treatments-xml/data/23/73/BB/2373BB7A3F7E3F0D8C7A1B7FFAA1F20E.xml

<document ID-DOI="http://doi.org/10.5281/zenodo.4762821" ID-GBIF-Dataset="97d9d905-b4c2-4be0-9886-eb21c200d2b8" ID-GBIF-Taxon="182073704" ID-ISSN="1854-0392" ID-Zenodo-Dep="4762821" checkinTime="1620936722352" checkinUser="felipe" docAuthor="Baumann, Richard W. &amp; Lee, Jonathan J." docDate="2009" docId="2373BB7A3F7E3F0D8C7A1B7FFAA1F20E" docLanguage="en" docName="Illiesia.5.1.1-10.pdf" docOrigin="Illiesia 5 (1)" docStyle="DocumentStyle:3ED4ECBC8873166DC296BE619562C54F.3:Illiesia.2009-.journal_article" docStyleId="3ED4ECBC8873166DC296BE619562C54F" docStyleName="Illiesia.2009-.journal_article" docStyleVersion="3" docTitle="Isoperla laucki Baumann &amp; Lee 2009, sp.n." docType="treatment" docVersion="3" lastPageId="4" lastPageNumber="5" masterDocId="DF4AC3023F7F3F098D0B1979FFB9F722" masterDocTitle="Two Interesting New Species Of Isoperla From Northern California (Plecoptera: Perlodidae)" masterLastPageNumber="10" masterPageNumber="1" pageId="1" pageNumber="2" updateTime="1643565812222" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title>Two Interesting New Species Of Isoperla From Northern California (Plecoptera: Perlodidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Baumann, Richard W.</mods:namePart>
<mods:affiliation>Monte L. Bean Life Science Museum, Department of Biology, Brigham Young University, Provo, UT 84602 E-mail: richard _ baumann @ byu. edu</mods:affiliation>
<mods:nameIdentifier type="email">richard_baumann@byu.edu</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lee, Jonathan J.</mods:namePart>
<mods:affiliation>2337 15 Street Eureka, California 95501 E-mail: jlee @ humboldt 1. com</mods:affiliation>
<mods:nameIdentifier type="email">jlee@humboldt</mods:nameIdentifier>
</mods:name>
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<mods:titleInfo>
<mods:title>Illiesia</mods:title>
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<mods:part>
<mods:date>2009</mods:date>
<mods:detail type="pubDate">
<mods:number>2009-01-26</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5</mods:number>
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<mods:number>1</mods:number>
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<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.4758990</mods:identifier>
<mods:identifier type="GBIF-Dataset">97d9d905-b4c2-4be0-9886-eb21c200d2b8</mods:identifier>
<mods:identifier type="ISSN">1854-0392</mods:identifier>
<mods:identifier type="Zenodo-Dep">4758990</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.4762821" ID-GBIF-Taxon="182073704" ID-Zenodo-Dep="4762821" LSID="urn:lsid:plazi:treatment:2373BB7A3F7E3F0D8C7A1B7FFAA1F20E" httpUri="http://treatment.plazi.org/id/2373BB7A3F7E3F0D8C7A1B7FFAA1F20E" lastPageId="4" lastPageNumber="5" pageId="1" pageNumber="2">
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<heading bold="true" box="[369,618,518,542]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="2" reason="2">
<emphasis bold="true" box="[369,618,518,542]" pageId="1" pageNumber="2">
<taxonomicName authority="Baumann &amp; Lee, 2009" authorityName="Baumann &amp; Lee" authorityYear="2009" box="[369,547,518,542]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="laucki" status="sp.n.">
<emphasis bold="true" box="[369,547,518,542]" italics="true" pageId="1" pageNumber="2">Isoperla laucki</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[560,618,518,542]" pageId="1" pageNumber="2" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<paragraph blockId="1.[409,577,554,578]" box="[409,577,554,578]" pageId="1" pageNumber="2">
(
<figureCitation box="[417,531,554,578]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="2.[189,240,1796,1820]" captionStartId-1="3.[189,240,1796,1820]" captionTargetBox-0="[249,1384,189,1747]" captionTargetBox-1="[249,1383,189,1748]" captionTargetId-0="figure-42@2.[249,1384,189,1749]" captionTargetId-1="figure-42@3.[249,1383,189,1748]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="Figs. 1‐8. Isoperla laucki: 1. Aedeagus, dorsal, 2. Dorsal spinule patch, aedeagus, close up, 3. Aedeagus, lateral, 4. Aedeagus, anterior, 5. Aedeagus, spinule patch, close up, 6. Stout spinules, aedeagus, posterior at base, 7. Large aedeagus spinules, posterior laterally, at base, 8. Digitate process on aedeagus, located in apical cleft." captionText-1="Figs. 9‐16. Isoperla laucki: 9. Male paraproct, right, 10. Subgenital plate, female, 11. Nymphal right mandible, ventral, flipped, 12. Nymphal right mandible, ventral, close up, 13. Nymphal right lacinia, ventral, 14. Nymphal right lacinia, ventral, close up, 15. Egg, lateral, 16. Egg, end view." figureDoi-0="http://doi.org/10.5281/zenodo.4758992" figureDoi-1="http://doi.org/10.5281/zenodo.4758994" httpUri-0="https://zenodo.org/record/4758992/files/figure.png" httpUri-1="https://zenodo.org/record/4758994/files/figure.png" pageId="1" pageNumber="2">Figs. 1‐16</figureCitation>
