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<document id="E86177A1985B5BF126A66E214B473861" ID-CLB-Dataset="29652" ID-DOI="http://dx.doi.org/10.3897/zookeys.777.23212" ID-GBIF-Dataset="516d998c-117e-4cbb-9e21-e38843778702" ID-PMC="PMC6079114" ID-Pensoft-Pub="1313-2970-777-57" ID-PubMed="30100790" ID-ZooBank="D9F353643DCD4BA6B70D62FB275DEB1B" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-777-57" ModsDocOrigin="ZooKeys 777" ModsDocTitle="A review of the Cholevinae from the island of Borneo (Coleoptera, Leiodidae)" checkinTime="1533035384019" checkinUser="pensoft" docAuthor="Schilthuizen, Menno, Perreau, Michel &amp; Njunjic, Iva" docDate="2018" docId="167D81E796E2CEE92F72E35F3216139B" docLanguage="en" docName="ZooKeys 777: 57-108" docOrigin="ZooKeys 777" docSource="http://dx.doi.org/10.3897/zookeys.777.23212" docTitle="Ptomaphaginus latimanus Schilthuizen &amp; Perreau 2008" docType="treatment" docVersion="6" lastPageNumber="73" masterDocId="F179FFE1BF6AFFEEEF68FFF0FFB5FFBD" masterDocTitle="A review of the Cholevinae from the island of Borneo (Coleoptera, Leiodidae)" masterLastPageNumber="108" masterPageNumber="57" pageNumber="71" updateTime="1701383097238" updateUser="ExternalLinkService">
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<mods:title id="3132784BCD64496ED9F6634FD8140838">A review of the Cholevinae from the island of Borneo (Coleoptera, Leiodidae)</mods:title>
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<mods:namePart id="6475041B4F0971635143DE01DC9C0E5E">Schilthuizen, Menno</mods:namePart>
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<mods:namePart id="4F8EC52F525AE3B24C4DE19A0D8C97C6">Perreau, Michel</mods:namePart>
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<mods:namePart id="E95072E1E59E5306FA1903352F63C36B">Njunjic, Iva</mods:namePart>
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<treatment id="167D81E796E2CEE92F72E35F3216139B" ID-GBIF-Taxon="146746179" LSID="urn:lsid:plazi:treatment:167D81E796E2CEE92F72E35F3216139B" httpUri="http://treatment.plazi.org/id/167D81E796E2CEE92F72E35F3216139B" lastPageId="16" lastPageNumber="73" pageId="14" pageNumber="71">
<subSubSection id="FA77F3175033766667870FAD4E70B026" pageId="14" pageNumber="71" type="nomenclature">
<paragraph id="F37B1878606960C7F38461533177A8FE" pageId="14" pageNumber="71">
<taxonomicName id="3CFA9C23482179E3A2A79EAA2FDEE59D" ID-CoL="78M3C" authority="Schilthuizen &amp; Perreau, 2008" authorityName="Schilthuizen &amp; Perreau" authorityYear="2008" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphaginus latimanus" order="Coleoptera" pageId="14" pageNumber="71" phylum="Arthropoda" rank="species" species="latimanus">Ptomaphaginus latimanus Schilthuizen &amp; Perreau, 2008</taxonomicName>
Figures 5i, 6g, h, 9j, 10h
</paragraph>
</subSubSection>
<subSubSection id="7DC0AB05D157203DEED05D19010359E5" pageId="15" pageNumber="72" type="reference_group">
<paragraph id="B745675EEC429312B4710762B822FA0D" pageId="15" pageNumber="72">
<taxonomicName id="24BC11390D2AEB18E897D81E4B16CF26" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphaginus latimanus" order="Coleoptera" pageId="15" pageNumber="72" phylum="Arthropoda" rank="species" species="latimanus">
<pageBreakToken id="6DF33412E69B983B6FB50873582547B6" pageId="15" pageNumber="72" start="start">Ptomaphaginus</pageBreakToken>
latimanus
</taxonomicName>
Schilthuizen &amp; Perreau, 2008: 196, figs 22-23; type from Gunung Trus Madi, Sabah, Borneo (in RMNH, RMNH.INS.1086293).
