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<mods:title id="033F9E39D52AF7A4B1DC7F00560CF88D">The sand shrimp genus Philocheras (Caridea: Crangonidae) from the continental margin of Western Australia including the description of a new species and a key to Australian species *</mods:title>
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<heading id="1E4D818AB1131E63FF44F5E1FDB02BA4" bold="true" box="[151,611,1730,1756]" fontSize="11" level="1" pageId="6" pageNumber="163" reason="1">
<taxonomicName id="82BA4D65B1131E63FF44F5E1FDB02BA4" ID-CoL="9D3KY" authority="(Doflein, 1902)" baseAuthorityName="Doflein" baseAuthorityYear="1902" box="[151,611,1730,1756]" class="Malacostraca" family="Crangonidae" genus="Philocheras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="163" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="77CEEAF4B1131E63FF44F5E1FDB02BA4" bold="true" box="[151,611,1730,1756]" pageId="6" pageNumber="163">
<emphasis id="77CEEAF4B1131E63FF44F5E1FE772BA4" bold="true" box="[151,420,1730,1756]" italics="true" pageId="6" pageNumber="163">Philocheras japonicus</emphasis>
(
<bibRefCitation id="212B4B17B1131E63FE60F5E1FD882BA4" author="Doflein, F." box="[435,603,1730,1756]" pageId="6" pageNumber="163" pagination="613 - 670" refId="ref6627" refString="Doflein, F. (1902) Ostasiatische Dekapoden. Abhandlungen der Bayerischen Akademie der Wissenschaften, Munchen, 21, 613 - 670, 6 pls." type="journal article" year="1902">Doflein, 1902</bibRefCitation>
)
</emphasis>
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<paragraph id="450536E6B1131E63FF44F5C6FF3C2B87" blockId="6.[151,611,1730,1791]" box="[151,239,1765,1791]" pageId="6" pageNumber="163">
(
<figureCitation id="DD812A63B1131E63FF73F5C6FF362B87" box="[160,229,1765,1791]" captionStart="FIGURE 3" captionStartId="7.[151,255,1886,1910]" captionTargetBox="[242,1349,155,1858]" captionTargetId="figure-100@7.[200,1369,155,1864]" captionTargetPageId="7" captionText="FIGURE 3. Carapace (dorsal view) of Philocheras species reported from Australian waters: P. angustirostris redrawn from De Man (1918: pl. 22 fig. 67); P. anthonyi reproduced from type illustration herein (fig. 1B); P. brucei redrawn from Komai (2004: fig. 1); P. flindersi redrawn from Poore (2004: fig. 36g); P. incisus redrawn from Kemp (1916: pl. 8 fig. 1); P. intermedius redrawn from Poore (2004: fig. 36h); P.japonicus redrawn from De Man (1918: pl. 23 fig. 69); P. lowisi redrawn from Kemp (1916: pl. 8 fig. 2)." figureDoi="http://doi.org/10.5281/zenodo.5305097" httpUri="https://zenodo.org/record/5305097/files/figure.png" pageId="6" pageNumber="163">Fig. 3</figureCitation>
)
</paragraph>
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<subSubSection id="0DA0656DB1131E63FF44F413FDB02AFF" pageId="6" pageNumber="163" type="reference_group">
<paragraph id="450536E6B1131E63FF44F413FCC92A1F" blockId="6.[151,1436,1840,1927]" pageId="6" pageNumber="163">
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<treatmentCitation id="C41B10F7B1131E63FF44F413FD8E2A3F" author="Doflein, F." box="[151,605,1840,1863]" page="621" pageId="6" pageNumber="163" year="1902">
