180 lines
22 KiB
XML
180 lines
22 KiB
XML
<document id="7E8A121910C3A642A3649A606CAC0E29" ID-CLB-Dataset="21316" ID-DOI="10.11646/zootaxa.4753.1.1" ID-GBIF-Dataset="f7974bcf-11e7-4d1b-b09d-530dc2dbd54f" ID-ISSN="1175-5326" ID-Zenodo-Dep="3983782" ID-ZooBank="AA9F66B3-EF8C-4F6B-8F35-0BCBEE5122ED" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1597351967001" checkinUser="felipe" docAuthor="Shear, William A., Richart, Casey H. & Wong, Victoria L." docDate="2020" docId="131D87EFFFEFFFDFFFDC5904FA63FE15" docLanguage="en" docName="zootaxa.4753.1.1.pdf" docOrigin="Zootaxa 4753 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Corypus kavanaughi Shelley 1997" docType="treatment" docVersion="9" lastPageNumber="71" masterDocId="EF24FF97FFA9FF98FF4B5F01FFFFFF9E" masterDocTitle="The millipede family Conotylidae in northwestern North America, with a complete bibliography of the family (Diplopoda, Chordeumatida, Heterochordeumatidea, Conotyloidea)" masterLastPageNumber="78" masterPageNumber="1" pageNumber="71" updateTime="1698844398316" updateUser="ExternalLinkService">
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<mods:title id="BB832C488996DE99DF1E421DBDD41F93">The millipede family Conotylidae in northwestern North America, with a complete bibliography of the family (Diplopoda, Chordeumatida, Heterochordeumatidea, Conotyloidea)</mods:title>
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<mods:namePart id="A91725E63E8DCC0892220F6958D50523">Shear, William A.</mods:namePart>
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<mods:namePart id="89CD4A633D625D2C79C68CEDF15EFAFA">Richart, Casey H.</mods:namePart>
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<mods:namePart id="726CEC5EBA539B1FED022F32BD2D4E5B">Wong, Victoria L.</mods:namePart>
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<mods:title id="21B798BEF44ACA96CC8E56AD6442BA2E">Zootaxa</mods:title>
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<taxonomicName id="5CB44D7AFFEFFFDEFFDC5904FDD3F981" ID-CoL="YST5" authority="Shelley 1997" authorityName="Shelley" authorityYear="1997" box="[151,556,1540,1567]" class="Diplopoda" family="Conotylidae" genus="Corypus" kingdom="Animalia" order="Chordeumatida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="kavanaughi">
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<emphasis id="A9C0EAEBFFEFFFDEFFDC5904FE74F981" box="[151,395,1541,1567]" italics="true" pageId="70" pageNumber="71">Corypus kavanaughi</emphasis>
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<bibRefCitation id="FF254B08FFEFFFDEFED95905FDD3F981" author="Shelley, R. M." box="[402,556,1540,1567]" pageId="70" pageNumber="71" pagination="121 - 126" refId="ref44643" refString="Shelley, R. M. (1997) Corypus kavanaughi, n. sp., a large-bodied chordeumatoid milliped from Idaho, with a review of the genus (Conotylidae). Myriapodologica, 4, 121 - 126." type="journal article" year="1997">Shelley 1997</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="D3AE6572FFEFFFDEFFDC5929FEC8F9DD" box="[151,311,1576,1603]" pageId="70" pageNumber="71" type="description">
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<paragraph id="9B0B36F9FFEFFFDEFFDC5929FEC8F9DD" blockId="70.[151,556,1540,1603]" box="[151,311,1576,1603]" pageId="70" pageNumber="71">
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<figureCitation id="038F2A7CFFEFFFDEFFDC5929FEFDF9DD" box="[151,258,1576,1603]" captionStart="FIGS" captionStartId="69.[151,212,1936,1961]" captionTargetBox="[151,1436,777,1909]" captionTargetId="figure@69.[151,1436,777,1910]" captionTargetPageId="69" captionText="FIGS. 251–253. Plumatyla humerosa and Corypus kavanaughi. Figs. 251, 252. Plumatyla humerosa. Fig. 251. Anterior gonopods, posterior view. Fig. 252. Posterior gonopod coxites, anterior view. Fig. 253. Corypus kavanaughi, posterior gonopods, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3983862" httpUri="https://zenodo.org/record/3983862/files/figure.png" pageId="70" pageNumber="71">Figs. 253</figureCitation>
