266 lines
26 KiB
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266 lines
26 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/asp.80.e90570" ID-Pensoft-Pub="1864-8312-80-649" ID-Pensoft-UUID="FAE6DFEF69735531A4CBA275E078320D" ID-ZooBank="AD0B1CC262414460B6B888E6B95F20D7" ModsDocID="1864-8312-80-649" checkinTime="1671114540108" checkinUser="pensoft" docAuthor="Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S. & Harms, Danilo" docDate="2022" docId="78D6D54E21FC5C548F4EDE39CB81241B" docLanguage="en" docName="ArthropodSystPhyl 80: 649-691" docOrigin="Arthropod Systematics & amp; Phylogeny 80" docPubDate="2022-12-14" docSource="http://dx.doi.org/10.3897/asp.80.e90570" docTitle="Toliaranella Lorenz & Loria & Harvey & Harms 2022, gen. nov." docType="treatment" docVersion="1" id="FAE6DFEF69735531A4CBA275E078320D" lastPageNumber="649" masterDocId="FAE6DFEF69735531A4CBA275E078320D" masterDocTitle="The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots" masterLastPageNumber="691" masterPageNumber="649" pageNumber="649" updateTime="1671114540108" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lorenz, Michelle</mods:namePart>
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<mods:affiliation>Museum of Nature Hamburg - Zoology, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Martin-Luther-King-Platz 3, 20146 Hamburg, Germany & University of Hamburg, Biology Department, Ohnhorststrasse 18, 22609 Hamburg, Germany</mods:affiliation>
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<mods:nameIdentifier type="email">michellelorenz@gmx.net</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Loria, Stephanie F.</mods:namePart>
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<mods:affiliation>Museum of Nature Hamburg - Zoology, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Martin-Luther-King-Platz 3, 20146 Hamburg, Germany</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Harvey, Mark S.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1482-0109</mods:nameIdentifier>
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<mods:affiliation>Western Australian Museum, Collections & Research, 49 Kew Street, Welshpool, WA 6106, Australia & University of Western Australia, School of Biological Sciences, Crawley WA 6009, Australia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Harms, Danilo</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7189-5345</mods:nameIdentifier>
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<mods:affiliation>Museum of Nature Hamburg - Zoology, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Martin-Luther-King-Platz 3, 20146 Hamburg, Germany</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Arthropod Systematics & amp; Phylogeny</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-12-14</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>80</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>649</mods:start>
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<mods:end>691</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/asp.80.e90570</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/asp.80.e90570</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1864-8312-80-649</mods:identifier>
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<mods:identifier type="ZooBank">AD0B1CC262414460B6B888E6B95F20D7</mods:identifier>
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<mods:identifier type="Pensoft-UUID">FAE6DFEF69735531A4CBA275E078320D</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:zoobank.org:act:NomEnClAturAlACts/F9EDCC8D-D7A5-41F4-B101-DA7320782EBA" httpUri="http://treatment.plazi.org/id/78D6D54E21FC5C548F4EDE39CB81241B" lastPageNumber="649" pageId="0" pageNumber="649">
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<subSubSection pageId="0" pageNumber="649" type="nomenclature">
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<paragraph pageId="0" pageNumber="649">
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<taxonomicName LSID="https://zoobank.org/NomenclaturalActs/F9EDCC8D-D7A5-41F4-B101-DA7320782EBA" authority="Lorenz & Loria & Harvey & Harms, 2022" authorityName="Lorenz & Loria & Harvey & Harms" authorityYear="2022" family="Feaellidae" genus="Toliaranella" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Toliaranella" order="Pseudoscorpiones" pageId="0" pageNumber="649" rank="genus" status="gen. nov.">Toliaranella</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="649">gen. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="649" type="type species">
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<paragraph pageId="0" pageNumber="649">Type species.</paragraph>
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<paragraph pageId="0" pageNumber="649">
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<taxonomicName authorityName="Lorenz & Loria & Harvey & Harms" authorityYear="2022" family="Feaellidae" genus="Toliaranella" kingdom="Animalia" lsidName="Toliaranella pumila" order="Pseudoscorpiones" pageId="0" pageNumber="649" rank="species" species="pumila">
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<emphasis italics="true" pageId="0" pageNumber="649">Toliaranella pumila</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="649" type="diagnosis">
