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<document id="477D046DF4F27383EBE6403D6AFCFA4C" ID-DOI="10.11646/zootaxa.3866.2.1" ID-GBIF-Dataset="f0c34243-cd3b-49fa-b4b7-ff17bc96c362" ID-ISSN="1175-5326" ID-Zenodo-Dep="249555" ID-ZooBank="2165AC20-3607-4CF2-A1A6-DB4CA45D7E22" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460916321666" checkinUser="plazi" docAuthor="Darby, Michael" docDate="2014" docId="F3388D2751343C3DE6EBFB5D494968EE" docLanguage="en" docName="zt03866p201.pdf" docOrigin="Zootaxa 3866 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Acrotrichis heissi Darby, 2014, sp. n." docType="treatment" docVersion="9" lastPageNumber="175" masterDocId="0F01F55F512C3C25E67CFF934A6A6F58" masterDocTitle="Studies of Madagascan Ptiliidae (Coleoptera) 3: The genus Acrotrichis including twenty seven new species" masterLastPageNumber="201" masterPageNumber="151" pageNumber="175" updateTime="1698361750748" updateUser="plazi">
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<mods:title id="6CF01E02A0B13DA3A1C12E6A48655799">Studies of Madagascan Ptiliidae (Coleoptera) 3: The genus Acrotrichis including twenty seven new species</mods:title>
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<mods:namePart id="1D4B01EC66DC9BF535516A3FDDC80F85">Darby, Michael</mods:namePart>
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<mods:date id="4CCB8D22B13F109554DDBDAB5506ADEC">2014</mods:date>
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<treatment id="F3388D2751343C3DE6EBFB5D494968EE" ID-DOI="http://doi.org/10.5281/zenodo.6131029" ID-GBIF-Taxon="119588822" ID-Zenodo-Dep="6131029" LSID="urn:lsid:plazi:treatment:F3388D2751343C3DE6EBFB5D494968EE" httpUri="http://treatment.plazi.org/id/F3388D2751343C3DE6EBFB5D494968EE" lastPageNumber="175" pageId="24" pageNumber="175">
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<heading id="20668B5D51343C3DE6EBFB5D4BD96BB0" bold="true" box="[151,435,1230,1256]" fontSize="11" level="1" pageId="24" pageNumber="175" reason="1">
<emphasis id="49E5E02351343C3DE6EBFB5D4BD96BB0" bold="true" box="[151,435,1230,1256]" pageId="24" pageNumber="175">
<taxonomicName id="BC9147B251343C3DE6EBFB5D4B006BB0" ID-CoL="9RK7" box="[151,362,1230,1256]" class="Insecta" family="Ptiliidae" genus="Acrotrichis" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="175" phylum="Arthropoda" rank="species" species="heissi" status="sp. nov.">
<emphasis id="49E5E02351343C3DE6EBFB5D4B006BB0" bold="true" box="[151,362,1230,1256]" italics="true" pageId="24" pageNumber="175">Acrotrichis heissi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="52D65D5851343C3DE70DFB5D4BD96BB0" box="[369,435,1230,1256]" pageId="24" pageNumber="175" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="7B2E3C3151343C3DE6EBFB624B456A52" blockId="24.[151,435,1230,1290]" box="[151,303,1265,1290]" pageId="24" pageNumber="175">
(
<figureCitation id="E3AA20B451343C3DE6E3FB624B656A52" box="[159,271,1265,1290]" captionStart="FIGURE 17. A" captionStartId="26.[151,250,1503,1525]" captionTargetBox="[231,1367,239,1439]" captionTargetId="figure@26.[151,1436,193,1482]" captionTargetPageId="26" captionText="FIGURE 17. A. heissi. A, habitus; B, side of pronotum viewed at 45 ˚; C, antennomeres 3 11; D, spermatheca; E, aedeagus; F, ventrite six; G, mesoventral collar x 535; H, metaventral posterior margin and metacoxal plates x 495; I, mentum and submentum showing chaetotaxy." httpUri="https://zenodo.org/record/249572/files/figure.png" pageId="24" pageNumber="175">Figs. 17A</figureCitation>
I)
</paragraph>
<paragraph id="7B2E3C3151343C3DE6EBFAAA49996ABA" blockId="24.[151,1437,1337,1974]" pageId="24" pageNumber="175">