</paragraph>
</subSubSection>
<subSubSection id="BB49D741F911B950147D6A93C61C0C28" pageId="15" pageNumber="72" type="description">
<paragraph id="23EEE59A9E3D0FC008040FD77A09E460" pageId="15" pageNumber="72">Description.</paragraph>
<paragraph id="13D2EDA15C53494BD3BA3BDDD29B6F56" pageId="15" pageNumber="72">
(Adapted from
<bibRefCitation id="26AD7D88AD4109BE0D09B8EA5EC0861F" author="Schilthuizen, M" journalOrPublisher="Zoologische Mededelingen Leiden" pageId="37" pageNumber="94" pagination="189 - 210" title="New species and new records of Ptomaphaginus Portevin from the Sunda region (Coleoptera: Leiodidae: Cholevinae)." volume="82" year="2008">Schilthuizen and Perreau (2008)</bibRefCitation>
). Length 2.3-2.9 mm. Habitus slender, ovoid. Pronotum 1.60-1.75 times as wide as long, as wide as the elytra. Elytra 1.15-1.25 times as long as their combined width (length measured from the caudal tip of the scutellum to the apex of the elytra). Winged. Aedeagus short and wide, with two elongated apical, laterally-directed
<normalizedToken id="74FEE202D9B2F122459111137C0B39D9" originalValue="wings">'wings'</normalizedToken>
and a short terminal processus. Spermatheca narrow, annulated, and bent over a rounded 90° angle, quite similar to that of
<taxonomicName id="2C582A8A7330C0D5ED76242C9BDA3F19" lsidName="P. kinabaluensis" pageId="15" pageNumber="72" rank="species" species="kinabaluensis">P. kinabaluensis</taxonomicName>
. Spermiduct long and narrow, with numerous coils. Antennae short, as long as the width of the head. Long setae on the ventral side of the male profemur and protibia absent. Male with broad and indistinct central notches on the 5th and 6th visible abdominal sternite. Male protarsi strongly dilated.
</paragraph>
</subSubSection>
<subSubSection id="4C5004F696A73F5837B26C0349AE31C7" pageId="15" pageNumber="72" type="differential diagnosis">
<paragraph id="670D06ACE4DD9F85C83EBE00F4AEADD8" pageId="15" pageNumber="72">Differential diagnosis.</paragraph>
<paragraph id="19802C164EF051B31552EE2F9CF1CA6A" pageId="15" pageNumber="72">
<taxonomicName id="61E0D8E25E17F29C645977B4C992D3B7" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphaginus latimanus" order="Coleoptera" pageId="15" pageNumber="72" phylum="Arthropoda" rank="species" species="latimanus">Ptomaphaginus latimanus</taxonomicName>
is closely related to
<taxonomicName id="CAFB6454D2967C1A51F48554C8394495" lsidName="P. kinabaluensis" pageId="15" pageNumber="72" rank="species" species="kinabaluensis">P. kinabaluensis</taxonomicName>
, but differs in the habitus, which is much more stocky in
<taxonomicName id="677A741407768100B268F1DDF177D2C3" lsidName="P. latimanus" pageId="15" pageNumber="72" rank="species" species="latimanus">P. latimanus</taxonomicName>
. Also,
<taxonomicName id="51196801032F19B06FED4A679F772018" lsidName="P. kinabaluensis" pageId="15" pageNumber="72" rank="species" species="kinabaluensis">P. kinabaluensis</taxonomicName>
has extended elytral apices in the female, less strongly dilated male protarsi, and a central extension on the male 4th abdominal sternite.