, 
<figureCitation box="[542,569,554,578]" captionStart="Figure" captionStartId="8.[189,269,873,897]" captionTargetBox="[249,1383,203,817]" captionTargetId="figure-42@8.[249,1383,203,817]" captionTargetPageId="8" captionText="Figure. 33. Isoperla laucki. Mature nymph, dorsal, habitus." figureDoi="http://doi.org/10.5281/zenodo.4759000" httpUri="https://zenodo.org/record/4759000/files/figure.png" pageId="1" pageNumber="2">33</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="2" type="materials_examined">
<paragraph blockId="1.[189,798,624,1928]" pageId="1" pageNumber="2">
<emphasis bold="true" box="[189,430,625,649]" pageId="1" pageNumber="2">Material examined.</emphasis>
<materialsCitation ID-GBIF-Occurrence="3111527308" collectingDate="2007-08-23" collectorName="J. J. Lee" county="Humboldt Co." latitude="1.23" location="Dragsaw Spring @ Rd 13 N 01" municipality="Rd" pageId="1" pageNumber="2" specimenCount="1" specimenCount-male="1" stateProvince="California" typeStatus="holotype">
<typeStatus box="[444,555,625,649]" pageId="1" pageNumber="2">Holotype</typeStatus>
<specimenCount box="[568,588,624,649]" pageId="1" pageNumber="2" type="male">♂</specimenCount>
from 
<collectingRegion box="[672,792,625,649]" pageId="1" pageNumber="2">California</collectingRegion>
, 
<collectingCounty box="[189,359,661,685]" pageId="1" pageNumber="2">Humboldt Co.</collectingCounty>
, 
<location LSID="urn:lsid:plazi:treatment:2373BB7A3F7E3F0D8C7A1B7FFAA1F20E:AE055CB73F7E3F088C7A1BECFD02F58F" box="[369,699,661,685]" county="Humboldt Co." latitude="1.23" municipality="Rd" name="Dragsaw Spring @ Rd 13 N 01" pageId="1" pageNumber="2" stateProvince="California">Dragsaw Spring @ Rd 13N01</location>
, 
<date pageId="1" pageNumber="2" value="2007-08-23">
<collectingDate pageId="1" pageNumber="2" value="2007-08-23">23‐VIII‐ 07</collectingDate>
</date>
, 
<collectorName box="[244,343,697,720]" pageId="1" pageNumber="2">J.J. Lee</collectorName>
</materialsCitation>
. 
<materialsCitation ID-GBIF-Occurrence="3111527301" collectorName="J. J. Lee &amp; Collector &amp; California &amp; Red Mountain Ck." county="Humboldt Co." latitude="12.4" location="Rd" pageId="1" pageNumber="2" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus box="[368,485,697,721]" pageId="1" pageNumber="2">Paratypes</typeStatus>
(
<collectorName box="[515,613,696,720]" pageId="1" pageNumber="2">J.J. Lee</collectorName>
, 
<collectorName box="[639,750,696,720]" pageId="1" pageNumber="2">Collector</collectorName>
): – 
<collectorName box="[189,309,732,756]" pageId="1" pageNumber="2">California</collectorName>
, 
<collectingCounty box="[322,496,732,756]" pageId="1" pageNumber="2">Humboldt Co.</collectingCounty>
, 
<collectorName box="[509,729,732,756]" pageId="1" pageNumber="2">Red Mountain Ck.</collectorName>
@ 
<location LSID="urn:lsid:plazi:treatment:2373BB7A3F7E3F0D8C7A1B7FFAA1F20E:AE055CB73F7E3F088FF01BA5FCA4F5D6" box="[763,797,732,756]" county="Humboldt Co." latitude="12.4" name="Rd" pageId="1" pageNumber="2">Rd</location>
10 
<geoCoordinate box="[217,293,767,791]" degrees="12, 4" direction="north" orientation="latitude" pageId="1" pageNumber="2" precision="5555" value="12.4">
N12, 
<date box="[280,386,767,791]" pageId="1" pageNumber="2" value="2005-08-04">4‐VIII‐05</date>
</geoCoordinate>
, 
<specimenCount box="[399,440,766,791]" pageId="1" pageNumber="2" type="female">1 ♀</specimenCount>
</materialsCitation>
; 10‐VIII‐05, 
<specimenCount box="[588,630,766,791]" pageId="1" pageNumber="2" type="female">1 ♀</specimenCount>
; 20‐X‐05, 
<specimenCount box="[749,790,766,791]" pageId="1" pageNumber="2" type="female">1 ♀</specimenCount>
; 14‐XII‐05, 
<specimenCount box="[311,408,803,827]" pageId="1" pageNumber="2" type="larva">9 larvae</specimenCount>
; 23‐VI‐06, 
<specimenCount box="[536,579,802,827]" pageId="1" pageNumber="2" type="female">3 ♀</specimenCount>
; 3‐VII‐06, 
<specimenCount box="[704,746,802,827]" pageId="1" pageNumber="2" type="female">2 ♀</specimenCount>
; 25‐ VIII‐06, 
<specimenCount box="[283,321,837,862]" pageId="1" pageNumber="2" type="male">6 ♂</specimenCount>
, 
<specimenCount box="[333,371,837,862]" pageId="1" pageNumber="2" type="female">1 ♀</specimenCount>
, 
<specimenCount box="[383,476,838,862]" pageId="1" pageNumber="2" type="larva">4 larvae</specimenCount>
; 25‐VII‐07, 
<specimenCount box="[606,644,837,862]" pageId="1" pageNumber="2" type="female">1 ♀</specimenCount>
