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<document id="F7E74CACC5938A8AAE48F74E916BCD1B" ID-DOI="10.5281/zenodo.173054" ID-GBIF-Dataset="cdda2aa6-ed47-4b05-a2de-8b853034f10c" ID-ISSN="1175­5326" ID-Zenodo-Dep="173054" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459768419171" checkinUser="plazi" docAuthor="Lowry, J. K." docDate="2006" docId="60705B7BFF93AB5A7C5FFA50FEAF818F" docLanguage="en" docName="zt01254p028.pdf" docOrigin="Zootaxa 1254" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Miramarassidae" docType="treatment" docVersion="9" lastPageNumber="18" masterDocId="9C492303FF9DAB4B7D57FFEEFFD28251" masterDocTitle="New families and subfamilies of amphipod crustaceans" masterLastPageNumber="28" masterPageNumber="1" pageNumber="15" updateTime="1698216993845" updateUser="plazi">
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<mods:title id="DA57FDCC2B3D8C94CB65883D5BD0790D">New families and subfamilies of amphipod crustaceans</mods:title>
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<mods:date id="E15E2B315263577EC41593BBB496B8C9">2006</mods:date>
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<treatment id="60705B7BFF93AB5A7C5FFA50FEAF818F" ID-DOI="http://doi.org/10.5281/zenodo.5669963" ID-GBIF-Taxon="119346617" ID-Zenodo-Dep="5669963" LSID="urn:lsid:plazi:treatment:60705B7BFF93AB5A7C5FFA50FEAF818F" httpUri="http://treatment.plazi.org/id/60705B7BFF93AB5A7C5FFA50FEAF818F" lastPageId="17" lastPageNumber="18" pageId="14" pageNumber="15">
<subSubSection id="A0C3B9E6FF93AB457C5FFA50FEB287AA" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="E866EA6DFF93AB457C5FFA50FD9D8789" blockId="14.[264,591,1470,1531]" box="[264,591,1470,1496]" pageId="14" pageNumber="15">
<heading id="B32E5D01FF93AB457C5FFA50FD9D8789" bold="true" box="[264,591,1470,1496]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
<emphasis id="DAAD367FFF93AB457C5FFA50FD9D8789" bold="true" box="[264,591,1470,1496]" pageId="14" pageNumber="15">
<taxonomicName id="2FD991EEFF93AB457C5FFA50FE078789" ID-CoL="7NJZM" box="[264,469,1470,1496]" class="Malacostraca" family="Miramarassidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="family">Miramarassidae</taxonomicName>
fam. nov.
</emphasis>
</heading>
</paragraph>
<paragraph id="E866EA6DFF93AB457C5FFA0FFEB287AA" blockId="14.[264,591,1470,1531]" box="[264,352,1505,1531]" pageId="14" pageNumber="15">(Fig. 4)</paragraph>
</subSubSection>
<subSubSection id="A0C3B9E6FF93AB457C5FF9DFFCC48422" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="E866EA6DFF93AB457C5FF9DFFE58841B" blockId="14.[264,790,1585,1651]" box="[264,394,1585,1610]" pageId="14" pageNumber="15">
<emphasis id="DAAD367FFF93AB457C5FF9DFFE58841B" box="[264,394,1585,1610]" italics="true" pageId="14" pageNumber="15">
<typeStatus id="376254CFFF93AB457C5FF9DFFEED841B" box="[264,319,1585,1610]" pageId="14" pageNumber="15">Type</typeStatus>
genus
</emphasis>
</paragraph>
<paragraph id="E866EA6DFF93AB457C6FF9B7FCC48422" blockId="14.[264,790,1585,1651]" box="[312,790,1625,1651]" pageId="14" pageNumber="15">
<taxonomicName id="2FD991EEFF93AB457C6FF9B7FCC08422" authority="Ortiz, Lalana &amp; Lio, 1999" authorityName="Ortiz, Lalana &amp; Lio" authorityYear="1999" box="[312,786,1625,1651]" class="Malacostraca" family="Miramarassidae" genus="Miramarassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="DAAD367FFF93AB457C6FF9B7FE078423" box="[312,469,1625,1650]" italics="true" pageId="14" pageNumber="15">Miramarassa</emphasis>
Ortiz, Lalana &amp; Lio, 1999
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="A0C3B9E6FF93AB5B7C5FF947FCC48411" lastPageId="16" lastPageNumber="17" pageId="14" pageNumber="15" type="description">
<paragraph id="E866EA6DFF93AB457C5FF947FE408493" blockId="14.[264,1323,1705,1891]" box="[264,402,1705,1730]" pageId="14" pageNumber="15">
<emphasis id="DAAD367FFF93AB457C5FF947FE408493" box="[264,402,1705,1730]" italics="true" pageId="14" pageNumber="15">Description</emphasis>
</paragraph>
<paragraph id="E866EA6DFF93AB447C6FF93FFC388097" blockId="14.[264,1323,1705,1891]" lastBlockId="15.[264,1324,284,1870]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
Head. Exposed; deeper than long; lateral cephalic lobe absent; anteroventral margin straight; anteroventral margin not recessed, not excavate; corner subquadrate; rostrum vestigial or absent; eyes poorly developed.
