treatments-xml/data/FD/63/B4/FD63B46680D75E579728BD979D857994.xml
2024-06-21 12:59:12 +02:00

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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1029.63940" ID-GBIF-Dataset="27e8615f-ba3f-4d59-99a1-ecb8493d10f2" ID-PMC="PMC8050014" ID-Pensoft-Pub="1313-2970-1029-155" ID-Pensoft-UUID="50846BA2C2A85039BEFA6C1E2E3526FB" ID-PubMed="33935554" ID-ZooBank="9AE12DC8CDA44BA481C2B655C649AEC5" ModsDocID="1313-2970-1029-155" checkinTime="1617901766197" checkinUser="pensoft" docAuthor="Hava, Jiri, Abdel-Dayem, Mahmoud S. &amp; Aldhafer, Hathal M." docDate="2021" docId="FD63B46680D75E579728BD979D857994" docLanguage="en" docName="ZooKeys 1029: 155-174" docOrigin="ZooKeys 1029" docPubDate="2021-04-08" docSource="http://dx.doi.org/10.3897/zookeys.1029.63940" docTitle="Thorictus sharafi Hava &amp; Abdel-Dayem 2021, sp. nov." docType="treatment" docUuid="75B52C5E-77B0-4BE7-9074-48B32BFE779C" docUuidSource="ZooBank" docVersion="4" id="50846BA2C2A85039BEFA6C1E2E3526FB" lastPageNumber="155" masterDocId="50846BA2C2A85039BEFA6C1E2E3526FB" masterDocTitle="The subfamily Thorictinae (Coleoptera, Dermestidae) from Saudi Arabia" masterLastPageNumber="174" masterPageNumber="155" pageNumber="155" updateTime="1668152078424" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The subfamily Thorictinae (Coleoptera, Dermestidae) from Saudi Arabia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hava, Jiri</mods:namePart>
<mods:affiliation>Forestry and Game Management Research Institute, Strnady 136, CZ- 252 02 Praha 5 - Zbraslav, Czech Republic</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Abdel-Dayem, Mahmoud S.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6276-1740</mods:nameIdentifier>
<mods:affiliation>King Saud University Museum of Arthropods (KSMA), Plant Protection Department, College of Food and Agricultral Sciences, King Saud University, P. O. Box 2460 Riyadh 11451, Saudi Arabia</mods:affiliation>
<mods:nameIdentifier type="email">mseleem@ksu.edu.sa</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Aldhafer, Hathal M.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4911-2332</mods:nameIdentifier>
<mods:affiliation>King Saud University Museum of Arthropods (KSMA), Plant Protection Department, College of Food and Agricultral Sciences, King Saud University, P. O. Box 2460 Riyadh 11451, Saudi Arabia</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-04-08</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1029</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>155</mods:start>
<mods:end>174</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1029.63940</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1029.63940</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1029-155</mods:identifier>
<mods:identifier type="ZooBank">9AE12DC8CDA44BA481C2B655C649AEC5</mods:identifier>
<mods:identifier type="Pensoft-UUID">50846BA2C2A85039BEFA6C1E2E3526FB</mods:identifier>
</mods:mods>
<subSection pageId="0" pageNumber="155" type="results">
<treatment ID-GBIF-Taxon="180881642" LSID="urn:lsid:zoobank.org:act:75B52C5E-77B0-4BE7-9074-48B32BFE779C" httpUri="http://treatment.plazi.org/id/FD63B46680D75E579728BD979D857994" lastPageNumber="155" pageId="0" pageNumber="155">
<subSubSection pageId="0" pageNumber="155" type="nomenclature">
<paragraph pageId="0" pageNumber="155">
<taxonomicName LSID="http://zoobank.org/75B52C5E-77B0-4BE7-9074-48B32BFE779C" authority="Háva &amp; Abdel-Dayem" authorityName="Hava &amp; Abdel-Dayem" authorityYear="2021" class="Insecta" family="Dermestidae" genus="Thorictus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thorictus sharafi" order="Coleoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="sharafi" status="sp. nov.">
Thorictus sharafi
<normalizedToken originalValue="Háva">Hava</normalizedToken>
&amp; Abdel-Dayem