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<paragraph pageId="0" pageNumber="649">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="649">
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<taxonomicName authorityName="Lorenz & Loria & Harvey & Harms" authorityYear="2022" family="Feaellidae" genus="Toliaranella" kingdom="Animalia" lsidName="Toliaranella" order="Pseudoscorpiones" pageId="0" pageNumber="649" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="649">Toliaranella</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="649">gen. nov.</emphasis>
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differs from all other Malagasy feaellids by the presence of a perforated organ above the coxal spines and smallest body size of any Malagasy groups. Like
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<taxonomicName authorityName="Lorenz & Loria & Harvey & Harms" authorityYear="2022" family="Feaellidae" genus="Antsirananaella" kingdom="Animalia" lsidName="Antsirananaella" order="Pseudoscorpiones" pageId="0" pageNumber="649" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="649">Antsirananaella</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="649">gen. nov.</emphasis>
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and
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<taxonomicName authorityName="Lorenz & Loria & Harvey & Harms" authorityYear="2022" family="Feaellidae" genus="Mahajanganella" kingdom="Animalia" lsidName="Mahajanganella" order="Pseudoscorpiones" pageId="0" pageNumber="649" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="649">Mahajanganella</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="649">gen. nov.</emphasis>
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, it differs from
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<taxonomicName authorityName="Judson" authorityYear="2017" class="Arachnida" family="Feaellidae" genus="Cybella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cybella" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="649">Cybella</emphasis>
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</taxonomicName>
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by having platelets on the pleural membrane (absent in
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<taxonomicName authorityName="Judson" authorityYear="2017" class="Arachnida" family="Feaellidae" genus="Cybella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cybella" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="649">Cybella</emphasis>
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</taxonomicName>
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), from
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<taxonomicName authorityName="Harvey, Andrade & Pinto-da-Rocha" authorityYear="2016" class="Arachnida" family="Feaellidae" genus="Iporangella" kingdom="Animalia" lsidName="Iporangella" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="649">Iporangella</emphasis>
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</taxonomicName>
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by the presence of specialized setae on the movable chelal finger (absent in
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<taxonomicName authorityName="Harvey, Andrade & Pinto-da-Rocha" authorityYear="2016" class="Arachnida" family="Feaellidae" genus="Iporangella" kingdom="Animalia" lsidName="Iporangella" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="649">Iporangella</emphasis>
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</taxonomicName>
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), from
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<taxonomicName authority="" class="Arachnida" family="Feaellidae" genus="Feaella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Feaella (Difeaella)" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="subGenus" subGenus="Difeaella">Feaella (Difeaella)</taxonomicName>
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and
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<taxonomicName authority="(Feaella)" class="Arachnida" family="Feaellidae" genus="Feaella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Feaella (Feaella)" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="subGenus" subGenus="Feaella">Feaella (Feaella)</taxonomicName>
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by the presence of four anterior carapaceal lobes (two and six, respectively), from
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<taxonomicName authority="" class="Arachnida" family="Feaellidae" genus="Feaella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Feaella (Tetrafeaella)" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="subGenus" subGenus="Tetrafeaella">Feaella (Tetrafeaella)</taxonomicName>
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in continental Africa by having a less pronounced depression on the base of coxa I and on top of coxa II (distinctly more pronounced in
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<taxonomicName authorityName="Tullgren" authorityYear="1907" class="Arachnida" family="Feaellidae" genus="Feaella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Feaella (T.) mucronata" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="species" species="mucronata" subGenus="T.">Feaella (T.) cf. mucronata</taxonomicName>