<taxonomicName id="82BA4D65B1131E63FF44F413FD8E2A3F" ID-CoL="9D59M" authority="Doflein, 1902: 621" authorityName="Doflein" authorityPageNumber="621" authorityYear="1902" box="[151,605,1840,1863]" class="Malacostraca" family="Crangonidae" genus="Pontophilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="163" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="77CEEAF4B1131E63FF44F413FE542A3F" box="[151,391,1840,1863]" italics="true" pageId="6" pageNumber="163">Pontophilus japonicus</emphasis>
<bibRefCitation id="212B4B17B1131E63FE5CF413FD8E2A3F" author="Doflein, F." box="[399,605,1840,1863]" pageId="6" pageNumber="163" pagination="613 - 670" refId="ref6627" refString="Doflein, F. (1902) Ostasiatische Dekapoden. Abhandlungen der Bayerischen Akademie der Wissenschaften, Munchen, 21, 613 - 670, 6 pls." type="journal article" year="1902">Doflein, 1902: 621</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 642, fig. B, pl. 3: fig. 6. — De
<treatmentCitation id="C41B10F7B1131E63FC6FF413FBB02A3F" author="Man, J. G. de" box="[956,1123,1840,1863]" page="286" pageId="6" pageNumber="163" year="1920">
<bibRefCitation id="212B4B17B1131E63FC6FF413FBB02A3F" author="Man, J. G. de" box="[956,1123,1840,1863]" pageId="6" pageNumber="163" pagination="1 - 318" refId="ref7083" refString="Man, J. G. de (1920) Pasiphaeidae, Stylodactylidae, Hoplophoridae, Nematocarcinidae, Thalassocaridae, Pandalidae, Psalidopodidae, Gnathophyllidae, Processidae, Glyphocranconidae and Crangonidae. Siboga Expeditie, 39 a (3), 1 - 318, pls 1 - 25." type="journal article" year="1920">Man 1920: 286</bibRefCitation>
</treatmentCitation>
, pl. 23: fig. 69ac, e, gi, pl. 24: fig 69d, f, j. —
<treatmentCitation id="C41B10F7B1131E63FE46F473FD112A1F" author="Fujino, T. &amp; Miyake, S." box="[405,706,1872,1895]" page="290" pageId="6" pageNumber="163" year="1970">
<bibRefCitation id="212B4B17B1131E63FE46F473FD112A1F" author="Fujino, T. &amp; Miyake, S." box="[405,706,1872,1895]" pageId="6" pageNumber="163" pagination="237 - 312" refId="ref6655" refString="Fujino, T. &amp; Miyake, S. (1970) Caridean and stenopodidean shrimps from the East China and the Yellow Seas (Crustacea, Decapoda, Natantia). Journal of the Faculty of Agriculture, Kyushu University, 16, 237 - 312." type="journal article" year="1970">Fujino &amp; Miyake 1970: 290</bibRefCitation>
</treatmentCitation>
, fig. 19.
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</paragraph>
<paragraph id="450536E6B1131E63FF44F453FDB02AFF" blockId="6.[151,1436,1840,1927]" box="[151,611,1904,1927]" pageId="6" pageNumber="163">
<treatmentCitationGroup id="65AA11C8B1131E63FF44F453FDB02AFF" box="[151,611,1904,1927]" pageId="6" pageNumber="163">
<taxonomicName id="82BA4D65B1131E63FF44F453FE562AFF" ID-CoL="9D3KY" baseAuthorityName="Doflein" baseAuthorityYear="1902" box="[151,389,1904,1927]" class="Malacostraca" family="Crangonidae" genus="Philocheras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="163" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="77CEEAF4B1131E63FF44F453FE562AFF" box="[151,389,1904,1927]" italics="true" pageId="6" pageNumber="163">Philocheras japonicus</emphasis>
</taxonomicName>
. —
<treatmentCitation id="C41B10F7B1131E63FE66F453FD8C2AFF" author="Chace, F. A." box="[437,607,1904,1927]" page="40" pageId="6" pageNumber="163" year="1984">