Habitus
<figureCitation id="E3AA20B451343C3DE684FAAA4B0B6A0A" box="[248,353,1337,1362]" captionStart="FIGURE 17. A" captionStartId="26.[151,250,1503,1525]" captionTargetBox="[231,1367,239,1439]" captionTargetId="figure@26.[151,1436,193,1482]" captionTargetPageId="26" captionText="FIGURE 17. A. heissi. A, habitus; B, side of pronotum viewed at 45 ˚; C, antennomeres 3 11; D, spermatheca; E, aedeagus; F, ventrite six; G, mesoventral collar x 535; H, metaventral posterior margin and metacoxal plates x 495; I, mentum and submentum showing chaetotaxy." httpUri="https://zenodo.org/record/249572/files/figure.png" pageId="24" pageNumber="175">Fig. 17A</figureCitation>
. Length
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. Colour brownish black, antennae and legs brownish yellow. Antennomeres 311
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long
<figureCitation id="E3AA20B451343C3DE70CFACE4BAB6A2E" box="[368,449,1373,1398]" captionStart="FIGURE 17. A" captionStartId="26.[151,250,1503,1525]" captionTargetBox="[231,1367,239,1439]" captionTargetId="figure@26.[151,1436,193,1482]" captionTargetPageId="26" captionText="FIGURE 17. A. heissi. A, habitus; B, side of pronotum viewed at 45 ˚; C, antennomeres 3 11; D, spermatheca; E, aedeagus; F, ventrite six; G, mesoventral collar x 535; H, metaventral posterior margin and metacoxal plates x 495; I, mentum and submentum showing chaetotaxy." httpUri="https://zenodo.org/record/249572/files/figure.png" pageId="24" pageNumber="175">Fig. 17</figureCitation>
C. Mentum and submentum chaetotaxy
<figureCitation id="E3AA20B451343C3DE503FACE49A76A2E" box="[895,973,1373,1398]" captionStart="FIGURE 17. A" captionStartId="26.[151,250,1503,1525]" captionTargetBox="[231,1367,239,1439]" captionTargetId="figure@26.[151,1436,193,1482]" captionTargetPageId="26" captionText="FIGURE 17. A. heissi. A, habitus; B, side of pronotum viewed at 45 ˚; C, antennomeres 3 11; D, spermatheca; E, aedeagus; F, ventrite six; G, mesoventral collar x 535; H, metaventral posterior margin and metacoxal plates x 495; I, mentum and submentum showing chaetotaxy." httpUri="https://zenodo.org/record/249572/files/figure.png" pageId="24" pageNumber="175">
Fig.
<date id="0F2F1AF151343C3DE5CEFACD49A76A2E" box="[946,973,1374,1398]" pageId="24" pageNumber="175">17</date>
</figureCitation>
I. Width across eyes
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. Pronotum
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wide,
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long. sides strongly rounded
<figureCitation id="E3AA20B451343C3DE577FA1249346AC2" box="[779,862,1409,1434]" captionStart="FIGURE 17. A" captionStartId="26.[151,250,1503,1525]" captionTargetBox="[231,1367,239,1439]" captionTargetId="figure@26.[151,1436,193,1482]" captionTargetPageId="26" captionText="FIGURE 17. A. heissi. A, habitus; B, side of pronotum viewed at 45 ˚; C, antennomeres 3 11; D, spermatheca; E, aedeagus; F, ventrite six; G, mesoventral collar x 535; H, metaventral posterior margin and metacoxal plates x 495; I, mentum and submentum showing chaetotaxy." httpUri="https://zenodo.org/record/249572/files/figure.png" pageId="24" pageNumber="175">Fig. 17</figureCitation>
B. Elytra
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long,
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wide. Posterior margin of mesoventral collar sloped at shoulders and with a shallow medial excision