<emphasis id="DAAD367FFF93AB457E5BF8CFFBD5856A" bold="true" box="[780,1031,1825,1851]" pageId="14" pageNumber="15">Body subcylindrical</emphasis>
; without setae;
<emphasis id="DAAD367FFF93AB457992F8CFFEBC8532" bold="true" pageId="14" pageNumber="15">dorsally carinate</emphasis>
.
<emphasis id="DAAD367FFF93AB457CDEF8A7FDDE8533" box="[393,524,1865,1890]" italics="true" pageId="14" pageNumber="15">Antenna 1</emphasis>
shorter than antenna 2; peduncle with sparse slender setae; peduncular article 1 shorter than article 2; article 2 longer than article 3; article 3 subequal to, or longer than article 1; peduncular articles not geniculate; accessory flagellum short, 1­articulate; primary flagellum 5­ or more articulate; callynophore absent. Antenna 2 peduncle with sparse slender setae or none; without hook­like process.
<emphasis id="DAAD367FFF92AB447932FE7AFB3C83FC" box="[1125,1262,404,429]" italics="true" pageId="15" pageNumber="16">Mouthparts</emphasis>
well developed.
<emphasis id="DAAD367FFF92AB447CD8FE52FDD18387" bold="true" box="[399,515,444,470]" italics="true" pageId="15" pageNumber="16">Mandible</emphasis>
incisor dentate; lacinia mobilis present on both sides; accessory setal row without distal tuft; molar
<emphasis id="DAAD367FFF92AB447FD1FE0DFC4A83AC" bold="true" box="[646,920,483,509]" pageId="15" pageNumber="16">dominating mandible</emphasis>
; palp 3­articulate.
<emphasis id="DAAD367FFF92AB4479D0FE0AFB3383AC" box="[1159,1249,484,509]" italics="true" pageId="15" pageNumber="16">Labium</emphasis>
outer lobes smooth.
<emphasis id="DAAD367FFF92AB447CE3FDE2FDF48074" box="[436,550,524,549]" italics="true" pageId="15" pageNumber="16">Maxilla 1</emphasis>
inner plate setose along medial margin; outer plate with 11 setalteeth; palp large, 2­articulate.
<emphasis id="DAAD367FFF92AB447F23FDDAFD3B801C" box="[628,745,564,589]" italics="true" pageId="15" pageNumber="16">Maxilla 2</emphasis>
inner and outer plates present; inner plate with oblique setal row, with weakly setose medial margin.
<emphasis id="DAAD367FFF92AB447ECEFDB2FBCA8024" box="[921,1048,604,629]" italics="true" pageId="15" pageNumber="16">Maxilliped</emphasis>
inner and outer plates well developed; palps well developed; outer plate longer than palp article 1, not longer than palp article 2, apical robust setae absent; palp 4­articulate.
</paragraph>
<paragraph id="E866EA6DFF92AB447C6FFD3AFC74816F" blockId="15.[264,1324,284,1870]" pageId="15" pageNumber="16">
Pereon.
<emphasis id="DAAD367FFF92AB447CCFFD3AFDCA80BC" box="[408,536,724,750]" italics="true" pageId="15" pageNumber="16">Coxae 17</emphasis>
well developed.
<emphasis id="DAAD367FFF92AB447FB5FD3AFCB080BC" box="[738,866,724,750]" italics="true" pageId="15" pageNumber="16">Coxae 14</emphasis>
longer than broad or as long as broad, overlapping, coxae not ventrally acute.
<emphasis id="DAAD367FFF92AB447FA7FD12FCA68144" box="[752,884,764,790]" italics="true" pageId="15" pageNumber="16">Coxae 13</emphasis>
similar in size, well developed, all subequal in size.