<bibRefCitation id="212B4B17B1131E63FE66F453FD8C2AFF" author="Chace, F. A." box="[437,607,1904,1927]" pageId="6" pageNumber="163" pagination="1 - 63" refId="ref6509" refString="Chace, F. A. (1984) The caridean shrimps (Crustacea; Decapoda) of the Albatross Philippine Expedition, 1907 - 1910, part 2: families Glyphocrangonidae and Crangonidae. Smithsonian Contributions to Zoology, 397, 1 - 63." type="journal article" year="1984">Chace 1984: 40</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="0DA0656DB1131E6DFF44F496FBCA2C3E" lastPageId="8" lastPageNumber="165" pageId="6" pageNumber="163" type="materials_examined">
<paragraph id="450536E6B1131E63FF44F496FC3A2A8C" blockId="6.[151,1436,1973,2036]" pageId="6" pageNumber="163">
<emphasis id="77CEEAF4B1131E63FF44F496FE582AB7" bold="true" box="[151,395,1973,1999]" pageId="6" pageNumber="163">Material examined.</emphasis>
<materialsCitation id="F5D23CBBB1131E63FE47F496FC362A8C" accessionNumber="J40941" collectingDate="1983-02-18" collectionCode="NWA" country="Australia" elevation="42" location="Northwest Shelf" pageId="6" pageNumber="163" specimenCode="NWA 1" specimenCount="2">
<collectingCountry id="3DAD7676B1131E63FE47F496FDD12AB7" box="[404,514,1973,1999]" name="Australia" pageId="6" pageNumber="163">Australia</collectingCountry>
, WA,
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,
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(19°59.0059.4'S, 117°51.450.4'E),
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,
<date id="31041026B1131E63FE6FF4F9FDE52A8C" box="[444,566,2010,2036]" pageId="6" pageNumber="163" value="1983-02-18">
<collectingDate id="2140E9CEB1131E63FE6FF4F9FDE52A8C" box="[444,566,2010,2036]" pageId="6" pageNumber="163" value="1983-02-18">18.II.1983</collectingDate>
</date>
(stn
<specimenCode id="151C9E9DB1131E63FDA2F4F9FD1B2A8C" box="[625,712,2010,2036]" collectionCode="NWA" pageId="6" pageNumber="163">NWA 1</specimenCode>
),
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(
<specimenCount id="53BCFD6FB1131E63FC94F4F9FC0E2A8C" box="[839,989,2010,2036]" pageId="6" pageNumber="163" type="generic">2 specimens</specimenCount>
)
</materialsCitation>
.
</paragraph>
<caption id="11C5666EB1121E62FF44F47DFD5B2A8D" ID-DOI="http://doi.org/10.5281/zenodo.5305097" ID-Zenodo-Dep="5305097" httpUri="https://zenodo.org/record/5305097/files/figure.png" pageId="7" pageNumber="164" startId="7.[151,255,1886,1910]" targetBox="[242,1349,155,1858]" targetPageId="7">
<paragraph id="450536E6B1121E62FF44F47DFD5B2A8D" blockId="7.[151,1440,1886,2038]" pageId="7" pageNumber="164">
<emphasis id="77CEEAF4B1121E62FF44F47DFEC92A0E" bold="true" box="[151,282,1886,1910]" pageId="7" pageNumber="164">FIGURE 3.</emphasis>
Carapace (dorsal view) of
<taxonomicName id="82BA4D65B1121E62FD90F47CFCC92A0D" box="[579,794,1886,1910]" class="Malacostraca" family="Crangonidae" genus="Philocheras" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="164" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="77CEEAF4B1121E62FD90F47CFD162A0E" box="[579,709,1887,1910]" italics="true" pageId="7" pageNumber="164">Philocheras</emphasis>
species
</taxonomicName>
reported from Australian waters:
