174 lines
19 KiB
XML
174 lines
19 KiB
XML
<document id="600EE7CBE9BC592CE6B92BD671217C14" ID-CLB-Dataset="28501" ID-DOI="http://dx.doi.org/10.3897/zookeys.826.26488" ID-GBIF-Dataset="9d62af83-e4f5-423b-8e12-7dd567f69921" ID-PMC="PMC6405737" ID-Pensoft-Pub="1313-2970-826-1" ID-PubMed="30858752" ID-ZooBank="B1550A3AE54744509A44BC4366D5E110" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1313-2970-826-1" ModsDocOrigin="ZooKeys 826" ModsDocTitle="Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records" checkinTime="1551446249571" checkinUser="pensoft" docAuthor="Chamorro, William, Marin-Armijos, Diego, senjo, Angelico & Vaz-De-Mello, Fernando Z." docDate="2019" docId="268C8D20FC16EC0C21A51E3B4603C61F" docLanguage="en" docName="ZooKeys 826: 1-343" docOrigin="ZooKeys 826" docSource="http://dx.doi.org/10.3897/zookeys.826.26488" docTitle="Dichotomius (Dichotomius) mamillatus Felsche 1901" docType="treatment" docVersion="7" lastPageNumber="112" masterDocId="FFECFFBAFFFC2D25D00B9843FFD67749" masterDocTitle="Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records" masterLastPageNumber="343" masterPageNumber="1" pageNumber="110" updateTime="1701379963039" updateUser="ExternalLinkService">
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<mods:title id="E4F9B62D344715E6FFFF8E6525730433">Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records</mods:title>
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<mods:namePart id="B76D6420FFFC1D3188DF86919BA1357D">Chamorro, William</mods:namePart>
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<mods:namePart id="4770625B05B38F8A9AED017AFA9EA883">Marin-Armijos, Diego</mods:namePart>
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<mods:namePart id="A0ECBC98207415BE1FEF72B247E90FFA">senjo, Angelico</mods:namePart>
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<mods:namePart id="98D05A692EAA78E9B2852A3716963BCF">Vaz-De-Mello, Fernando Z.</mods:namePart>
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<mods:typeOfResource id="97F5583A3433F71A72195D9ED511E8B0">text</mods:typeOfResource>
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<mods:date id="37473F46843377627DB1330A78473B3C">2019</mods:date>
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<mods:detail id="2388355E248EC6A5708412FB80ECF0AE" type="volume">
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<mods:number id="36BF0EFF5C16A540E818CEBB2915B123">826</mods:number>
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<mods:classification id="DDB4E1C37A2CDD75264CC4BEDEE757A1">journal article</mods:classification>
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<mods:identifier id="E4727A312EC1F1A041CF9B8049D5EF9B" type="DOI">http://dx.doi.org/10.3897/zookeys.826.26488</mods:identifier>
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<treatment id="268C8D20FC16EC0C21A51E3B4603C61F" ID-GBIF-Taxon="154127767" LSID="urn:lsid:plazi:treatment:268C8D20FC16EC0C21A51E3B4603C61F" httpUri="http://treatment.plazi.org/id/268C8D20FC16EC0C21A51E3B4603C61F" lastPageId="111" lastPageNumber="112" pageId="109" pageNumber="110">
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<subSubSection id="76F240A3260678F3F9868987299FCEC9" pageId="109" pageNumber="110" type="nomenclature">
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<paragraph id="726D38DE903181AC3D1B65650561FE3C" pageId="109" pageNumber="110">
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<taxonomicName id="BECB74F40E725B9BC531B13D375449E5" ID-CoL="7ZN4F" authority="Felsche, 1901" authorityName="Felsche" authorityYear="1901" class="Insecta" family="Scarabaeidae" genus="Dichotomius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dichotomius (Dichotomius) mamillatus" order="Coleoptera" pageId="109" pageNumber="110" phylum="Arthropoda" rank="species" species="mamillatus" subGenus="Dichotomius">Dichotomius (Dichotomius) mamillatus (Felsche, 1901)</taxonomicName>
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Plate 25B
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</paragraph>
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</subSubSection>
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<subSubSection id="334F6ED457188C41694B1AC15C23EF95" pageId="110" pageNumber="111" type="reference_group">
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<paragraph id="87F86120E4B3BB54B51240B769E5D8A0" pageId="110" pageNumber="111">
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<taxonomicName id="42464ED778ECA2C69789C84A94B0D0D1" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pinotus mamillatus" order="Coleoptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="species" species="mamillatus">
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<pageBreakToken id="AC7B196A19F7E0B43D640499CEDAFF5C" pageId="110" pageNumber="111" start="start">Pinotus</pageBreakToken>
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mamillatus
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</taxonomicName>
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Felsche, 1901: 143 (original description. Type locality: Ecuador).