<emphasis id="DAAD367FFF92AB447C85FCCAFD82816C" box="[466,592,804,830]" italics="true" pageId="15" pageNumber="16">Coxae 24</emphasis>
none extensively broadened.
</paragraph>
<paragraph id="E866EA6DFF92AB447C6FFCA2FD1A87C7" blockId="15.[264,1324,284,1870]" pageId="15" pageNumber="16">
<emphasis id="DAAD367FFF92AB447C6FFCA2FE068134" box="[312,468,844,869]" italics="true" pageId="15" pageNumber="16">Gnathopod 1</emphasis>
smaller (or weaker) than gnathopod 2; similar in form to gnathopod 2; subchelate; coxa subequal to coxa 2; merus and carpus not rotated; carpus/propodus not cantilevered; carpus longer than propodus, not produced along posterior margin of propodus; dactylus large.
<emphasis id="DAAD367FFF92AB447F6FFC2AFCE7818F" bold="true" box="[568,821,964,990]" pageId="15" pageNumber="16">
<emphasis id="DAAD367FFF92AB447F6FFC2AFD04818F" bold="true" box="[568,726,964,990]" italics="true" pageId="15" pageNumber="16">Gnathopod 2</emphasis>
chelate
</emphasis>
; coxa smaller than but not hidden by coxa 3;
<emphasis id="DAAD367FFF92AB447C7CFC02FE1D8657" bold="true" box="[299,463,1004,1030]" pageId="15" pageNumber="16">ischium long</emphasis>
; merus not fused along, or produced away from, posterior margin of carpus; carpus long, longer than propodus, not produced along posterior margin of propodus; dactylus well developed.
<emphasis id="DAAD367FFF92AB447F83FBD2FC838604" box="[724,849,1084,1109]" italics="true" pageId="15" pageNumber="16">Pereopods</emphasis>
none prehensile.
<emphasis id="DAAD367FFF92AB44796CFBD2FB658604" box="[1083,1207,1084,1109]" italics="true" pageId="15" pageNumber="16">Pereopods</emphasis>
34 not glandular.
<emphasis id="DAAD367FFF92AB447CD4FB8AFDDB862C" box="[387,521,1124,1149]" italics="true" pageId="15" pageNumber="16">Pereopod 3</emphasis>
coxa well developed, as long as broad; carpus longer than propodus, not produced; dactylus well developed.
<emphasis id="DAAD367FFF92AB447FB7FB62FCBB86F4" box="[736,873,1164,1189]" italics="true" pageId="15" pageNumber="16">Pereopod 4</emphasis>
coxa larger than coxa 3, not ventrally acute, without posteroventral lobe; carpus longer than propodus, not produced; dactylus well developed.
<emphasis id="DAAD367FFF92AB447C9FFB32FDAA86A4" box="[456,632,1244,1270]" italics="true" pageId="15" pageNumber="16">Pereopods 57</emphasis>
with few robust or slender setae; coxa smaller than coxa 4, with large anteroventral lobe; basis expanded, subrectangular, without posteroventral lobe.
<emphasis id="DAAD367FFF92AB447C5FFAC2FE468714" box="[264,404,1324,1349]" italics="true" pageId="15" pageNumber="16">Pereopod 6</emphasis>
shorter than, extending beyond propodus of, pereopod 7; basis expanded; dactylus without setae.
<emphasis id="DAAD367FFF92AB447F7FFABAFD61873C" box="[552,691,1364,1389]" italics="true" pageId="15" pageNumber="16">Pereopod 7</emphasis>
similar in structure to pereopod 6; basis expanded, subovate, without dense slender setae.
</paragraph>
<paragraph id="E866EA6DFF92AB447C6FFA4AFB05851F" blockId="15.[264,1324,284,1870]" pageId="15" pageNumber="16">
Pleon.
<emphasis id="DAAD367FFF92AB447CDFFA4AFDF887EF" bold="true" box="[392,554,1444,1470]" italics="true" pageId="15" pageNumber="16">Pleonites 13</emphasis>
without transverse dorsal serrations,
<emphasis id="DAAD367FFF92AB447E8BFA4AFAF187EF" bold="true" box="[988,1315,1444,1470]" pageId="15" pageNumber="16">12 each with dorsal spine</emphasis>
, without lateral teeth or ridging, without dorsal carina; without slender or robust dorsal setae.
