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<document id="F7A72BD2F3D9BD0DC81CD479B972F54A" ID-DOI="10.11646/zootaxa.3746.1.3" ID-GBIF-Dataset="def67b1f-7898-42ac-bf47-0c350748dfd0" ID-ISSN="1175-5326" ID-Zenodo-Dep="219621" ID-ZooBank="A34057EB-13C1-4E5D-BF39-0AD1EE10AB12" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460706961445" checkinUser="plazi" docAuthor="Huemer, Peter, Elsner, Gustav &amp; Karsholt, Ole" docDate="2013" docId="131748077A06FFEFE98CF8ED18A8E1EA" docLanguage="en" docName="zt03746p100.pdf" docOrigin="Zootaxa 3746 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eulamprotes kailai Karsholt &amp; Huemer, sp. nov." docType="treatment" docVersion="8" lastPageNumber="95" masterDocId="EF2E307F7A1EFFF4E91BFFD91A40E13F" masterDocTitle="Review of the Eulamprotes wilkella species-group based on morphology and DNA barcodes, with descriptions of new taxa (Lepidoptera, Gelechiidae)" masterLastPageNumber="100" masterPageNumber="69" pageNumber="93" updateTime="1698357352301" updateUser="plazi">
<mods:mods id="3B2FF53723F88124812741981C0FD472" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="047FE3C2E2C40D356D648DFB18401136">
<mods:title id="1A3FD9AA868B0CFFA58B23586B680504">Review of the Eulamprotes wilkella species-group based on morphology and DNA barcodes, with descriptions of new taxa (Lepidoptera, Gelechiidae)</mods:title>
</mods:titleInfo>
<mods:name id="7F55D59B03C2E1FDFF94742ECBF1DCAD" type="personal">
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<mods:roleTerm id="5B5AAA32C141F660E03A272169DE6AAF">Author</mods:roleTerm>
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<mods:namePart id="489098744E7E0B145BB2E86E02ACA11C">Huemer, Peter</mods:namePart>
</mods:name>
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<mods:namePart id="5BA6683584FAA12C3A5676E97845BE80">Elsner, Gustav</mods:namePart>
</mods:name>
<mods:name id="ECD72DE0ABF086520EE23761AB172778" type="personal">
<mods:role id="64DD88B74860AA1DE2471548B7C63AE5">
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<mods:namePart id="766978CD696EF9353950EF122118C500">Karsholt, Ole</mods:namePart>
</mods:name>
<mods:typeOfResource id="5E0CB95D10B470EB3B990CA3B6171C75">text</mods:typeOfResource>
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<mods:title id="79E49C80E50128DDA1A83168FA55B85D">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="85F73DB23943128131F5210F17C41469">
<mods:date id="E0F0546E1F78A6426179F90E2A158038">2013</mods:date>
<mods:detail id="A406AF57ABF8C5B490626BD700C32BF1" type="volume">
<mods:number id="36F7902433DDC21460EF37F85D620EF2">3746</mods:number>
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<mods:detail id="9A799C6EBAB2B9285D5758B7B3C31C3A" type="issue">
<mods:number id="9D6AB0860277368BD5EF3177E37612EF">1</mods:number>
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<mods:start id="05144456FB79CC91119B80ACF0C4814D">69</mods:start>
<mods:end id="307E8C5EFCFBEE812F620FC9139C0CD4">100</mods:end>
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<mods:classification id="1B58878AC2B1024D3375CBCCB2D2A102">journal article</mods:classification>
<mods:identifier id="94927318D766703DA4C9A41FC361110B" type="DOI">10.11646/zootaxa.3746.1.3</mods:identifier>
<mods:identifier id="3DFDA7AA4DE0A852385F089F560B52D8" type="GBIF-Dataset">def67b1f-7898-42ac-bf47-0c350748dfd0</mods:identifier>
<mods:identifier id="B4F3B7FA9D8A6880EE936E1E27E3860F" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="4295162D92CB666C026C2B4E62869754" type="Zenodo-Dep">219621</mods:identifier>
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</mods:mods>
<treatment id="131748077A06FFEFE98CF8ED18A8E1EA" ID-DOI="http://doi.org/10.5281/zenodo.6146596" ID-GBIF-Taxon="119584439" ID-Zenodo-Dep="6146596" LSID="urn:lsid:plazi:treatment:131748077A06FFEFE98CF8ED18A8E1EA" httpUri="http://treatment.plazi.org/id/131748077A06FFEFE98CF8ED18A8E1EA" lastPageId="27" lastPageNumber="95" pageId="24" pageNumber="93">
<subSubSection id="D3A4AA9A7A06FFEDE98CF8ED1E0EE0CB" lastPageId="25" lastPageNumber="94" pageId="24" pageNumber="93" type="nomenclature">
<paragraph id="9B01F9117A06FFECE98CF8ED18A6E671" blockId="24.[151,742,1844,1870]" box="[151,742,1844,1870]" pageId="24" pageNumber="93">
<heading id="C0494E7D7A06FFECE98CF8ED18A6E671" bold="true" box="[151,742,1844,1870]" fontSize="11" level="1" pageId="24" pageNumber="93" reason="1">
<emphasis id="A9CA25037A06FFECE98CF8ED18A6E671" bold="true" box="[151,742,1844,1870]" pageId="24" pageNumber="93">
<taxonomicName id="5CBE82927A06FFECE98CF8ED18C0E671" ID-CoL="93GGR" authority="Karsholt &amp; Huemer" authorityName="Karsholt &amp; Huemer" box="[151,640,1844,1870]" class="Insecta" family="Gelechiidae" genus="Eulamprotes" kingdom="Animalia" order="Lepidoptera" pageId="24" pageNumber="93" phylum="Arthropoda" rank="species" species="kailai" status="sp. nov.">
<emphasis id="A9CA25037A06FFECE98CF8ED1B3DE671" bold="true" box="[151,381,1844,1870]" italics="true" pageId="24" pageNumber="93">Eulamprotes kailai</emphasis>
Karsholt &amp; Huemer
</taxonomicName>
<taxonomicNameLabel id="B2F998787A06FFECEB93F8ED18A6E671" box="[648,742,1844,1870]" pageId="24" pageNumber="93" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="9B01F9117A06FFECE98CF8A319CCE688" blockId="24.[151,1436,1914,1975]" pageId="24" pageNumber="93">
Examined material.
<typeStatus id="440547B37A06FFECE862F8A31BA5E6AC" box="[377,485,1914,1939]" pageId="24" pageNumber="93" type="holotype">Holotype</typeStatus>
. ♂.“USSR,
<collectingCountry id="E3A9B9817A06FFECEB77F8A31886E6AC" box="[620,710,1914,1939]" name="Kyrgyzstan" pageId="24" pageNumber="93">Kirgisia</collectingCountry>
41˚30’N, 75˚35´N [
<emphasis id="A9CA25037A06FFECEAB2F8A51988E6AC" box="[937,968,1916,1939]" italics="true" pageId="24" pageNumber="93">sic</emphasis>
]
<quantity id="5C4654F47A06FFECEAC3F8A21E60E6AC" box="[984,1056,1914,1939]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="24" pageNumber="93" unit="km" value="10.0">10 km</quantity>
SE Lake Song Köl” “steppe/river bed
<date id="EF00DFD17A06FFECE9DDF8471B76E688" box="[198,310,1950,1975]" pageId="24" pageNumber="93" value="1990-07-26">26.7.1990</date>
L. Kaila leg.” “GU 13/1361 ♂ P. Huemer” (ZMUH).
</paragraph>
<paragraph id="9B01F9117A07FFEDE9DCFF411E0EE0CB" blockId="25.[151,1436,151,1653]" pageId="25" pageNumber="94">
<typeStatus id="440547B37A07FFEDE9DCFF411B78E18F" box="[199,312,152,176]" pageId="25" pageNumber="94" type="paratype">Paratypes</typeStatus>
. Kyrgizia:
<specimenCount id="8DB832987A07FFEDE8A6FF411BA2E18F" box="[445,482,152,176]" pageId="25" pageNumber="94" type="generic">3♂</specimenCount>
,
<quantity id="5C4654F47A07FFEDE8E8FF411800E18F" box="[499,576,151,176]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="25" pageNumber="94" unit="km" value="30.0">30 km</quantity>
E Naryn, 41˚25’N, 76˚20E,
<quantity id="5C4654F47A07FFEDEA80FF4E19B8E190" box="[923,1016,151,176]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="25" pageNumber="94" unit="m" value="2500.0">2500 m</quantity>
,
<date id="EF00DFD17A07FFEDED12FF4E1ECAE18F" box="[1033,1162,151,176]" pageId="25" pageNumber="94" value="1990-07-29">29.vii.1990</date>
, leg. Kaila &amp; Mikkola (ZMUH).
<collectingCountry id="E3A9B9817A07FFEDE810FF651BD2E1EA" box="[267,402,188,213]" name="Kazakhstan" pageId="25" pageNumber="94">Kazakhstan</collectingCountry>
:
<specimenCount id="8DB832987A07FFEDE885FF651B83E1EB" box="[414,451,188,212]" pageId="25" pageNumber="94" type="generic">2♂</specimenCount>
, Zailiskiy, Alatau, Alma-Atinskij Nat. Par., 43˚05’N, 77˚15E,
<quantity id="5C4654F47A07FFEDED9DFF651E9FE1EB" box="[1158,1247,188,213]" metricMagnitude="3" metricUnit="m" metricValue="1.85" pageId="25" pageNumber="94" unit="m" value="1850.0">1850 m</quantity>
,
<date id="EF00DFD17A07FFEDEDF0FF651F26E1EA" box="[1259,1382,188,213]" pageId="25" pageNumber="94" value="1990-06-25">25.vi.1990</date>
, leg. Kaila, genitalia slide GU 13/1358 ♂ P. Huemer (TLMF, ZMUH).
<collectingCountry id="E3A9B9817A07FFEDEA96FF06199CE1C7" box="[909,988,223,248]" name="Russia" pageId="25" pageNumber="94">Russia</collectingCountry>
: Tuva Republic,
<specimenCount id="8DB832987A07FFEDEDBBFF391E93E1C7" box="[1184,1235,224,248]" pageId="25" pageNumber="94" type="generic">10♂</specimenCount>
, Lake Tere-Khol, 50˚01’N, 95 ˚03’E,
<quantity id="5C4654F47A07FFEDE862FEDD1B91E023" box="[377,465,260,285]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="25" pageNumber="94" unit="m" value="1150.0">1150 m</quantity>
, sand dunes,
<date id="EF00DFD17A07FFEDEB77FEDD1948E022" box="[620,776,260,285]" pageId="25" pageNumber="94" value="1995-06-09" valueMax="1995-06-12" valueMin="1995-06-09">9.–12.vi.1995</date>
, leg. Jalava &amp; Kullberg (RCGE, ZMUH, ZMUC);
<specimenCount id="8DB832987A07FFEDEC4AFEDD1F36E023" box="[1361,1398,260,284]" pageId="25" pageNumber="94" type="generic">3♂</specimenCount>
, E. Tannu-Ola Mts., Irbitel r., 50˚44’N, 93 ˚08’E,
<quantity id="5C4654F47A07FFEDEB86FEF118B4E000" box="[669,756,296,320]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="25" pageNumber="94" unit="m" value="1000.0">1000 m</quantity>
, stony steppe slopes,
<date id="EF00DFD17A07FFEDEAFCFEFE1ED3E07F" box="[999,1171,295,320]" pageId="25" pageNumber="94" value="1995-06-13" valueMax="1995-06-16" valueMin="1995-06-13">13.–16.vi.1995</date>
, leg. Jalava &amp; Kullberg (ZMUH);
<specimenCount id="8DB832987A07FFEDE80AFE951B75E05B" box="[273,309,332,356]" pageId="25" pageNumber="94" type="generic">1♂</specimenCount>
, E. Tannu-Ola Mts.,