<taxonomicName id="82BA4D65B1121E62FB5FF47CFAEF2A0E" baseAuthorityName="De Man" baseAuthorityYear="1918" box="[1164,1340,1887,1910]" class="Malacostraca" family="Crangonidae" genus="Philocheras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="164" phylum="Arthropoda" rank="species" species="angustirostris">
<emphasis id="77CEEAF4B1121E62FB5FF47CFAEF2A0E" box="[1164,1340,1887,1910]" italics="true" pageId="7" pageNumber="164">P. angustirostris</emphasis>
</taxonomicName>
redrawn from De
<bibRefCitation id="212B4B17B1121E62FF28F45DFEAB2AED" author="Man, J. G. de" box="[251,376,1918,1941]" pageId="7" pageNumber="164" pagination="159 - 166" refId="ref7052" refString="Man, J. G. de (1918) Diagnosis of new species of macrurous decapod Crustacea from the Siboga-Expedition. Zoologische Mededeelingen, 4, 159 - 166." type="journal article" year="1918">Man (1918</bibRefCitation>
: pl. 22 fig. 67);
<taxonomicName id="82BA4D65B1121E62FDF9F45CFD772AEE" authorityName="Taylor" authorityYear="2010" box="[554,676,1919,1942]" class="Malacostraca" family="Crangonidae" genus="Philocheras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="164" phylum="Arthropoda" rank="species" species="anthonyi">
<emphasis id="77CEEAF4B1121E62FDF9F45CFD772AEE" box="[554,676,1919,1942]" italics="true" pageId="7" pageNumber="164">P. anthonyi</emphasis>
</taxonomicName>
reproduced from type illustration herein (fig. 1B);
<taxonomicName id="82BA4D65B1121E62FB09F45CFAE82AEE" authorityName="Komai" authorityYear="2004" box="[1242,1339,1919,1942]" class="Malacostraca" family="Crangonidae" genus="Philocheras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="164" phylum="Arthropoda" rank="species" species="brucei">
<emphasis id="77CEEAF4B1121E62FB09F45CFAE82AEE" box="[1242,1339,1919,1942]" italics="true" pageId="7" pageNumber="164">P. brucei</emphasis>
</taxonomicName>
redrawn from
<bibRefCitation id="212B4B17B1121E62FF01F4BDFEB02ACD" author="Komai, T." box="[210,355,1950,1973]" pageId="7" pageNumber="164" pagination="665 - 673" refId="ref6909" refString="Komai, T. (2004) A new species of the crangonid genus Philocheras S tebbing (Crustacea: Decapoda: Caridea) from northeastern Australia. Memoirs of the Queensland Museum, 49, 665 - 673." type="journal article" year="2004">Komai (2004</bibRefCitation>
: fig. 1);
<taxonomicName id="82BA4D65B1121E62FE6CF4BCFDE02ACE" baseAuthorityName="Fulton &amp; Grant" baseAuthorityYear="1902" box="[447,563,1951,1974]" class="Malacostraca" family="Crangonidae" genus="Philocheras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="164" phylum="Arthropoda" rank="species" species="flindersi">
<emphasis id="77CEEAF4B1121E62FE6CF4BCFDE02ACE" box="[447,563,1951,1974]" italics="true" pageId="7" pageNumber="164">P. flindersi</emphasis>
</taxonomicName>
redrawn from
<bibRefCitation id="212B4B17B1121E62FD07F4BDFC882ACD" author="Poore, G. C. B." box="[724,859,1950,1973]" pageId="7" pageNumber="164" refId="ref7133" refString="Poore, G. C. B. (2004) Marine decapod Crustacea of southern Australia. A guide to identification (with chapter on Stomatopoda by Shane Ahyong). CSIRO Publishing, Melbourne, 574 pp." type="book" year="2004">Poore (2004</bibRefCitation>
: fig. 36g);
<taxonomicName id="82BA4D65B1121E62FC02F4BCFBE52ACE" baseAuthorityName="Kemp" baseAuthorityYear="1916" box="[977,1078,1951,1974]" class="Malacostraca" family="Crangonidae" genus="Philocheras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="164" phylum="Arthropoda" rank="species" species="incisus">
<emphasis id="77CEEAF4B1121E62FC02F4BCFBE52ACE" box="[977,1078,1951,1974]" italics="true" pageId="7" pageNumber="164">P. incisus</emphasis>
</taxonomicName>
redrawn from
<bibRefCitation id="212B4B17B1121E62FB04F4BDFAB22ACD" author="Kemp, S." box="[1239,1377,1950,1973]" pageId="7" pageNumber="164" pagination="355 - 384" refId="ref6876" refString="Kemp, S. (1916) Notes on Crustacea Decapoda in the Indian Museum. VI. Indian Crangonidae. Records of the Indian Museum, 12, 355 - 384." type="journal article" year="1916">Kemp (1916</bibRefCitation>
: pl. 8 fig. 1);
<taxonomicName id="82BA4D65B1121E62FF34F49CFE532AAE" baseAuthorityName="Bate" baseAuthorityYear="1863" box="[231,384,1983,2006]" class="Malacostraca" family="Crangonidae" genus="Philocheras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="164" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="77CEEAF4B1121E62FF34F49CFE532AAE" box="[231,384,1983,2006]" italics="true" pageId="7" pageNumber="164">P. intermedius</emphasis>
</taxonomicName>
redrawn from
<bibRefCitation id="212B4B17B1121E62FDF2F49DFD7B2AAD" author="Poore, G. C. B." box="[545,680,1982,2005]" pageId="7" pageNumber="164" refId="ref7133" refString="Poore, G. C. B. (2004) Marine decapod Crustacea of southern Australia. A guide to identification (with chapter on Stomatopoda by Shane Ahyong). CSIRO Publishing, Melbourne, 574 pp." type="book" year="2004">Poore (2004</bibRefCitation>
: fig. 36h);
<taxonomicName id="82BA4D65B1121E62FCCDF49CFC4A2AAE" baseAuthorityName="Doflein" baseAuthorityYear="1902" box="[798,921,1983,2006]" class="Malacostraca" family="Crangonidae" genus="Philocheras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="164" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="77CEEAF4B1121E62FCCDF49CFC4A2AAE" box="[798,921,1983,2006]" italics="true" pageId="7" pageNumber="164">P.japonicus</emphasis>
</taxonomicName>
redrawn from De
<bibRefCitation id="212B4B17B1121E62FBB2F49DFB082AAD" author="Man, J. G. de" box="[1121,1243,1982,2005]" pageId="7" pageNumber="164" pagination="159 - 166" refId="ref7052" refString="Man, J. G. de (1918) Diagnosis of new species of macrurous decapod Crustacea from the Siboga-Expedition. Zoologische Mededeelingen, 4, 159 - 166." type="journal article" year="1918">Man (1918</bibRefCitation>
: pl. 23 fig. 69);
<taxonomicName id="82BA4D65B1121E62FA5BF49CFF042A8E" baseAuthorityName="Kemp" baseAuthorityYear="1916" class="Malacostraca" family="Crangonidae" genus="Philocheras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="164" phylum="Arthropoda" rank="species" species="lowisi">
<emphasis id="77CEEAF4B1121E62FA5BF49CFF042A8E" italics="true" pageId="7" pageNumber="164">P. lowisi</emphasis>
</taxonomicName>
redrawn from
<bibRefCitation id="212B4B17B1121E62FEA5F4FDFE2C2A8D" author="Kemp, S." box="[374,511,2014,2037]" pageId="7" pageNumber="164" pagination="355 - 384" refId="ref6876" refString="Kemp, S. (1916) Notes on Crustacea Decapoda in the Indian Museum. VI. Indian Crangonidae. Records of the Indian Museum, 12, 355 - 384." type="journal article" year="1916">Kemp (1916</bibRefCitation>
: pl. 8 fig. 2).