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) and from the Australian species presently attributed to
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<taxonomicName authority="" class="Arachnida" family="Feaellidae" genus="Feaella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Feaella (Tetrafeaella)" order="Pseudoscorpiones" pageId="0" pageNumber="649" phylum="Arthropoda" rank="subGenus" subGenus="Tetrafeaella">Feaella (Tetrafeaella)</taxonomicName>
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by having fewer coxal spines (one pair versus three or four in the Australian taxa).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="649" type="etymology">
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<paragraph pageId="0" pageNumber="649">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="649">This genus is named after the former Toliara Province, where all specimens were collected. The gender is feminine.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="649" type="description">
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<paragraph pageId="0" pageNumber="649">Description.</paragraph>
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<paragraph pageId="0" pageNumber="649">
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The following description is based on holotype and allotype. -
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<emphasis italics="true" pageId="0" pageNumber="649">Carapace</emphasis>
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(Figs
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<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Toliaranella gen. nov., scanning electron micrographs. Specimens used for imaging: A T. griswoldi sp. nov. male paratype (ZMH-A 0016740); B, C T. meridionalis sp. nov. male paratype (ZMH-A 0016753). A carapace, dorsal view; B sternum ventral overview; C coxal spines and perforated organ in detail (highlighted in red). Scale bars: 0.2 mm (A, B); 0.1 mm (C)." figureDoi="10.3897/asp.80.e90570.figure18" httpUri="https://binary.pensoft.net/fig/782010" pageId="0" pageNumber="649">18A</figureCitation>
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,
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<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Drawings of Toliaranella pumila sp. nov.: A carapace from dorsal; B leg IV; C left chela from dorsal with trichobothrial pattern; D right pedipalp from dorsal. Scale bars: 0.2 mm (A - C); 0.4 mm (D)." figureDoi="10.3897/asp.80.e90570.figure20" httpUri="https://binary.pensoft.net/fig/782012" pageId="0" pageNumber="649">20A</figureCitation>
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,
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<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Drawings of Toliaranella fisheri sp. nov.: A carapace from dorsal; B leg IV; C left chela from dorsal with trichobothrial pattern; D right pedipalp from dorsal. Scale bars: 0.2 mm (A - C); 0.4 mm (D)." figureDoi="10.3897/asp.80.e90570.figure21" httpUri="https://binary.pensoft.net/fig/782013" pageId="0" pageNumber="649">21A</figureCitation>
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,
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<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Drawings of Toliaranella griswoldi sp. nov.: A carapace from dorsal; B leg IV; C left chela from dorsal with trichobothrial pattern; D right pedipalp from dorsal. Scale bars: 0.2 mm (A - C); 0.4 mm (D)." figureDoi="10.3897/asp.80.e90570.figure22" httpUri="https://binary.pensoft.net/fig/782014" pageId="0" pageNumber="649">22A</figureCitation>
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,
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<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Drawings of Toliaranella mahnerti sp. nov.: A carapace from dorsal; B leg IV; C left chela from dorsal with trichobothrial pattern; D right pedipalp from dorsal. Scale bars: 0.2 mm (A - C); 0.4 mm (D)." figureDoi="10.3897/asp.80.e90570.figure23" httpUri="https://binary.pensoft.net/fig/782015" pageId="0" pageNumber="649">23A</figureCitation>
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,
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<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Drawings of Toliaranella meridionalis sp. nov.: A carapace from dorsal; B leg IV; C left chela from dorsal with trichobothrial pattern; D right pedipalp from dorsal. Scale bars: 0.2 mm (A - C); 0.4 mm (D)." figureDoi="10.3897/asp.80.e90570.figure24" httpUri="https://binary.pensoft.net/fig/782016" pageId="0" pageNumber="649">24A</figureCitation>
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): Strongly granulate, all four anterior lobes with same distance to each other and rounded at tips. -
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<emphasis italics="true" pageId="0" pageNumber="649">Pedipalps</emphasis>
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(Figs