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,
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<figureCitation id="038F2A7CFFEFFFDEFE465929FEC8F9DD" box="[269,311,1576,1603]" captionStart="FIGS" captionStartId="71.[151,212,1758,1783]" captionTargetBox="[151,1436,418,1732]" captionTargetId="figure@71.[151,1436,417,1733]" captionTargetPageId="71" captionText="FIGS. 254–257. Corypus kavanaughi and Lophomus laxus. Fig. 254, Corypus kavanaughi, Right anterior gonopod, posterior view. Figs. 255–257. Lophomus laxus. Fig. 255. Anterior gonopods, anterior view. Fig. 256. Anterior gonopods, anterior view. Fig. 257. Posterior gonopods, posterior view." figureDoi="http://doi.org/10.5281/zenodo.3983864" httpUri="https://zenodo.org/record/3983864/files/figure.png" pageId="70" pageNumber="71">254</figureCitation>
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</paragraph>
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<subSubSection id="D3AE6572FFEFFFDEFFDC596CFEBBF83B" pageId="70" pageNumber="71" type="reference_group">
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<paragraph id="9B0B36F9FFEFFFDEFFDC596CFDBEF91B" blockId="70.[151,577,1644,1669]" box="[151,577,1644,1669]" pageId="70" pageNumber="71">
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<taxonomicName id="5CB44D7AFFEFFFDEFFDC596CFDC1F91B" ID-CoL="YST5" authority="Shelley 1997, p. 123" authorityName="Shelley" authorityPageNumber="123" authorityYear="1997" box="[151,574,1644,1669]" class="Diplopoda" family="Conotylidae" genus="Corypus" kingdom="Animalia" order="Chordeumatida" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="kavanaughi">
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<emphasis id="A9C0EAEBFFEFFFDEFFDC596CFE96F91B" box="[151,361,1645,1669]" italics="true" pageId="70" pageNumber="71">Corypus kavanaughi</emphasis>
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<bibRefCitation id="FF254B08FFEFFFDEFE3B596DFE08F91B" author="Shelley, R. M." box="[368,503,1644,1669]" pageId="70" pageNumber="71" pagination="121 - 126" refId="ref44643" refString="Shelley, R. M. (1997) Corypus kavanaughi, n. sp., a large-bodied chordeumatoid milliped from Idaho, with a review of the genus (Conotylidae). Myriapodologica, 4, 121 - 126." type="journal article" year="1997">Shelley 1997</bibRefCitation>
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, p.123
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="9B0B36F9FFEFFFDEFFDC59B2FEBBF83B" blockId="70.[151,1437,1714,2030]" pageId="70" pageNumber="71">
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<emphasis id="A9C0EAEBFFEFFFDEFFDC59B2FED2F953" bold="true" box="[151,301,1715,1741]" pageId="70" pageNumber="71">New records</emphasis>
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:
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<collectingRegion id="5970F81BFFEFFFDEFE7059B2FE65F953" box="[315,410,1715,1741]" country="United States of America" name="Idaho" pageId="70" pageNumber="71">IDAHO</collectingRegion>
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:
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<emphasis id="A9C0EAEBFFEFFFDEFEEF59B3FDE9F953" bold="true" box="[420,534,1714,1741]" italics="true" pageId="70" pageNumber="71">
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<collectingRegion id="5970F81BFFEFFFDEFEEF59B3FE17F953" box="[420,488,1714,1741]" country="United States of America" name="Idaho" pageId="70" pageNumber="71">Idaho</collectingRegion>
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Co.