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</paragraph>
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<paragraph id="87014950AA3BE35C7AB23DB1917577F3" pageId="110" pageNumber="111">
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<taxonomicName id="3433EFD5CEA4E9470DE19FE79CC587D6" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pinotus mamillatus" order="Coleoptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="species" species="mamillatus">Pinotus mamillatus</taxonomicName>
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:
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<bibRefCitation id="05E210EC077891B8904335243CF2DF36" author="Gillet, JJE" journalOrPublisher="IW Junk, Berlin, Germany" pageId="268" pageNumber="269" title="Coleopterorum Catalogus. Pars 38: Scarabaeidae: Coprinae." year="1911 a">Gillet 1911a</bibRefCitation>
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: 61 (complete list of species);
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<bibRefCitation id="6897C46D5B9565DF672AF4012B923866" author="Luederwaldt, H" journalOrPublisher="Revista do Museu Paulista" pageId="272" pageNumber="273" pagination="1 - 174" title="As especies brasileiras do genero Pinotus, com algumas consideractes tambem sobra outras especies." volume="16" year="1929">Luederwaldt 1929</bibRefCitation>
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: 53 (characters in key);
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<bibRefCitation id="61C6F36BCB1CB00071FF00A514936C71" author="Balthasar, V" journalOrPublisher="Beitrag Fauna Perus, Jena" pageId="262" pageNumber="263" pagination="337 - 358" title="Scarabaeidae laparostictae." volume="1" year="1941">Balthasar 1941</bibRefCitation>
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: 349 (cited for Peru);
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<bibRefCitation id="5DFEC533CA155EFE92E891E58D33E699" author="Blackwelder, RE" journalOrPublisher="National Museum Bulletin" pageId="262" pageNumber="263" pagination="189 - 341" title="Checklist of the coleopterous insects of Mexico, Central America, West Indies and South America. Part 2." volume="185" year="1944">Blackwelder 1944</bibRefCitation>
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: 207 (list of species from Latin America);
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<bibRefCitation id="29405A2706C6C2C15CD5C30A681309EC" author="Balthasar, V" journalOrPublisher="Beitraege zur Fauna Perus, Jena" pageId="262" pageNumber="263" pagination="322 - 344" title="Scarabaeidae laparostictae." volume="2" year="1951">Balthasar 1951</bibRefCitation>
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: 334 (cited for Peru).
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</paragraph>
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<paragraph id="BB2535729BC25F880C37A61312655CCC" pageId="110" pageNumber="111">
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<taxonomicName id="A361B124137B7EF65684BAF3E137F4B5" class="Insecta" family="Scarabaeidae" genus="Dichotomius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dichotomius mamillatus" order="Coleoptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="species" species="mamillatus">Dichotomius mamillatus</taxonomicName>
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:
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<bibRefCitation id="2C36A2790DFA378DDE86DC493DD52492" author="Vulcano, MA" journalOrPublisher="Atas do Simposio sobre a Biota Amazonica" pageId="279" pageNumber="280" pagination="533 - 603" title="Sinopse dos Passalidae e Scarabaeidae s. str. da regiao amazonica (Insecta, Coleoptera)." volume="5" year="1967">Vulcano and Pereira 1967</bibRefCitation>
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: 585 (characters in key);
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<bibRefCitation id="D881BCC5D1B5E50501E9D603E7110058" author="Vaz-de-Mello, FZ" editor="Martin-Piera, F" journalOrPublisher="Sociedad Entomologica Aragonesa" pageId="278" pageNumber="279" pagination="183 - 195" title="Estado atual de conhecimentos dos Scarabaeidae s. str. (Coleoptera: Scarabaeoidea) do Brasil." volume="1" volumeTitle="Hacia un proyecto CYTED para el inventario y estimacion de la diversidad entomologica en Iberoamerica: PRIBES- 2000. Monografias Tercer Milenio m 3 m." year="2000">Vaz-de-Mello 2000</bibRefCitation>
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: 193 (cited for Brazil);
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<bibRefCitation id="456D03F5ED969F5946B710D74FD0CED0" author="Medina, CA" journalOrPublisher="Biota Colombiana" pageId="274" pageNumber="275" pagination="131 - 144" title="Escarabajos coprofagos (Coleoptera: Scarabaeinae) de Colombia." volume="2" year="2001">Medina et al. 2001</bibRefCitation>