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<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Toliaranella gen. nov.. Specimen used for imaging: T. griswoldi sp. nov. holotype (CAS 9071500). A dorsal overview; B ventral overview; C lateral overview. Scale bar: 0.2 mm." figureDoi="10.3897/asp.80.e90570.figure17" httpUri="https://binary.pensoft.net/fig/782009" pageId="0" pageNumber="649">17A, B</figureCitation>
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,
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<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Toliaranella, gen. nov., scanning electron micrographs. Specimen used for imaging: T. pumila sp. nov. male paratype (ZMH-A 0016722). A right chela from ventral with specialized setae in a group in detail (highlighted in red); B left chela with double trichobothrium dt in detail (highlighted in red); C left chela from medial with terminal teeth in a compact group in detail. Scale bar: 0.2 mm." figureDoi="10.3897/asp.80.e90570.figure19" httpUri="https://binary.pensoft.net/fig/782011" pageId="0" pageNumber="649">19A-C</figureCitation>
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,
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<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Drawings of Toliaranella pumila sp. nov.: A carapace from dorsal; B leg IV; C left chela from dorsal with trichobothrial pattern; D right pedipalp from dorsal. Scale bars: 0.2 mm (A - C); 0.4 mm (D)." figureDoi="10.3897/asp.80.e90570.figure20" httpUri="https://binary.pensoft.net/fig/782012" pageId="0" pageNumber="649">20C, D</figureCitation>
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,
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<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Drawings of Toliaranella fisheri sp. nov.: A carapace from dorsal; B leg IV; C left chela from dorsal with trichobothrial pattern; D right pedipalp from dorsal. Scale bars: 0.2 mm (A - C); 0.4 mm (D)." figureDoi="10.3897/asp.80.e90570.figure21" httpUri="https://binary.pensoft.net/fig/782013" pageId="0" pageNumber="649">21C, D</figureCitation>
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,
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<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Drawings of Toliaranella griswoldi sp. nov.: A carapace from dorsal; B leg IV; C left chela from dorsal with trichobothrial pattern; D right pedipalp from dorsal. Scale bars: 0.2 mm (A - C); 0.4 mm (D)." figureDoi="10.3897/asp.80.e90570.figure22" httpUri="https://binary.pensoft.net/fig/782014" pageId="0" pageNumber="649">22C, D</figureCitation>
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,
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<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Drawings of Toliaranella mahnerti sp. nov.: A carapace from dorsal; B leg IV; C left chela from dorsal with trichobothrial pattern; D right pedipalp from dorsal. Scale bars: 0.2 mm (A - C); 0.4 mm (D)." figureDoi="10.3897/asp.80.e90570.figure23" httpUri="https://binary.pensoft.net/fig/782015" pageId="0" pageNumber="649">23C, D</figureCitation>
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,
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<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Drawings of Toliaranella meridionalis sp. nov.: A carapace from dorsal; B leg IV; C left chela from dorsal with trichobothrial pattern; D right pedipalp from dorsal. Scale bars: 0.2 mm (A - C); 0.4 mm (D)." figureDoi="10.3897/asp.80.e90570.figure24" httpUri="https://binary.pensoft.net/fig/782016" pageId="0" pageNumber="649">24C, D</figureCitation>
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): Terminal teeth with 7-8 teeth on both the fixed and the movable fingers (including 1 large tooth on fixed finger). -
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<emphasis italics="true" pageId="0" pageNumber="649">Coxal region</emphasis>
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(Figs
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<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Toliaranella gen. nov.. Specimen used for imaging: T. griswoldi sp. nov. holotype (CAS 9071500). A dorsal overview; B ventral overview; C lateral overview. Scale bar: 0.2 mm." figureDoi="10.3897/asp.80.e90570.figure17" httpUri="https://binary.pensoft.net/fig/782009" pageId="0" pageNumber="649">17B</figureCitation>
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,
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<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Toliaranella gen. nov., scanning electron micrographs. Specimens used for imaging: A T. griswoldi sp. nov. male paratype (ZMH-A 0016740); B, C T. meridionalis sp. nov. male paratype (ZMH-A 0016753). A carapace, dorsal view; B sternum ventral overview; C coxal spines and perforated organ in detail (highlighted in red). Scale bars: 0.2 mm (A, B); 0.1 mm (C)." figureDoi="10.3897/asp.80.e90570.figure18" httpUri="https://binary.pensoft.net/fig/782010" pageId="0" pageNumber="649">18B, C</figureCitation>
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): Pedipalpal coxae each with one spike laterally near base; coxae I with basal depression, each side with one small coxal spine; slightly more cranial of coxal spine one novel organ on each side: rose-shaped, not granulate but perforated organ (Fig.
