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<document id="6D16E12EF0B0B92DD3ECF5F7CA7F303C" ID-DOI="10.5281/zenodo.185959" ID-GBIF-Dataset="813a57fd-0abc-4ebf-ab00-4c71ec46f634" ID-ISSN="1175-5326" ID-Zenodo-Dep="185959" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460031728588" checkinUser="plazi" docAuthor="OLoughlin, Mark, Manjón-Cabeza, Eugenia &amp; Ruiz, Francina Moya" docDate="2009" docId="F31587C27950FF979FDEFF221343151B" docLanguage="en" docName="zt02016p016.pdf" docOrigin="Zootaxa 2016" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Crucella scotiae Vaney, syn. nov." docType="treatment" docVersion="9" lastPageNumber="10" masterDocId="0F2CFFBA7959FF9E9F49FFC3113E1167" masterDocTitle="Antarctic holothuroids from the Bellingshausen Sea, with descriptions of new species (Echinodermata: Holothuroidea)" masterLastPageNumber="16" masterPageNumber="1" pageNumber="10" updateTime="1698234235397" updateUser="plazi">
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<mods:title id="6E32F1C834B5C945B072A44EB0944CA3">Antarctic holothuroids from the Bellingshausen Sea, with descriptions of new species (Echinodermata: Holothuroidea)</mods:title>
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<mods:namePart id="2F0FAF16962DD9BF714A5B1A1BBAACA1">OLoughlin, Mark</mods:namePart>
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<mods:namePart id="72F6357C0322AE1E4786EF7F74C30CB7">Manjón-Cabeza, Eugenia</mods:namePart>
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<mods:namePart id="5C6DAABC5AD6F65D15F869D5D9449C78">Ruiz, Francina Moya</mods:namePart>
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<mods:date id="8E91CA8B2460456184987EB70F28B0C3">2009</mods:date>
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<treatment id="F31587C27950FF979FDEFF221343151B" ID-DOI="http://doi.org/10.5281/zenodo.6219742" ID-GBIF-Taxon="119382495" ID-Zenodo-Dep="6219742" LSID="urn:lsid:plazi:treatment:F31587C27950FF979FDEFF221343151B" httpUri="http://treatment.plazi.org/id/F31587C27950FF979FDEFF221343151B" lastPageNumber="10" pageId="9" pageNumber="10">
<subSubSection id="33A6655F7950FF979FDEFF2211CB1079" pageId="9" pageNumber="10" type="nomenclature">
<paragraph id="7B0336D47950FF979FDEFF2213F4119D" blockId="9.[151,714,224,286]" box="[151,714,224,251]" pageId="9" pageNumber="10">
<heading id="204B81B87950FF979FDEFF2213F4119D" bold="true" box="[151,714,224,251]" fontSize="11" level="1" pageId="9" pageNumber="10" reason="1">
<emphasis id="49C8EAC67950FF979FDEFF2213F4119D" bold="true" box="[151,714,224,251]" pageId="9" pageNumber="10">
<taxonomicName id="BCBC4D577950FF979FDEFF2210FF119D" ID-CoL="ZRMN" authority="Vaney" authorityName="Vaney" box="[151,449,224,251]" class="Holothuroidea" family="Paracucumidae" genus="Crucella" kingdom="Animalia" order="Dendrochirotida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="species" species="scotiae">
<emphasis id="49C8EAC67950FF979FDEFF221067119C" bold="true" box="[151,345,225,251]" italics="true" pageId="9" pageNumber="10">Crucella scotiae</emphasis>
(Vaney)
</taxonomicName>
<taxonomicNameLabel id="52FB57BD7950FF979E80FF231308119D" box="[457,566,224,250]" pageId="9" pageNumber="10">syn. nov.</taxonomicNameLabel>
, comb. nov.