, 
<specimenCount box="[656,749,838,862]" pageId="1" pageNumber="2" type="larva">2 larvae</specimenCount>
, 
<specimenCount box="[759,797,837,862]" pageId="1" pageNumber="2" type="male">1 ♂</specimenCount>
(reared 9‐VIII‐07); 8‐VIII‐07, 
<specimenCount box="[519,557,873,898]" pageId="1" pageNumber="2" type="male">1 ♂</specimenCount>
, 
<specimenCount box="[569,648,874,898]" pageId="1" pageNumber="2" type="larva">1 larva</specimenCount>
; 23‐VIII‐07, 
<specimenCount pageId="1" pageNumber="2" type="female">1 ♀</specimenCount>
, 
<specimenCount box="[223,263,908,933]" pageId="1" pageNumber="2" type="male">1 ♂</specimenCount>
&amp; 
<specimenCount box="[297,337,908,933]" pageId="1" pageNumber="2" type="female">1 ♀</specimenCount>
(reared 26‐VIII‐07), 
<materialsCitation ID-GBIF-Occurrence="3111527311" collectingDate="2007-08-27" collectorName="J. J. Lee &amp; Collector &amp; California &amp; Red Mountain Ck." county="Humboldt Co." isTailEnumeration="true" latitude="12.4" location="Rd" pageId="1" pageNumber="2" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount box="[575,608,908,933]" pageId="1" pageNumber="2" type="male">1♂</specimenCount>
(reared 
<date pageId="1" pageNumber="2" value="2007-08-27">
<collectingDate pageId="1" pageNumber="2" value="2007-08-27">27‐VIII‐ 07</collectingDate>
</date>
)
</materialsCitation>
, 
<specimenCount box="[241,284,944,969]" pageId="1" pageNumber="2" type="female">1 ♀</specimenCount>
(reared 28‐VIII‐07); 29‐VIII‐07, 
<specimenCount box="[663,748,945,969]" pageId="1" pageNumber="2" type="larva">1 larva</specimenCount>
; 11‐ VIII‐08, 
<specimenCount box="[287,386,980,1004]" pageId="1" pageNumber="2" type="larva">3 larvae</specimenCount>
, 
<specimenCount box="[401,444,979,1004]" pageId="1" pageNumber="2" type="female">1 ♀</specimenCount>
(reared 29‐VIII‐08); Dragsaw Spring @ Rd 13 
<geoCoordinate box="[366,454,1016,1040]" degrees="01, 25" direction="north" orientation="latitude" pageId="1" pageNumber="2" precision="555" value="1.25">N01, 25</geoCoordinate>
‐VII‐07, 
<specimenCount box="[548,588,1015,1040]" pageId="1" pageNumber="2" type="male">1 ♂</specimenCount>
, 
<specimenCount box="[601,641,1015,1040]" pageId="1" pageNumber="2" type="female">1 ♀</specimenCount>
; 29‐VIII‐07, 
<specimenCount pageId="1" pageNumber="2" type="female">1 ♀</specimenCount>
; 11‐VIII‐08, 
<specimenCount box="[350,388,1050,1075]" pageId="1" pageNumber="2" type="female">1 ♀</specimenCount>
; 28‐VIII‐08, 
<specimenCount box="[528,566,1050,1075]" pageId="1" pageNumber="2" type="male">1 ♂</specimenCount>
; 
<materialsCitation ID-GBIF-Occurrence="3111527309" collectingDate="2007-03-21" location="Boise Ck." municipality="Hwy." pageId="1" pageNumber="2" specimenCount="1" specimenCount-larva="1" typeStatus="paratype">
spring east of 
<location LSID="urn:lsid:plazi:treatment:2373BB7A3F7E3F0D8C7A1B7FFAA1F20E:AE055CB73F7E3F088FEB1D62FF51F375" municipality="Hwy." name="Boise Ck." pageId="1" pageNumber="2">Boise Ck.</location>
, 
<collectingMunicipality box="[244,309,1087,1111]" pageId="1" pageNumber="2">Hwy.</collectingMunicipality>
299, 
<date box="[371,471,1087,1111]" pageId="1" pageNumber="2" value="2007-03-21">
<collectingDate box="[371,471,1087,1111]" pageId="1" pageNumber="2" value="2007-03-21">21‐III‐07</collectingDate>
</date>
, 
<specimenCount box="[483,566,1087,1111]" pageId="1" pageNumber="2" type="larva">1 larva</specimenCount>
</materialsCitation>
; 
<materialsCitation ID-GBIF-Occurrence="3111527307" collectingDate="2007-07-03" location="East Fork Willow Creek Campground" pageId="1" pageNumber="2" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
spring @ 
<location LSID="urn:lsid:plazi:treatment:2373BB7A3F7E3F0D8C7A1B7FFAA1F20E:AE055CB73F7E3F088FA51D46FDBDF358" name="East Fork Willow Creek Campground" pageId="1" pageNumber="2">East Fork Willow Creek Campground</location>
, 
<date box="[526,623,1122,1146]" pageId="1" pageNumber="2" value="2007-07-03">
<collectingDate box="[526,623,1122,1146]" pageId="1" pageNumber="2" value="2007-07-03">3‐VII‐07</collectingDate>
</date>
, 
<specimenCount box="[633,671,1121,1146]" pageId="1" pageNumber="2" type="male">1 ♂</specimenCount>
</materialsCitation>
; 10‐VII‐07, 
<specimenCount box="[189,232,1157,1182]" pageId="1" pageNumber="2" type="male">1 ♂</specimenCount>
, 
<specimenCount box="[248,291,1157,1182]" pageId="1" pageNumber="2" type="female">1 ♀</specimenCount>
; 28‐V‐08, 
<specimenCount box="[414,457,1157,1182]" pageId="1" pageNumber="2" type="female">1 ♀</specimenCount>
; 9‐VI‐08, 
<specimenCount box="[575,673,1158,1182]" pageId="1" pageNumber="2" type="larva">2 larvae</specimenCount>
. 