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<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Toliaranella gen. nov., scanning electron micrographs. Specimens used for imaging: A T. griswoldi sp. nov. male paratype (ZMH-A 0016740); B, C T. meridionalis sp. nov. male paratype (ZMH-A 0016753). A carapace, dorsal view; B sternum ventral overview; C coxal spines and perforated organ in detail (highlighted in red). Scale bars: 0.2 mm (A, B); 0.1 mm (C)." figureDoi="10.3897/asp.80.e90570.figure18" httpUri="https://binary.pensoft.net/fig/782010" pageId="0" pageNumber="649">18B, C</figureCitation>
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) filling the base of the depression; coxae II with irregular shaped spines framing depression of coxae I.
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</paragraph>
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<caption doi="10.3897/asp.80.e90570.figure17" httpUri="https://binary.pensoft.net/fig/782009" pageId="0" pageNumber="649" start="Figure 17" startId="F17">
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<paragraph pageId="0" pageNumber="649">
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<emphasis bold="true" pageId="0" pageNumber="649">Figure 17.</emphasis>
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<taxonomicName authorityName="Lorenz & Loria & Harvey & Harms" authorityYear="2022" family="Feaellidae" genus="Toliaranella" kingdom="Animalia" lsidName="Toliaranella" order="Pseudoscorpiones" pageId="0" pageNumber="649" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="649">Toliaranella</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="649">gen. nov.</emphasis>
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|
. Specimen used for imaging:
|
|
<taxonomicName lsidName="T. griswoldi" pageId="0" pageNumber="649" rank="species" species="griswoldi">
|
|
<emphasis italics="true" pageId="0" pageNumber="649">T. griswoldi</emphasis>
|
|
</taxonomicName>
|
|
<emphasis bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
|
|
holotype (CAS 9071500).
|
|
<emphasis bold="true" pageId="0" pageNumber="649">A</emphasis>
|
|
dorsal overview;
|
|
<emphasis bold="true" pageId="0" pageNumber="649">B</emphasis>
|
|
ventral overview;
|
|
<emphasis bold="true" pageId="0" pageNumber="649">C</emphasis>
|
|
lateral overview. Scale bar: 0.2 mm.
|
|
</paragraph>
|
|
</caption>
|
|
<caption doi="10.3897/asp.80.e90570.figure18" httpUri="https://binary.pensoft.net/fig/782010" pageId="0" pageNumber="649" start="Figure 18" startId="F18">
|
|
<paragraph pageId="0" pageNumber="649">
|
|
<emphasis bold="true" pageId="0" pageNumber="649">Figure 18.</emphasis>
|
|
<taxonomicName authorityName="Lorenz & Loria & Harvey & Harms" authorityYear="2022" family="Feaellidae" genus="Toliaranella" kingdom="Animalia" lsidName="Toliaranella" order="Pseudoscorpiones" pageId="0" pageNumber="649" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="649">Toliaranella</emphasis>
|
|
</taxonomicName>
|
|
<emphasis bold="true" pageId="0" pageNumber="649">gen. nov.</emphasis>
|
|
, scanning electron micrographs. Specimens used for imaging:
|
|
<emphasis bold="true" pageId="0" pageNumber="649">A</emphasis>
|
|
<taxonomicName lsidName="T. griswoldi" pageId="0" pageNumber="649" rank="species" species="griswoldi">
|
|
<emphasis italics="true" pageId="0" pageNumber="649">T. griswoldi</emphasis>
|
|
</taxonomicName>
|
|
<emphasis bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
|
|
male paratype (ZMH-A0016740);
|
|
<emphasis bold="true" pageId="0" pageNumber="649">B</emphasis>
|
|
,
|
|
<emphasis bold="true" pageId="0" pageNumber="649">C</emphasis>
|
|
<taxonomicName lsidName="T. meridionalis" pageId="0" pageNumber="649" rank="species" species="meridionalis">
|
|
<emphasis italics="true" pageId="0" pageNumber="649">T. meridionalis</emphasis>
|
|
</taxonomicName>
|
|
<emphasis bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
|
|
male paratype (ZMH-A0016753).