</emphasis>
</heading>
</paragraph>
<paragraph id="7B0336D47950FF979FDEFEC711CB1079" blockId="9.[151,714,224,286]" box="[151,245,260,286]" pageId="9" pageNumber="10">Table 1.</paragraph>
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<subSubSection id="33A6655F7950FF979FDEFE9212B210AF" pageId="9" pageNumber="10" type="reference_group">
<paragraph id="7B0336D47950FF979FDEFE92124110EF" blockId="9.[151,1160,337,456]" pageId="9" pageNumber="10">
<treatmentCitationGroup id="5BAC11FA7950FF979FDEFE92124110EF" pageId="9" pageNumber="10">
<taxonomicName id="BCBC4D577950FF979FDEFE921339100F" authority="Vaney, 1906a: 400" authorityName="Vaney" authorityPageNumber="400" authorityYear="1906" box="[151,519,337,360]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="species" species="scotiae">
<emphasis id="49C8EAC67950FF979FDEFE921009100F" box="[151,311,337,360]" italics="true" pageId="9" pageNumber="10">Thyone scotiae</emphasis>
<treatmentCitation id="FA1D10C57950FF979E77FE921339100F" author="Vaney" box="[318,519,337,360]" page="400" pageId="9" pageNumber="10" year="1906">
<bibRefCitation id="1F2D4B257950FF979E77FE9210EF100F" author="Vaney" box="[318,465,337,360]" pageId="9" pageNumber="16" refString="Vaney, C. (1906 a) Deux nouvelles holothuries du genre Thyone provenant des Orcades du Sud. Bulletin du Museum d'Histoire naturelle, Paris, 12, 400 - 402." type="journal article" year="1906" yearSuffix="a">Vaney, 1906a</bibRefCitation>
: 400
</treatmentCitation>
</taxonomicName>
-401, fig. 1.—
<treatmentCitation id="FA1D10C57950FF979DDCFE921264100F" author="Ekman" box="[661,858,337,360]" page="106" pageId="9" pageNumber="10" year="1925">
<bibRefCitation id="1F2D4B257950FF979DDCFE921219100F" author="Ekman" box="[661,807,337,360]" pageId="9" pageNumber="14" refString="Ekman, S. (1925) Holothurien. Further zoological results of the Swedish Antarctic Expedition 1901 - 1903, 1 (6), 1 - 194." type="journal article" year="1925">Ekman 1925</bibRefCitation>
: 106
</treatmentCitation>
108, fig. 24.
<taxonomicName id="BCBC4D577950FF979FDEFEB210BE10EF" box="[151,384,369,392]" class="Holothuroidea" family="Ypsilothuriidae" genus="Ypsilocucumis" kingdom="Animalia" order="Dendrochirotida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="species" species="scotiae">
<emphasis id="49C8EAC67950FF979FDEFEB210BE10EF" box="[151,384,369,392]" italics="true" pageId="9" pageNumber="10">Ypsilocucumis scotiae</emphasis>
</taxonomicName>
.—
<treatmentCitation id="FA1D10C57950FF979EE9FEB2134D10EF" author="Panning" box="[416,627,369,392]" page="455" pageId="9" pageNumber="10" year="1949">
<bibRefCitation id="1F2D4B257950FF979EE9FEB2130110EF" author="Panning" box="[416,575,369,392]" pageId="9" pageNumber="15" refString="Panning, A. (1949) Versuch einer Neuordnung der Familie Cucumariidae (Holothurioidea, Dendrochirota). Zoologische Jahrbucher Abtheilung fur Systematik, Geographie und Biologie der Thiere, 78, 404 - 470." type="journal article" year="1949">Panning, 1949</bibRefCitation>
: 455
</treatmentCitation>
.—
<bibRefCitation id="1F2D4B257950FF979DDBFEB2121210EF" author="Pawson" box="[658,812,369,392]" pageId="9" pageNumber="15" refString="Pawson, D. L. (1969) Holothuroidea. In: Bushnell, V. C. &amp; J. W. Hedgpeth, J. W. (Eds.), Distribution of Selected Groups of Marine Invertebrates in Waters South of 35 º S Latitude. Antarctic Map Folio Series (II). American Geographical Society, New York, pp. 36 - 38, 1 pl." type="journal article" year="1969">Pawson, 1969</bibRefCitation>
: map 3.