<materialsCitation ID-GBIF-Occurrence="3111527303" location="California Academy of Sciences" pageId="1" pageNumber="2" specimenCount="1" typeStatus="holotype">
<typeStatus box="[686,797,1158,1182]" pageId="1" pageNumber="2">Holotype</typeStatus>
deposited at the California Academy of Sciences
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="5" pageId="1" pageNumber="2" type="description">
<paragraph blockId="1.[189,798,624,1928]" lastBlockId="1.[835,1444,234,1928]" pageId="1" pageNumber="2">
<emphasis bold="true" box="[189,258,1229,1253]" pageId="1" pageNumber="2">Male.</emphasis>
Macropterous. Length of forewings 10.0‐11.0 mm; length of body 9.0‐10.0 mm. General body color golden yellow. Head yellow, light U‐ shaped pattern forward of anterior ocellus, orange brown patch inside “U” extends to frontal margin; interocellar area light orange brown; occiput orange brown mesally behind lateral ocelli, occipital suture lines light; posterior corners behind eyes with brown rugose patch and scattered short spinules. Antennae yellow at base, infuscate distally. Pronotum widest anteriorly, with narrow band of light brown rugosities lateral to median, yellow stripe. Wings slightly hyaline yellow, veins brown, and fumose yellow in costal space. Legs yellow, covered with short spines, tibiae with incomplete dorsal brown band in proximal one‐ fourth. Abdomen bright yellow, vesicle on eighth sternum yellow orange, with scattered short setae, longer setae on posterolateral margins, shallowly rounded posteriorly and appearing oval, width approximately 
<subSection lastPageId="4" lastPageNumber="5" pageId="1" pageNumber="2" type="multiple">
<heading box="[835,1443,198,223]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="2" reason="4">1.5 to 2 times length. Tenth tergum with</heading>
posteromedian depression and anteromedian unsclerotized area. Cerci golden brown with scattered short setae, one or two ventral setae approaching 1/2 segment length distally on each segment. Paraprocts bluntly pointed, curving slightly over tergum 10 (
<figureCitation box="[1030,1096,411,435]" captionStart="Figs" captionStartId="3.[189,240,1796,1820]" captionTargetBox="[249,1383,189,1748]" captionTargetId="figure-42@3.[249,1383,189,1748]" captionTargetPageId="3" captionText="Figs. 9‐16. Isoperla laucki: 9. Male paraproct, right, 10. Subgenital plate, female, 11. Nymphal right mandible, ventral, flipped, 12. Nymphal right mandible, ventral, close up, 13. Nymphal right lacinia, ventral, 14. Nymphal right lacinia, ventral, close up, 15. Egg, lateral, 16. Egg, end view." figureDoi="http://doi.org/10.5281/zenodo.4758994" httpUri="https://zenodo.org/record/4758994/files/figure.png" pageId="1" pageNumber="2">Fig. 9</figureCitation>
). Aedeagus (
<figureCitation box="[1249,1374,411,435]" captionStart="Figs" captionStartId="2.[189,240,1796,1820]" captionTargetBox="[249,1384,189,1747]" captionTargetId="figure-42@2.[249,1384,189,1749]" captionTargetPageId="2" captionText="Figs. 1‐8. Isoperla laucki: 1. Aedeagus, dorsal, 2. Dorsal spinule patch, aedeagus, close up, 3. Aedeagus, lateral, 4. Aedeagus, anterior, 5. Aedeagus, spinule patch, close up, 6. Stout spinules, aedeagus, posterior at base, 7. Large aedeagus spinules, posterior laterally, at base, 8. Digitate process on aedeagus, located in apical cleft." figureDoi="http://doi.org/10.5281/zenodo.4758992" httpUri="https://zenodo.org/record/4758992/files/figure.png" pageId="1" pageNumber="2">Figs. 1, 3‐4</figureCitation>
) with an elliptical‐ shaped patch of stout, brown spinules, located dorsally, patch ranging from 2 to 3 times as long as wide (
<figureCitation box="[994,1114,518,542]" captionStart="Figs" captionStartId="2.[189,240,1796,1820]" captionTargetBox="[249,1384,189,1747]" captionTargetId="figure-42@2.[249,1384,189,1749]" captionTargetPageId="2" captionText="Figs. 1‐8. Isoperla laucki: 1. Aedeagus, dorsal, 2. Dorsal spinule patch, aedeagus, close up, 3. Aedeagus, lateral, 4. Aedeagus, anterior, 5. Aedeagus, spinule patch, close up, 6. Stout spinules, aedeagus, posterior at base, 7. Large aedeagus spinules, posterior laterally, at base, 8. Digitate process on aedeagus, located in apical cleft." figureDoi="http://doi.org/10.5281/zenodo.4758992" httpUri="https://zenodo.org/record/4758992/files/figure.png" pageId="1" pageNumber="2">Figs. 2 &amp; 5</figureCitation>
); posterobasal patch of short stout, orange brown spinules, forming sinuate rows (
