphilus" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="185" phylum="Arthropoda" rank="species" species="aequatorialis">
<emphasis id="89EB1020FF8BFFDD3D1CFF20A5BA43D5" box="[1060,1233,187,213]" italics="true" pageId="9" pageNumber="185">I. aequatorialis</emphasis>
</taxonomicName>
, but the posterior edge is almost straight in
<taxonomicName id="7C9FB7B1FF8BFFDD388EFF44A37343F9" authority="Sáfián &amp; Collins, 2022" authorityName="Sáfián &amp; Collins" authorityYear="2022" box="[438,536,223,249]" class="Insecta" family="Lycaenidae" genus="Iolaus" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="185" phylum="Arthropoda" rank="species" species="brazza" status="sp. nov.">
<emphasis id="89EB1020FF8BFFDD388EFF44A37343F9" box="[438,536,223,249]" italics="true" pageId="9" pageNumber="185">I. brazza</emphasis>
</taxonomicName>
<emphasis id="89EB1020FF8BFFDD3B27FF44A33443F9" bold="true" box="[543,607,223,249]" pageId="9" pageNumber="185">
<taxonomicNameLabel id="92D8AD5BFF8BFFDD3B27FF44A33443F9" box="[543,607,223,249]" pageId="9" pageNumber="185" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, and slightly undulate in
<taxonomicName id="7C9FB7B1FF8BFFDD3A43FF44A54C43F9" baseAuthorityName="Stempffer &amp; Bennett" baseAuthorityYear="1958" box="[891,1063,223,249]" class="Insecta" family="Lycaenidae" genus="Iolaphilus" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="185" phylum="Arthropoda" rank="species" species="aequatorialis">
<emphasis id="89EB1020FF8BFFDD3A43FF44A54C43F9" box="[891,1063,223,249]" italics="true" pageId="9" pageNumber="185">I. aequatorialis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="F3859FB9FF8BFFDD39FFFE98A34D4241" pageId="9" pageNumber="185" type="etymology">
<paragraph id="BB20CC32FF8BFFDD39FFFE98A34D4241" blockId="9.[151,1437,151,429]" pageId="9" pageNumber="185">
<emphasis id="89EB1020FF8BFFDD39FFFE98A025421D" bold="true" box="[199,334,259,285]" pageId="9" pageNumber="185">Etymology.</emphasis>
The name of the new species refers to Brazzaville, the capital city of the
<collectingCountry id="C3888CA2FF8BFFDD3DACFE98A406421D" box="[1172,1389,259,285]" name="Republic of the Congo" pageId="9" pageNumber="185">Republic of Congo</collectingCountry>
, the broader
<typeStatus id="64247290FF8BFFDD39CCFEBCA04F4241" box="[244,292,295,321]" pageId="9" pageNumber="185">type</typeStatus>
locality of the species.
</paragraph>
</subSubSection>
<subSubSection id="F3859FB9FF8BFFDD39FFFED1A2D042AD" pageId="9" pageNumber="185" type="discussion">
<paragraph id="BB20CC32FF8BFFDD39FFFED1A2D042AD" blockId="9.[151,1437,151,429]" pageId="9" pageNumber="185">
<emphasis id="89EB1020FF8BFFDD39FFFED1A0274264" bold="true" box="[199,332,330,356]" pageId="9" pageNumber="185">Discussion.</emphasis>
So far, the species is confirmed from the broader Brazzaville area of the
<collectingCountry id="C3888CA2FF8BFFDD3DBBFED0A4324265" box="[1155,1369,331,357]" name="Republic of the Congo" pageId="9" pageNumber="185">Republic of Congo</collectingCountry>
south of the Congolian Forest Zone, in the transition toward the southern African woodland and savanna areas. Further records from the forest zone proper need confirmation (see also below).
</paragraph>
</subSubSection>
</treatment>
</document>
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