</treatmentCitationGroup>
</paragraph>
<paragraph id="7B0336D47950FF979FDEFE5212B210AF" blockId="9.[151,1160,337,456]" pageId="9" pageNumber="10">
<treatmentCitationGroup id="5BAC11FA7950FF979FDEFE5212B210AF" pageId="9" pageNumber="10">
<taxonomicName id="BCBC4D577950FF979FDEFE52130610CF" ID-CoL="PB45" authority="Gutt, 1990: 105" authorityName="Gutt" authorityPageNumber="105" authorityYear="1990" box="[151,568,401,424]" class="Holothuroidea" family="Paracucumidae" genus="Caespitugo" higherTaxonomySource="CoL" kingdom="Animalia" order="Dendrochirotida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="species" species="citriformis">
<emphasis id="49C8EAC67950FF979FDEFE5210B910CF" box="[151,391,401,424]" italics="true" pageId="9" pageNumber="10">Caespitugo citriformis</emphasis>
<treatmentCitation id="FA1D10C57950FF979EC7FE52130610CF" author="Gutt" box="[398,568,401,424]" page="105" pageId="9" pageNumber="10" year="1990">
<bibRefCitation id="1F2D4B257950FF979EC7FE52133A10CF" author="Gutt" box="[398,516,401,424]" pageId="9" pageNumber="15" refString="Gutt, J. (1990) New Antarctic holothurians (Echinodermata) - I. Five new species with four new genera of the order Dendrochirotida. Zoologica Scripta, 19 (1), 119 - 127." type="journal article" year="1990">Gutt, 1990</bibRefCitation>
: 105
</treatmentCitation>
</taxonomicName>
, figs 1, 57, table 2.—
<treatmentCitation id="FA1D10C57950FF979C63FE5212ED10CF" author="Gutt" box="[810,979,401,424]" page="324" pageId="9" pageNumber="10" year="1991">
<bibRefCitation id="1F2D4B257950FF979C63FE52129F10CF" author="Gutt" box="[810,929,401,424]" pageId="9" pageNumber="15" refString="Gutt, J. (1991) Are Weddell Sea holothurians typical representatives of the Antarctic benthos? Meeresforschung, 33 (4), 312 - 329." type="journal article" year="1991">Gutt, 1991</bibRefCitation>
: 324
</treatmentCitation>
(new synonym).
<taxonomicName id="BCBC4D577950FF979FDEFE7210E310AF" ID-CoL="ZRMM" authority="Gutt, 1990" authorityName="Gutt" authorityYear="1990" box="[151,477,433,456]" class="Holothuroidea" family="Paracucumidae" genus="Crucella" kingdom="Animalia" order="Dendrochirotida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="species" species="hystrix">
<emphasis id="49C8EAC67950FF979FDEFE7211CA10AF" box="[151,244,433,456]" italics="true" pageId="9" pageNumber="10">Crucella</emphasis>
cf.