<figureCitation box="[843,914,589,613]" captionStart="Figs" captionStartId="2.[189,240,1796,1820]" captionTargetBox="[249,1384,189,1747]" captionTargetId="figure-42@2.[249,1384,189,1749]" captionTargetPageId="2" captionText="Figs. 1‐8. Isoperla laucki: 1. Aedeagus, dorsal, 2. Dorsal spinule patch, aedeagus, close up, 3. Aedeagus, lateral, 4. Aedeagus, anterior, 5. Aedeagus, spinule patch, close up, 6. Stout spinules, aedeagus, posterior at base, 7. Large aedeagus spinules, posterior laterally, at base, 8. Digitate process on aedeagus, located in apical cleft." figureDoi="http://doi.org/10.5281/zenodo.4758992" httpUri="https://zenodo.org/record/4758992/files/figure.png" pageId="1" pageNumber="2">Fig. 6</figureCitation>
); a stronger patch of posteromedial orange brown spinules (
<figureCitation box="[1056,1133,624,649]" captionStart="Figs" captionStartId="2.[189,240,1796,1820]" captionTargetBox="[249,1384,189,1747]" captionTargetId="figure-42@2.[249,1384,189,1749]" captionTargetPageId="2" captionText="Figs. 1‐8. Isoperla laucki: 1. Aedeagus, dorsal, 2. Dorsal spinule patch, aedeagus, close up, 3. Aedeagus, lateral, 4. Aedeagus, anterior, 5. Aedeagus, spinule patch, close up, 6. Stout spinules, aedeagus, posterior at base, 7. Large aedeagus spinules, posterior laterally, at base, 8. Digitate process on aedeagus, located in apical cleft." figureDoi="http://doi.org/10.5281/zenodo.4758992" httpUri="https://zenodo.org/record/4758992/files/figure.png" pageId="1" pageNumber="2">Fig. 7</figureCitation>
) becoming smaller and scattered laterally; aedeagus narrowing apically, cleft at apex, with a short, digitate, lightly sclerotized process arising from cleft (
<figureCitation box="[1146,1271,731,755]" captionStart="Figs" captionStartId="2.[189,240,1796,1820]" captionTargetBox="[249,1384,189,1747]" captionTargetId="figure-42@2.[249,1384,189,1749]" captionTargetPageId="2" captionText="Figs. 1‐8. Isoperla laucki: 1. Aedeagus, dorsal, 2. Dorsal spinule patch, aedeagus, close up, 3. Aedeagus, lateral, 4. Aedeagus, anterior, 5. Aedeagus, spinule patch, close up, 6. Stout spinules, aedeagus, posterior at base, 7. Large aedeagus spinules, posterior laterally, at base, 8. Digitate process on aedeagus, located in apical cleft." figureDoi="http://doi.org/10.5281/zenodo.4758992" httpUri="https://zenodo.org/record/4758992/files/figure.png" pageId="1" pageNumber="2">Figs. 4 &amp; 8</figureCitation>
). 
<paragraph blockId="1.[835,1444,234,1928]" pageId="1" pageNumber="2">
<emphasis bold="true" box="[835,928,767,791]" pageId="1" pageNumber="2">Female.</emphasis>
Macropterous. Length of forewings 
<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.25" metricValueMin="1.15" pageId="1" pageNumber="2" unit="mm" value="12.0" valueMax="12.5" valueMin="11.5">11.5‐12.5 mm</quantity>
; length of body 10.0‐11.0 mm. General body color, head and pronotal pigmentation patterns similar to male. Subgenital plate broadly triangular but rounded apically, extending at least over one‐ half length of ninth sternum (
<figureCitation box="[1129,1210,944,969]" captionStart="Figs" captionStartId="3.[189,240,1796,1820]" captionTargetBox="[249,1383,189,1748]" captionTargetId="figure-42@3.[249,1383,189,1748]" captionTargetPageId="3" captionText="Figs. 9‐16. Isoperla laucki: 9. Male paraproct, right, 10. Subgenital plate, female, 11. Nymphal right mandible, ventral, flipped, 12. Nymphal right mandible, ventral, close up, 13. Nymphal right lacinia, ventral, 14. Nymphal right lacinia, ventral, close up, 15. Egg, lateral, 16. Egg, end view." figureDoi="http://doi.org/10.5281/zenodo.4758994" httpUri="https://zenodo.org/record/4758994/files/figure.png" pageId="1" pageNumber="2">Fig. 10</figureCitation>
).
</paragraph>
<paragraph blockId="1.[835,1444,234,1928]" pageId="1" pageNumber="2">
<emphasis bold="true" box="[835,881,980,1004]" pageId="1" pageNumber="2">Egg</emphasis>
. Shape nearly triangular in cross section, short and angular, with five nearly equal sides (
<figureCitation captionStart="Figs" captionStartId="3.[189,240,1796,1820]" captionTargetBox="[249,1383,189,1748]" captionTargetId="figure-42@3.[249,1383,189,1748]" captionTargetPageId="3" captionText="Figs. 9‐16. Isoperla laucki: 9. Male paraproct, right, 10. Subgenital plate, female, 11. Nymphal right mandible, ventral, flipped, 12. Nymphal right mandible, ventral, close up, 13. Nymphal right lacinia, ventral, 14. Nymphal right lacinia, ventral, close up, 15. Egg, lateral, 16. Egg, end view." figureDoi="http://doi.org/10.5281/zenodo.4758994" httpUri="https://zenodo.org/record/4758994/files/figure.png" pageId="1" pageNumber="2">Figs. 15‐ 16</figureCitation>
). Collar poorly developed and appearing as a flattened cap. Chorion with thick ridges interspersed with deep angular depressions that cover the remainder of the outer surface. Micropyles set on ridges near bottom third of egg. Mature eggs light brown and almost translucent, appearing like small jewels in preserved specimens.