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</emphasis>
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, O’Hara Creek Campground, Selway River Road
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<quantity id="5C4C9B1CFFEFFFDEFB0359B2FB6FF953" box="[1096,1168,1715,1741]" metricMagnitude="4" metricUnit="m" metricValue="1.07826048" pageId="70" pageNumber="71" unit="mi" value="6.7">6.7 mi</quantity>
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SE of US-12 at Lowell, Clearwater National Forest,
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, -
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<geoCoordinate id="FE80503EFFEFFFDEFD2659D7FD05F96F" box="[621,762,1750,1777]" degrees="115.5129" direction="west" orientation="longitude" pageId="70" pageNumber="71" precision="5" value="-115.5129">115.5129°W</geoCoordinate>
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, elev.
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<specimenCount id="8DB2FD70FFEFFFDEFC0F59D6FC6AF96C" box="[836,917,1751,1778]" pageId="70" pageNumber="71" type="generic">
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.
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</specimenCount>
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,
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<date id="EF0A1039FFEFFFDEFCE859D7FBD3F96F" box="[931,1068,1750,1777]" pageId="70" pageNumber="71" value="2008-07-05">5 July 2008</date>
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, C. Richart, S. Derkarabetian, J. Underwood, from forest litter and woody debris, m f;
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<emphasis id="A9C0EAEBFFEFFFDEFDA159FBFC75F88B" bold="true" box="[746,906,1786,1813]" italics="true" pageId="70" pageNumber="71">Shoshone Co.</emphasis>
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, Eagle Creek, FS-1214,
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<quantity id="5C4C9B1CFFEFFFDEFBD159FAFB1EF88B" box="[1178,1249,1787,1813]" metricMagnitude="2" metricUnit="m" metricValue="1.609344" pageId="70" pageNumber="71" unit="mi" value="0.1">0.1 mi</quantity>
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N of FS-50,
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<collectingRegion id="5970F81BFFEFFFDEFA2459FAFF4CF8A7" country="United States of America" name="Idaho" pageId="70" pageNumber="71">Idaho</collectingRegion>
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Panhandle National Forests,
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<geoCoordinate id="FE80503EFFEFFFDEFEB6581EFD81F8A7" box="[509,638,1823,1849]" degrees="47.2122" direction="west" orientation="longitude" pageId="70" pageNumber="71" precision="5" value="-47.2122">47.2122°W</geoCoordinate>
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, -
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<geoCoordinate id="FE80503EFFEFFFDEFDD8581FFCDEF8A7" box="[659,801,1822,1849]" degrees="115.5484" direction="west" orientation="longitude" pageId="70" pageNumber="71" precision="5" value="-115.5484">115.5484°W</geoCoordinate>
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, elev.