<emphasis id="49C8EAC67950FF979E54FE72105B10AF" box="[285,357,433,456]" italics="true" pageId="9" pageNumber="10">hystrix</emphasis>
<bibRefCitation id="1F2D4B257950FF979E25FE7210E310AF" author="Gutt" box="[364,477,433,456]" pageId="9" pageNumber="15" refString="Gutt, J. (1990) New Antarctic holothurians (Echinodermata) - I. Five new species with four new genera of the order Dendrochirotida. Zoologica Scripta, 19 (1), 119 - 127." type="journal article" year="1990">Gutt, 1990</bibRefCitation>
</taxonomicName>
.—
<treatmentCitation id="FA1D10C57950FF979EB4FE72120D10AF" author="O'Loughlin" box="[509,819,433,456]" page="552" pageId="9" pageNumber="10" year="1994">
<bibRefCitation id="1F2D4B257950FF979EB4FE7213C010AF" author="O'Loughlin" box="[509,766,433,456]" pageId="9" pageNumber="15" refString="O'Loughlin, P. M., Bardsley, T. M. &amp; O'Hara, T. D. (1994) A preliminary analysis of diversity and distribution of Holothurioidea from Prydz Bay and the MacRobertson Shelf, eastern Antarctica. In: David, B., Guille, A., Feral, J- P. &amp; Roux, M. (Eds.), Echinoderms through Time. Proceedings of the Eighth International Echinoderm Conference, Dijon, France, 6 - 10 September, 1993. Balkema: Rotterdam, pp. 549 - 555, 1 fig., 2 tables." type="book chapter" year="1994">OLoughlin et al., 1994</bibRefCitation>
: 552
</treatmentCitation>
, table 2.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="33A6655F7950FF979FDEFE341343151B" pageId="9" pageNumber="10" type="discussion">
<paragraph id="7B0336D47950FF979FDEFE341343151B" blockId="9.[151,1437,503,1148]" pageId="9" pageNumber="10">
<emphasis id="49C8EAC67950FF979FDEFE34102A1376" bold="true" box="[151,276,503,529]" pageId="9" pageNumber="10">Remarks.</emphasis>
<taxonomicName id="BCBC4D577950FF979E55FE3B134C1375" authority="Vaney, 1906" authorityName="Vaney" authorityYear="1906" box="[284,626,504,530]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="species" species="scotiae">
<emphasis id="49C8EAC67950FF979E55FE3B10EA1376" box="[284,468,504,529]" italics="true" pageId="9" pageNumber="10">Thyone scotiae</emphasis>
<bibRefCitation id="1F2D4B257950FF979E95FE3B134C1375" author="Vaney" box="[476,626,504,530]" pageId="9" pageNumber="16" refString="Vaney, C. (1906 a) Deux nouvelles holothuries du genre Thyone provenant des Orcades du Sud. Bulletin du Museum d'Histoire naturelle, Paris, 12, 400 - 402." type="journal article" year="1906" yearSuffix="a">Vaney, 1906</bibRefCitation>
</taxonomicName>
a has
<bibRefCitation id="1F2D4B257950FF979DF3FE3B13F91375" author="Vaney" box="[698,711,504,530]" pageId="9" pageNumber="16" refString="Vaney, C. (1906 a) Deux nouvelles holothuries du genre Thyone provenant des Orcades du Sud. Bulletin du Museum d'Histoire naturelle, Paris, 12, 400 - 402." type="journal article" year="1906" yearSuffix="a">a</bibRefCitation>
circum-polar distribution (M. OLoughlin, pers. obs.). The distinguishing characters of
<taxonomicName id="BCBC4D577950FF979EA4FDDC127E135F" authority="Vaney, 1906" authorityName="Vaney" authorityYear="1906" box="[493,832,542,568]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="species" species="scotiae">
<emphasis id="49C8EAC67950FF979EA4FDDC139C135F" box="[493,674,543,568]" italics="true" pageId="9" pageNumber="10">Thyone scotiae</emphasis>
<bibRefCitation id="1F2D4B257950FF979DE2FDDD127E135F" author="Vaney" box="[683,832,542,568]" pageId="9" pageNumber="16" refString="Vaney, C. (1906 a) Deux nouvelles holothuries du genre Thyone provenant des Orcades du Sud. Bulletin du Museum d'Histoire naturelle, Paris, 12, 400 - 402." type="journal article" year="1906" yearSuffix="a">Vaney, 1906</bibRefCitation>
</taxonomicName>
a and
<taxonomicName id="BCBC4D577950FF979CC0FDDC141D135F" authority="Gutt, 1990" authorityName="Gutt" authorityYear="1990" box="[905,1315,542,568]" class="Holothuroidea" family="Paracucumidae" genus="Caespitugo" higherTaxonomySource="CoL" kingdom="Animalia" order="Dendrochirotida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="species" species="citriformis">
<emphasis id="49C8EAC67950FF979CC0FDDC15A7135F" box="[905,1177,543,568]" italics="true" pageId="9" pageNumber="10">Caespitugo citriformis</emphasis>
<bibRefCitation id="1F2D4B257950FF979BE8FDDD141D135F" author="Gutt" box="[1185,1315,542,568]" pageId="9" pageNumber="15" refString="Gutt, J. (1990) New Antarctic holothurians (Echinodermata) - I. Five new species with four new genera of the order Dendrochirotida. Zoologica Scripta, 19 (1), 119 - 127." type="journal article" year="1990">Gutt, 1990</bibRefCitation>
</taxonomicName>
are: body with narrowed tail; hard to parchment-like body wall; 10 tentacles with ventral pair small; cucumariid not phyllophorid calcareous ring; tube feet evenly distributed around body; body wall ossicles small knobbed plates, large irregular thick knobbed perforated plates with central secondary mesh thickening; posterior body with multi-layered ossicles.
<taxonomicName id="BCBC4D577950FF979EBFFD7A12A513B4" authority="Gutt, 1990" authorityName="Gutt" authorityYear="1990" box="[502,923,697,723]" class="Holothuroidea" family="Paracucumidae" genus="Caespitugo" higherTaxonomySource="CoL" kingdom="Animalia" order="Dendrochirotida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="species" species="citriformis">
<emphasis id="49C8EAC67950FF979EBFFD7A123013B5" box="[502,782,697,722]" italics="true" pageId="9" pageNumber="10">Caespitugo citriformis</emphasis>
<bibRefCitation id="1F2D4B257950FF979C5FFD7A12A513B4" author="Gutt" box="[790,923,697,723]" pageId="9" pageNumber="15" refString="Gutt, J. (1990) New Antarctic holothurians (Echinodermata) - I. Five new species with four new genera of the order Dendrochirotida. Zoologica Scripta, 19 (1), 119 - 127." type="journal article" year="1990">Gutt, 1990</bibRefCitation>
</taxonomicName>
is judged here to be a junior synonym of
<taxonomicName id="BCBC4D577950FF979FDEFD2310D1139D" authority="Vaney, 1906" authorityName="Vaney" authorityYear="1906" box="[151,495,736,762]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="species" species="scotiae">
<emphasis id="49C8EAC67950FF979FDEFD231070139E" box="[151,334,736,761]" italics="true" pageId="9" pageNumber="10">Thyone scotiae</emphasis>
<bibRefCitation id="1F2D4B257950FF979E1FFD2310D1139D" author="Vaney" box="[342,495,736,762]" pageId="9" pageNumber="16" refString="Vaney, C. (1906 a) Deux nouvelles holothuries du genre Thyone provenant des Orcades du Sud. Bulletin du Museum d'Histoire naturelle, Paris, 12, 400 - 402." type="journal article" year="1906" yearSuffix="a">Vaney, 1906</bibRefCitation>
</taxonomicName>
a.
<taxonomicName id="BCBC4D577950FF979D40FD231395139D" authority="Oken" authorityName="Oken" box="[521,683,736,762]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="genus">
<emphasis id="49C8EAC67950FF979D40FD23135C139E" box="[521,610,736,761]" italics="true" pageId="9" pageNumber="10">Thyone</emphasis>
Oken
</taxonomicName>
is an inappropriate genus for
<taxonomicName id="BCBC4D577950FF979B51FD23141A139D" authority="Vaney" authorityName="Vaney" box="[1048,1316,736,762]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="species" species="scotiae">
<emphasis id="49C8EAC67950FF979B51FD2315F1139E" box="[1048,1231,736,761]" italics="true" pageId="9" pageNumber="10">Thyone scotiae</emphasis>
Vaney
</taxonomicName>
, since the calcareous rings of
<taxonomicName id="BCBC4D577950FF979ECFFCC410E11247" box="[390,479,775,800]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="genus">
<emphasis id="49C8EAC67950FF979ECFFCC410E11247" box="[390,479,775,800]" italics="true" pageId="9" pageNumber="10">Thyone</emphasis>
</taxonomicName>
species have elongate composite posterior projections characteristic of family
<taxonomicName id="BCBC4D577950FF979FDEFCEE106E1220" box="[151,336,813,839]" class="Holothuroidea" family="Phyllophoridae" kingdom="Animalia" order="Dendrochirotida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="family">Phyllophoridae</taxonomicName>
.