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: 138 (cited for Colombia);
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<bibRefCitation id="120EA1CE95920B83833133DFBD52F85C" author="Sarmiento-Garces, R" journalOrPublisher="Revista de la Academia Colombiana de Ciencias Exactas, Fisicas y Naturales" pageId="276" pageNumber="277" pagination="285 - 296" title="Escarabajos del genero Dichotomius Hope, 1838 (Scarabaeidae: Scarabaeinae) en la amazonia colombiana." volume="33" year="2009">
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<normalizedToken id="5BBFD0B3C7C79A2E7ED6C997AB8F2192" originalValue="Sarmiento-Garcés">Sarmiento-Garces</normalizedToken>
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and
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<normalizedToken id="1160D643E0A19C1DCB14EB6F27CAA3A4" originalValue="Amat-García">Amat-Garcia</normalizedToken>
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2009
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</bibRefCitation>
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: 291 (diagnosis), 295 (characters in key);
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<bibRefCitation id="7A350CA936F5AEC4368E458DFFC85075" author="Carvajal, V" journalOrPublisher="Quito, Ecuado" pageId="264" pageNumber="265" title="Escarabajos del Ecuador. Principales generos. Instituto de Ciencias Biologicas. Escuela Politecnica Nacional. Serie Entomologia, Nro. 1." year="2011">Carvajal et al. 2011</bibRefCitation>
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: 320-321 (cited for Ecuador);
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<bibRefCitation id="59958330DC9C615423277E7E943EF714" pageId="110" pageNumber="111">Krajcik 2012</bibRefCitation>
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: 91 (complete list of species);
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<bibRefCitation id="74353F9759C9FBB9046BCE534B8E973E" author="Sarmiento-Garces, R" journalOrPublisher="Universidad Nacional de Colombia, Facultad de Ciencias, Instituto de Ciencias Naturales" pageId="276" pageNumber="277" title="Escarabajos del genero Dichotomius Hope, 1838 (Scarabaeidae: Scarabaeinae) en Colombia. Fauna de Colombia Monografia No. 4." year="2014">
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<normalizedToken id="B18EF44B4BC5B2F4778A79964B0B9937" originalValue="Sarmiento-Garcés">Sarmiento-Garces</normalizedToken>
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and
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<normalizedToken id="6F4136A3F613BDE58AAA6EEB7DD0EE9E" originalValue="Amat-García">Amat-Garcia</normalizedToken>
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2014
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</bibRefCitation>
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: 53 (characters in key), 73 (diagnosis);
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<bibRefCitation id="01322747088253DCFDBA7772E375B7D8" author="Ratcliffe, BC" journalOrPublisher="Journal of the Kansas Entomological Society" pageId="276" pageNumber="277" pagination="186 - 207" title="Beetles (Coleoptera) of Peru: A Survey of the Families." url="https://doi.org/10.2317/kent-88-02-186-207.1" volume="88" year="2015">Ratcliffe et al. 2015</bibRefCitation>
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: 196 (cited for Peru).
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</paragraph>
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<paragraph id="B4DFFBFF288CDEC79F0266C33EB013A0" pageId="110" pageNumber="111">
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<taxonomicName id="C73253652BF68C239C3E2C5929332D60" class="Insecta" family="Scarabaeidae" genus="Dichotomius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dichotomius (Dichotomius) mamillatus" order="Coleoptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="species" species="mamillatus" subGenus="Dichotomius">Dichotomius (Dichotomius) mamillatus</taxonomicName>
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:
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<bibRefCitation id="2D4B53CBEFBE2008CD7900147806D5B5" author="Chamorro, W" journalOrPublisher="Revista Colombiana de Entomologi ́ a" pageId="264" pageNumber="265" pagination="72 - 100" title="Listado de especies y clave de ge ́ neros y subge ́ neros de escarabajos estercoleros (Coleoptera: Scarabaei- dae: Scarabaeinae) presentes y presuntos para Ecuador." url="https://doi.org/10.25100/socolen.v44i1.6545" volume="44" year="2018">Chamorro et al. 2018</bibRefCitation>
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: 94 (cited for Ecuador).
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</paragraph>
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</subSubSection>
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<subSubSection id="5D346F6F7A0E6B7388C846D5875DCA42" lastPageId="111" lastPageNumber="112" pageId="110" pageNumber="111" type="types specimens">
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<paragraph id="2D7B3300AD921B0DFF78B107B8DBAF2B" pageId="110" pageNumber="111">Types specimens.</paragraph>
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<paragraph id="F93A4BAA5A382304EA8DA2008F697022" pageId="110" pageNumber="111">
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<taxonomicName id="6675B76BE766433220AC5FFCE3A831CA" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pinotus mamillatus" order="Coleoptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="species" species="mamillatus">Pinotus mamillatus</taxonomicName>
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Felsche, 1901. One syntype (♂) examined deposited at the SMTD. Lectotype to be designated in a future work on this species group.
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</paragraph>
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<paragraph id="663BED57C0FD6999F3642AF97E22173B" pageId="111" pageNumber="112">
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<pageBreakToken id="BF263330F91410A5D474D5689F2A03D5" pageId="111" pageNumber="112" start="start">Syntype</pageBreakToken>
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(♂): "Ecuador / Baron [p]", "Typus [p, red label, black margin]", "Coll. C. Felsche / Kauf 20, 1918 [p, green label, black margin]", "mamillatus / Felsche / Ecuador [hw, purple label]", "Staatl. Museum
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<normalizedToken id="F491B581CAFCDD5DA54C799ADFC2DEB9" originalValue="für">fuer</normalizedToken>
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/ Tierkunde D esden [p]".
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</paragraph>
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</subSubSection>
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<subSubSection id="E949AC2510378606738F46E5E75AF24A" pageId="111" pageNumber="112" type="distribution">
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<paragraph id="06B7A9B92D97E81589850CD2257C8C79" pageId="111" pageNumber="112">Distribution.</paragraph>
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<paragraph id="6D0521580C4080308EA6A446C21CF3A0" pageId="111" pageNumber="112">Brazil, Colombia, Ecuador, and Peru.</paragraph>
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</subSubSection>
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<subSubSection id="D8F7B32A08E3617D077829F92CF83FCC" pageId="111" pageNumber="112" type="records examined">
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<paragraph id="2B844C5D7A3A664AB1452029D38C0086" pageId="111" pageNumber="112">Records examined.</paragraph>
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<paragraph id="AD2E29DE20F9A1382B7CAF6A0DB173A6" pageId="111" pageNumber="112">
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MORONA SANTIAGO: Cumpi, Cordillera del
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<normalizedToken id="A1163FDF89F79B8E025DF52010266F44" originalValue="Kutukú">Kutuku</normalizedToken>
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(1 specimen MUTPL); road Mendez-Paute, 1250 m (2 specimens CEMT; 5 specimens MQCAZ); NAPO: Nor Oeste de Puerto Napo, 470 m, Pungarayacu (1 specimen MQCAZ); Tena (3 specimens CEMT). ORELLANA: Bloque 31 Parque Nacional
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<normalizedToken id="10FDF6356861A7F2149642FBB559EEF4" originalValue="Yasuní">Yasuni</normalizedToken>
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(5 specimens MECN); Dayuma Campo Hormiguero, plataforma Hormiguero, 320 m (1 specimen MUTPL); Dayuma Campo Palanda-Yuca Sur plataforma Yuca 13, 255 m (1 specimen MGO-UC); Dayuma plataforma Ungurahua, 300 m (1 specimen MUTPL); El Dorado plataforma Guarango, 300 m (1 specimen MUTPL);
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<normalizedToken id="DA17E20C048A2C30A173C0BF10A4183B" originalValue="Estación">Estacion</normalizedToken>
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<normalizedToken id="894A1058645E1923DEAE051C4B64AF1F" originalValue="Científica">Cientifica</normalizedToken>
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<normalizedToken id="A05607253C5155B855D4B82E0C000533" originalValue="Yasuní">Yasuni</normalizedToken>