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<quantity id="5C4C9B1CFFEFFFDEFC2C581EFC4EF8A4" box="[871,945,1823,1850]" metricMagnitude="2" metricUnit="m" metricValue="8.97" pageId="70" pageNumber="71" unit="m" value="897.0">897 m</quantity>
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,
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<date id="EF0A1039FFEFFFDEFCF6581EFBAEF8A7" box="[957,1105,1823,1849]" pageId="70" pageNumber="71" value="2008-07-27">27 July 2008</date>
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,
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<taxonomicName id="5CB44D7AFFEFFFDEFB17581EFB0CF8A7" ID-CoL="56NTV" box="[1116,1267,1823,1849]" class="Pinopsida" family="Cupressaceae" genus="Thuja" kingdom="Plantae" order="Pinales" pageId="70" pageNumber="71" phylum="Tracheophyta" rank="species" species="plicata">
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<emphasis id="A9C0EAEBFFEFFFDEFB17581EFB0CF8A7" box="[1116,1267,1823,1849]" italics="true" pageId="70" pageNumber="71">Thuja plicata</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="5CB44D7AFFEFFFDEFA4A581EFA62F8A7" ID-CoL="8KCY" box="[1281,1437,1823,1849]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="70" pageNumber="71" phylum="Tracheophyta" rank="species" species="grandis">
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||
<emphasis id="A9C0EAEBFFEFFFDEFA4A581EFA62F8A7" box="[1281,1437,1823,1849]" italics="true" pageId="70" pageNumber="71">Abies grandis</emphasis>
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</taxonomicName>
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||
forest, stream-side woody debris, mm f; Prospector Creek at FS-752,
|
||
<quantity id="5C4C9B1CFFEFFFDEFCE85842FC14F8C3" box="[931,1003,1859,1885]" metricMagnitude="2" metricUnit="m" metricValue="1.609344" pageId="70" pageNumber="71" unit="mi" value="0.1">0.1 mi</quantity>
|
||
S of FS-50,
|
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<collectingRegion id="5970F81BFFEFFFDEFB325842FB46F8C3" box="[1145,1209,1859,1885]" country="United States of America" name="Idaho" pageId="70" pageNumber="71">Idaho</collectingRegion>
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||
Panhandle National Forests,
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||
<geoCoordinate id="FE80503EFFEFFFDEFFB35866FE8AF81F" box="[248,373,1895,1921]" degrees="47.2236" direction="north" orientation="latitude" pageId="70" pageNumber="71" precision="5" value="47.2236">47.2236°N</geoCoordinate>
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||
, -
|
||
<geoCoordinate id="FE80503EFFEFFFDEFEC05867FDE6F81F" box="[395,537,1894,1921]" degrees="115.6072" direction="west" orientation="longitude" pageId="70" pageNumber="71" precision="5" value="-115.6072">115.6072°W</geoCoordinate>
|
||
, elev.
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||
<quantity id="5C4C9B1CFFEFFFDEFD285866FD51F81C" box="[611,686,1895,1922]" metricMagnitude="2" metricUnit="m" metricValue="8.6" pageId="70" pageNumber="71" unit="m" value="860.0">860 m</quantity>
|
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,
|
||
<date id="EF0A1039FFEFFFDEFDF05866FCADF81F" box="[699,850,1895,1921]" pageId="70" pageNumber="71" value="2008-07-26">26 July 2008</date>
|
||
,
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||
<taxonomicName id="5CB44D7AFFEFFFDEFC145866FC08F81F" ID-CoL="56NTV" box="[863,1015,1895,1921]" class="Pinopsida" family="Cupressaceae" genus="Thuja" kingdom="Plantae" order="Pinales" pageId="70" pageNumber="71" phylum="Tracheophyta" rank="species" species="plicata">
|
||
<emphasis id="A9C0EAEBFFEFFFDEFC145866FC08F81F" box="[863,1015,1895,1921]" italics="true" pageId="70" pageNumber="71">Thuja plicata</emphasis>
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||