<bibRefCitation id="1F2D4B257950FF979E12FCEE132D1220" author="Panning" box="[347,531,813,839]" pageId="9" pageNumber="15" refString="Panning, A. (1949) Versuch einer Neuordnung der Familie Cucumariidae (Holothurioidea, Dendrochirota). Zoologische Jahrbucher Abtheilung fur Systematik, Geographie und Biologie der Thiere, 78, 404 - 470." type="journal article" year="1949">Panning (1949)</bibRefCitation>
inappropriately referred
<taxonomicName id="BCBC4D577950FF979C75FCEE12951221" box="[828,939,813,838]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="species" species="scotiae">
<emphasis id="49C8EAC67950FF979C75FCEE12951221" box="[828,939,813,838]" italics="true" pageId="9" pageNumber="10">T. scotiae</emphasis>
</taxonomicName>
to his new genus
<taxonomicName id="BCBC4D577950FF979BCCFCEE142A1221" box="[1157,1300,813,838]" class="Holothuroidea" family="Ypsilothuriidae" genus="Ypsilothuria" kingdom="Animalia" order="Dendrochirotida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="genus">
<emphasis id="49C8EAC67950FF979BCCFCEE142A1221" box="[1157,1300,813,838]" italics="true" pageId="9" pageNumber="10">Ypsilothuria</emphasis>
</taxonomicName>
, diagnosed as having 10 equal tentacles.
<taxonomicName id="BCBC4D577950FF979EA4FC9713D31209" authority="Vaney" authorityName="Vaney" box="[493,749,852,878]" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="species" species="scotiae">
<emphasis id="49C8EAC67950FF979EA4FC9713A0120A" box="[493,670,852,877]" italics="true" pageId="9" pageNumber="10">Thyone scotiae</emphasis>
Vaney
</taxonomicName>
is referred here to
<taxonomicName id="BCBC4D577950FF979C82FC9715881209" authority="Gutt, 1990" authorityName="Gutt" authorityYear="1990" box="[971,1206,852,878]" class="Holothuroidea" family="Paracucumidae" genus="Crucella" kingdom="Animalia" order="Dendrochirotida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="genus">
<emphasis id="49C8EAC67950FF979C82FC97150F120A" box="[971,1073,852,877]" italics="true" pageId="9" pageNumber="10">Crucella</emphasis>
<bibRefCitation id="1F2D4B257950FF979B71FC9715881209" author="Gutt" box="[1080,1206,852,878]" pageId="9" pageNumber="15" refString="Gutt, J. (1990) New Antarctic holothurians (Echinodermata) - I. Five new species with four new genera of the order Dendrochirotida. Zoologica Scripta, 19 (1), 119 - 127." type="journal article" year="1990">Gutt, 1990</bibRefCitation>
</taxonomicName>
that has: body with narrowed tail; hard to parchment-like body wall; 10 tentacles with ventral pair small; cucumariid not phyllophorid calcareous ring; tube feet evenly distributed around body; body wall ossicles crosses with four perforated arms, with or without central secondary mesh thickening and spine.