<figureCitation id="E3AA20B451343C3DE228FA364EC86AE6" box="[1108,1186,1445,1470]" captionStart="FIGURE 17. A" captionStartId="26.[151,250,1503,1525]" captionTargetBox="[231,1367,239,1439]" captionTargetId="figure@26.[151,1436,193,1482]" captionTargetPageId="26" captionText="FIGURE 17. A. heissi. A, habitus; B, side of pronotum viewed at 45 ˚; C, antennomeres 3 11; D, spermatheca; E, aedeagus; F, ventrite six; G, mesoventral collar x 535; H, metaventral posterior margin and metacoxal plates x 495; I, mentum and submentum showing chaetotaxy." httpUri="https://zenodo.org/record/249572/files/figure.png" pageId="24" pageNumber="175">Fig. 17</figureCitation>
G. Posterior margin of metaventrum between metacoxae short
<figureCitation id="E3AA20B451343C3DE428FA5A48C96ABA" box="[596,675,1481,1506]" captionStart="FIGURE 17. A" captionStartId="26.[151,250,1503,1525]" captionTargetBox="[231,1367,239,1439]" captionTargetId="figure@26.[151,1436,193,1482]" captionTargetPageId="26" captionText="FIGURE 17. A. heissi. A, habitus; B, side of pronotum viewed at 45 ˚; C, antennomeres 3 11; D, spermatheca; E, aedeagus; F, ventrite six; G, mesoventral collar x 535; H, metaventral posterior margin and metacoxal plates x 495; I, mentum and submentum showing chaetotaxy." httpUri="https://zenodo.org/record/249572/files/figure.png" pageId="24" pageNumber="175">Fig. 17</figureCitation>
H; metacoxal plates
<figureCitation id="E3AA20B451343C3DE5F5FA5A49B26ABA" box="[905,984,1481,1506]" captionStart="FIGURE 17. A" captionStartId="26.[151,250,1503,1525]" captionTargetBox="[231,1367,239,1439]" captionTargetId="figure@26.[151,1436,193,1482]" captionTargetPageId="26" captionText="FIGURE 17. A. heissi. A, habitus; B, side of pronotum viewed at 45 ˚; C, antennomeres 3 11; D, spermatheca; E, aedeagus; F, ventrite six; G, mesoventral collar x 535; H, metaventral posterior margin and metacoxal plates x 495; I, mentum and submentum showing chaetotaxy." httpUri="https://zenodo.org/record/249572/files/figure.png" pageId="24" pageNumber="175">Fig. 17</figureCitation>
H.
</paragraph>
</subSubSection>
<subSubSection id="338B6FBA51343C3DE6BBFA7E4E846972" pageId="24" pageNumber="175" type="description">
<paragraph id="7B2E3C3151343C3DE6BBFA7E4F7F695E" blockId="24.[151,1437,1337,1974]" box="[199,1301,1517,1542]" pageId="24" pageNumber="175">
<emphasis id="49E5E02351343C3DE6BBFA7E4B6B695E" box="[199,257,1517,1542]" italics="true" pageId="24" pageNumber="175">Male</emphasis>
: ventrite six with +/- 13 spines in two rows
<figureCitation id="E3AA20B451343C3DE493FA7E4955695E" box="[751,831,1517,1542]" captionStart="FIGURE 17. A" captionStartId="26.[151,250,1503,1525]" captionTargetBox="[231,1367,239,1439]" captionTargetId="figure@26.[151,1436,193,1482]" captionTargetPageId="26" captionText="FIGURE 17. A. heissi. A, habitus; B, side of pronotum viewed at 45 ˚; C, antennomeres 3 11; D, spermatheca; E, aedeagus; F, ventrite six; G, mesoventral collar x 535; H, metaventral posterior margin and metacoxal plates x 495; I, mentum and submentum showing chaetotaxy." httpUri="https://zenodo.org/record/249572/files/figure.png" pageId="24" pageNumber="175">Fig. 17</figureCitation>
F; aedeagus restricted before tip
<figureCitation id="E3AA20B451343C3DE2D1FA7E4E96695E" box="[1197,1276,1517,1542]" captionStart="FIGURE 17. A" captionStartId="26.[151,250,1503,1525]" captionTargetBox="[231,1367,239,1439]" captionTargetId="figure@26.[151,1436,193,1482]" captionTargetPageId="26" captionText="FIGURE 17. A. heissi. A, habitus; B, side of pronotum viewed at 45 ˚; C, antennomeres 3 11; D, spermatheca; E, aedeagus; F, ventrite six; G, mesoventral collar x 535; H, metaventral posterior margin and metacoxal plates x 495; I, mentum and submentum showing chaetotaxy." httpUri="https://zenodo.org/record/249572/files/figure.png" pageId="24" pageNumber="175">Fig. 17</figureCitation>
E.