</paragraph>
</subSubSection>
<subSubSection id="A481DA1545836A15BF89B0950BBC2802" pageId="16" pageNumber="73" type="habitat">
<paragraph id="A613AF5D94C639F745E7DF7E35695B10" pageId="16" pageNumber="73">
<pageBreakToken id="BF58DCB1F41090919D3D5B07EC89B363" pageId="16" pageNumber="73" start="start">Habitat</pageBreakToken>
and distribution.
</paragraph>
<paragraph id="F651FB0B35EE25E8DC10851EACE44A0B" pageId="16" pageNumber="73">
Only known from montane forest at Gunung Trusmadi in Sabah, at 1400 m elev. One
<taxonomicName id="394A2D2B668517CC88F6D000FAA348AA" class="Magnoliopsida" family="Malvaceae" genus="Staphylinoidea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Staphylinoidea bryanti" order="Malvales" pageId="16" pageNumber="73" phylum="Tracheophyta" rank="species" species="bryanti">bryanti</taxonomicName>
-group female (RMNH.INS.555611) from Sayap substation on Gunung Kinabalu is genetically unique (BOLD BIN: ACK0183) and might also belong to this species.
</paragraph>
<caption id="4F2677A6055103CEA4A58B7BAC29BBC3" pageId="16" pageNumber="73">
<paragraph id="4435E077413E30465C5EA26C98FFF53F" pageId="16" pageNumber="73">
Figure 10.
<taxonomicName id="AD0694EC85219B25F0557B5586472FA3" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphaginus" order="Coleoptera" pageId="16" pageNumber="73" phylum="Arthropoda" rank="genus">Ptomaphaginus</taxonomicName>
, female spermiduct and spermatheca. a
<taxonomicName id="B4D489BFD384C94F1EE5D6E13C1B94F6" lsidName="P. bryantioides" pageId="16" pageNumber="73" rank="species" species="bryantioides">P. bryantioides</taxonomicName>
Schilthuizen &amp; Perreau, Gunung Mulu (NHMUK) b
<taxonomicName id="E522AF93612E658A6AD256B92332C112" lsidName="P. grandis" pageId="16" pageNumber="73" rank="species" species="grandis">P. grandis</taxonomicName>
sp. n., Gunung Mulu, paratype (NHMUK) c
<taxonomicName id="86D40656D83F696FE32FF1B378D362D6" lsidName="P. isabellarossellini" pageId="16" pageNumber="73" rank="species" species="isabellarossellini">P. isabellarossellini</taxonomicName>
sp. n., Gunung Kinabalu, paratype (RMNH.INS.1086161) d
<taxonomicName id="CB229DA35E14BC49397F3138E76BB2CA" lsidName="P. louis" pageId="16" pageNumber="73" rank="species" species="louis">P. louis</taxonomicName>
sp. n., Gunung Mulu, paratype (NHMUK) e
<taxonomicName id="A19AA659DFD1681FA70CB4CED586AF65" lsidName="P. muluensis" pageId="16" pageNumber="73" rank="species" species="muluensis">P. muluensis</taxonomicName>
sp. n., Gunung Mulu, paratype (NHMUK) f
<taxonomicName id="28EDE44371197B7454A68AA974DBAE1C" lsidName="P. scaphaner" pageId="16" pageNumber="73" rank="species" species="scaphaner">P. scaphaner</taxonomicName>
Szymczakowski, Gunung Mulu (NHMUK) g
<taxonomicName id="ACC51E6A8404AFF86BF137610F19E9D0" lsidName="P. kinabaluensis" pageId="16" pageNumber="73" rank="species" species="kinabaluensis">P. kinabaluensis</taxonomicName>
Schilthuizen &amp; Perreau, Gunung Kinabalu (TXEX) h
<taxonomicName id="CB9B210C455A9917CD355FCF3B7B31A9" lsidName="P. latimanus" pageId="16" pageNumber="73" rank="species" species="latimanus">P. latimanus</taxonomicName>
Schilthuizen &amp; Perreau, Gunung Trus Madi, paratype (RMNH).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>