<emphasis id="DAAD367FFF92AB447C04FA1AFE0F845C" box="[339,477,1524,1549]" italics="true" pageId="15" pageNumber="16">Epimeron 1</emphasis>
well developed.
<emphasis id="DAAD367FFF92AB447FFDFA1AFCC6845C" box="[682,788,1524,1549]" italics="true" pageId="15" pageNumber="16">Urosome</emphasis>
not dorsoventrally flattened; urosomites 13 free; not carinate;
<emphasis id="DAAD367FFF92AB447CB9F9F2FDAC8464" box="[494,638,1564,1589]" italics="true" pageId="15" pageNumber="16">urosomite 1</emphasis>
longer than urosomite 2.
<emphasis id="DAAD367FFF92AB447E93F9F2FD61840F" bold="true" pageId="15" pageNumber="16">
<emphasis id="DAAD367FFF92AB447E93F9F2FBB68467" bold="true" box="[964,1124,1564,1590]" italics="true" pageId="15" pageNumber="16">Uropods 12</emphasis>
apices of rami without robust or embedded setae
</emphasis>
.
<emphasis id="DAAD367FFF92AB447F94F9AAFC8E840C" box="[707,860,1604,1630]" italics="true" pageId="15" pageNumber="16">Uropods 13</emphasis>
similar in structure and size.
<emphasis id="DAAD367FFF92AB4479EDF9AAFAF8840C" box="[1210,1322,1604,1629]" italics="true" pageId="15" pageNumber="16">Uropod 1</emphasis>
peduncle without long plumose setae, without basofacial robust seta (robust or simple), without ventromedial spine.
<emphasis id="DAAD367FFF92AB447F0CF97AFD1E84FC" box="[603,716,1684,1709]" italics="true" pageId="15" pageNumber="16">Uropod 2</emphasis>
without dorsal flange, without ventromedial spine; inner ramus shorter than outer ramus.
<emphasis id="DAAD367FFF92AB447F83F952FC958484" box="[724,839,1724,1749]" italics="true" pageId="15" pageNumber="16">Uropod 3</emphasis>
without medial process; biramous; rami styliform; outer ramus longer than peduncle, without recurved spines; inner ramus subequal to outer ramus, inner ramus not apically setose.
<emphasis id="DAAD367FFF92AB447E80F8E2FBF18577" bold="true" box="[983,1059,1804,1830]" italics="true" pageId="15" pageNumber="16">Telson</emphasis>
laminar; longer than broad;
<emphasis id="DAAD367FFF92AB447C0DF8DAFE3A851F" bold="true" box="[346,488,1844,1870]" pageId="15" pageNumber="16">deeply cleft</emphasis>
; dorsal or lateral robust setae absent; apical robust setae absent.
</paragraph>
<paragraph id="E866EA6DFF8DAB5B7CC6FA5DFAD8879B" blockId="16.[401,1290,1459,1482]" box="[401,1290,1459,1482]" pageId="16" pageNumber="17">
<taxonomicName id="2FD991EEFF8DAB5B7CC6FA5DFDEE879B" box="[401,572,1459,1482]" class="Malacostraca" family="Miramarassidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="family">Miramarassidae</taxonomicName>
(
<taxonomicName id="2FD991EEFF8DAB5B7FEDFA5DFC7E879B" box="[698,940,1459,1482]" class="Malacostraca" family="Miramarassidae" genus="Miramarassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="sanchezi">
<emphasis id="DAAD367FFF8DAB5B7FEDFA5DFC7E879B" box="[698,940,1459,1482]" italics="true" pageId="16" pageNumber="17">Miramarassa sanchezi</emphasis>
</taxonomicName>
after Ortiz, Lalana &amp; Lio 1999).