</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="155">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Habitus dorsal aspect of Thorictus sharafi Hava &amp; Abdel-Dayem, sp. nov." figureDoi="10.3897/zookeys.1029.63940.figure15" httpUri="https://binary.pensoft.net/fig/529732" pageId="0" pageNumber="155">Figs 15</figureCitation>
<figureCitation captionStart="Figures 16" captionStartId="F16" captionText="Figures 16. Thorictus sharafi Hava &amp; Abdel-Dayem, sp. nov. a head lateral eye b ventral setation on pronotum and metepisternum, T dohrni John, 1965 c head lateral eye d ventral setation on pronotum and metepisternum (according to John 1963)." figureDoi="10.3897/zookeys.1029.63940.figures16" httpUri="https://binary.pensoft.net/fig/529733" pageId="0" pageNumber="155">, 16</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="155" type="materials_examined">
<paragraph pageId="0" pageNumber="155">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="155">
<materialsCitation ID-GBIF-Occurrence="3069288309" collectingDate="2010-05-17" collectorName="Dr M. R. Sharaf" country="Saudi Arabia" elevation="1931" latitude="19.806784" location="Belgershi" longLatPrecision="1" longitude="41.711967" municipality="Alqamh Park" specimenCount="1" stateProvince="Baha Region" typeStatus="Holotype">
<emphasis bold="true" pageId="0" pageNumber="155">
<emphasis italics="true" pageId="0" pageNumber="155">
<typeStatus>Holotype</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry name="Saudi Arabia">Saudi Arabia</collectingCountry>
<specimenCount type="generic">1 ex</specimenCount>
;
<collectingRegion country="Saudi Arabia" name="Al Bahah">Baha Region</collectingRegion>
,
<collectingMunicipality>Alqamh Park</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:FD63B46680D75E579728BD979D857994:49931C9F7D73F6622EE36DD10BE5FC45" country="Saudi Arabia" latitude="19.806784" longLatPrecision="1" longitude="41.711967" municipality="Alqamh Park" name="Belgershi" stateProvince="Baha Region">Belgershi</location>
;
<geoCoordinate degrees="19" direction="north" minutes="48.407" orientation="latitude" precision="1" value="19.806784">19°48.407'N</geoCoordinate>
,
<geoCoordinate degrees="41" direction="east" minutes="42.718" orientation="longitude" precision="1" value="41.711967">41°42.718'E</geoCoordinate>
; alt.
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.9309999999999998" unit="m" value="1931.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.9309999999999998" unit="m" value="1931.0">1931 m</elevation>
</quantity>
;
<collectingDate value="2010-05-17">17 May 2010</collectingDate>
;
<collectorName>Dr M.R. Sharaf</collectorName>
leg.; KSMA
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3069288301" collectingDate="2010-05-14" collectingDateMax="2010-05-19" collectingDateMin="2010-05-14" collectorName="Dr M. R. Sharaf" country="Saudi Arabia" county="Amadan" elevation="1881" latitude="20.045383" location="Shohba forest" longLatPrecision="1" longitude="41.47608" municipality="Mandaq" specimenCount="2" stateProvince="Baha Region" typeStatus="Paratypes">
<emphasis bold="true" pageId="0" pageNumber="155">
<emphasis italics="true" pageId="0" pageNumber="155">
<typeStatus>Paratypes</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry name="Saudi Arabia">Saudi Arabia</collectingCountry>
<specimenCount type="generic">1 ex</specimenCount>
;
<collectingRegion country="Saudi Arabia" name="Al Bahah">Baha Region</collectingRegion>
,
<collectingCounty>Amadan</collectingCounty>
,
<collectingMunicipality>Mandaq</collectingMunicipality>
;
<geoCoordinate degrees="20" direction="north" minutes="12.163" orientation="latitude" precision="1" value="20.202717">20°12.163'N</geoCoordinate>
,
<geoCoordinate degrees="41" direction="east" minutes="13.906" orientation="longitude" precision="1" value="41.231766">41°13.906'E</geoCoordinate>
; alt.
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.8809999999999998" unit="m" value="1881.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.8809999999999998" unit="m" value="1881.0">1881 m</elevation>
</quantity>
;
<collectingDate value="2010-05-19">19 May 2010</collectingDate>
;
<collectorName>Dr M.R. Sharaf</collectorName>
leg.; KSMA •
<specimenCount type="generic">1 ex</specimenCount>
;
<collectingRegion country="Saudi Arabia" name="Al Bahah">Baha Region</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:FD63B46680D75E579728BD979D857994:13C684795FD0A44BD6A10813A477E3EA" country="Saudi Arabia" county="Amadan" latitude="20.045383" longLatPrecision="1" longitude="41.47608" municipality="Mandaq" name="Shohba forest" stateProvince="Baha Region">Shohba forest</location>
;
<geoCoordinate degrees="20" direction="north" minutes="02.723" orientation="latitude" precision="1" value="20.045383">20°02.723'N</geoCoordinate>
,
<geoCoordinate degrees="41" direction="east" minutes="28.565" orientation="longitude" precision="1" value="41.47608">41°28.565'E</geoCoordinate>
; alt.
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.324" unit="m" value="2324.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="2.324" unit="m" value="2324.0">2324 m</elevation>
</quantity>
;
<collectingDate value="2010-05-14">14 May 2010</collectingDate>
;
<collectorName>Dr M.R. Sharaf</collectorName>
leg. JHAC
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="155" type="description">
<paragraph pageId="0" pageNumber="155">Description of holotype.</paragraph>
<paragraph pageId="0" pageNumber="155">Body small, brown covered by short yellow setae on dorsal surfaces and short setae on ventral surfaces. Measurements (mm): TL 2.1, PW 1.1, EW 1.2. Head finely punctate with short yellow setae. Labial palpi entirely brown. Antennae brown, with 11 antennomeres, antennal club compact with three antennomeres. Pronotum as finely punctate as head, covered by short yellow setation. Lateral margin of pronotum not dentate. Pronotum in posterior part near scutellum without bumps. Ventral posterior pronotal angles without long yellow setation. Scutellum not visible from above. Elytra very finely punctate covered by short yellow setation. Each elytron in anterior part near humeri with one very small bump. Epipleuron finely punctate; anterior angles with short yellow setation. Prosternum finely punctate. Mesosternum finely punctate, mesosternal bulge. Metasternum finely punctate. Visible abdominal ventrites very finely punctate, covered by short yellow setae. First abdominal ventrite with anterior, longitudinal striation. Legs brown, covered by yellow setae.</paragraph>
<caption doi="10.3897/zookeys.1029.63940.figure15" httpUri="https://binary.pensoft.net/fig/529732" pageId="0" pageNumber="155" start="Figure 15" startId="F15">
<paragraph pageId="0" pageNumber="155">
<emphasis bold="true" pageId="0" pageNumber="155">Figure 15.</emphasis>
Habitus dorsal aspect of
<taxonomicName authorityName="Hava &amp; Abdel-Dayem" authorityYear="2021" class="Insecta" family="Dermestidae" genus="Thorictus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thorictus sharafi" order="Coleoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="sharafi">
<emphasis italics="true" pageId="0" pageNumber="155">Thorictus sharafi</emphasis>
</taxonomicName>
<normalizedToken originalValue="Háva">Hava</normalizedToken>
&amp; Abdel-Dayem, sp. nov.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="155" type="variability">
<paragraph pageId="0" pageNumber="155">Variability.</paragraph>
<paragraph pageId="0" pageNumber="155">Body measurements (mm): TL 2.1-2.2, PW 1.0-1.1, EW 1.1-1.2.</paragraph>
<caption doi="10.3897/zookeys.1029.63940.figures16" httpUri="https://binary.pensoft.net/fig/529733" pageId="0" pageNumber="155" start="Figures 16" startId="F16">
<paragraph pageId="0" pageNumber="155">
<emphasis bold="true" pageId="0" pageNumber="155">Figures 16.</emphasis>
<taxonomicName authorityName="Hava &amp; Abdel-Dayem" authorityYear="2021" class="Insecta" family="Dermestidae" genus="Thorictus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thorictus sharafi" order="Coleoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="sharafi">
<emphasis italics="true" pageId="0" pageNumber="155">Thorictus sharafi</emphasis>
</taxonomicName>
<normalizedToken originalValue="Háva">Hava</normalizedToken>
&amp; Abdel-Dayem, sp. nov.
<emphasis bold="true" pageId="0" pageNumber="155">a</emphasis>
head lateral eye
<emphasis bold="true" pageId="0" pageNumber="155">b</emphasis>
ventral setation on pronotum and metepisternum,
<taxonomicName class="Insecta" family="Dermestidae" genus="Thorictus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="T dohrni" order="Coleoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="dohrni">
<emphasis italics="true" pageId="0" pageNumber="155">T dohrni</emphasis>
</taxonomicName>
John, 1965
<emphasis bold="true" pageId="0" pageNumber="155">c</emphasis>
head lateral eye
<emphasis bold="true" pageId="0" pageNumber="155">d</emphasis>
ventral setation on pronotum and metepisternum (according to
<bibRefCitation author="John, H" journalOrPublisher="Entomologische Abhandlungen und Berichte aus dem Staatlichen Museum fuer Tierkunde in Dresden" pageId="0" pageNumber="155" pagination="177 - 255" refId="B16" refString="John, H, 1963. Revision der Gattung Thorictus Germar (Thorictidae, Clavicornia Col.). Eine Bestandaufnahme, Kritik und Neubeschreibungen. Entomologische Abhandlungen und Berichte aus dem Staatlichen Museum fuer Tierkunde in Dresden 28: 177 - 255" title="Revision der Gattung Thorictus Germar (Thorictidae, Clavicornia Col.). Eine Bestandaufnahme, Kritik und Neubeschreibungen." volume="28" year="1963">John 1963</bibRefCitation>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="155" type="differential diagnosis">
<paragraph pageId="0" pageNumber="155">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="155">
This new species is similar to
<taxonomicName class="Insecta" family="Dermestidae" genus="Thorictus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="T dohrni" order="Coleoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="dohrni">
<emphasis italics="true" pageId="0" pageNumber="155">T dohrni</emphasis>
</taxonomicName>
John, 1965 (Ethiopia), but differs from it by the abovementioned characteristics.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="155" type="etymology">
<paragraph pageId="0" pageNumber="155">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="155">The specific epithet is a Latinized noun in the genitive case in the masculine form based on the honorific name &quot;Dr Mostafa Sharaf,&quot; who collected the specimens of this species.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="155" type="ecological notes">
<paragraph pageId="0" pageNumber="155">Ecological notes.</paragraph>
<paragraph pageId="0" pageNumber="155">
This new species inhabits areas at elevations of 1881-2324 m within the African pencil cedar forest,
<taxonomicName class="Pinopsida" family="Cupressaceae" genus="Juniperus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Juniperus procera" order="Pinales" pageId="0" pageNumber="155" phylum="Coniferophyta" rank="species" species="procera">
<emphasis italics="true" pageId="0" pageNumber="155">Juniperus procera</emphasis>
</taxonomicName>
Hochst. ex Endl. (
<taxonomicName class="Pinopsida" family="Cupressaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Pinales" pageId="0" pageNumber="155" phylum="Coniferophyta" rank="family">Cupressaceae</taxonomicName>
) in the Baha Province (Figs
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Photo of African pencil cedar forest, Juniperus procera Hochst. ex Endl. (Cupressaceae), living habitat of Thorictus sharafi Hava &amp; Abdel-Dayem, sp. nov. holotype at Alqamh Park, Belgershi, Baha Province, in the mountains of southwestern Saudi Arabia, at 1931 m elevation." figureDoi="10.3897/zookeys.1029.63940.figure17" httpUri="https://binary.pensoft.net/fig/529734" pageId="0" pageNumber="155">17</figureCitation>
,
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Living habitat of Thorictus sharafi Hava &amp; Abdel-Dayem sp. nov. paratype in the mountains of southwestern Saudi Arabia, Baha Province, within the African pencil cedar forest, Juniperus procera Hochst. ex Endl. (Cupressaceae) of Shohba forest at 2324 m elevation." figureDoi="10.3897/zookeys.1029.63940.figure18" httpUri="https://binary.pensoft.net/fig/529735" pageId="0" pageNumber="155">18</figureCitation>
). All specimens were collected by hand-picking under stones during May. Host unknown.