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Puce, 250 m (19 specimens MQCAZ);
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<normalizedToken id="14BAACB947A16F3B2EB05A398851A926" originalValue="Estación">Estacion</normalizedToken>
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de Biodiversidad Tiputini, 285 m, Parque Nacional Yasuni (7 specimes MUTPL); Rodrigo Borja IAMOE (1 specimen CEMT; 9 specimens MQCAZ); San Sebastian del Coca, 345 m, Comuna Guataraco, Campo Pata (1 specimen MUTPL); San
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<normalizedToken id="8CEA8730702C35542E8D7D62461D4475" originalValue="Sebastián">Sebastian</normalizedToken>
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del Coca, 345 m, Comuna Shamanal, Campo Palo Azul (1 specimen MUTPL). PASTAZA: Bosque Protector
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<normalizedToken id="98DB598FD2D496BE494C175936AE9C5E" originalValue="Oglán">Oglan</normalizedToken>
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Alto, 660 m (1 specimen MUTPL); Chuyayacu, 200 m, Oleoducto km 25 (1 specimen MUTPL); Nuevo San Jose del Curaray, 245 m (2 specimens MUTPL); Pandanuque, 420 m (1 specimen MUTPL); San Virgilio (1 specimen MUTPL); Villano (3 specimens MECN).
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<normalizedToken id="62DF6344ED7D53B0E14ACFF41A72EB02" originalValue="SUCUMBÍOS">SUCUMBIOS</normalizedToken>
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: Limoncocha, 250 m (2 speciemens CEMT); 6 km de Dureno, Precooperativa Los Vergeles, 290 m (4 specimens MGO-UC); Aucayacu
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<normalizedToken id="95DBC71D839C5AF6132171C159BB05BC" originalValue="Río">Rio</normalizedToken>
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El Eno, 275 m, 16 km de Lago Agrio (2 specimens MGO-UC); Nueva Loja plataforma Iguana, 310 m (1 specimen MUTPL); Pacayacu, 265 m, Campo Libertador, Tapi (1 specimen MUTPL); Tarapoa Campo Marian, plataforma Fanny 5, 260 m (1 specimen MUTPL); Tarapoa, Nuevo
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<normalizedToken id="75CBDEB5065DE6C5AA35E18CBF8705A6" originalValue="Manabí">Manabi</normalizedToken>
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, 270 m (1 specimen MUTPL); Zabalo, 520 m (1 specimen CEMT). ZAMORA CHINCHIPE: Tundayme campamento, 800 m (2 specimens MUTPL); Tundayme campamento Mirador, San Marcos, 900 m (1 specimen MUTPL); Tundayme campamento Mirador, Las Maravillas, 1060 m (1 specimen COSEC); Zurmi, Comunidad La Wants, 1010 m (2 specimens MEPN; 1 specimen MUTPL); Zurmi Las Orquideas
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<normalizedToken id="67A07F538D56AE3D9B1841D6C365675F" originalValue="Río">Rio</normalizedToken>
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Nangaritza, 870 m (1 specimen MUTPL). UNDETERMINED PROVINCE: without specific locality (1 specimen SMTD).
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</paragraph>
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</subSubSection>
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<subSubSection id="14A94089BC55970AFAE40774AD4D6766" pageId="111" pageNumber="112" type="temporal data">
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<paragraph id="CDDF4FCC1023D62D03A7C4DE1E1BFCA4" pageId="111" pageNumber="112">Temporal data.</paragraph>
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<paragraph id="3E3E23BEE984C0F6D72F80CF4280BE54" pageId="111" pageNumber="112">Collected every month of the year.</paragraph>
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</subSubSection>
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<subSubSection id="0362CBADEF2C9F9372D4EFA57A7FC331" pageId="111" pageNumber="112" type="remarks">
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<paragraph id="690DCA194A394EAD4C64338C81B7CE95" pageId="111" pageNumber="112">Remarks.</paragraph>
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<paragraph id="937A832B749B1FC1C97A480F117AA590" pageId="111" pageNumber="112">Inhabits the lowland evergreen forests, varzea forests and the foothill evergreen forests of the Amazon region from 200-1060 m a.s.l. Collected with flight interception traps and pitfall traps baited with carrion and human feces.</paragraph>
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</subSubSection>
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</treatment>
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</document> |