</paragraph>
<paragraph id="E866EA6DFF8DAB5B7C5FFA10FE2E8446" blockId="16.[264,790,1534,1600]" box="[264,508,1534,1559]" pageId="16" pageNumber="17">
<emphasis id="DAAD367FFF8DAB5B7C5FFA10FE2E8446" box="[264,508,1534,1559]" italics="true" pageId="16" pageNumber="17">Generic composition</emphasis>
</paragraph>
<paragraph id="E866EA6DFF8DAB5B7C6FF9C8FCC48411" blockId="16.[264,790,1534,1600]" box="[312,790,1574,1600]" pageId="16" pageNumber="17">
<taxonomicName id="2FD991EEFF8DAB5B7C6FF9C8FCC08411" authority="Ortiz, Lalana &amp; Lio, 1999" authorityName="Ortiz, Lalana &amp; Lio" authorityYear="1999" box="[312,786,1574,1600]" class="Malacostraca" family="Miramarassidae" genus="Miramarassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="DAAD367FFF8DAB5B7C6FF9C8FE07846E" box="[312,469,1574,1599]" italics="true" pageId="16" pageNumber="17">Miramarassa</emphasis>
Ortiz, Lalana &amp; Lio, 1999
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="A0C3B9E6FF8DAB5A7C5FF998FBA48137" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="discussion">
<paragraph id="E866EA6DFF8DAB5B7C5FF998FEBC84DE" blockId="16.[264,1323,1654,1880]" box="[264,366,1654,1679]" pageId="16" pageNumber="17">
<emphasis id="DAAD367FFF8DAB5B7C5FF998FEBC84DE" box="[264,366,1654,1679]" italics="true" pageId="16" pageNumber="17">Remarks</emphasis>
</paragraph>
<paragraph id="E866EA6DFF8DAB5A7C6FF970FD90830F" blockId="16.[264,1323,1654,1880]" lastBlockId="17.[264,1323,284,870]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">There are several distinctive characteristics of the miramarassids: left and right lacinia mobilis present; setae along the entire medial margin of the inner plate of maxilla 1; a well developed oblique setal row on the inner plate of maxilla 2; an elongate ischium of gnathopod 2; absence of a posteroventral lobe on the coxa of pereopod 4; absence of apical robust setae on the rami of uropods 1 and 2; and a deeply cleft telson. Melphidippids have these characteristics, but melphidippids have differently shaped coxae and gnathopods and strongly different uropods.</paragraph>
<paragraph id="E866EA6DFF8CAB5A7C6FFE82FC9C8097" blockId="17.[264,1323,284,870]" pageId="17" pageNumber="18">
<taxonomicName id="2FD991EEFF8CAB5A7C6FFE82FE0183D7" box="[312,467,364,390]" class="Malacostraca" family="Iphimediidae" kingdom="Animalia" order="Amphipoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="family">Iphimediidae</taxonomicName>
are similar to miramarassids in the characters above except for the well developed oblique setal row on the inner plate of maxilla 2 which is usually absent in iphimediids although a few facial setae are sometimes present. The second gnathopods are very distinctive and similar between these two groups. Iphimediids differ from miramarassids in having styliform mouthparts, a small molar, only a left lacinia mobilis, acuminate coxae and never with a deeply cleft telson. Based on the similarities between these taxa it is possible to think of miramarassids as a primitive sister taxon to the iphimediids the way an iphimediid may have looked before it developed a highly modified mouthpart bundle and acuminate coxae.
</paragraph>
<paragraph id="E866EA6DFF8CAB5A7C6FFD3AFBC6816F" blockId="17.[264,1323,284,870]" pageId="17" pageNumber="18">There are certain characteristics which remain unknown, such as the possibility of a callynophore in the adult male, the relative lengths of antennae 1 and 2 and the surface of the molar which appears to be non­triturative, but is not described.</paragraph>
<paragraph id="E866EA6DFF8CAB5A7C6FFCA2FBA48137" blockId="17.[264,1323,284,870]" box="[312,1142,844,870]" pageId="17" pageNumber="18">The relationship of this taxon within the amphipods remains uncertain.</paragraph>
</subSubSection>
<subSubSection id="A0C3B9E6FF8CAB5A7C5FFC72FEAF818F" box="[264,407,924,990]" pageId="17" pageNumber="18" type="distribution">
<paragraph id="E866EA6DFF8CAB5A7C5FFC72FE4581E4" blockId="17.[264,407,924,990]" box="[264,407,924,949]" pageId="17" pageNumber="18">
<emphasis id="DAAD367FFF8CAB5A7C5FFC72FE4581E4" box="[264,407,924,949]" italics="true" pageId="17" pageNumber="18">Distribution</emphasis>
</paragraph>
<paragraph id="E866EA6DFF8CAB5A7C6FFC2AFEAF818F" blockId="17.[264,407,924,990]" box="[312,381,964,990]" pageId="17" pageNumber="18">
<collectingCountry id="90CEAAFDFF8CAB5A7C6FFC2AFEA8818F" box="[312,378,964,990]" name="Cuba" pageId="17" pageNumber="18">Cuba</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>