</paragraph>
<caption doi="10.3897/zookeys.1029.63940.figure17" httpUri="https://binary.pensoft.net/fig/529734" pageId="0" pageNumber="155" start="Figure 17" startId="F17">
<paragraph pageId="0" pageNumber="155">
<emphasis bold="true" pageId="0" pageNumber="155">Figure 17.</emphasis>
Photo of African pencil cedar forest,
<taxonomicName class="Pinopsida" family="Cupressaceae" genus="Juniperus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Juniperus procera" order="Pinales" pageId="0" pageNumber="155" phylum="Coniferophyta" rank="species" species="procera">
<emphasis italics="true" pageId="0" pageNumber="155">Juniperus procera</emphasis>
</taxonomicName>
Hochst. ex Endl. (
<taxonomicName class="Pinopsida" family="Cupressaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Pinales" pageId="0" pageNumber="155" phylum="Coniferophyta" rank="family">Cupressaceae</taxonomicName>
), living habitat of
<taxonomicName authorityName="Hava &amp; Abdel-Dayem" authorityYear="2021" class="Insecta" family="Dermestidae" genus="Thorictus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thorictus sharafi" order="Coleoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="sharafi">
<emphasis italics="true" pageId="0" pageNumber="155">Thorictus sharafi</emphasis>
</taxonomicName>
<normalizedToken originalValue="Háva">Hava</normalizedToken>
&amp; Abdel-Dayem, sp. nov. holotype at Alqamh Park, Belgershi, Baha Province, in the mountains of southwestern Saudi Arabia, at 1931 m elevation.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1029.63940.figure18" httpUri="https://binary.pensoft.net/fig/529735" pageId="0" pageNumber="155" start="Figure 18" startId="F18">
<paragraph pageId="0" pageNumber="155">
<emphasis bold="true" pageId="0" pageNumber="155">Figure 18.</emphasis>
Living habitat of
<taxonomicName authorityName="Hava &amp; Abdel-Dayem" authorityYear="2021" class="Insecta" family="Dermestidae" genus="Thorictus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Thorictus sharafi" order="Coleoptera" pageId="0" pageNumber="155" phylum="Arthropoda" rank="species" species="sharafi">
<emphasis italics="true" pageId="0" pageNumber="155">Thorictus sharafi</emphasis>
</taxonomicName>
<normalizedToken originalValue="Háva">Hava</normalizedToken>
&amp; Abdel-Dayem sp. nov. paratype in the mountains of southwestern Saudi Arabia, Baha Province, within the African pencil cedar forest,
<taxonomicName class="Pinopsida" family="Cupressaceae" genus="Juniperus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Juniperus procera" order="Pinales" pageId="0" pageNumber="155" phylum="Coniferophyta" rank="species" species="procera">
<emphasis italics="true" pageId="0" pageNumber="155">Juniperus procera</emphasis>
</taxonomicName>
Hochst. ex Endl. (
<taxonomicName class="Pinopsida" family="Cupressaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Pinales" pageId="0" pageNumber="155" phylum="Coniferophyta" rank="family">Cupressaceae</taxonomicName>
) of Shohba forest at 2324 m elevation.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="155" type="distribution">
<paragraph pageId="0" pageNumber="155">Geographical distribution.</paragraph>
<paragraph pageId="0" pageNumber="155">
This species is known only from its type locality in the mountains of southwestern Saudi Arabia, at Alqamh Park, Amadan, and Shohba Forest in the Baha Province (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Map of Saudi Arabian provinces and distribution of Thorictinae species." figureDoi="10.3897/zookeys.1029.63940.figure1" httpUri="https://binary.pensoft.net/fig/529719" pageId="0" pageNumber="155">1</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>