|
|
<emphasis bold="true" pageId="0" pageNumber="649">A</emphasis>
|
|
carapace, dorsal view;
|
|
<emphasis bold="true" pageId="0" pageNumber="649">B</emphasis>
|
|
sternum ventral overview;
|
|
<emphasis bold="true" pageId="0" pageNumber="649">C</emphasis>
|
|
coxal spines and perforated organ in detail (highlighted in red). Scale bars: 0.2 mm (
|
|
<emphasis bold="true" pageId="0" pageNumber="649">A</emphasis>
|
|
,
|
|
<emphasis bold="true" pageId="0" pageNumber="649">B</emphasis>
|
|
); 0.1 mm (
|
|
<emphasis bold="true" pageId="0" pageNumber="649">C</emphasis>
|
|
).
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="649" type="included species">
|
|
<paragraph pageId="0" pageNumber="649">Included species.</paragraph>
|
|
<paragraph pageId="0" pageNumber="649">
|
|
<taxonomicName authorityName="Lorenz & Loria & Harvey & Harms" authorityYear="2022" family="Feaellidae" genus="Toliaranella" kingdom="Animalia" lsidName="Toliaranella fisheri" order="Pseudoscorpiones" pageId="0" pageNumber="649" rank="species" species="fisheri">
|
|
<emphasis italics="true" pageId="0" pageNumber="649">Toliaranella fisheri</emphasis>
|
|
</taxonomicName>
|
|
<emphasis bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
|
|
;
|
|
<taxonomicName authorityName="Lorenz & Loria & Harvey & Harms" authorityYear="2022" family="Feaellidae" genus="Toliaranella" kingdom="Animalia" lsidName="Toliaranella griswoldi" order="Pseudoscorpiones" pageId="0" pageNumber="649" rank="species" species="griswoldi">
|
|
<emphasis italics="true" pageId="0" pageNumber="649">Toliaranella griswoldi</emphasis>
|
|
</taxonomicName>
|
|
<emphasis bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
|
|
;
|
|
<taxonomicName authorityName="Lorenz & Loria & Harvey & Harms" authorityYear="2022" family="Feaellidae" genus="Toliaranella" kingdom="Animalia" lsidName="Toliaranella mahnerti" order="Pseudoscorpiones" pageId="0" pageNumber="649" rank="species" species="mahnerti">
|
|
<emphasis italics="true" pageId="0" pageNumber="649">Toliaranella mahnerti</emphasis>
|
|
</taxonomicName>
|
|
<emphasis bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
|
|
;
|
|
<taxonomicName authorityName="Lorenz & Loria & Harvey & Harms" authorityYear="2022" family="Feaellidae" genus="Toliaranella" kingdom="Animalia" lsidName="Toliaranella meridionalis" order="Pseudoscorpiones" pageId="0" pageNumber="649" rank="species" species="meridionalis">
|
|
<emphasis italics="true" pageId="0" pageNumber="649">Toliaranella meridionalis</emphasis>
|
|
</taxonomicName>
|
|
<emphasis bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
|
|
; and
|
|
<taxonomicName authorityName="Lorenz & Loria & Harvey & Harms" authorityYear="2022" family="Feaellidae" genus="Toliaranella" kingdom="Animalia" lsidName="Toliaranella pumila" order="Pseudoscorpiones" pageId="0" pageNumber="649" rank="species" species="pumila">
|
|
<emphasis italics="true" pageId="0" pageNumber="649">Toliaranella pumila</emphasis>
|
|
</taxonomicName>
|
|
<emphasis bold="true" pageId="0" pageNumber="649">sp. nov.</emphasis>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="649" type="distribution">
|
|
<paragraph pageId="0" pageNumber="649">Distribution.</paragraph>
|
|
<paragraph pageId="0" pageNumber="649">Members of this genus are localized to the southern part of Madagascar in the Androy, Anosy and Atsimo-Andrefana Regions (formerly Toliara Province) where the main landscape is covered by arid spiny bush vegetation.</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |