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<document id="1B718484EA4C448BD3E7DFD154925EBB" ID-DOI="10.11646/zootaxa.3926.1.2" ID-GBIF-Dataset="fec3fda7-4f6d-4ca1-b15c-0353d7b6ce22" ID-ISSN="1175-5326" ID-Zenodo-Dep="288010" ID-ZooBank="20666BE9-1457-41A6-9727-AC0077203595" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461005957897" checkinUser="plazi" docAuthor="Randall, Zachary S. &amp; Page, Lawrence M." docDate="2015" docId="2573D038A904971CFF4EFD1DA4EE0A8D" docLanguage="en" docName="zt03926p086.pdf" docOrigin="Zootaxa 3926 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Homalopterula Fowler 1940" docType="treatment" docVersion="7" lastPageNumber="72" masterDocId="D94AA840A90A9713FFD9FF92A7710D34" masterDocTitle="On the paraphyly of Homaloptera (Teleostei: Balitoridae) and description of a new genus of hillstream loaches from the Western Ghats of India" masterLastPageNumber="86" masterPageNumber="57" pageNumber="71" updateTime="1698565552551" updateUser="plazi">
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<mods:title id="D664A8CDFE950219F6CECB71A64C95A2">On the paraphyly of Homaloptera (Teleostei: Balitoridae) and description of a new genus of hillstream loaches from the Western Ghats of India</mods:title>
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<mods:date id="EB23340C717966B4EA649381365BB896">2015</mods:date>
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<treatment id="2573D038A904971CFF4EFD1DA4EE0A8D" ID-DOI="http://doi.org/10.5281/zenodo.5622840" ID-GBIF-Taxon="127671915" ID-Zenodo-Dep="5622840" LSID="urn:lsid:plazi:treatment:2573D038A904971CFF4EFD1DA4EE0A8D" httpUri="http://treatment.plazi.org/id/2573D038A904971CFF4EFD1DA4EE0A8D" lastPageId="15" lastPageNumber="72" pageId="14" pageNumber="71">
<subSubSection id="E5C032A5A904971DFF4EFD1DA6EC0FFF" pageId="14" pageNumber="71" type="nomenclature">
<paragraph id="AD65612EA904971DFF4EFD1DA6810F9E" blockId="14.[151,496,655,715]" box="[151,496,655,682]" pageId="14" pageNumber="71">
<heading id="F62DD642A904971DFF4EFD1DA6810F9E" bold="true" box="[151,496,655,682]" fontSize="11" level="1" pageId="14" pageNumber="71" reason="1">
<taxonomicName id="6ADA1AADA904971DFF4EFD1DA6810F9E" ID-CoL="6377J" ID-ENA="1590167" authority="Fowler 1940" authorityName="Fowler" authorityYear="1940" box="[151,496,655,682]" family="Balitoridae" genus="Homalopterula" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="71" phylum="Chordata" rank="genus">
<emphasis id="9FAEBD3CA904971DFF4EFD1DA6810F9E" bold="true" box="[151,496,655,682]" pageId="14" pageNumber="71">
<emphasis id="9FAEBD3CA904971DFF4EFD1DA63E0F9D" bold="true" box="[151,335,655,681]" italics="true" pageId="14" pageNumber="71">Homalopterula</emphasis>
<bibRefCitation id="C94B1CDFA904971DFE8FFD02A6810F9E" author="Fowler" box="[342,496,656,682]" pageId="14" pageNumber="71" refString="Fowler, H. W. (1940) Zoological results of the George Vanderbilt Sumatran Expedition, 1936 - 1939. Part II. - The fishes." type="book chapter" year="1940">Fowler 1940</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="AD65612EA904971DFF4EFD20A6EC0FFF" blockId="14.[151,496,655,715]" box="[151,413,690,715]" pageId="14" pageNumber="71">
(
<figureCitation id="35E17DABA904971DFF47FD20A6780FFF" box="[158,265,690,715]" captionStart="FIGURE 3" captionStartId="8.[151,250,1858,1880]" captionTargetBox="[211,1328,347,1828]" captionTargetId="figure@8.[151,1436,336,1848]" captionTargetPageId="8" captionText="FIGURE 3. Lateral views of living (A) Homaloptera orthogoniata, not cataloged, Kalimantan, Borneo, Indonesia. Strong reddish tints on body likely breeding colors; (B) Homalopteroides smithi, UF 235740, 45.0 mm SL, Khlong Tasae, Salui Subdistrict, Chumphon, Thailand; (C) Homalopterula cf. ripleyi, not cataloged, Sumatra, Indonesia; (D) Balitoropsis zollingeri, not cataloged, Kalimantan, Borneo, Indonesia; (E) Pseudohomaloptera leonardi, UF 235735, 34.2 mm SL, Ta Pi River, Nakhon Si Thammarat, Thailand. Photos (A) &amp; (D) by Renny Hadiaty. Photo (C) by Daniel Lumbantobing. Specimens not to scale." httpUri="https://zenodo.org/record/288013/files/figure.png" pageId="14" pageNumber="71">Figures 3</figureCitation>
C, 4E, 5C, 9)
</paragraph>
</subSubSection>
<subSubSection id="E5C032A5A904971DFF4EFD68A6590E1B" pageId="14" pageNumber="71" type="reference_group">
<paragraph id="AD65612EA904971DFF4EFD68A6590E1B" blockId="14.[151,1436,762,815]" pageId="14" pageNumber="71">
<treatmentCitationGroup id="8DCA4600A904971DFF4EFD68A6590E1B" pageId="14" pageNumber="71">
<taxonomicName id="6ADA1AADA904971DFF4EFD68A6870E24" ID-CoL="6377J" authority="Fowler, 1940:379" authorityName="Fowler" authorityPageNumber="379" authorityYear="1940" box="[151,502,762,784]" family="Balitoridae" genus="Homalopterula" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="71" phylum="Chordata" rank="genus">
<emphasis id="9FAEBD3CA904971DFF4EFD68A6430E24" box="[151,306,762,784]" italics="true" pageId="14" pageNumber="71">Homalopterula</emphasis>
<treatmentCitation id="2C7B473FA904971DFEE4FD68A6870E24" author="Fowler" box="[317,502,762,784]" page="379" pageId="14" pageNumber="71" year="1940">
<bibRefCitation id="C94B1CDFA904971DFEE4FD68A6BD0E24" author="Fowler" box="[317,460,762,784]" pageId="14" pageNumber="71" refString="Fowler, H. W. (1940) Zoological results of the George Vanderbilt Sumatran Expedition, 1936 - 1939. Part II. - The fishes." type="book chapter" year="1940">Fowler, 1940</bibRefCitation>
:379
</treatmentCitation>
</taxonomicName>
(
<typeStatus id="7261DF8CA904971DFDD3FD68A5460E24" box="[522,567,762,784]" pageId="14" pageNumber="71">type</typeStatus>
species:
<taxonomicName id="6ADA1AADA904971DFD79FD68A3440E24" ID-CoL="3MDWQ" authority="Fowler 1940:379" authorityName="Fowler" authorityPageNumber="379" authorityYear="1940" box="[672,1077,762,784]" class="Actinopterygii" family="Balitoridae" genus="Homaloptera" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="71" phylum="Chordata" rank="species" species="ripleyi">
<emphasis id="9FAEBD3CA904971DFD79FD68A4040E24" box="[672,885,762,784]" italics="true" pageId="14" pageNumber="71">Homaloptera ripleyi</emphasis>
<treatmentCitation id="2C7B473FA904971DFC58FD68A3440E24" author="Fowler" box="[897,1077,762,784]" page="379" pageId="14" pageNumber="71" year="1940">
<bibRefCitation id="C94B1CDFA904971DFC58FD68A37B0E24" author="Fowler" box="[897,1034,762,784]" pageId="14" pageNumber="71" refString="Fowler, H. W. (1940) Zoological results of the George Vanderbilt Sumatran Expedition, 1936 - 1939. Part II. - The fishes." type="book chapter" year="1940">Fowler 1940</bibRefCitation>
:379
</treatmentCitation>
</taxonomicName>
) by original designation. Gender feminine.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E5C032A5A904971DFF4EFCCEA5B20859" pageId="14" pageNumber="71" type="discussion">
<paragraph id="AD65612EA904971DFF4EFCCEA377091C" blockId="14.[151,1436,860,2001]" pageId="14" pageNumber="71">
<emphasis id="9FAEBD3CA904971DFF4EFCCEA67A0E41" bold="true" box="[151,267,860,885]" pageId="14" pageNumber="71">Remarks.</emphasis>
<taxonomicName id="6ADA1AADA904971DFECAFCCEA6CF0E40" box="[275,446,860,884]" family="Balitoridae" genus="Homalopterula" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="71" phylum="Chordata" rank="genus">
<emphasis id="9FAEBD3CA904971DFECAFCCEA6CF0E40" box="[275,446,860,884]" italics="true" pageId="14" pageNumber="71">Homalopterula</emphasis>
</taxonomicName>
was created for the new species
<taxonomicName id="6ADA1AADA904971DFCE1FCCEA3BC0E40" authority="Fowler 1940" authorityName="Fowler" authorityYear="1940" box="[824,1229,860,885]" family="Balitoridae" genus="Homalopterula" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="71" phylum="Chordata" rank="species" species="ripleyi">
<emphasis id="9FAEBD3CA904971DFCE1FCCEA3440E40" box="[824,1077,860,884]" italics="true" pageId="14" pageNumber="71">Homalopterula ripleyi</emphasis>
<bibRefCitation id="C94B1CDFA904971DFBE4FCCEA3BC0E40" author="Fowler" box="[1085,1229,860,885]" pageId="14" pageNumber="71" refString="Fowler, H. W. (1940) Zoological results of the George Vanderbilt Sumatran Expedition, 1936 - 1939. Part II. - The fishes." type="book chapter" year="1940">Fowler 1940</bibRefCitation>
</taxonomicName>
and distinguished from other balitorids by the “peculiar shape of its jaws, in combination with its truncated caudal and entirely naked medial under surface of the abdomen” (
<bibRefCitation id="C94B1CDFA904971DFDB1FC36A4420E88" author="Fowler" box="[616,819,932,957]" pageId="14" pageNumber="71" refString="Fowler, H. W. (1940) Zoological results of the George Vanderbilt Sumatran Expedition, 1936 - 1939. Part II. - The fishes." type="book chapter" year="1940">Fowler 1940:379</bibRefCitation>
).
<taxonomicName id="6ADA1AADA904971DFC90FC36A4850E88" box="[841,1012,932,956]" family="Balitoridae" genus="Homalopterula" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="71" phylum="Chordata" rank="genus">
<emphasis id="9FAEBD3CA904971DFC90FC36A4850E88" box="[841,1012,932,956]" italics="true" pageId="14" pageNumber="71">Homalopterula</emphasis>
</taxonomicName>
was treated as a junior synonym of
<taxonomicName id="6ADA1AADA904971DFF4EFC5AA65D0ED4" box="[151,300,968,992]" class="Actinopterygii" family="Balitoridae" genus="Homaloptera" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="71" phylum="Chordata" rank="genus">
<emphasis id="9FAEBD3CA904971DFF4EFC5AA65D0ED4" box="[151,300,968,992]" italics="true" pageId="14" pageNumber="71">Homaloptera</emphasis>
</taxonomicName>
based on the illustration provided in Fowler’s
<typeStatus id="7261DF8CA904971DFCEAFC5AA4120ED4" box="[819,867,968,992]" pageId="14" pageNumber="71">type</typeStatus>
description and the variation that “exists regarding the nature of the caudal and presence or absence of scales on the ventral surface of the abdomen in species of
<taxonomicName id="6ADA1AADA904971DFF4EFB82A65D091C" box="[151,300,1040,1064]" class="Actinopterygii" family="Balitoridae" genus="Homaloptera" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="71" phylum="Chordata" rank="genus">
<emphasis id="9FAEBD3CA904971DFF4EFB82A65D091C" box="[151,300,1040,1064]" italics="true" pageId="14" pageNumber="71">Homaloptera</emphasis>
</taxonomicName>
” by
<bibRefCitation id="C94B1CDFA904971DFEBAFB9DA69F091C" author="Silas" box="[355,494,1039,1064]" pageId="14" pageNumber="71" refString="Silas, E. G. (1953) Classification, zoogeography and evolution of the fishes of the cyprinoid families Homalopteridae and Gastromyzonidae. Records of the Indian Museum, 50, 173 - 263." type="journal article" year="1953">Silas (1953)</bibRefCitation>
. This decision was followed by
<bibRefCitation id="C94B1CDFA904971DFC8FFB9DA370091C" author="Roberts" box="[854,1025,1039,1064]" pageId="14" pageNumber="71" refString="Roberts, T. R. (1989) The freshwater fishes of western Borneo (Kalimantan Barat, Indonesia). Memoirs of the California Academy of Sciences, 14, 1 - 210." type="journal article" year="1989">Roberts (1989)</bibRefCitation>
.
</paragraph>
<paragraph id="AD65612EA904971DFF1EFBA6A5B20859" blockId="14.[151,1436,860,2001]" pageId="14" pageNumber="71">
<bibRefCitation id="C94B1CDFA904971DFF1EFBA6A6060978" author="Kottelat" box="[199,375,1076,1101]" pageId="14" pageNumber="71" refString="Kottelat, M. (1998) Homaloptera yuwonoi, a new species of hillstream loach from Borneo, with a new generic name for H. thamicola (Teleostei: Balitoridae). Ichthyological Exploration of Freshwaters, 9, 267 - 272." type="journal article" year="1998">Kottelat (1998)</bibRefCitation>
recognized
<taxonomicName id="6ADA1AADA904971DFDD1FBA6A4920979" authority="Bleeker 1853" authorityName="Bleeker" authorityYear="1853" box="[520,995,1076,1101]" class="Actinopterygii" family="Balitoridae" genus="Homaloptera" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="71" phylum="Chordata" rank="species" species="gymnogaster">
<emphasis id="9FAEBD3CA904971DFDD1FBA6A44B0978" box="[520,826,1076,1100]" italics="true" pageId="14" pageNumber="71">Homaloptera gymnogaster</emphasis>
Bleeker 1853
</taxonomicName>
,
<taxonomicName id="6ADA1AADA904971DFC2BFBA7A3960979" authority="Weber" authorityName="Weber" box="[1010,1255,1076,1101]" class="Actinopterygii" family="Balitoridae" genus="Homaloptera" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="71" phylum="Chordata" rank="species" species="heterolepis">
<emphasis id="9FAEBD3CA904971DFC2BFBA7A3E20978" box="[1010,1171,1076,1100]" italics="true" pageId="14" pageNumber="71">H. heterolepis</emphasis>
Weber
</taxonomicName>
&amp; de Beaufort 1916,
<taxonomicName id="6ADA1AADA904971DFF05FBCBA6810944" authority="Fowler 1940" authorityName="Fowler" authorityYear="1940" box="[220,496,1111,1136]" family="Balitoridae" genus="Homalopterula" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="71" phylum="Chordata" rank="species" species="ripleyi">
<emphasis id="9FAEBD3CA904971DFF05FBCBA6360944" box="[220,327,1112,1136]" italics="true" pageId="14" pageNumber="71">H. ripleyi</emphasis>
(
<bibRefCitation id="C94B1CDFA904971DFE8CFBC5A6980944" author="Fowler" box="[341,489,1111,1136]" pageId="14" pageNumber="71" refString="Fowler, H. W. (1940) Zoological results of the George Vanderbilt Sumatran Expedition, 1936 - 1939. Part II. - The fishes." type="book chapter" year="1940">Fowler 1940</bibRefCitation>
)
</taxonomicName>
, and
<taxonomicName id="6ADA1AADA904971DFDF3FBCBA42A0944" authority="Fowler 1940" authorityName="Fowler" authorityYear="1940" box="[554,859,1111,1136]" class="Actinopterygii" family="Balitoridae" genus="Homaloptera" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="71" phylum="Chordata" rank="species" species="vanderbilti">
<emphasis id="9FAEBD3CA904971DFDF3FBCBA5B60944" box="[554,711,1112,1136]" italics="true" pageId="14" pageNumber="71">H. vanderbilti</emphasis>
<bibRefCitation id="C94B1CDFA904971DFD14FBC5A42A0944" author="Fowler" box="[717,859,1111,1136]" pageId="14" pageNumber="71" refString="Fowler, H. W. (1940) Zoological results of the George Vanderbilt Sumatran Expedition, 1936 - 1939. Part II. - The fishes." type="book chapter" year="1940">Fowler 1940</bibRefCitation>
</taxonomicName>
as possibly forming a clade (
<taxonomicName id="6ADA1AADA904971DFB78FBCAA23D0944" box="[1185,1356,1112,1136]" family="Balitoridae" genus="Homalopterula" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="71" phylum="Chordata" rank="genus">
<emphasis id="9FAEBD3CA904971DFB78FBCAA23D0944" box="[1185,1356,1112,1136]" italics="true" pageId="14" pageNumber="71">Homalopterula</emphasis>
</taxonomicName>
) based on having “a more cylindrical body, a relatively wide mouth, short paired fins, and a truncated or slightly emarginated caudal fin.” Based on this description, Tan &amp; Ng (2005) and
<bibRefCitation id="C94B1CDFA904971DFC3EFB32A32F098C" author="Ott" box="[999,1118,1184,1208]" pageId="14" pageNumber="71" refString="Ott, G. (2009) Redescription of Homaloptera ripleyi (Fowler, 1940) from Sumatra, Indonesia (Teleostei: Balitoridae). Bulletin of Fish Biology, 11, 73 - 86." type="journal article" year="2009">Ott (2009)</bibRefCitation>
treated
<taxonomicName id="6ADA1AADA904971DFB65FB32A216098C" box="[1212,1383,1184,1208]" family="Balitoridae" genus="Homalopterula" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="71" phylum="Chordata" rank="genus">
<emphasis id="9FAEBD3CA904971DFB65FB32A216098C" box="[1212,1383,1184,1208]" italics="true" pageId="14" pageNumber="71">Homalopterula</emphasis>
</taxonomicName>
as a subgenus of
<taxonomicName id="6ADA1AADA904971DFEF5FB56A6B009E8" box="[300,449,1220,1244]" class="Actinopterygii" family="Balitoridae" genus="Homaloptera" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="71" phylum="Chordata" rank="genus">
<emphasis id="9FAEBD3CA904971DFEF5FB56A6B009E8" box="[300,449,1220,1244]" italics="true" pageId="14" pageNumber="71">Homaloptera</emphasis>
</taxonomicName>
.
<bibRefCitation id="C94B1CDFA904971DFE0DFB56A59209E8" author="Randall" box="[468,739,1220,1245]" pageId="14" pageNumber="71" refString="Randall, Z. S. &amp; Page, L. M. (2012) Resurrection of the genus Homalopteroides (Teleostei: Balitoridae) with a redescription of H. modestus (Vinciguerra 1890). Zootaxa, 3586, 329 - 346." type="journal article" year="2012">Randall &amp; Page (2012)</bibRefCitation>
recognized
<taxonomicName id="6ADA1AADA904971DFCAAFB56A36F09E8" box="[883,1054,1220,1244]" family="Balitoridae" genus="Homalopterula" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="71" phylum="Chordata" rank="genus">
<emphasis id="9FAEBD3CA904971DFCAAFB56A36F09E8" box="[883,1054,1220,1244]" italics="true" pageId="14" pageNumber="71">Homalopterula</emphasis>
</taxonomicName>
as a subgenus based on “mouth morphology, dorsal-fin origin over pelvic fin, ≥ 60 lateral-line scales, and ≥ 30 predorsal scales.” They did not include
<taxonomicName id="6ADA1AADA904971DFF29FA9FA6E20810" box="[240,403,1292,1316]" family="Balitoridae" genus="Homalopterula" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="71" phylum="Chordata" rank="species" species="modiglianii">
<emphasis id="9FAEBD3CA904971DFF29FA9FA6E20810" box="[240,403,1292,1316]" italics="true" pageId="14" pageNumber="71">H. modiglianii</emphasis>
</taxonomicName>
, then recognized as a junior synonym of
<taxonomicName id="6ADA1AADA904971DFC84FA9FA3B30810" authority="Kottelat 1993" authorityName="Kottelat" authorityYear="1993" box="[861,1218,1292,1317]" class="Actinopterygii" family="Balitoridae" genus="Homaloptera" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="71" phylum="Chordata" rank="species" species="gymnogaster">
<emphasis id="9FAEBD3CA904971DFC84FA9FA4000810" box="[861,881,1293,1316]" italics="true" pageId="14" pageNumber="71">H</emphasis>
.
<emphasis id="9FAEBD3CA904971DFCA6FA9FA3610810" box="[895,1040,1293,1316]" italics="true" pageId="14" pageNumber="71">gymnogaster</emphasis>
(
<bibRefCitation id="C94B1CDFA904971DFBC7FA9EA3C80810" author="Kottelat" box="[1054,1209,1292,1317]" pageId="14" pageNumber="71" refString="Kottelat, M., Whitten, A. J., Kartikasari, S. N. &amp; Wirjoatmodjo, S. (1993) Freshwater Fishes of Western Indonesia and Sulawesi. Periplus Editions, Hong Kong, 259 pp." type="book" year="1993">Kottelat 1993</bibRefCitation>
)
</taxonomicName>
. Including it in this study following
<bibRefCitation id="C94B1CDFA904971DFEB8FABDA561087C" author="Kottelat" box="[353,528,1327,1352]" pageId="14" pageNumber="71" refString="Kottelat, M. (2012) Conspectus cobitidum: an inventory of the loaches of the world (Teleostei: Cypriniformes: Cobitoidei). The Raffles Bulletin of Zoology, Supplement 26, 1 - 199." type="book chapter" year="2012">Kottelat (2012</bibRefCitation>
,
<bibRefCitation id="C94B1CDFA904971DFDFCFAA2A52E087C" author="Kottelat" box="[549,607,1328,1352]" pageId="14" pageNumber="71" refString="Kottelat, M. (2013) The fishes of the inland waters of southeast Asia: a catalogue and core biography of the fishes known to occur in freshwaters, mangroves and estuaries. The Raffles Bulletin of Zoology Supplement, 27, 1 - 663." type="journal article" year="2013">2013</bibRefCitation>
) changes the lateral-line and predorsal scale count range found in
<taxonomicName id="6ADA1AADA904971DFF4EFAC6A6330858" box="[151,322,1364,1388]" family="Balitoridae" genus="Homalopterula" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="71" phylum="Chordata" rank="genus">
<emphasis id="9FAEBD3CA904971DFF4EFAC6A6330858" box="[151,322,1364,1388]" italics="true" pageId="14" pageNumber="71">Homalopterula</emphasis>
</taxonomicName>
to 57–75 and 28–56, respectively.
</paragraph>
</subSubSection>
<subSubSection id="E5C032A5A904971CFF1EFAE5A6A40B8A" lastPageId="15" lastPageNumber="72" pageId="14" pageNumber="71" type="diagnosis">
<paragraph id="AD65612EA904971DFF1EFAE5A4FA0B5C" blockId="14.[151,1436,860,2001]" pageId="14" pageNumber="71">
<emphasis id="9FAEBD3CA904971DFF1EFAE5A63008A4" bold="true" box="[199,321,1399,1424]" pageId="14" pageNumber="71">Diagnosis.</emphasis>
Distinguishing characters are given in
<tableCitation id="E0585495A904971DFCF5FAE5A4F608A4" box="[812,903,1399,1424]" captionStart="TABLE 4" captionStartId="7.[125,207,600,621]" captionTargetBox="[125,1823,686,815]" captionTargetPageId="7" captionText="TABLE 4. Characters distinguishing Homaloptera, Homalopteroides, Homalopterula, Balitoropsis, Pseudohomaloptera and Ghatsa. Values are given as ranges; numbers in parentheses represent modal values. For Ghatsa, data from original descriptions are included; values in parentheses are for G. montana (CAS-SU 39871)." httpUri="http://table.plazi.org/id/F9A531A6A90D9714FFA4FDCAA1230FBC" pageId="14" pageNumber="71" tableUuid="F9A531A6A90D9714FFA4FDCAA1230FBC">Table 4</tableCitation>
and shown in
<figureCitation id="35E17DABA904971DFB97FAE5A3CE08A4" box="[1102,1215,1399,1424]" captionStart="FIGURE 3" captionStartId="8.[151,250,1858,1880]" captionTargetBox="[211,1328,347,1828]" captionTargetId="figure@8.[151,1436,336,1848]" captionTargetPageId="8" captionText="FIGURE 3. Lateral views of living (A) Homaloptera orthogoniata, not cataloged, Kalimantan, Borneo, Indonesia. Strong reddish tints on body likely breeding colors; (B) Homalopteroides smithi, UF 235740, 45.0 mm SL, Khlong Tasae, Salui Subdistrict, Chumphon, Thailand; (C) Homalopterula cf. ripleyi, not cataloged, Sumatra, Indonesia; (D) Balitoropsis zollingeri, not cataloged, Kalimantan, Borneo, Indonesia; (E) Pseudohomaloptera leonardi, UF 235735, 34.2 mm SL, Ta Pi River, Nakhon Si Thammarat, Thailand. Photos (A) &amp; (D) by Renny Hadiaty. Photo (C) by Daniel Lumbantobing. Specimens not to scale." httpUri="https://zenodo.org/record/288013/files/figure.png" pageId="14" pageNumber="71">Figures 3</figureCitation>
C, 4E, 5C and 9.
<taxonomicName id="6ADA1AADA904971DFF4EFA0EA6330880" box="[151,322,1436,1460]" family="Balitoridae" genus="Homalopterula" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="71" phylum="Chordata" rank="genus">
<emphasis id="9FAEBD3CA904971DFF4EFA0EA6330880" box="[151,322,1436,1460]" italics="true" pageId="14" pageNumber="71">Homalopterula</emphasis>
</taxonomicName>
is distinguished by the following combination of characters: without reddish tints on fins in life (
<figureCitation id="35E17DABA904971DFF46FA2DA79008EC" box="[159,225,1471,1496]" captionStart="FIGURE 3" captionStartId="8.[151,250,1858,1880]" captionTargetBox="[211,1328,347,1828]" captionTargetId="figure@8.[151,1436,336,1848]" captionTargetPageId="8" captionText="FIGURE 3. Lateral views of living (A) Homaloptera orthogoniata, not cataloged, Kalimantan, Borneo, Indonesia. Strong reddish tints on body likely breeding colors; (B) Homalopteroides smithi, UF 235740, 45.0 mm SL, Khlong Tasae, Salui Subdistrict, Chumphon, Thailand; (C) Homalopterula cf. ripleyi, not cataloged, Sumatra, Indonesia; (D) Balitoropsis zollingeri, not cataloged, Kalimantan, Borneo, Indonesia; (E) Pseudohomaloptera leonardi, UF 235735, 34.2 mm SL, Ta Pi River, Nakhon Si Thammarat, Thailand. Photos (A) &amp; (D) by Renny Hadiaty. Photo (C) by Daniel Lumbantobing. Specimens not to scale." httpUri="https://zenodo.org/record/288013/files/figure.png" pageId="14" pageNumber="71">Fig. 3</figureCitation>
C); dorsal-fin origin posterior to pelvic-fin origin; 5½ and 7½, 7½ (M) branched dorsal-fin rays; 7 pelvic-fin rays; truncated or emarginated caudal fin; smooth scales (
<figureCitation id="35E17DABA904971DFCF8FA76A41408C8" box="[801,869,1508,1533]" captionStart="FIGURE 4" captionStartId="9.[151,250,1660,1682]" captionTargetBox="[155,1431,193,1638]" captionTargetId="figure@9.[155,1432,193,1638]" captionTargetPageId="9" captionText="FIGURE 4. Scale sizes and keel patterns of predorsal region: Large, and keeled scales, (A) Balitoropsis zollingeri, UF 166094, 52.1 mm SL; (B) Pseudohomaloptera tatereganii, RMNH 7632 (holotype), 64.6 mm SL. Large, and wart-like / keeled scales (C) Homalopteroides wassinkii, UF 183330, 43.6 mm SL. Medium, and keeled scales, (D) Homaloptera cf. ocellata, UF 161718, 64.1 mm SL. Small, and smooth scales, (E) Homalopterula ripleyi, ANSP 188908, 43.1 mm SL; (F) Ghatsa montana, CAS-SU 39871 (holotype), 46.6 mm SL. Scale bar represents 1 mm. Left side of photo is anterior." httpUri="https://zenodo.org/record/288014/files/figure.png" pageId="14" pageNumber="71">Fig. 4</figureCitation>
E), 57–75 total lateral-line scales, 28–56 predorsal scales; anus closer to anal-fin origin than to pelvic-fin base; adipose keel on caudal peduncle; small rostral cap; 2 thick and widely separated rostral barbels; thick crescentic upper lip; 2 fleshy lobes between lateral portions of lower lip (
<figureCitation id="35E17DABA904971DFED2F9DDA63C0B5C" box="[267,333,1615,1640]" captionStart="FIGURE 5" captionStartId="10.[151,250,1357,1379]" captionTargetBox="[150,1436,267,1335]" captionTargetId="figure@10.[150,1436,267,1335]" captionTargetPageId="10" captionText="FIGURE 5. Oral morphology of (A) Homaloptera cf. ocellata, UF 161718, 64.1 mm SL; (B) Homalopteroides wassinkii, UMMZ 155660, 46.2 mm SL; (C) Homalopterula ripleyi, ANSP 188908, 43.1 mm SL; (D) Balitoropsis zollingeri, UF 166094, 52.1 mm SL; (E) Pseudohomaloptera tatereganii, RMNH 7632 (holotype), 64.6 mm SL; (F) Ghatsa montana, (holotype) CAS- SU 39871, 46.4 mm SL. Abbreviations: CF, central furrow; FL, fleshy lobe; FP, fleshy pad; LJ, lower jaw; LL, lower lip; LRB, lateral-rostral barbel; MB, maxillary barbel; MRB, medial-rostral barbel; RC, rostral cap; UJ, upper jaw; UL, upper lip." httpUri="https://zenodo.org/record/288015/files/figure.png" pageId="14" pageNumber="71">Fig. 5</figureCitation>
C); and presence of a central furrow at the isthmus.
</paragraph>
<paragraph id="AD65612EA904971DFF1EF9E6A2390BE0" blockId="14.[151,1436,860,2001]" pageId="14" pageNumber="71">
<emphasis id="9FAEBD3CA904971DFF1EF9E6A6D10BB9" bold="true" box="[199,416,1652,1677]" pageId="14" pageNumber="71">Species included.</emphasis>
<taxonomicName id="6ADA1AADA904971DFE6DF9E6A4A30BB9" authority="Bleeker 1853" authorityName="Bleeker" authorityYear="1853" box="[436,978,1652,1677]" family="Balitoridae" genus="Homalopterula" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="71" phylum="Chordata" rank="species" species="gymnogaster">
<emphasis id="9FAEBD3CA904971DFE6DF9E6A4740BB8" box="[436,773,1652,1676]" italics="true" pageId="14" pageNumber="71">Homalopterula gymnogaster</emphasis>
(Bleeker 1853)
</taxonomicName>
,
<taxonomicName id="6ADA1AADA904971DFC32F9E7A2160BB8" authority="Perugia 1893" authorityName="Perugia" authorityYear="1893" box="[1003,1383,1652,1677]" family="Balitoridae" genus="Homalopterula" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="71" phylum="Chordata" rank="species" species="modiglianii">
<emphasis id="9FAEBD3CA904971DFC32F9E7A3ED0BB8" box="[1003,1180,1652,1676]" italics="true" pageId="14" pageNumber="71">H. modiglianii</emphasis>
(Perugia 1893)
</taxonomicName>
,
<taxonomicName id="6ADA1AADA904971DFA58F9E7A50D0B84" authority="Weber &amp; Beaufort 1916" authorityName="Weber &amp; Beaufort" authorityYear="1916" family="Balitoridae" genus="Homalopterula" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="71" phylum="Chordata" rank="species" species="amphisquamata">
<emphasis id="9FAEBD3CA904971DFA58F9E7A63B0B84" italics="true" pageId="14" pageNumber="71">H. amphisquamata</emphasis>
(Weber &amp; Beaufort 1916)
</taxonomicName>
,
<taxonomicName id="6ADA1AADA904971DFD53F90BA4580B84" box="[650,809,1688,1712]" class="Actinopterygii" family="Balitoridae" genus="Homaloptera" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="71" phylum="Chordata" rank="species" species="heterolepis">
<emphasis id="9FAEBD3CA904971DFD53F90BA4580B84" box="[650,809,1688,1712]" italics="true" pageId="14" pageNumber="71">H. heterolepis</emphasis>
</taxonomicName>
(Weber &amp; de Beaufort 1916),
<taxonomicName id="6ADA1AADA904971DFB54F90BA2E90B84" authority="Fowler 1940" authorityName="Fowler" authorityYear="1940" box="[1165,1432,1687,1712]" family="Balitoridae" genus="Homalopterula" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="71" phylum="Chordata" rank="species" species="ripleyi">
<emphasis id="9FAEBD3CA904971DFB54F90BA38A0B84" box="[1165,1275,1688,1712]" italics="true" pageId="14" pageNumber="71">H. ripleyi</emphasis>
<bibRefCitation id="C94B1CDFA904971DFADDF905A2E90B84" author="Fowler" box="[1284,1432,1687,1712]" pageId="14" pageNumber="71" refString="Fowler, H. W. (1940) Zoological results of the George Vanderbilt Sumatran Expedition, 1936 - 1939. Part II. - The fishes." type="book chapter" year="1940">Fowler 1940</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="6ADA1AADA904971DFF1FF92FA57F0BE0" authority="Fowler 1940" authorityName="Fowler" authorityYear="1940" box="[198,526,1724,1749]" class="Actinopterygii" family="Balitoridae" genus="Homaloptera" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="71" phylum="Chordata" rank="species" species="vanderbilti">
<emphasis id="9FAEBD3CA904971DFF1FF92FA6120BE0" box="[198,355,1724,1748]" italics="true" pageId="14" pageNumber="71">H. vanderbilti</emphasis>
(
<bibRefCitation id="C94B1CDFA904971DFEABF92EA5770BE0" author="Fowler" box="[370,518,1724,1749]" pageId="14" pageNumber="71" refString="Fowler, H. W. (1940) Zoological results of the George Vanderbilt Sumatran Expedition, 1936 - 1939. Part II. - The fishes." type="book chapter" year="1940">Fowler 1940</bibRefCitation>
)
</taxonomicName>
. The
<typeStatus id="7261DF8CA904971DFD95F92FA50D0BE1" box="[588,636,1725,1749]" pageId="14" pageNumber="71">type</typeStatus>
localities for species of
<taxonomicName id="6ADA1AADA904971DFC54F92EA3490BE0" box="[909,1080,1724,1748]" family="Balitoridae" genus="Homalopterula" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="71" phylum="Chordata" rank="genus">
<emphasis id="9FAEBD3CA904971DFC54F92EA3490BE0" box="[909,1080,1724,1748]" italics="true" pageId="14" pageNumber="71">Homalopterula</emphasis>
</taxonomicName>
are shown in
<figureCitation id="35E17DABA904971DFB0CF92EA2350BE0" box="[1237,1348,1724,1749]" captionStart="FIGURE 10" captionStartId="16.[151,250,1729,1751]" captionTargetBox="[244,1337,204,1695]" captionTargetId="figure@16.[237,1350,195,1707]" captionTargetPageId="16" captionText="FIGURE 10. Type localities for species of Homalopterula. Asterisk represents the type species of the genus." httpUri="https://zenodo.org/record/288020/files/figure.png" pageId="14" pageNumber="71">Figure 10</figureCitation>
.
</paragraph>
<paragraph id="AD65612EA904971DFF1EF94DA5150AE5" blockId="14.[151,1436,860,2001]" pageId="14" pageNumber="71">
<emphasis id="9FAEBD3CA904971DFF1EF94DA6130BCC" bold="true" box="[199,354,1759,1784]" pageId="14" pageNumber="71">Comparison.</emphasis>
<taxonomicName id="6ADA1AADA904971DFEB3F972A5640BCC" box="[362,533,1760,1784]" family="Balitoridae" genus="Homalopterula" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="71" phylum="Chordata" rank="genus">
<emphasis id="9FAEBD3CA904971DFEB3F972A5640BCC" box="[362,533,1760,1784]" italics="true" pageId="14" pageNumber="71">Homalopterula</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="6ADA1AADA904971DFCCFF972A4DA0BCC" box="[790,939,1760,1784]" class="Actinopterygii" family="Balitoridae" genus="Homaloptera" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="71" phylum="Chordata" rank="genus">
<emphasis id="9FAEBD3CA904971DFCCFF972A4DA0BCC" box="[790,939,1760,1784]" italics="true" pageId="14" pageNumber="71">Homaloptera</emphasis>
</taxonomicName>
by absence vs. presence of reddish tints on fins in life; dorsal-fin origin posterior vs. anterior to pelvic-fin origin; 5½ and 7½, 7½ (M) vs. 7–8½, 8½ (M) branched dorsal-fin rays; 28–56 vs. 20–27 predorsal scales; smooth vs. keeled scales; truncated or emarginated vs. forked caudal fin; presence vs. absence of adipose keel on caudal peduncle; small vs. large rostral cap; medial- and lateral-rostral barbels widely separated from one another at base vs. barbels in close proximity to one another; crescentic rather than triangular upper lip; presence vs. absence of a central furrow at the isthmus; fleshy lobes vs. pad between lateral portions of lower lip.
</paragraph>
<caption id="F9A531A6A905971CFF4EFB2AA5B909D8" httpUri="https://zenodo.org/record/288019/files/figure.png" pageId="15" pageNumber="72" targetBox="[243,1346,206,1174]" targetPageId="15">
<paragraph id="AD65612EA905971CFF4EFB2AA5B909D8" blockId="15.[151,1436,1208,1260]" pageId="15" pageNumber="72">
<emphasis id="9FAEBD3CA905971CFF4EFB2AA66409F9" bold="true" box="[151,277,1208,1230]" pageId="15" pageNumber="72">FIGURE 9.</emphasis>
Dorsal, lateral, and ventral views of preserved
<taxonomicName id="6ADA1AADA905971CFCD2FB2AA48009FA" box="[779,1009,1208,1230]" family="Balitoridae" genus="Homalopterula" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="72" phylum="Chordata" rank="species" species="ripleyi">
<emphasis id="9FAEBD3CA905971CFCD2FB2AA48009FA" box="[779,1009,1208,1230]" italics="true" pageId="15" pageNumber="72">Homalopterula ripleyi</emphasis>
</taxonomicName>
, ANSP 188908, 43.1 mm SL, Kampung Bassam, Sumatera Utara Province, Sumatra, Indonesia.
</paragraph>
</caption>
<paragraph id="AD65612EA905971CFF1EFA8BA54408AA" blockId="15.[151,1437,1305,1978]" pageId="15" pageNumber="72">
<taxonomicName id="6ADA1AADA905971CFF1EFA8BA6030805" box="[199,370,1305,1329]" family="Balitoridae" genus="Homalopterula" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="72" phylum="Chordata" rank="genus">
<emphasis id="9FAEBD3CA905971CFF1EFA8BA6030805" box="[199,370,1305,1329]" italics="true" pageId="15" pageNumber="72">Homalopterula</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="6ADA1AADA905971CFDAAFA8BA4450805" box="[627,820,1305,1329]" class="Actinopterygii" family="Balitoridae" genus="Homalopteroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="72" phylum="Chordata" rank="genus">
<emphasis id="9FAEBD3CA905971CFDAAFA8BA4450805" box="[627,820,1305,1329]" italics="true" pageId="15" pageNumber="72">Homalopteroides</emphasis>
</taxonomicName>
by having truncated or emarginated vs. forked caudal fin; smooth vs. wart-like/keeled scales; 28–56 vs. 14–25 predorsal scales; 57–75 vs. 33–52 total lateral-line scales; presence vs. absence of adipose keel on caudal peduncle; thick vs. thin lips; presence vs. absence of fleshy lobes between lateral portions of lower lip.
</paragraph>
<paragraph id="AD65612EA905971CFF1EFA3BA6A40B8A" blockId="15.[151,1437,1305,1978]" pageId="15" pageNumber="72">
<taxonomicName id="6ADA1AADA905971CFF1EFA3BA60308F5" box="[199,370,1449,1473]" family="Balitoridae" genus="Homalopterula" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="72" phylum="Chordata" rank="genus">
<emphasis id="9FAEBD3CA905971CFF1EFA3BA60308F5" box="[199,370,1449,1473]" italics="true" pageId="15" pageNumber="72">Homalopterula</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="6ADA1AADA905971CFD48FA3BA46808F5" box="[657,793,1449,1473]" family="Balitoridae" genus="Balitoropsis" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="72" phylum="Chordata" rank="genus">
<emphasis id="9FAEBD3CA905971CFD48FA3BA46808F5" box="[657,793,1449,1473]" italics="true" pageId="15" pageNumber="72">Balitoropsis</emphasis>
</taxonomicName>
and
<taxonomicName id="6ADA1AADA905971CFCBBFA3BA33508F5" box="[866,1092,1449,1473]" family="Balitoridae" genus="Pseudohomaloptera" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="72" phylum="Chordata" rank="genus">
<emphasis id="9FAEBD3CA905971CFCBBFA3BA33508F5" box="[866,1092,1449,1473]" italics="true" pageId="15" pageNumber="72">Pseudohomaloptera</emphasis>
</taxonomicName>
by having dorsal-fin origin posterior vs. anterior to or above pelvic-fin origin; 5½ and 7½, 7½ (M) vs. 8½ branched dorsal-fin rays; 7 vs. 7–9, 8 (M) branched pelvic-fin rays; truncated or emarginated vs. forked caudal fin; smooth vs. keeled scales; 28–56 vs. 13–15 and 13–19 predorsal scales, respectively; presence vs. absence of adipose keel on caudal peduncle; small vs. large rostral cap; medial- and lateral-rostral barbels widely separated from one another at the base vs. barbels in close proximity to one another; presence of fleshy lobes vs. pad between lateral portions of lower lip. It is further distinguished from
<taxonomicName id="6ADA1AADA905971CFEABF913A68B0BAD" box="[370,506,1665,1689]" family="Balitoridae" genus="Balitoropsis" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="72" phylum="Chordata" rank="genus">
<emphasis id="9FAEBD3CA905971CFEABF913A68B0BAD" box="[370,506,1665,1689]" italics="true" pageId="15" pageNumber="72">Balitoropsis</emphasis>
</taxonomicName>
by having the anus closer to anal-fin origin than to pelvic-fin insertion and 57–75 vs. 42–55 lateral-line scales.
</paragraph>
</subSubSection>
<subSubSection id="E5C032A5A905971CFF1EF95BA4EE0A8D" pageId="15" pageNumber="72" type="materials_examined">
<paragraph id="AD65612EA905971CFF1EF95BA4EE0A8D" blockId="15.[151,1437,1305,1978]" pageId="15" pageNumber="72">
<emphasis id="9FAEBD3CA905971CFF1EF95BA4610BD6" bold="true" box="[199,784,1737,1762]" pageId="15" pageNumber="72">
Material examined.
<taxonomicName id="6ADA1AADA905971CFE62F95BA4790BD4" box="[443,776,1737,1762]" family="Balitoridae" genus="Homalopterula" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="72" phylum="Chordata" rank="species" species="gymnogaster">
<emphasis id="9FAEBD3CA905971CFE62F95BA4790BD4" bold="true" box="[443,776,1737,1762]" italics="true" pageId="15" pageNumber="72">Homalopterula gymnogaster</emphasis>
</taxonomicName>
:
</emphasis>
Sumatra:
<collectionCode id="CBCBF9EBA905971CFC5EF958A4AA0BD5" box="[903,987,1738,1761]" pageId="15" pageNumber="72">BMNH</collectionCode>
1866.5.2.49 (
<typeStatus id="7261DF8CA905971CFBA3F95BA3AF0BD6" box="[1146,1246,1737,1762]" pageId="15" pageNumber="72" type="holotype">holotype</typeStatus>
of
<taxonomicName id="6ADA1AADA905971CFADFF95BA6590A31" class="Actinopterygii" family="Balitoridae" genus="Homaloptera" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="72" phylum="Chordata" rank="species" species="gymnogaster">
<emphasis id="9FAEBD3CA905971CFADFF95BA6590A31" italics="true" pageId="15" pageNumber="72">Homaloptera gymnogaster</emphasis>
</taxonomicName>
);
<collectionCode id="CBCBF9EBA905971CFE9EF97CA6F40A32" box="[327,389,1774,1798]" httpUri="http://grbio.org/cool/a65z-wv6r" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="15" pageNumber="72">ZMA</collectionCode>
<date id="D96447EEA905971CFE4DF97FA6990A32" box="[404,488,1773,1798]" pageId="15" pageNumber="72" value="56-10-02">100256</date>
(
<typeStatus id="7261DF8CA905971CFE26F97CA5120A32" box="[511,611,1774,1798]" pageId="15" pageNumber="72" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="6ADA1AADA905971CFD4EF97FA4B90A31" box="[663,968,1773,1797]" class="Actinopterygii" family="Balitoridae" genus="Homaloptera" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="72" phylum="Chordata" rank="species" species="lepidogaster">
<emphasis id="9FAEBD3CA905971CFD4EF97FA4B90A31" box="[663,968,1773,1797]" italics="true" pageId="15" pageNumber="72">Homaloptera lepidogaster</emphasis>
</taxonomicName>
) (3); UF 185031 (1).
<emphasis id="9FAEBD3CA905971CFB3AF97CA2ED0A32" bold="true" box="[1251,1436,1773,1798]" pageId="15" pageNumber="72">
<taxonomicName id="6ADA1AADA905971CFB3AF97CA2E30A32" box="[1251,1426,1773,1798]" family="Balitoridae" genus="Homalopterula" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="72" phylum="Chordata" rank="species" species="modiglianii">
<emphasis id="9FAEBD3CA905971CFB3AF97CA2E30A32" bold="true" box="[1251,1426,1773,1798]" italics="true" pageId="15" pageNumber="72">H. modiglianii</emphasis>
</taxonomicName>
:
</emphasis>
Sumatra:
<collectionCode id="CBCBF9EBA905971CFEDAF880A6260A1D" box="[259,343,1810,1833]" pageId="15" pageNumber="72">BMNH</collectionCode>
1931.10.29.1-2 (
<typeStatus id="7261DF8CA905971CFDCEF883A50A0A1D" box="[535,635,1809,1833]" pageId="15" pageNumber="72" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="6ADA1AADA905971CFD79F883A4CF0A1D" box="[672,958,1809,1833]" class="Actinopterygii" family="Balitoridae" genus="Homaloptera" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="72" phylum="Chordata" rank="species" species="modiglianii">
<emphasis id="9FAEBD3CA905971CFD79F883A4CF0A1D" box="[672,958,1809,1833]" italics="true" pageId="15" pageNumber="72">Homaloptera modiglianii</emphasis>
</taxonomicName>
) (2).
<emphasis id="9FAEBD3CA905971CFC24F883A3920A1E" bold="true" box="[1021,1251,1809,1834]" pageId="15" pageNumber="72">
<taxonomicName id="6ADA1AADA905971CFC24F883A3AB0A1E" box="[1021,1242,1809,1834]" family="Balitoridae" genus="Homalopterula" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="72" phylum="Chordata" rank="species" species="amphisquamata">
<emphasis id="9FAEBD3CA905971CFC24F883A3AB0A1E" bold="true" box="[1021,1242,1809,1834]" italics="true" pageId="15" pageNumber="72">H. amphisquamata</emphasis>
</taxonomicName>
:
</emphasis>
Sumatra:
<collectionCode id="CBCBF9EBA905971CFA8FF883A2E90A1D" box="[1366,1432,1809,1833]" httpUri="http://grbio.org/cool/a65z-wv6r" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="15" pageNumber="72">ZMA</collectionCode>
. 100.998 (
<typeStatus id="7261DF8CA905971CFEDAF8A4A62D0A7A" box="[259,348,1846,1870]" pageId="15" pageNumber="72" type="syntype">syntype</typeStatus>
of
<taxonomicName id="6ADA1AADA905971CFE5CF8A7A5A70A79" box="[389,726,1845,1869]" class="Actinopterygii" family="Balitoridae" genus="Homaloptera" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="72" phylum="Chordata" rank="species" species="amphisquamata">
<emphasis id="9FAEBD3CA905971CFE5CF8A7A5A70A79" box="[389,726,1845,1869]" italics="true" pageId="15" pageNumber="72">Homaloptera amphisquamata</emphasis>
</taxonomicName>
), 100.994 (
<typeStatus id="7261DF8CA905971CFC85F8A4A4B10A7A" box="[860,960,1846,1870]" pageId="15" pageNumber="72" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="6ADA1AADA905971CFC33F8A7A24D0A79" box="[1002,1340,1845,1869]" class="Actinopterygii" family="Balitoridae" genus="Homaloptera" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="72" phylum="Chordata" rank="species" species="amphisquamata">
<emphasis id="9FAEBD3CA905971CFC33F8A7A24D0A79" box="[1002,1340,1845,1869]" italics="true" pageId="15" pageNumber="72">Homaloptera amphisquamata</emphasis>
</taxonomicName>
) (2).
<emphasis id="9FAEBD3CA905971CFAA6F8A4A66F0A46" bold="true" pageId="15" pageNumber="72">
<taxonomicName id="6ADA1AADA905971CFAA6F8A4A6640A46" class="Actinopterygii" family="Balitoridae" genus="Homaloptera" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="72" phylum="Chordata" rank="species" species="heterolepis">
<emphasis id="9FAEBD3CA905971CFAA6F8A4A6640A46" bold="true" italics="true" pageId="15" pageNumber="72">H. heterolepis</emphasis>
</taxonomicName>
:
</emphasis>
<collectionCode id="CBCBF9EBA905971CFEF7F8CBA61D0A45" box="[302,364,1881,1905]" httpUri="http://grbio.org/cool/a65z-wv6r" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="15" pageNumber="72">ZMA</collectionCode>
100.999 (
<typeStatus id="7261DF8CA905971CFE36F8CBA5220A45" box="[495,595,1881,1905]" pageId="15" pageNumber="72" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="6ADA1AADA905971CFD50F8CBA4D80A45" box="[649,937,1881,1905]" class="Actinopterygii" family="Balitoridae" genus="Homaloptera" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="72" phylum="Chordata" rank="species" species="heterolepis">
<emphasis id="9FAEBD3CA905971CFD50F8CBA4D80A45" box="[649,937,1881,1905]" italics="true" pageId="15" pageNumber="72">Homaloptera heterolepis</emphasis>
</taxonomicName>
) (3).
<emphasis id="9FAEBD3CA905971CFC22F8CBA3080A46" bold="true" box="[1019,1145,1881,1906]" pageId="15" pageNumber="72">
<taxonomicName id="6ADA1AADA905971CFC22F8CBA3010A46" box="[1019,1136,1881,1906]" family="Balitoridae" genus="Homalopterula" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="72" phylum="Chordata" rank="species" species="ripleyi">
<emphasis id="9FAEBD3CA905971CFC22F8CBA3010A46" bold="true" box="[1019,1136,1881,1906]" italics="true" pageId="15" pageNumber="72">H. ripleyi</emphasis>
</taxonomicName>
:
</emphasis>
Sumatra:
<collectionCode id="CBCBF9EBA905971CFB27F8CBA2370A45" LSID="urn:lsid:biocol.org:col:34784" box="[1278,1350,1881,1905]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="15" pageNumber="72">ANSP</collectionCode>
68713 (
<typeStatus id="7261DF8CA905971CFF46F8EFA6760AA2" box="[159,263,1917,1942]" pageId="15" pageNumber="72" type="holotype">holotype</typeStatus>
), 187003 (2), 187004 (1), 188907 (1), 188908 (4).
<emphasis id="9FAEBD3CA905971CFC89F8ECA48A0AA2" bold="true" box="[848,1019,1917,1942]" pageId="15" pageNumber="72">
<taxonomicName id="6ADA1AADA905971CFC89F8ECA4800AA2" box="[848,1009,1917,1942]" class="Actinopterygii" family="Balitoridae" genus="Homaloptera" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="72" phylum="Chordata" rank="species" species="vanderbilti">
<emphasis id="9FAEBD3CA905971CFC89F8ECA4800AA2" bold="true" box="[848,1009,1917,1942]" italics="true" pageId="15" pageNumber="72">H. vanderbilti</emphasis>
</taxonomicName>
:
</emphasis>
Sumatra:
<collectionCode id="CBCBF9EBA905971CFBABF8ECA3C80AA2" LSID="urn:lsid:biocol.org:col:34784" box="[1138,1209,1918,1942]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="15" pageNumber="72">ANSP</collectionCode>
68688 (
<typeStatus id="7261DF8CA905971CFAC1F8EFA20D0AA2" box="[1304,1404,1917,1942]" pageId="15" pageNumber="72" type="holotype">holotype</typeStatus>
of
<taxonomicName id="6ADA1AADA905971CFF4EF833A6DC0A8D" box="[151,429,1953,1977]" class="Actinopterygii" family="Balitoridae" genus="Homaloptera" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="72" phylum="Chordata" rank="species" species="vanderbilti">
<emphasis id="9FAEBD3CA905971CFF4EF833A6DC0A8D" box="[151,429,1953,1977]" italics="true" pageId="15" pageNumber="72">Homaloptera vanderbilti</emphasis>
</taxonomicName>
), 68700 (
<typeStatus id="7261DF8CA905971CFDC0F833A5F20A8E" box="[537,643,1953,1978]" pageId="15" pageNumber="72" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="6ADA1AADA905971CFD70F833A4FF0A8D" box="[681,910,1953,1977]" class="Actinopterygii" family="Balitoridae" genus="Homaloptera" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="72" phylum="Chordata" rank="species" species="ulmeri">
<emphasis id="9FAEBD3CA905971CFD70F833A4FF0A8D" box="[681,910,1953,1977]" italics="true" pageId="15" pageNumber="72">Homaloptera ulmeri</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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