</taxonomicName>
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||
forest, stream-side woody debris, C. Richart, mm ff.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="D3AE6572FFEFFFDFFF8C58AEFA63FE15" lastPageId="71" lastPageNumber="72" pageId="70" pageNumber="71" type="description">
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<paragraph id="9B0B36F9FFEFFFDFFF8C58AEFA63FE15" blockId="70.[151,1437,1714,2030]" lastBlockId="71.[151,1437,153,395]" lastPageId="71" lastPageNumber="72" pageId="70" pageNumber="71">
|
||
<emphasis id="A9C0EAEBFFEFFFDEFF8C58AEFEF5F857" bold="true" box="[199,266,1967,1993]" pageId="70" pageNumber="71">Notes</emphasis>
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||
: This species was exhaustively described by
|
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<bibRefCitation id="FF254B08FFEFFFDEFC5958AEFC42F857" author="Shelley, R. M." box="[786,957,1967,1993]" pageId="70" pageNumber="71" pagination="121 - 126" refId="ref44643" refString="Shelley, R. M. (1997) Corypus kavanaughi, n. sp., a large-bodied chordeumatoid milliped from Idaho, with a review of the genus (Conotylidae). Myriapodologica, 4, 121 - 126." type="journal article" year="1997">Shelley (1997)</bibRefCitation>
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. Our SEMs show details not noted on the drawings in that paper. The filaments on the posterior side of the anterior gonopods are seen here (
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<emphasis id="A9C0EAEBFFEFFFDEFBB658D3FAF2F873" bold="true" box="[1277,1293,2002,2029]" pageId="70" pageNumber="71">f,</emphasis>
|
||
<figureCitation id="038F2A7CFFEFFFDEFA5D58D2FA88F873" box="[1302,1399,2002,2029]" captionStart="FIGS" captionStartId="71.[151,212,1758,1783]" captionTargetBox="[151,1436,418,1732]" captionTargetId="figure@71.[151,1436,417,1733]" captionTargetPageId="71" captionText="FIGS. 254–257. Corypus kavanaughi and Lophomus laxus. Fig. 254, Corypus kavanaughi, Right anterior gonopod, posterior view. Figs. 255–257. Lophomus laxus. Fig. 255. Anterior gonopods, anterior view. Fig. 256. Anterior gonopods, anterior view. Fig. 257. Posterior gonopods, posterior view." figureDoi="http://doi.org/10.5281/zenodo.3983864" httpUri="https://zenodo.org/record/3983864/files/figure.png" pageId="70" pageNumber="71">Fig. 254</figureCitation>
|
||
) to be finely serrate, and representing a continuation of previously undetected fimbriate lamellae that border a distinct channel (
|
||
<emphasis id="A9C0EAEBFFEEFFDFFE4B5FBCFEF3FF49" bold="true" box="[256,268,189,215]" pageId="71" pageNumber="72">c</emphasis>
|
||
,
|
||
<figureCitation id="038F2A7CFFEEFFDFFE575FBCFE83FF49" box="[284,380,188,215]" captionStart="FIGS" captionStartId="71.[151,212,1758,1783]" captionTargetBox="[151,1436,418,1732]" captionTargetId="figure@71.[151,1436,417,1733]" captionTargetPageId="71" captionText="FIGS. 254–257. Corypus kavanaughi and Lophomus laxus. Fig. 254, Corypus kavanaughi, Right anterior gonopod, posterior view. Figs. 255–257. Lophomus laxus. Fig. 255. Anterior gonopods, anterior view. Fig. 256. Anterior gonopods, anterior view. Fig. 257. Posterior gonopods, posterior view." figureDoi="http://doi.org/10.5281/zenodo.3983864" httpUri="https://zenodo.org/record/3983864/files/figure.png" pageId="71" pageNumber="72">Fig. 254</figureCitation>
|
||
) leading up the posterior surface of the gonopod from a similar region at its base. This basal fimbriate area may partially conceal a pore (
|
||
<emphasis id="A9C0EAEBFFEEFFDFFDCD5FE0FD5CFF65" bold="true" box="[646,675,225,251]" pageId="71" pageNumber="72">p?</emphasis>
|
||
,
|
||