</paragraph>
<paragraph id="7B2E3C3151343C3DE6BBF9824E846972" blockId="24.[151,1437,1337,1974]" box="[199,1262,1553,1578]" pageId="24" pageNumber="175">
<emphasis id="49E5E02351343C3DE6BBF9824B716972" box="[199,283,1553,1578]" italics="true" pageId="24" pageNumber="175">Female</emphasis>
: spermatheca
<figureCitation id="E3AA20B451343C3DE7C0F98248606972" box="[444,522,1553,1578]" captionStart="FIGURE 17. A" captionStartId="26.[151,250,1503,1525]" captionTargetBox="[231,1367,239,1439]" captionTargetId="figure@26.[151,1436,193,1482]" captionTargetPageId="26" captionText="FIGURE 17. A. heissi. A, habitus; B, side of pronotum viewed at 45 ˚; C, antennomeres 3 11; D, spermatheca; E, aedeagus; F, ventrite six; G, mesoventral collar x 535; H, metaventral posterior margin and metacoxal plates x 495; I, mentum and submentum showing chaetotaxy." httpUri="https://zenodo.org/record/249572/files/figure.png" pageId="24" pageNumber="175">Fig. 17</figureCitation>
D (if twisted slightly, as in some preparations, appears straighter).
</paragraph>
</subSubSection>
<subSubSection id="338B6FBA51343C3DE6BBF9A64E176916" box="[199,1149,1589,1614]" pageId="24" pageNumber="175" type="etymology">
<paragraph id="7B2E3C3151343C3DE6BBF9A64E176916" blockId="24.[151,1437,1337,1974]" box="[199,1149,1589,1614]" pageId="24" pageNumber="175">
<emphasis id="49E5E02351343C3DE6BBF9A64B2C6916" bold="true" box="[199,326,1589,1614]" pageId="24" pageNumber="175">Etymology</emphasis>
. Johnson ms name in honour of his friend Herr Ernst Heiss of Innsbruck.
</paragraph>
</subSubSection>
<subSubSection id="338B6FBA51343C3DE6BBF9CA498E69CE" pageId="24" pageNumber="175" type="discussion">
<paragraph id="7B2E3C3151343C3DE6BBF9CA498E69CE" blockId="24.[151,1437,1337,1974]" pageId="24" pageNumber="175">
<emphasis id="49E5E02351343C3DE6BBF9CA4B5E692A" bold="true" box="[199,308,1625,1650]" pageId="24" pageNumber="175">Remarks</emphasis>
. A rather variable species similar externally to
<taxonomicName id="BC9147B251343C3DE52DF9C949DA692A" box="[849,944,1626,1650]" class="Insecta" family="Ptiliidae" genus="Acrotrichis" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="175" phylum="Arthropoda" rank="species" species="africana">
<emphasis id="49E5E02351343C3DE52DF9C949DA692A" box="[849,944,1626,1650]" italics="true" pageId="24" pageNumber="175">africana</emphasis>
</taxonomicName>
but smaller and narrower, and the pronotal sides are more strongly curved. The spermatheca is very similar to
<emphasis id="49E5E02351343C3DE5FAF9EE498E69CE" box="[902,996,1661,1686]" italics="true" pageId="24" pageNumber="175">
<taxonomicName id="BC9147B251343C3DE5FAF9EE498B69CE" box="[902,993,1661,1686]" class="Insecta" family="Ptiliidae" genus="Acrotrichis" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="175" phylum="Arthropoda" rank="species" species="cuspida">cuspida</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="338B6FBA51343C3DE6BBF932494968EE" pageId="24" pageNumber="175" type="materials_examined">
<paragraph id="7B2E3C3151343C3DE6BBF932494968EE" blockId="24.[151,1437,1337,1974]" pageId="24" pageNumber="175">
<emphasis id="49E5E02351343C3DE6BBF9324B5469E2" bold="true" box="[199,318,1697,1722]" pageId="24" pageNumber="175">
<typeStatus id="A42A829351343C3DE6BBF9324B6869E2" box="[199,258,1697,1722]" pageId="24" pageNumber="175">Type</typeStatus>
data
</emphasis>
.