<quantity id="5C4654F47A07FFEDEB22FE951838E05A" box="[569,632,332,357]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="25" pageNumber="94" unit="km" value="5.0">5 km</quantity>
ENE Khol-Oozha, 50˚45’N, 94˚29’E,
<quantity id="5C4654F47A07FFEDED57FE951EE9E05B" box="[1100,1193,332,357]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="25" pageNumber="94" unit="m" value="1250.0">1250 m</quantity>
, steppe slopes,
<date id="EF00DFD17A07FFEDEC70FE951B51E0B7" pageId="25" pageNumber="94" value="1995-06-16" valueMax="1995-06-19" valueMin="1995-06-16">16.– 19.vi.1995</date>
, leg. Jalava &amp; Kullberg (ZMUH); Buryatia,
<specimenCount id="8DB832987A07FFEDEA1DFEA9196AE0B7" box="[774,810,368,392]" pageId="25" pageNumber="94" type="generic">1♂</specimenCount>
, Svyaloj Nos pns., Monahovo, 53˚40’N, 109˚00’E,
<quantity id="5C4654F47A07FFEDEC69FEB61AEFE093" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="25" pageNumber="94" unit="m" value="450.0">450 m</quantity>
, mixed forest,
<date id="EF00DFD17A07FFEDE846FE4D1847E092" box="[349,519,404,429]" pageId="25" pageNumber="94" value="1996-06-27" valueMax="1996-06-30" valueMin="1996-06-27">27.–30.vi.1996</date>
, leg. Jalava &amp; Kullberg (ZMUH);
<specimenCount id="8DB832987A07FFEDEA80FE4D19FFE093" box="[923,959,404,428]" pageId="25" pageNumber="94" type="generic">1♂</specimenCount>
, Barguzin range, Olso r. valley, 54˚52’N, 110˚55’E,
<quantity id="5C4654F47A07FFEDE80FFE611B25E0F0" box="[276,357,440,464]" metricMagnitude="2" metricUnit="m" metricValue="9.2" pageId="25" pageNumber="94" unit="m" value="920.0">920 m</quantity>
, taiga,
<date id="EF00DFD17A07FFEDE8D9FE6E1877E0EF" box="[450,567,439,464]" pageId="25" pageNumber="94" value="1996-07-07">7.vii.1996</date>
, leg. Jalava &amp; Kullberg (ZMUH);
<specimenCount id="8DB832987A07FFEDEAF8FE611E48E0EF" box="[995,1032,440,464]" pageId="25" pageNumber="94" type="generic">1♂</specimenCount>
, Barguzin valley, Malsky village, 54˚35’N, 110 ˚48’E,
<quantity id="5C4654F47A07FFEDE89AFE051B8AE0CB" box="[385,458,476,501]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="25" pageNumber="94" unit="m" value="500.0">500 m</quantity>
, sandy yard,
<date id="EF00DFD17A07FFEDEB45FE051893E0CA" box="[606,723,476,501]" pageId="25" pageNumber="94" value="1996-07-07">7.vii.1996</date>
, leg. Jalava &amp; Kullberg (ZMUH).
</paragraph>
</subSubSection>
<subSubSection id="D3A4AA9A7A07FFEDE9DCFDD9185AE5D7" pageId="25" pageNumber="94" type="description">
<paragraph id="9B01F9117A07FFEDE9DCFDD919C5E3F2" blockId="25.[151,1436,151,1653]" pageId="25" pageNumber="94">
DESCRIPTION.—Adult. (
<figureCitation id="0385E5947A07FFEDE8C3FE26186EE327" box="[472,558,511,536]" captionStart="FIGURES 13 – 21" captionStartId="6.[151,263,1418,1441]" captionTargetBox="[188,1396,493,1383]" captionTargetId="figure@6.[183,1404,483,1394]" captionTargetPageId="6" captionText="FIGURES 13 – 21. Adults of Eulamprotes spp. 13, E. occidentella Huemer &amp; Karsholt, paratype, ♂ (Italy); 14, ditto, paratype, ♀ (Italy); 15, E. atrifrontella sp. nov., paratype, ♂ (Turkey); 16, E. wieseri sp. nov., holotype, ♂ (Kirgizia); 17, E. altaicella sp. nov., holotype, ♂ (Russia, Altai); 18, E. kailai sp. nov., holotype, ♂ (Kirgizia); 19, E. libertinella (Zeller), ♂ (Italy); 20, ditto, ♀ (Switzerland); 21, E. gemerensis sp. nov., paratype, ♂ (Slovakia). Scale bar = 2 mm." httpUri="https://zenodo.org/record/219623/files/figure.png" pageId="25" pageNumber="94">Fig. 18</figureCitation>
). Male. Wingspan
<quantity id="5C4654F47A07FFEDEA13FDD91938E328" box="[776,888,512,536]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="25" pageNumber="94" unit="mm" value="9.5" valueMax="10.0" valueMin="9.0">9–10 mm</quantity>
. Labial palpus white; segment 3 with black tip. Antenna ringed black and whitish to tip. Head whitish; thorax and tegula black. Forewing black with white markings with only a slight silvery shine: an oblique fascia from 1/6 at costa to 1/5 at fold; a fascia from middle of costa reaching two-thirds towards dorsum; pre-apical costal spot close to tornal spot; apex with silvery scales around apex; cilia blackish grey, whitish grey at tip of apex. Hindwing almost as broad as forewing, grey. Abdomen yellowish grey in basal third, then dark grey with yellow-white tip.
</paragraph>
<paragraph id="9B01F9117A07FFEDE9DCFD0E1BDDE3CF" blockId="25.[151,1436,151,1653]" box="[199,413,727,752]" pageId="25" pageNumber="94">Female. Unknown.</paragraph>
<paragraph id="9B01F9117A07FFEDE9DCFD25195EE262" blockId="25.[151,1436,151,1653]" pageId="25" pageNumber="94">Variation. The pre-apical and tornal spots may be fused or divided. The colour of the head varies from pure white to whitish yellow, and the colour of the labial palpi varies from pure white to cream-white. Some specimens have dark scales at base of segment 2 of the labial palpus.</paragraph>
<paragraph id="9B01F9117A07FFEDE9DCFCBE1E73E5FA" blockId="25.[151,1436,151,1653]" pageId="25" pageNumber="94">
Male genitalia (
<figureCitation id="0385E5947A07FFEDE86DFCBE1BB5E2BF" box="[374,501,871,896]" captionStart="FIGURES 40 – 45" captionStartId="15.[151,263,1759,1782]" captionTargetBox="[172,1416,199,1726]" captionTargetId="figure@15.[151,1436,195,1735]" captionTargetPageId="15" captionText="FIGURES 40 – 45. Male genitalia of Eulamprotes spp. 40, E. altaicella sp. nov., paratype, (Russa, Altai) gen. slide GU 13 / 1359 P. Huemer; 41, ditto, phallus; 42, E. kailai sp. nov., holotype, (Kirgizia) gen. slide GEL 1196 P. Huemer; 43, ditto, phallus; 44, E. kailai sp. nov., paratype, (Kazakhstan) gen. slide GU 13 / 1358 P. Huemer; 45, ditto, phallus." httpUri="https://zenodo.org/record/219627/files/figure.png" pageId="25" pageNumber="94">Figs 42–45</figureCitation>
).—Segment VIII with two pairs of coremata in intersegmental membrane, grouped into short tufts of moderately broad and lanceolate scales, respectively. Uncus a short digitate process with three apical setae; tegumen short, sub-rectangular, a sclerotized belt (gnathos) connects the dorsolateral corners of the tegumen, from these corners two very long and rather stout setae are arising, anterior margin nearly straight, pedunculi very small; valva weakly bird’s head-shaped, moderately broad, with strongly convex mediodorsal (outer) and weakly concave ventral (inner) margin, distal part strongly tapered; separate plate-like sclerite at base of valva, covered with few setae; sacculus a short setose lobe; saccus almost as long as distance from anterior margin of vinculum to tip of valva, basally moderately broad, distally evenly tapered; phallus cone-shaped, moderately broad and short, distal part gradually constricted towards distinctly tapered apical fifth, about onequarter of maximum width of its anterior part; vesica with number of small grains.
</paragraph>
<paragraph id="9B01F9117A07FFEDE9DCFB16185AE5D7" blockId="25.[151,1436,151,1653]" box="[199,538,1231,1256]" pageId="25" pageNumber="94">Female genitalia.—Unknown.</paragraph>
</subSubSection>
<subSubSection id="D3A4AA9A7A07FFEDE9DCFB2D18F6E4FF" pageId="25" pageNumber="94" type="diagnosis">
<paragraph id="9B01F9117A07FFEDE9DCFB2D19F5E4A2" blockId="25.[151,1436,151,1653]" pageId="25" pageNumber="94">
DIAGNOSIS.—
<taxonomicName id="5CBE82927A07FFEDE87DFB2C1B85E432" box="[358,453,1268,1293]" class="Insecta" family="Gelechiidae" genus="Eulamprotes" kingdom="Animalia" order="Lepidoptera" pageId="25" pageNumber="94" phylum="Arthropoda" rank="species" species="kailai" status="sp. nov.">
<emphasis id="A9CA25037A07FFEDE87DFB2C1B85E432" box="[358,453,1268,1293]" italics="true" pageId="25" pageNumber="94">E. kailai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B2F998787A07FFEDE8D7FB2C1863E433" box="[460,547,1269,1292]" pageId="25" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
is characterized by being rather small, by having the antennae ringed black and whitish to tip, and by the almost pure white markings in the forewing with only a slight silvery shine. It is most similar to
<taxonomicName id="5CBE82927A07FFEDE81CFAE41BE2E46A" box="[263,418,1340,1365]" class="Insecta" family="Gelechiidae" genus="Eulamprotes" kingdom="Animalia" order="Lepidoptera" pageId="25" pageNumber="94" phylum="Arthropoda" rank="species" species="libertinella">
<emphasis id="A9CA25037A07FFEDE81CFAE41BE2E46A" box="[263,418,1340,1365]" italics="true" pageId="25" pageNumber="94">E. libertinella</emphasis>
</taxonomicName>
, but that species is larger (
<quantity id="5C4654F47A07FFEDEBD0FAE51909E46B" box="[715,841,1340,1364]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="25" pageNumber="94" unit="mm" value="12.5" valueMax="13.0" valueMin="12.0">12–13 mm</quantity>
), the markings of the forewings more silvery shining, and the pre-apical and tornal spots are more separated. The male genitalia match several species of
<taxonomicName id="5CBE82927A07FFEDEC15FA861FDCE447" box="[1294,1436,1375,1400]" class="Insecta" family="Gelechiidae" genus="Eulamprotes" kingdom="Animalia" order="Lepidoptera" pageId="25" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="A9CA25037A07FFEDEC15FA861FDCE447" box="[1294,1436,1375,1400]" italics="true" pageId="25" pageNumber="94">Eulamprotes</emphasis>
</taxonomicName>
with short uncus but differ from all these taxa in the shape of the valva.
</paragraph>
<paragraph id="9B01F9117A07FFEDE9DCFA7118F6E4FF" blockId="25.[151,1436,151,1653]" box="[199,694,1447,1472]" pageId="25" pageNumber="94">GENETIC DATA.—No genetic data available.</paragraph>
</subSubSection>
<subSubSection id="D3A4AA9A7A07FFEDE9DCFA151F58E4DA" box="[199,1304,1484,1509]" pageId="25" pageNumber="94" type="distribution">
<paragraph id="9B01F9117A07FFEDE9DCFA151F58E4DA" blockId="25.[151,1436,151,1653]" box="[199,1304,1484,1509]" pageId="25" pageNumber="94">
DISTRIBUTION.—Central Asia to Siberia (
<collectingCountry id="E3A9B9817A07FFEDEB82FA151960E4DA" box="[665,800,1484,1509]" name="Kazakhstan" pageId="25" pageNumber="94">Kazakhstan</collectingCountry>
, Kyrgizia,
<collectingCountry id="E3A9B9817A07FFEDEA80FA1519AAE4DA" box="[923,1002,1484,1509]" name="Russia" pageId="25" pageNumber="94">Russia</collectingCountry>
: Buryatia, Tuva Republic).
</paragraph>
</subSubSection>
<subSubSection id="D3A4AA9A7A07FFEDE9DCFA29185EE713" pageId="25" pageNumber="94" type="biology_ecology">
<paragraph id="9B01F9117A07FFEDE9DCFA29185EE713" blockId="25.[151,1436,151,1653]" pageId="25" pageNumber="94">
BIOLOGY.—Host-plant and early stages unknown. Adults were collected in June in different kinds of biotope at altitudes between 450 and
<quantity id="5C4654F47A07FFEDE8DAF9CD1859E713" box="[449,537,1556,1581]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="25" pageNumber="94" unit="m" value="2500.0">2500 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="D3A4AA9A7A07FFEDE9DCF9E11825E7DE" pageId="25" pageNumber="94" type="etymology">
<paragraph id="9B01F9117A07FFEDE9DCF9E1197EE74A" blockId="25.[151,1436,151,1653]" pageId="25" pageNumber="94">
ETYMOLOGY.—The name (a noun in the genitive case) is dedicated to Lauri Kaila (ZMUH) who collected the
<typeStatus id="440547B37A07FFEDE98CF9841A87E74A" box="[151,199,1629,1653]" pageId="25" pageNumber="94">type</typeStatus>
series and numerous additional
<taxonomicName id="5CBE82927A07FFEDEB2AF98518FFE74A" box="[561,703,1628,1653]" class="Insecta" family="Gelechiidae" genus="Eulamprotes" kingdom="Animalia" order="Lepidoptera" pageId="25" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="A9CA25037A07FFEDEB2AF98518FFE74A" box="[561,703,1628,1653]" italics="true" pageId="25" pageNumber="94">Eulamprotes</emphasis>
</taxonomicName>
from Asia.
</paragraph>
<paragraph id="9B01F9117A07FFEDE98CF91E1825E7DE" blockId="25.[151,613,1735,1761]" box="[151,613,1735,1761]" pageId="25" pageNumber="94">
<taxonomicName id="5CBE82927A07FFEDE98CF91E1825E7DE" authority="Zeller, 1872" authorityName="Zeller" authorityYear="1872" box="[151,613,1735,1761]" class="Insecta" family="Gelechiidae" genus="Eulamprotes" kingdom="Animalia" order="Lepidoptera" pageId="25" pageNumber="94" phylum="Arthropoda" rank="species" species="libertinella">
<emphasis id="A9CA25037A07FFEDE98CF91E1BFDE7DE" bold="true" box="[151,445,1735,1761]" italics="true" pageId="25" pageNumber="94">Eulamprotes libertinella</emphasis>
(Zeller, 1872)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="D3A4AA9A7A07FFEEE98CF8D718BCE5A0" lastPageId="26" lastPageNumber="95" pageId="25" pageNumber="94" type="description">
<paragraph id="9B01F9117A07FFEDE98CF8D718CEE61B" blockId="25.[151,654,1806,1828]" box="[151,654,1806,1828]" pageId="25" pageNumber="94">
<taxonomicName id="5CBE82927A07FFEDE98CF8D718CAE61B" authority="Zeller 1872: 112" authorityName="Zeller" authorityPageNumber="112" authorityYear="1872" box="[151,650,1806,1828]" class="Insecta" family="Gelechiidae" genus="Gelechia" kingdom="Animalia" order="Lepidoptera" pageId="25" pageNumber="94" phylum="Arthropoda" rank="species" species="libertinella" subGenus="Argyritis">
<emphasis id="A9CA25037A07FFEDE98CF8D71AB3E61B" box="[151,243,1806,1828]" italics="true" pageId="25" pageNumber="94">Gelechia</emphasis>
(
<emphasis id="A9CA25037A07FFEDE81AF8D61B1AE61B" box="[257,346,1807,1828]" italics="true" pageId="25" pageNumber="94">Argyritis</emphasis>
)
<emphasis id="A9CA25037A07FFEDE872F8D71B9AE61B" box="[361,474,1806,1828]" italics="true" pageId="25" pageNumber="94">libertinella</emphasis>
Zeller 1872: 112
</taxonomicName>
.
</paragraph>
<paragraph id="9B01F9117A07FFEDE98CF8881AB1E6B1" blockId="25.[151,1436,1873,2006]" pageId="25" pageNumber="94">
Examined material.
<collectingCountry id="E3A9B9817A07FFEDE863F8881B8AE655" box="[376,458,1873,1898]" name="Austria" pageId="25" pageNumber="94">Austria</collectingCountry>
(n=58),
<collectingCountry id="E3A9B9817A07FFEDEB31F88B1836E656" box="[554,630,1874,1897]" name="France" pageId="25" pageNumber="94">France</collectingCountry>
(n=68),
<collectingCountry id="E3A9B9817A07FFEDEBCCF888194AE655" box="[727,778,1873,1898]" name="Italy" pageId="25" pageNumber="94">Italy</collectingCountry>
(n=136),
<collectingCountry id="E3A9B9817A07FFEDEA63F88819BEE655" box="[888,1022,1873,1898]" name="Switzerland" pageId="25" pageNumber="94">Switzerland</collectingCountry>
(n=14) (see also Huemer &amp; Karsholt (2011)).
</paragraph>
<paragraph id="9B01F9117A07FFEEE9DCF8431ABCE07F" blockId="25.[151,1436,1873,2006]" lastBlockId="26.[151,1437,151,2013]" lastPageId="26" lastPageNumber="95" pageId="25" pageNumber="94">
DESCRIPTION.—Adult (
<figureCitation id="0385E5947A07FFEDE8D5F8401812E68D" box="[462,594,1945,1970]" captionStart="FIGURES 13 – 21" captionStartId="6.[151,263,1418,1441]" captionTargetBox="[188,1396,493,1383]" captionTargetId="figure@6.[183,1404,483,1394]" captionTargetPageId="6" captionText="FIGURES 13 – 21. Adults of Eulamprotes spp. 13, E. occidentella Huemer &amp; Karsholt, paratype, ♂ (Italy); 14, ditto, paratype, ♀ (Italy); 15, E. atrifrontella sp. nov., paratype, ♂ (Turkey); 16, E. wieseri sp. nov., holotype, ♂ (Kirgizia); 17, E. altaicella sp. nov., holotype, ♂ (Russia, Altai); 18, E. kailai sp. nov., holotype, ♂ (Kirgizia); 19, E. libertinella (Zeller), ♂ (Italy); 20, ditto, ♀ (Switzerland); 21, E. gemerensis sp. nov., paratype, ♂ (Slovakia). Scale bar = 2 mm." httpUri="https://zenodo.org/record/219623/files/figure.png" pageId="25" pageNumber="94">Figs 19–20</figureCitation>
). Male. Wingspan
<quantity id="5C4654F47A07FFEDEA3FF84319E0E68E" box="[804,928,1946,1970]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="25" pageNumber="94" unit="mm" value="12.5" valueMax="13.0" valueMin="12.0">12–13 mm</quantity>
. Labial palpus white; basal half of segment 2 black on outer surface; tip of segment 3 black. Antenna black, distal part ringed with light grey and whitish grey towards tip. Head whitish; thorax and tegula black. Forewing black with silvery white markings: an oblique streak from 1/6 at costa to 1/5 near dorsum; a streak from middle of costa reaching two-thirds towards dorsum; a silvery white tornal spot well separated from a cream-white pre-apical costal spot; some silvery white scales in apex and along termen; cilia blackish grey, whitish grey at tip of apex. Hindwing about as broad as forewing, grey. Abdomen blackish.
</paragraph>
<paragraph id="9B01F9117A04FFEEE9DCFE951F11E0EF" blockId="26.[151,1437,151,2013]" pageId="26" pageNumber="95">
Female. Wingspan
<quantity id="5C4654F47A04FFEEE8BFFE95184DE05B" box="[420,525,332,357]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="26" pageNumber="95" unit="mm" value="10.5">10.5 mm</quantity>
. Labial palpus cream-white; distal part of segment 3 black. Head cream-white. Forewing broadest towards base, tapering towards tip; silvery white streaks from costa slender; apex without silvery white scales. Hindwing reduced, no longer than breadth of forewing, light grey. Abdomen black, lighter dorsally on basal segments; posterior part of each segment on dorsal side silvery. Otherwise similar to male.
</paragraph>
<paragraph id="9B01F9117A04FFEEE9DCFE051E65E35F" blockId="26.[151,1437,151,2013]" pageId="26" pageNumber="95">
Variation. There is some variation in the amount of dark scales in the labial palpi. The basal part of the antennae can be black for a shorter or longer distance. In the forewings the silvery white markings can be more or less whitish, and the apex can be almost without silvery scales. Some of this variation also depends on the condition of the specimens. The geographical variation in
<taxonomicName id="5CBE82927A04FFEEEA38FD9019FFE35F" box="[803,959,583,608]" class="Insecta" family="Gelechiidae" genus="Gelechia" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="95" phylum="Arthropoda" rank="species" species="libertinella">
<emphasis id="A9CA25037A04FFEEEA38FD9019FFE35F" box="[803,959,583,608]" italics="true" pageId="26" pageNumber="95">E. libertinella</emphasis>
</taxonomicName>
is slight.
</paragraph>
<paragraph id="9B01F9117A04FFEEE9DCFDB51893E2F7" blockId="26.[151,1437,151,2013]" pageId="26" pageNumber="95">
Male genitalia (
<figureCitation id="0385E5947A04FFEEE86DFDB51BB6E3BA" box="[374,502,620,645]" captionStart="FIGURES 46 – 49" captionStartId="16.[151,263,1276,1299]" captionTargetBox="[160,1415,221,1227]" captionTargetId="figure@16.[151,1436,195,1253]" captionTargetPageId="16" captionText="FIGURES 46 – 49. Male genitalia of Eulamprotes spp. 46, E. libertinella (Zeller), (Italy) gen. slide GEL 1154 P. Huemer; 47, ditto, phallus; 48, E. gemerensis sp. nov., paratype, (Slovakia) gen. slide GU 13 / 1344 P. Huemer; 49, ditto, phallus." httpUri="https://zenodo.org/record/219628/files/figure.png" pageId="26" pageNumber="95">Figs 46–47</figureCitation>
).—Segment VIII with two pairs of coremata in intersegmental membrane, grouped into short tufts of moderately broad and lanceolate scales, respectively. Uncus a comparatively long digitate process with several apical setae; tegumen short, sub-rectangular, a sclerotized belt (gnathos) connects the dorsolateral corners of the tegumen, from these corners two long and rather stout setae are arising, anterior margin nearly straight, pedunculi very small; valva moderately broad, with strongly convex mediodorsal (outer) and almost straight ventral (inner) margin, apically tapered; separate plate-like sclerite at base of valva, covered with few setae; sacculus a distinct, setose lobe; saccus almost as long as distance from anterior margin of vinculum to tip of valva, basally moderately broad, distally gradually tapered; phallus cone-shaped, basally broad and moderately short, distal part gradually constricted to distinctly tapered apical fifth, about one-quarter of maximum width of its anterior part; vesica with a number of small grains.
</paragraph>
<paragraph id="9B01F9117A04FFEEE9DCFC0D18BCE5A0" blockId="26.[151,1437,151,2013]" pageId="26" pageNumber="95">
Female genitalia (
<figureCitation id="0385E5947A04FFEEE888FC0D1BA3E2D2" box="[403,483,980,1005]" captionStart="FIGURES 54 – 55" captionStartId="19.[151,263,1720,1743]" captionTargetBox="[410,1183,326,1689]" captionTargetId="figure@19.[394,1192,303,1696]" captionTargetPageId="19" captionText="FIGURES 54 – 55. Female genitalia of Eulamprotes spp. 54, E. occidentella Huemer &amp; Karsholt sp. nov., paratype, (Italy) gen. slide GEL 1141 P. Huemer; 55, E. libertinella (Zeller), (Austria) gen. slide GEL 1142 P. Huemer." httpUri="https://zenodo.org/record/219631/files/figure.png" pageId="26" pageNumber="95">Fig. 55</figureCitation>
).—Apophysis posterioris slender, rod-shaped, about three times as long as segment VIII; segment VIII with weakly sclerotized posteriolateral part, otherwise membranous; ostium bursae laterally with sclerotized folds; apophysis anterioris slender, rod-shaped, about two times as long as segment VIII; ductus bursae long and slender, posterior half with slender, oblong sclerite extending distinctly beyond apex of apophysis anterioris, anterior half membranous, evenly expanded; corpus bursae suboval, small; signum a large suboval plate, anterior and posterior edge with a short spinal process.
</paragraph>
</subSubSection>
<subSubSection id="D3A4AA9A7A04FFEEE9DCFB751F0EE4DA" pageId="26" pageNumber="95" type="diagnosis">
<paragraph id="9B01F9117A04FFEEE9DCFB7518B0E447" blockId="26.[151,1437,151,2013]" pageId="26" pageNumber="95">
DIAGNOSIS.—
<taxonomicName id="5CBE82927A04FFEEE87DFB741846E5FA" box="[358,518,1196,1221]" class="Insecta" family="Gelechiidae" genus="Gelechia" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="95" phylum="Arthropoda" rank="species" species="libertinella">
<emphasis id="A9CA25037A04FFEEE87DFB741846E5FA" box="[358,518,1196,1221]" italics="true" pageId="26" pageNumber="95">E. libertinella</emphasis>
</taxonomicName>
resembles
<taxonomicName id="5CBE82927A04FFEEEB92FB741975E5FA" box="[649,821,1196,1221]" class="Insecta" family="Gelechiidae" genus="Eulamprotes" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="95" phylum="Arthropoda" rank="species" species="occidentella">
<emphasis id="A9CA25037A04FFEEEB92FB741975E5FA" box="[649,821,1196,1221]" italics="true" pageId="26" pageNumber="95">E. occidentella</emphasis>
</taxonomicName>
,
<taxonomicName id="5CBE82927A04FFEEEA5EFB7419E8E5FA" box="[837,936,1196,1221]" class="Insecta" family="Gelechiidae" genus="Eulamprotes" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="95" phylum="Arthropoda" rank="species" species="kailai" status="sp. nov.">
<emphasis id="A9CA25037A04FFEEEA5EFB7419E8E5FA" box="[837,936,1196,1221]" italics="true" pageId="26" pageNumber="95">E. kailai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B2F998787A04FFEEEAAAFB741E4BE5FB" box="[945,1035,1197,1220]" pageId="26" pageNumber="95" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="5CBE82927A04FFEEED5CFB741E94E5FA" box="[1095,1236,1196,1221]" class="Insecta" family="Gelechiidae" genus="Eulamprotes" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="95" phylum="Arthropoda" rank="species" species="mirusella" status="sp. nov.">
<emphasis id="A9CA25037A04FFEEED5CFB741E94E5FA" box="[1095,1236,1196,1221]" italics="true" pageId="26" pageNumber="95">E. mirusella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B2F998787A04FFEEEDC6FB741F6DE5FB" box="[1245,1325,1197,1220]" pageId="26" pageNumber="95" rank="species">sp.nov.</taxonomicNameLabel>
It is also similar to
<taxonomicName id="5CBE82927A04FFEEE811FB081BC2E5D7" box="[266,386,1231,1256]" class="Insecta" family="Gelechiidae" genus="Phalaena" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="95" phylum="Arthropoda" rank="species" species="wilkella">
<emphasis id="A9CA25037A04FFEEE811FB081BC2E5D7" box="[266,386,1231,1256]" italics="true" pageId="26" pageNumber="95">E. wilkella</emphasis>
</taxonomicName>
. The latter has almost normally winged females, has the distal part of the antenna white, and has the tornal and pre-apical costal spot of the forewing connected by an oblique fascia. The male genitalia are overall similar to several other species but differ by collectively diagnostic features, particularly the distinct digitate uncus and the shape of the valva. The female genitalia differ from other species by the shape of the signum with a single spine on the anterior and posterior edge.
</paragraph>
<paragraph id="9B01F9117A04FFEEE9DCFA5D1F0EE4DA" blockId="26.[151,1437,151,2013]" pageId="26" pageNumber="95">
GENETIC DATA.—
<taxonomicName id="5CBE82927A04FFEEE888FA5C1867E4A2" box="[403,551,1412,1437]" class="Insecta" family="Gelechiidae" genus="Gelechia" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="95" phylum="Arthropoda" rank="species" species="libertinella">
<emphasis id="A9CA25037A04FFEEE888FA5C1867E4A2" box="[403,551,1412,1437]" italics="true" pageId="26" pageNumber="95">E. libertinella</emphasis>
</taxonomicName>
shows an exceptional intraspecific divergence of the barcode region, ranging from 2.18%–5.43% (n=16) with geographically separated major haplotypes. Within these genetic clades divergence is low with a maximum of 1.08%. The distance of the barcode to the nearest neighbour
<taxonomicName id="5CBE82927A04FFEEEAEDFA141ECAE4DB" box="[1014,1162,1485,1508]" class="Insecta" family="Gelechiidae" genus="Eulamprotes" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="95" phylum="Arthropoda" rank="species" species="gemerensis" status="sp. nov.">
<emphasis id="A9CA25037A04FFEEEAEDFA141ECAE4DB" box="[1014,1162,1485,1508]" italics="true" pageId="26" pageNumber="95">E. gemerensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B2F998787A04FFEEED8AFA141EA3E4DB" box="[1169,1251,1485,1508]" pageId="26" pageNumber="95" rank="species">sp. nov.</taxonomicNameLabel>
is 6.14%.
</paragraph>
</subSubSection>
<subSubSection id="D3A4AA9A7A04FFEEE9DCFA291E29E74A" pageId="26" pageNumber="95" type="distribution">
<paragraph id="9B01F9117A04FFEEE9DCFA291E29E74A" blockId="26.[151,1437,151,2013]" pageId="26" pageNumber="95">
DISTRIBUTION.—Known from the major part of the Alpine arc, ranging from the Maritime Alps in the southwest up to the north-eastern Austrian Limestone Alps in the north-east and the Julian Alps in the south-east. Records from
<collectingCountry id="E3A9B9817A04FFEEE857F9EE1BCCE76F" box="[332,396,1591,1616]" name="Spain" pageId="26" pageNumber="95">Spain</collectingCountry>
(Seebold 1899), the
<collectingCountry id="E3A9B9817A04FFEEEB85F9EE191AE76F" box="[670,858,1591,1616]" name="Czech Republic" pageId="26" pageNumber="95">Czech Republic</collectingCountry>
(Sterneck &amp; Zimmermann 1933) and Bosnia-
<collectingCountry id="E3A9B9817A04FFEEE98CF9851B66E74A" box="[151,294,1628,1653]" name="Bosnia and Herzegovina" pageId="26" pageNumber="95">Herzegovina</collectingCountry>
(Rebel 1904) are unconfirmed and most likely belong to different species.
</paragraph>
</subSubSection>
<subSubSection id="D3A4AA9A7A04FFEEE9DCF9591F25E64F" pageId="26" pageNumber="95" type="biology_ecology">
<paragraph id="9B01F9117A04FFEEE9DCF9591F25E64F" blockId="26.[151,1437,151,2013]" pageId="26" pageNumber="95">
BIOLOGY.—Host-plant and early stages unknown. One female was bred from moss and lichens collected on rocks in NW
<collectingCountry id="E3A9B9817A04FFEEE837F97D1B1FE782" box="[300,351,1700,1725]" name="Italy" pageId="26" pageNumber="95">Italy</collectingCountry>
together with specimens of
<taxonomicName id="5CBE82927A04FFEEEB82F97D18BEE782" box="[665,766,1700,1725]" class="Insecta" family="Tineidae" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="95" phylum="Arthropoda" rank="family">Tineidae</taxonomicName>
. It ran rapidly around using legs and tips of the forewings in combination, being unable to fly (Baldizzone 2007). The adults have been collected from June to early September, depending on the altitude of the habitat. The males are easily attracted to light but can also be found during the day. The brachypterous female was observed baiting males in the evening at sunset (Huemer &amp; Karsholt 2011). Females are rare in collections.
<taxonomicName id="5CBE82927A04FFEEE882F8EC1875E672" box="[409,565,1844,1869]" class="Insecta" family="Gelechiidae" genus="Gelechia" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="95" phylum="Arthropoda" rank="species" species="libertinella">
<emphasis id="A9CA25037A04FFEEE882F8EC1875E672" box="[409,565,1844,1869]" italics="true" pageId="26" pageNumber="95">E. libertinella</emphasis>
</taxonomicName>
prefers rocky habitats and scree with scattered vegetation both on calcareous and siliceous soil. It prefers altitudes between
<date id="EF00DFD17A04FFEEEBBFF881190AE64F" box="[676,842,1879,1904]" pageId="26" pageNumber="95" value="1000" valueMax="2600">
1000 and
<quantity id="5C4654F47A04FFEEEA09F88E190AE64F" box="[786,842,1879,1904]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="26" pageNumber="95" unit="m" value="2600.0">2600</quantity>
</date>
m, but is occasionally found at lower elevation.
</paragraph>
</subSubSection>
<subSubSection id="D3A4AA9A7A04FFEFE9DCF8A518A8E1EA" lastPageId="27" lastPageNumber="96" pageId="26" pageNumber="95" type="discussion">
<paragraph id="9B01F9117A04FFEFE9DCF8A518A8E1EA" blockId="26.[151,1437,151,2013]" lastBlockId="27.[151,1436,151,213]" lastPageId="27" lastPageNumber="96" pageId="26" pageNumber="95">
REMARKS.—
<taxonomicName id="5CBE82927A04FFEEE841F8A41BB8E6AA" box="[346,504,1916,1941]" class="Insecta" family="Gelechiidae" genus="Gelechia" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="95" phylum="Arthropoda" rank="species" species="libertinella">
<emphasis id="A9CA25037A04FFEEE841F8A41BB8E6AA" box="[346,504,1916,1941]" italics="true" pageId="26" pageNumber="95">E. libertinella</emphasis>
</taxonomicName>
was described from an unspecified number of specimens collected in the area of Bergün (Graubünden,
<collectingCountry id="E3A9B9817A04FFEEE88CF879185FE686" box="[407,543,1952,1977]" name="Switzerland" pageId="26" pageNumber="95">Switzerland</collectingCountry>
) and four specimens from the Hochschwab area and one from the Tragösstal near Bruck an der Muhr (Styria,
<collectingCountry id="E3A9B9817A04FFEEEB18F81D1817E6E2" box="[515,599,1988,2013]" name="Austria" pageId="26" pageNumber="95">Austria</collectingCountry>
) (Zeller 1872). Huemer &amp; Karsholt (2011) selected a male from Bergün as the
<typeStatus id="440547B37A05FFEFE9D9FF4E1B6CE18F" box="[194,300,151,176]" pageId="27" pageNumber="96" type="lectotype">lectotype</typeStatus>
. The remarkable genetic divergence may reflect cryptic diversity but so far we have been unable to find substantial diagnostic characters in morphology.
</paragraph>
</subSubSection>
</treatment>
</document>
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