</paragraph>
<paragraph blockId="1.[835,1444,234,1928]" lastBlockId="4.[189,798,198,1928]" lastPageId="4" lastPageNumber="5" pageId="1" pageNumber="2">
<emphasis bold="true" box="[835,910,1300,1324]" pageId="1" pageNumber="2">Larva.</emphasis>
Body length of mature nymph 9.0‐10.0 mm. General body color brown, with dark clothing hairs. Dorsum of head brown, light M‐ shaped pattern forward of median ocellus, frontal margin pale, small oval, light patch between frontal pale band and peaks of “M” pattern, similar light patch between median ocellus and compound eyes; interocellar area brown, occiput with light, oval patch adjacent to eyes; light patch posteriorly on occiput broken by dark reticulate lines; sinuate band of occipital spinules separating the two light patches; posterior corners behind compound eyes brown with scattered spinules (
<figureCitation box="[948,1029,1726,1751]" captionStart="Figure" captionStartId="8.[189,269,873,897]" captionTargetBox="[249,1383,203,817]" captionTargetId="figure-42@8.[249,1383,203,817]" captionTargetPageId="8" captionText="Figure. 33. Isoperla laucki. Mature nymph, dorsal, habitus." figureDoi="http://doi.org/10.5281/zenodo.4759000" httpUri="https://zenodo.org/record/4759000/files/figure.png" pageId="1" pageNumber="2">Fig. 33</figureCitation>
). Antennae light brown with apical circlets of short setae; dark clothing hairs on basal segments. Lacinia triangular, bidentate; subapical tooth ca. 1/2 length of apical tooth; 1 or 2 axillary setae, 6‐8 stout marginal setae below subapical tooth, setae 7 and/or 8 small; 2 fine submarginal setae below subapical tooth; sparse, irregularly spaced fine marginal and submarginal setae below stout marginal setae (
<figureCitation box="[375,501,269,293]" captionStart="Figs" captionStartId="3.[189,240,1796,1820]" captionTargetBox="[249,1383,189,1748]" captionTargetId="figure-42@3.[249,1383,189,1748]" captionTargetPageId="3" captionText="Figs. 9‐16. Isoperla laucki: 9. Male paraproct, right, 10. Subgenital plate, female, 11. Nymphal right mandible, ventral, flipped, 12. Nymphal right mandible, ventral, close up, 13. Nymphal right lacinia, ventral, 14. Nymphal right lacinia, ventral, close up, 15. Egg, lateral, 16. Egg, end view." figureDoi="http://doi.org/10.5281/zenodo.4758994" httpUri="https://zenodo.org/record/4758994/files/figure.png" pageId="4" pageNumber="5">Figs. 13‐14</figureCitation>
). Mandibles with 6 cusps in 2 groups of 3; at least middle cusp in each group serrate; median row of long medium stout setae below outer apical cusp to near mandibular base; inner marginal fringe of 40+ long stout setae; right mandible with row of short stout setae below inner 3rd apical cusp (
<figureCitation box="[382,461,482,506]" captionStart="Figs" captionStartId="3.[189,240,1796,1820]" captionTargetBox="[249,1383,189,1748]" captionTargetId="figure-42@3.[249,1383,189,1748]" captionTargetPageId="3" captionText="Figs. 9‐16. Isoperla laucki: 9. Male paraproct, right, 10. Subgenital plate, female, 11. Nymphal right mandible, ventral, flipped, 12. Nymphal right mandible, ventral, close up, 13. Nymphal right lacinia, ventral, 14. Nymphal right lacinia, ventral, close up, 15. Egg, lateral, 16. Egg, end view." figureDoi="http://doi.org/10.5281/zenodo.4758994" httpUri="https://zenodo.org/record/4758994/files/figure.png" pageId="4" pageNumber="5">Fig. 11</figureCitation>
), setae longer and row more extensive on left mandible. Pronotum oval, width slightly narrower than head, brown with reticulate light markings and dark brown border, slightly lighter brown laterally; margin fringed with small stout setae, occasionally longer setae at upper and lower angles and lateral margin. Meso and metanota brown with reticulate markings. Legs light brown with scattered variable sized stout setae, with sparse dorsal fringe of silky setae (
<figureCitation box="[515,597,802,826]" captionStart="Figure" captionStartId="8.[189,269,873,897]" captionTargetBox="[249,1383,203,817]" captionTargetId="figure-42@8.[249,1383,203,817]" captionTargetPageId="8" captionText="Figure. 33. Isoperla laucki. Mature nymph, dorsal, habitus." figureDoi="http://doi.org/10.5281/zenodo.4759000" httpUri="https://zenodo.org/record/4759000/files/figure.png" pageId="4" pageNumber="5">Fig. 33</figureCitation>
). Thoracic sterna with distinct mesal sclera with dark clothing hairs; mesosternal Y‐ ridge arms extend to posterior corners of furcal pits; with band of erect bristles scattered across anterior surface of mesosternum. Abdominal segments dark brown; terga with paired mesal and lateral light spots; sometimes a third light spot evident forming a mesal triangle with anteromedial pair; lateral light spots often obscured posterior to first few tergites. Cerci with ca. 28 segments; each with posterior circlet of setae, lacking setal fringe.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4758992" ID-Zenodo-Dep="4758992" httpUri="https://zenodo.org/record/4758992/files/figure.png" pageId="2" pageNumber="3" startId="2.[189,240,1796,1820]" targetBox="[249,1384,189,1747]" targetPageId="2">
<paragraph blockId="2.[189,1444,1796,1892]" pageId="2" pageNumber="3">
Figs. 1‐8. 
<taxonomicName authorityName="Baumann &amp; Lee" authorityYear="2009" box="[296,450,1796,1820]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="laucki">
<emphasis box="[296,450,1796,1820]" italics="true" pageId="2" pageNumber="3">Isoperla laucki</emphasis>
</taxonomicName>
: 1. Aedeagus, dorsal, 2. Dorsal spinule patch, aedeagus, close up, 3. Aedeagus, lateral, 4. Aedeagus, anterior, 5. Aedeagus, spinule patch, close up, 6. Stout spinules, aedeagus, posterior at base, 7. Large aedeagus spinules, posterior laterally, at base, 8. Digitate process on aedeagus, located in apical cleft.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4758994" ID-Zenodo-Dep="4758994" httpUri="https://zenodo.org/record/4758994/files/figure.png" pageId="3" pageNumber="4" startId="3.[189,240,1796,1820]" targetBox="[249,1383,189,1748]" targetPageId="3">
<paragraph blockId="3.[189,1443,1796,1892]" pageId="3" pageNumber="4">
Figs. 9‐16. 
<taxonomicName authorityName="Baumann &amp; Lee" authorityYear="2009" box="[312,467,1796,1820]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="laucki">
<emphasis box="[312,467,1796,1820]" italics="true" pageId="3" pageNumber="4">Isoperla laucki</emphasis>
</taxonomicName>
: 9. Male paraproct, right, 10. Subgenital plate, female, 11. Nymphal right mandible, ventral, flipped, 12. Nymphal right mandible, ventral, close up, 13. Nymphal right lacinia, ventral, 14. Nymphal right lacinia, ventral, close up, 15. Egg, lateral, 16. Egg, end view.
</paragraph>
</caption>
<subSubSection pageId="4" pageNumber="5" type="etymology">
<paragraph blockId="4.[189,798,198,1928]" pageId="4" pageNumber="5">
<emphasis bold="true" box="[189,327,1193,1217]" pageId="4" pageNumber="5">Etymology.</emphasis>
We are proud to name this species in honor of our friend, teacher, and colleague David R. Lauck, who taught for many years at Humboldt State University in Arcata, California. He instilled in us the excitement and desire to study the stonefly fauna of the Willow Creek area and the greater TrinityKlamath River drainage. Humboldt Stripetail is suggested as the common name.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="5" type="diagnosis">
<paragraph blockId="4.[189,798,198,1928]" lastBlockId="4.[835,1443,198,1324]" pageId="4" pageNumber="5">
<emphasis bold="true" box="[189,311,1477,1501]" pageId="4" pageNumber="5">Diagnosis</emphasis>
. This species is unique among described western 
<taxonomicName box="[287,370,1513,1537]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis box="[287,370,1513,1537]" italics="true" pageId="4" pageNumber="5">Isoperla</emphasis>
</taxonomicName>
. Adults in the field can be recognized by the golden yellow color, superficially resembling robust 
<taxonomicName authorityName="Ricker" authorityYear="1943" box="[276,359,1584,1608]" class="Insecta" family="Chloroperlidae" genus="Sweltsa" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis box="[276,359,1584,1608]" italics="true" pageId="4" pageNumber="5">Sweltsa</emphasis>
</taxonomicName>
. Males can be distinguished by the oval patch of strong spinules dorsally on the aedeagus and the unique lightly sclerotized digitate apical process (
<figureCitation box="[410,531,1691,1715]" captionStart="Figs" captionStartId="2.[189,240,1796,1820]" captionTargetBox="[249,1384,189,1747]" captionTargetId="figure-42@2.[249,1384,189,1749]" captionTargetPageId="2" captionText="Figs. 1‐8. Isoperla laucki: 1. Aedeagus, dorsal, 2. Dorsal spinule patch, aedeagus, close up, 3. Aedeagus, lateral," figureDoi="http://doi.org/10.5281/zenodo.4758992" httpUri="https://zenodo.org/record/4758992/files/figure.png" pageId="4" pageNumber="5">Figs. 1‐4</figureCitation>
). Nymphs can be distinguished from other western 
<taxonomicName box="[588,762,1726,1750]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis box="[588,672,1726,1750]" italics="true" pageId="4" pageNumber="5">Isoperla</emphasis>
species
</taxonomicName>
by the presence of an M‐ shaped pattern on the head, the lack of abdominal striping (
<figureCitation box="[529,614,1797,1821]" captionStart="Figure" captionStartId="8.[189,269,873,897]" captionTargetBox="[249,1383,203,817]" captionTargetId="figure-42@8.[249,1383,203,817]" captionTargetPageId="8" captionText="Figure. 33. Isoperla laucki. Mature nymph, dorsal, habitus." figureDoi="http://doi.org/10.5281/zenodo.4759000" httpUri="https://zenodo.org/record/4759000/files/figure.png" pageId="4" pageNumber="5">Fig. 33</figureCitation>
), and the erect bristles anteriorly on the mesosternum. Erect bristles are evident in early instars but may be worn off in mature later instars. Nymphs superficially resemble small 
<taxonomicName authority="(Claassen)" baseAuthorityName="Claassen" baseAuthorityYear="1936" box="[904,1286,198,222]" class="Insecta" family="Perlodidae" genus="Chernokrilus" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="misnomus">
<emphasis box="[904,1160,198,222]" italics="true" pageId="4" pageNumber="5">Chernokrilus misnomus</emphasis>
(Claassen)
</taxonomicName>
and 
<taxonomicName authority="(Banks)" baseAuthorityName="Banks" baseAuthorityYear="1906" class="Insecta" family="Perlodidae" genus="Calliperla" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="luctuosa">
<emphasis italics="true" pageId="4" pageNumber="5">Calliperla luctuosa</emphasis>
(Banks)
</taxonomicName>
, however, the small size of late instars, combination of lack of abdominal striping and mesosternal “Y”‐arms meeting the posterior corners of the furcal pits, and erect mesosternal bristles distinguishes them. Females have a rounded subgenital plate that extends over half of sternum nine (
<figureCitation box="[905,988,447,471]" captionStart="Figs" captionStartId="3.[189,240,1796,1820]" captionTargetBox="[249,1383,189,1748]" captionTargetId="figure-42@3.[249,1383,189,1748]" captionTargetPageId="3" captionText="Figs. 9‐16. Isoperla laucki: 9. Male paraproct, right, 10. Subgenital plate, female, 11. Nymphal right mandible, ventral, flipped, 12. Nymphal right mandible, ventral, close up, 13. Nymphal right lacinia, ventral, 14. Nymphal right lacinia, ventral, close up, 15. Egg, lateral, 16. Egg, end view." figureDoi="http://doi.org/10.5281/zenodo.4758994" httpUri="https://zenodo.org/record/4758994/files/figure.png" pageId="4" pageNumber="5">Fig. 10</figureCitation>
). The eggs have a distinctive angular shape, with five nearly equal sides (
<figureCitation box="[1256,1382,482,506]" captionStart="Figs" captionStartId="3.[189,240,1796,1820]" captionTargetBox="[249,1383,189,1748]" captionTargetId="figure-42@3.[249,1383,189,1748]" captionTargetPageId="3" captionText="Figs. 9‐16. Isoperla laucki: 9. Male paraproct, right, 10. Subgenital plate, female, 11. Nymphal right mandible, ventral, flipped, 12. Nymphal right mandible, ventral, close up, 13. Nymphal right lacinia, ventral, 14. Nymphal right lacinia, ventral, close up, 15. Egg, lateral, 16. Egg, end view." figureDoi="http://doi.org/10.5281/zenodo.4758994" httpUri="https://zenodo.org/record/4758994/files/figure.png" pageId="4" pageNumber="5">Figs. 15‐16</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="5" type="discussion">
<paragraph blockId="4.[835,1443,198,1324]" pageId="4" pageNumber="5">
<emphasis bold="true" box="[835,952,518,542]" pageId="4" pageNumber="5">Remarks.</emphasis>
This species was found in small, perennial spring creeks between 400 and 900 meters elevation. Summer water temperatures ranged from 8 
<emphasis bold="true" box="[849,860,624,649]" pageId="4" pageNumber="5">°</emphasis>
C to 12 
<emphasis bold="true" box="[958,969,624,649]" pageId="4" pageNumber="5">°</emphasis>
C, summer air temperature regularly exceeded 27 
<emphasis bold="true" box="[987,998,659,684]" pageId="4" pageNumber="5">°</emphasis>
C. Females with mature eggs were found from June to October indicating an extended emergence. Early instar nymphs were collected in December; mid to late instar nymphs in March through August, suggesting a univoltine life cycle. Other 
<taxonomicName box="[916,1046,838,862]" class="Insecta" family="Perlodidae" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="family">Perlodidae</taxonomicName>
found with 
<taxonomicName authorityName="Baumann &amp; Lee" authorityYear="2009" box="[1207,1300,838,862]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="laucki">
<emphasis box="[1207,1300,838,862]" italics="true" pageId="4" pageNumber="5">I. laucki</emphasis>
</taxonomicName>
include: 
<taxonomicName baseAuthorityName="Banks" baseAuthorityYear="1906" class="Insecta" family="Perlodidae" genus="Calliperla" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="luctuosa">
<emphasis italics="true" pageId="4" pageNumber="5">C. luctuosa</emphasis>
</taxonomicName>
, 
<taxonomicName baseAuthorityName="Claassen" baseAuthorityYear="1936" box="[953,1109,874,897]" class="Insecta" family="Perlodidae" genus="Chernokrilus" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="misnomus">
<emphasis box="[953,1109,874,897]" italics="true" pageId="4" pageNumber="5">C. misnomus</emphasis>
</taxonomicName>
, and 
<taxonomicName authority="Baumann &amp; Lauck." authorityName="Baumann &amp; Lauck." class="Insecta" family="Perlodidae" genus="Salmoperla" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="sylvanica">
<emphasis box="[1198,1442,873,897]" italics="true" pageId="4" pageNumber="5">Salmoperla sylvanica</emphasis>
Baumann &amp; Lauck.
</taxonomicName>
</paragraph>
<paragraph blockId="4.[835,1443,198,1324]" pageId="4" pageNumber="5">
“Drumming” behavior was observed in 
<specimenCount pageId="4" pageNumber="5" type="male">one male</specimenCount>
specimen. A laboratory reared female was placed in a container with a field caught male. After approximately 25 hours the female came out of hiding and mating occurred within 15 minutes. The male was observed to touch his vesicle to the substrate 3 to 5 times then raise his abdomen above horizontal and vibrate it before rapidly bringing it back down. This apparent “tremulation” has not been previously reported for 
<taxonomicName authority="(Stewart 2001)" baseAuthorityName="Stewart" baseAuthorityYear="2001" box="[1182,1443,1264,1288]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis box="[1182,1269,1264,1288]" italics="true" pageId="4" pageNumber="5">Isoperla</emphasis>
(
<bibRefCitation author="Stewart, K. W." box="[1282,1435,1264,1288]" pageId="4" pageNumber="5" refId="ref4113" refString="Stewart, K. W. 2001. Vibrational communication (drumming) and mate searching behavior of stoneflies; Evolutionary considerations. pp. 217 - 226 in Trends in research in Ephemeroptera and Plecoptera. Dominguez E., (ed.) Kluwer Academic / Plenum Publishers, NY." type="book" year="2001">Stewart 2001</bibRefCitation>
)
</taxonomicName>
and further investigation is warranted.
</paragraph>
</subSubSection>
</subSection>
</paragraph>
</subSubSection>
</treatment>
</document>