</paragraph>
</caption>
<paragraph id="450536E6B11D1E6DFF15F3B4FBCA2C3E" blockId="8.[151,1436,151,327]" pageId="8" pageNumber="165">
<emphasis id="77CEEAF4B11D1E6DFF15F3B4FEBF2DC9" bold="true" box="[198,364,151,177]" pageId="8" pageNumber="165">
<typeStatus id="9A018844B11D1E6DFF15F3B4FED72DC9" box="[198,260,151,177]" pageId="8" pageNumber="165">Type</typeStatus>
locality.
</emphasis>
Southern Sagami Nada,
<collectingCountry id="3DAD7676B11D1E6DFD43F3BBFD052DCA" box="[656,726,152,178]" name="Japan" pageId="8" pageNumber="165">Japan</collectingCountry>
.
<emphasis id="77CEEAF4B11D1E6DFF15F39EFEB42DAF" bold="true" box="[198,359,189,215]" pageId="8" pageNumber="165">Distribution.</emphasis>
<collectingCountry id="3DAD7676B11D1E6DFEBEF39EFE632DAF" box="[365,432,189,215]" name="Japan" pageId="8" pageNumber="165">Japan</collectingCountry>
and
<collectingCountry id="3DAD7676B11D1E6DFE3AF39EFDA32DAF" box="[489,624,189,215]" name="Philippines" pageId="8" pageNumber="165">Philippines</collectingCountry>
;
<collectingCountry id="3DAD7676B11D1E6DFDAFF39EFD392DAF" box="[636,746,189,215]" name="Australia" pageId="8" pageNumber="165">Australia</collectingCountry>
,
<collectingRegion id="877EF804B11D1E6DFD24F39EFC1F2DAF" box="[759,972,189,215]" country="Australia" name="Western Australia" pageId="8" pageNumber="165">Western Australia</collectingRegion>
;
<quantity id="82429B03B11D1E6DFC0BF39EFB822DAF" box="[984,1105,189,215]" metricMagnitude="2" metricUnit="m" metricValue="2.82" metricValueMax="5.22" metricValueMin="0.42" pageId="8" pageNumber="165" unit="m" value="282.0" valueMax="522.0" valueMin="42.0">
42
<specimenCount id="53BCFD6FB11D1E6DFBD7F39EFB822DAF" box="[1028,1105,189,215]" pageId="8" pageNumber="165" type="generic">522 m</specimenCount>
</quantity>
.
<emphasis id="77CEEAF4B11D1E6DFF15F3C1FEEC2D84" bold="true" box="[198,319,226,252]" pageId="8" pageNumber="165">Remarks.</emphasis>
This species shares the minutely dentate lateral margin of the antennal scale and a broad rostrum with
<emphasis id="77CEEAF4B11D1E6DFEEEF22BFE632C59" box="[317,432,264,289]" italics="true" pageId="8" pageNumber="165">
<taxonomicName id="82BA4D65B11D1E6DFEEEF22BFE7F2C59" baseAuthorityName="Kemp" baseAuthorityYear="1916" box="[317,428,264,289]" class="Malacostraca" family="Crangonidae" genus="Philocheras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="165" phylum="Arthropoda" rank="species" species="lowisi">P. lowisi</taxonomicName>
.
</emphasis>
It can be readily distinguished by the lack of a lateral spine on the antennal scale (present in
<taxonomicName id="82BA4D65B11D1E6DFEC8F20EFEAE2C3E" baseAuthorityName="Kemp" baseAuthorityYear="1916" box="[283,381,301,326]" class="Malacostraca" family="Crangonidae" genus="Philocheras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="165" phylum="Arthropoda" rank="species" species="lowisi">
<emphasis id="77CEEAF4B11D1E6DFEC8F20EFEAE2C3E" box="[283,381,301,326]" italics="true" pageId="8" pageNumber="165">P.lowisi</emphasis>
</taxonomicName>
) and the placement of spines and carina on the carapace.
</paragraph>
</subSubSection>
</treatment>
</document>