<typeStatus id="A42A829351343C3DE731F9324BD069E2" box="[333,442,1697,1722]" pageId="24" pageNumber="175" type="holotype">Holotype</typeStatus>
: ♀, Ranomafana NP, Vatoharanana RNF
<date id="0F2F1AF151343C3DE5F7F931499769E2" box="[907,1021,1698,1722]" pageId="24" pageNumber="175" value="2012-09">Sept 2012</date>
GPS 209, sifting litter, Winkler app. extraction, S.21˚1659.7” E.47˚25 30.3”,
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,
<date id="0F2F1AF151343C3DE4AAF95649516986" box="[726,827,1733,1758]" pageId="24" pageNumber="175" value="2012-10-05">5.x.2012</date>
, L.S.Rahanitriniaina (
<collectionCode id="1D80A4F451343C3DE24EF9554EE66986" box="[1074,1164,1734,1758]" pageId="24" pageNumber="175">BMNH</collectionCode>
).
<typeStatus id="A42A829351343C3DE2DCF9554F786986" box="[1184,1298,1734,1758]" pageId="24" pageNumber="175" type="paratype">Paratypes</typeStatus>
: 8 examples same data as
<typeStatus id="A42A829351343C3DE731F97A4BD9685A" box="[333,435,1769,1794]" pageId="24" pageNumber="175" type="holotype">holotype</typeStatus>
; 1♀, Ankarafantsika N.P., AKF/05/2011, sifting forest litter, Winkler app. extr., S.16˚1805.1” E.46˚4908.3”,
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,
<date id="0F2F1AF151343C3DE44CF89E48CD687E" box="[560,679,1805,1830]" pageId="24" pageNumber="175" value="2011-04-23">23.iv.2011</date>
, L.S.Rahanitriniaina &amp; R. Raveloson (
<collectionCode id="1D80A4F451343C3DE22AF89D4EDA687E" box="[1110,1200,1806,1830]" pageId="24" pageNumber="175">BMNH</collectionCode>
); 1♀, Ambohitantely Spec. Res., ABT/12/2011, sifting plant residues under palm tree, Winkler app. extr., S.18˚1148.9” E.47˚1710.5”,
<quantity id="BC6991D451343C3DE6EBF8C64A9A6835" box="[151,240,1877,1902]" metricMagnitude="3" metricUnit="m" metricValue="1.6019999999999999" pageId="24" pageNumber="175" unit="m" value="1602.0">1602 m</quantity>
,
<date id="0F2F1AF151343C3DE682F8C64B1F6836" box="[254,373,1877,1902]" pageId="24" pageNumber="175" value="2011-04-19">19.iv.2011</date>
, L.S.Rhanitriniaina &amp; R. Raveloson (
<collectionCode id="1D80A4F451343C3DE558F8C549E36836" box="[804,905,1878,1902]" pageId="24" pageNumber="175">BMNH</collectionCode>
);1♀, Montagne dAmbre,
<date id="0F2F1AF151343C3DE2D0F8C54E966836" box="[1196,1276,1878,1902]" pageId="24" pageNumber="175" value="1970-10">x.1970</date>
, P. Hammond (
<collectionCode id="1D80A4F451343C3DE6E3F8E94A9368CA" box="[159,249,1914,1938]" pageId="24" pageNumber="175">BMNH</collectionCode>
);
<specimenCount id="6D97F7B851343C3DE770F8E94B4568CA" box="[268,303,1914,1938]" pageId="24" pageNumber="175" type="generic">4♂</specimenCount>
9♀, Mt. dAmbre, Mt. du Petit Lac,
<quantity id="BC6991D451343C3DE4B5F8E9490E68C9" box="[713,868,1914,1938]" metricMagnitude="3" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="24" pageNumber="175" unit="m" value="1200.0" valueMax="1300.0" valueMin="1100.0">11001300 m</quantity>
, forest,
<date id="0F2F1AF151343C3DE5C1F8EA4E4768CA" box="[957,1069,1913,1938]" pageId="24" pageNumber="175" value="1969-05-21">21.v.1969</date>
, leg H.Franz (MM); 2♀, Perinet,
<date id="0F2F1AF151343C3DE6EBF80E4B7A68EE" box="[151,272,1949,1974]" pageId="24" pageNumber="175" value="1969-04-15">15.iv.1969</date>
, forest of transition,
<quantity id="BC6991D451343C3DE784F80D485068EE" box="[504,570,1950,1974]" metricMagnitude="2" metricUnit="m" metricValue="9.28" pageId="24" pageNumber="175" unit="m" value="928.0">928m</quantity>
, leg. H.Franz (MM).
</paragraph>
</subSubSection>
</treatment>
</document>