<bibRefCitation id="1F2D4B257950FF979B0CFC0B15EE1285" author="Gutt" box="[1093,1232,968,994]" pageId="9" pageNumber="15" refString="Gutt, J. (1990) New Antarctic holothurians (Echinodermata) - I. Five new species with four new genera of the order Dendrochirotida. Zoologica Scripta, 19 (1), 119 - 127." type="journal article" year="1990">Gutt (1990)</bibRefCitation>
referred his now junior synonym species
<taxonomicName id="BCBC4D577950FF979EF8FC2C1371156F" box="[433,591,1007,1032]" class="Holothuroidea" family="Paracucumidae" genus="Caespitugo" higherTaxonomySource="CoL" kingdom="Animalia" order="Dendrochirotida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="species" species="citriformis">
<emphasis id="49C8EAC67950FF979EF8FC2C1371156F" box="[433,591,1007,1032]" italics="true" pageId="9" pageNumber="10">C. citriformis</emphasis>
</taxonomicName>
to his genus
<taxonomicName id="BCBC4D577950FF979DAEFC2C1252156F" box="[743,876,1007,1032]" class="Holothuroidea" family="Paracucumidae" genus="Caespitugo" higherTaxonomySource="CoL" kingdom="Animalia" order="Dendrochirotida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="genus">
<emphasis id="49C8EAC67950FF979DAEFC2C1252156F" box="[743,876,1007,1032]" italics="true" pageId="9" pageNumber="10">Caespitugo</emphasis>
</taxonomicName>
, but it is judged here that the senior synonym
<taxonomicName id="BCBC4D577950FF979ACFFC2C11D51549" class="Holothuroidea" family="Phyllophoridae" genus="Thyone" kingdom="Animalia" order="Dendrochirotida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="species" species="scotiae">
<emphasis id="49C8EAC67950FF979ACFFC2C11D51549" italics="true" pageId="9" pageNumber="10">T. scotiae</emphasis>
</taxonomicName>
is more appropriately referred to
<taxonomicName id="BCBC4D577950FF979DCDFBD612481548" authority="Gutt, 1990" authorityName="Gutt" authorityYear="1990" box="[644,886,1045,1071]" class="Holothuroidea" family="Paracucumidae" genus="Crucella" kingdom="Animalia" order="Dendrochirotida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="genus">
<emphasis id="49C8EAC67950FF979DCDFBD613D21549" box="[644,748,1045,1070]" italics="true" pageId="9" pageNumber="10">Crucella</emphasis>
<bibRefCitation id="1F2D4B257950FF979DBDFBD612481548" author="Gutt" box="[756,886,1045,1071]" pageId="9" pageNumber="15" refString="Gutt, J. (1990) New Antarctic holothurians (Echinodermata) - I. Five new species with four new genera of the order Dendrochirotida. Zoologica Scripta, 19 (1), 119 - 127." type="journal article" year="1990">Gutt, 1990</bibRefCitation>
</taxonomicName>
because of the multi-layering in the centre of some perforated plates, and some plates cross-shaped or with irregular arms.
<taxonomicName id="BCBC4D577950FF979B6AFBFF14131531" authority="Gutt, 1990" authorityName="Gutt" authorityYear="1990" box="[1059,1325,1084,1110]" class="Holothuroidea" family="Paracucumidae" genus="Caespitugo" higherTaxonomySource="CoL" kingdom="Animalia" order="Dendrochirotida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="genus">
<emphasis id="49C8EAC67950FF979B6AFBFF15961532" box="[1059,1192,1084,1109]" italics="true" pageId="9" pageNumber="10">Caespitugo</emphasis>
<bibRefCitation id="1F2D4B257950FF979BF9FBFF14131531" author="Gutt" box="[1200,1325,1084,1110]" pageId="9" pageNumber="15" refString="Gutt, J. (1990) New Antarctic holothurians (Echinodermata) - I. Five new species with four new genera of the order Dendrochirotida. Zoologica Scripta, 19 (1), 119 - 127." type="journal article" year="1990">Gutt, 1990</bibRefCitation>
</taxonomicName>
becomes a junior synonym of
<taxonomicName id="BCBC4D577950FF979EC2FBA01347151B" authority="Gutt, 1990" authorityName="Gutt" authorityYear="1990" box="[395,633,1122,1148]" class="Holothuroidea" family="Paracucumidae" genus="Crucella" kingdom="Animalia" order="Dendrochirotida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="genus">
<emphasis id="49C8EAC67950FF979EC2FBA010CC151B" box="[395,498,1123,1148]" italics="true" pageId="9" pageNumber="10">Crucella</emphasis>
<bibRefCitation id="1F2D4B257950FF979EB1FBA11347151B" author="Gutt" box="[504,633,1122,1148]" pageId="9" pageNumber="15" refString="Gutt, J. (1990) New Antarctic holothurians (Echinodermata) - I. Five new species with four new genera of the order Dendrochirotida. Zoologica Scripta, 19 (1), 119 - 127." type="journal article" year="1990">Gutt, 1990</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>