<figureCitation id="038F2A7CFFEEFFDFFDFA5FE0FCEFFF65" box="[689,784,224,251]" captionStart="FIGS" captionStartId="71.[151,212,1758,1783]" captionTargetBox="[151,1436,418,1732]" captionTargetId="figure@71.[151,1436,417,1733]" captionTargetPageId="71" captionText="FIGS. 254–257. Corypus kavanaughi and Lophomus laxus. Fig. 254, Corypus kavanaughi, Right anterior gonopod, posterior view. Figs. 255–257. Lophomus laxus. Fig. 255. Anterior gonopods, anterior view. Fig. 256. Anterior gonopods, anterior view. Fig. 257. Posterior gonopods, posterior view." figureDoi="http://doi.org/10.5281/zenodo.3983864" httpUri="https://zenodo.org/record/3983864/files/figure.png" pageId="71" pageNumber="72">Fig. 254</figureCitation>
|
||
) homologous to the large pores seen posterior on the anterior gonopods in many of the other conotylids illustrated in this paper. The posterior gonopods, as
|
||
<bibRefCitation id="FF254B08FFEEFFDFFB935E04FA82FE81" author="Shelley, R. M." box="[1240,1405,261,287]" pageId="71" pageNumber="72" pagination="121 - 126" refId="ref44643" refString="Shelley, R. M. (1997) Corypus kavanaughi, n. sp., a large-bodied chordeumatoid milliped from Idaho, with a review of the genus (Conotylidae). Myriapodologica, 4, 121 - 126." type="journal article" year="1997">Shelley (1997)</bibRefCitation>
|
||
illustrated, bear coxites seemingly arising from a reduced coxa fused to the sternum (
|
||
<emphasis id="A9C0EAEBFFEEFFDFFB795E28FBA8FEDD" bold="true" box="[1074,1111,297,323]" pageId="71" pageNumber="72">cxs</emphasis>
|
||
,
|
||
<figureCitation id="038F2A7CFFEEFFDFFB2E5E28FB3BFEDD" box="[1125,1220,296,323]" captionStart="FIGS" captionStartId="69.[151,212,1936,1961]" captionTargetBox="[151,1436,777,1909]" captionTargetId="figure@69.[151,1436,777,1910]" captionTargetPageId="69" captionText="FIGS. 251–253. Plumatyla humerosa and Corypus kavanaughi. Figs. 251, 252. Plumatyla humerosa. Fig. 251. Anterior gonopods, posterior view. Fig. 252. Posterior gonopod coxites, anterior view. Fig. 253. Corypus kavanaughi, posterior gonopods, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3983862" httpUri="https://zenodo.org/record/3983862/files/figure.png" pageId="71" pageNumber="72">Fig. 253</figureCitation>
|
||
), and with a strong, setose anterior lobe (
|
||
<emphasis id="A9C0EAEBFFEEFFDFFE355E4CFE6BFEF9" bold="true" box="[382,404,333,359]" pageId="71" pageNumber="72">al</emphasis>
|
||
,
|
||
<figureCitation id="038F2A7CFFEEFFDFFEEA5E4CFE00FEF9" box="[417,511,332,359]" captionStart="FIGS" captionStartId="69.[151,212,1936,1961]" captionTargetBox="[151,1436,777,1909]" captionTargetId="figure@69.[151,1436,777,1910]" captionTargetPageId="69" captionText="FIGS. 251–253. Plumatyla humerosa and Corypus kavanaughi. Figs. 251, 252. Plumatyla humerosa. Fig. 251. Anterior gonopods, posterior view. Fig. 252. Posterior gonopod coxites, anterior view. Fig. 253. Corypus kavanaughi, posterior gonopods, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3983862" httpUri="https://zenodo.org/record/3983862/files/figure.png" pageId="71" pageNumber="72">Fig. 253</figureCitation>
|
||
) that is tightly appresed to the coxite on the lateral side. As in species of
|
||
<emphasis id="A9C0EAEBFFEEFFDFFA695E4CFA63FEF9" box="[1314,1436,333,359]" italics="true" pageId="71" pageNumber="72">
|
||
<taxonomicName id="5CB44D7AFFEEFFDFFA695E4CFA67FEF9" authorityName="Causey" authorityYear="1961" box="[1314,1432,333,359]" class="Diplopoda" family="Conotylidae" genus="Austrotyla" kingdom="Animalia" order="Chordeumatida" pageId="71" pageNumber="72" phylum="Arthropoda" rank="genus">Austrotyla</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
the posterior gonopod coxites are cupped anteriorly and more or less conceal the smaller anterior gonopods
|
||
<emphasis id="A9C0EAEBFFEEFFDFFA045E73FA63FE15" box="[1359,1436,370,395]" italics="true" pageId="71" pageNumber="72">in situ.</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |