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<document id="CC124D33BF384F3928B1033E122BFAA0" ID-CLB-Dataset="34608" ID-DOI="http://dx.doi.org/10.3897/zookeys.478.8057" ID-GBIF-Dataset="f3493113-35ec-4c31-b7cc-b32cd9c717e2" ID-PMC="PMC4319052" ID-Pensoft-Pub="1313-2970-478-45" ID-PubMed="25685010" ID-ZooBank="EC360FAF6BF94FEF96DAF336302D1789" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-478-45" ModsDocOrigin="ZooKeys 478" ModsDocTitle="Phylogeny and biogeography of Asthenopodinae with a revision of Asthenopus, reinstatement of Asthenopodes, and the description of the new genera Hubbardipes and Priasthenopus (Ephemeroptera, Polymitarcyidae)" checkinTime="1451244725713" checkinUser="pensoft" docAuthor="Molineri, Carlos, Salles, Frederico F. &amp; Peters, Janice G." docDate="2015" docId="90882BE28039CC26BF665B801AF15661" docLanguage="en" docName="ZooKeys 478: 45-128" docOrigin="ZooKeys 478" docSource="http://dx.doi.org/10.3897/zookeys.478.8057" docTitle="Priasthenopus gilliesi, comb. n." docType="treatment" docVersion="6" lastPageNumber="68" masterDocId="FFBFFF8FFFA98B3E3C274356FFB0EE6C" masterDocTitle="Phylogeny and biogeography of Asthenopodinae with a revision of Asthenopus, reinstatement of Asthenopodes, and the description of the new genera Hubbardipes and Priasthenopus (Ephemeroptera, Polymitarcyidae)" masterLastPageNumber="128" masterPageNumber="45" pageNumber="67" updateTime="1732718156974" updateUser="ExternalLinkService">
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<mods:title id="99EDF031149CEB2D20787EBE598C5397">Phylogeny and biogeography of Asthenopodinae with a revision of Asthenopus, reinstatement of Asthenopodes, and the description of the new genera Hubbardipes and Priasthenopus (Ephemeroptera, Polymitarcyidae)</mods:title>
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<mods:namePart id="F854BCB40482FD0EA1D56FD3CAB59289">Molineri, Carlos</mods:namePart>
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<mods:namePart id="020003F4EBD0D19485974E365BB00EE6">Salles, Frederico F.</mods:namePart>
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<mods:namePart id="78D4FAFF605FDE13E8A04406AA6451AC">Peters, Janice G.</mods:namePart>
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<mods:classification id="E6A2C65B6D61CFBF36C8B64B2A58DF6D">journal article</mods:classification>
<mods:identifier id="E1B0A00A0B8685D1056BC3FB90ABA4E6" type="DOI">http://dx.doi.org/10.3897/zookeys.478.8057</mods:identifier>
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<treatment id="90882BE28039CC26BF665B801AF15661" ID-GBIF-Taxon="152058051" LSID="urn:lsid:plazi:treatment:90882BE28039CC26BF665B801AF15661" httpUri="http://treatment.plazi.org/id/90882BE28039CC26BF665B801AF15661" lastPageId="23" lastPageNumber="68" pageId="22" pageNumber="67" scope_family="Polymitarcyidae" scope_order="Ephemeroptera" scope_subFamily="Campsurinae">
<subSubSection id="250F70C60C2FABCB796AA4685656E760" pageId="22" pageNumber="67" type="multiple">
<paragraph id="E1FA89935B224EDAF2F6755D205545DF" pageId="22" pageNumber="67">Taxon classification Animalia Ephemeroptera Polymitarcyidae</paragraph>
</subSubSection>
<subSubSection id="AAB8E3206D3D615D6F4DEB983B6CF48F" pageId="22" pageNumber="67" type="nomenclature">
<paragraph id="852605DD185D66638D48529460E8FB40" pageId="22" pageNumber="67">
<taxonomicName id="594161A73F62568F9BC080EF7B6813B4" authority="Dominguez" genus="Priasthenopus" lsidName="Priasthenopus gilliesi" pageId="22" pageNumber="67" rank="species" species="gilliesi">
Priasthenopus gilliesi (
<normalizedToken id="88ECD6242586909F80472B97ECE7B6EC" originalValue="Domínguez">Dominguez</normalizedToken>
)
</taxonomicName>
<taxonomicNameLabel id="CB05B4C1B8F0D85853E0086E902EA184" pageId="22" pageNumber="67">comb. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="D1851A2E37FA13C868212113317691F0" pageId="22" pageNumber="67" type="reference_group">
<paragraph id="B6E0F3E711B27452D6B6644CAA789528" pageId="22" pageNumber="67">
<taxonomicName id="DCF67F5F38995AED23F4C82E74AF00B3" class="Insecta" family="Polymitarcyidae" genus="Asthenopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Asthenopus gilliesi" order="Ephemeroptera" pageId="22" pageNumber="67" phylum="Arthropoda" rank="species" species="gilliesi">Asthenopus gilliesi</taxonomicName>
<bibRefCitation id="E7C15CA7643CC65218A0AB73A4D47103" author="Dominguez, E" journalOrPublisher="Anales del Museo de Historia Natural de Valparaiso" pageId="51" pageNumber="96" pagination="21 - 26" title="Asthenopusgilliesi sp. n. y su importancia en la taxonomia de la subfamilia Asthenopodinae (Ephemeroptera: Polymitarcyidae)." volume="19" year="1988">
<normalizedToken id="7939582F1A7CE63E724FAF50B3ABF6E6" originalValue="Domínguez">Dominguez</normalizedToken>
1988
</bibRefCitation>
: 21;
<bibRefCitation id="8D6202B9D13698075A07E4B8B32F6C2C" author="Hubbard, MD" journalOrPublisher="The Florida Entomologist" pageId="52" pageNumber="97" pagination="207 - 210" title="Synonymy of the neotropical mayfly genera Asthenopus and Asthenopodes (Ephemeroptera: Polymitarcyidae: Asthenopodinae)." url="10.2307/3495369" volume="71" year="1988">
Hubbard and
<normalizedToken id="E3A0755C5BD980985EC73CB630EE526A" originalValue="Domínguez">Dominguez</normalizedToken>
1988
</bibRefCitation>
: 207;
<bibRefCitation id="FD2B1B677049C62ED64EFFB04C2CB4A4" author="Dominguez, E" journalOrPublisher="Pensoft" pageId="51" pageNumber="96" title="Ephemeroptera of South America. Aquatic Biodiversity in Latin America" year="2006">
<normalizedToken id="7836D05C3BEC81F25E2FB25D9486E369" originalValue="Domínguez">Dominguez</normalizedToken>
et al. 2006
</bibRefCitation>
: 562
</paragraph>
</subSubSection>
<subSubSection id="E9DE2CD8E200C2217C48D37B68B8C290" pageId="22" pageNumber="67" type="material">
<paragraph id="2CA430B0696B1BF6FEBBE72F4A2B0C23" pageId="22" pageNumber="67">Material.</paragraph>
<paragraph id="3379199933016202E60C2E301DFD7E8A" pageId="22" pageNumber="67">
Paratype male (IFML TEPH095, slide 041) from URUGUAY, Artigas, San Gregorio, orillas
<normalizedToken id="00E7E5D989E44218D6A055587DF69D80" originalValue="río">rio</normalizedToken>
Uruguay, 29.xi.1959, light trap, C.S. Carbonell col.; 5 male imagos (slide IBN141CM) and 1 male and 1 female (slide IBN471CM) subimagos (IBN) from COLOMBIA, Amazonas, P. N. Amacayacu,
<normalizedToken id="15C2A797A40B0503B645BE267E8768FD" originalValue="río">rio</normalizedToken>
Amacayacu, 93 m,
<geoCoordinate id="BB4606CB97D9BF88CFD80AC0D2FBB6B6" direction="south" orientation="latitude" precision="15" value="-3.8077776">S 3°48'28&quot;</geoCoordinate>
<geoCoordinate id="BA25C5490F8B6017A4BC626A96802149" direction="west" orientation="longitude" precision="15" value="-70.25584">W 70°15'21&quot;</geoCoordinate>
, 3.ii.1999, light trap 1820 h PM, E.
<normalizedToken id="AAE86A6ECAF3DD8C8A0F1EF886C75883" originalValue="Domínguez">Dominguez</normalizedToken>
, M.C.
<normalizedToken id="4D418417DA95E939E59FE043048AC63C" originalValue="Zúñiga">Zuniga</normalizedToken>
&amp; C. Molineri cols.; 5 male imagos and 1 female subimago (IBN) from BOLIVIA, Santa Cruz, near Once Por Ciento,
<normalizedToken id="22D735CCCE675216D15BD0B0B65A65A2" originalValue="río">rio</normalizedToken>
Blanco, 250 m,
<geoCoordinate id="B840C85C40312FD8D9FF14E09A3C08B0" direction="south" orientation="latitude" precision="1" value="-15.361028">S 15°21'39.7&quot;</geoCoordinate>
<geoCoordinate id="792FBC19823ADA143DF9D648B32CA921" direction="west" orientation="longitude" precision="1" value="-63.291332">W 63°17'28.8&quot;</geoCoordinate>
, 14.vi.2000, light trap, E.
<normalizedToken id="03EDC0BEB56AC3AC33BF6556795696D9" originalValue="Domínguez">Dominguez</normalizedToken>
col.; 1 female adult (CZNC) from BRAZIL, Rio N.
<normalizedToken id="41C4C15DFB1E40056ADFC750968C88A1" originalValue="Aripuanã">Aripuana</normalizedToken>
, Rio Juma, Ig. Campineiro Gde., 89.ix.2004,
<normalizedToken id="F01B2E752A7D81C18B1CABA354C4F5D7" originalValue="Pennsylvãnia">Pennsylvania</normalizedToken>
light trap; and 2 male imagos (CZNC) from Amazonas, Barcelos, rio Demene,
<normalizedToken id="C65F991E8FDA584EF8123279D3BF8B91" originalValue="´boca´barco">'boca'barco</normalizedToken>
, 89.viii.2009, Pennsylvania light trap.
</paragraph>
</subSubSection>
<subSubSection id="8300089531B01B2D198009B8D39E9BAC" lastPageId="23" lastPageNumber="68" pageId="22" pageNumber="67" type="diagnosis">
<paragraph id="3A7931A79EE3CED5A96E3C1BCA7A36D4" pageId="22" pageNumber="67">Diagnosis.</paragraph>
<paragraph id="6841D0CBC12D4D922040BC1F25827773" pageId="22" pageNumber="67">
<taxonomicName id="93845FC7114E93F45406627718C50F10" genus="Priasthenopus" lsidName="Priasthenopus gilliesi" pageId="22" pageNumber="67" rank="species" species="gilliesi">Priasthenopus gilliesi</taxonomicName>
is known from adults of both sexes and eggs, and is the only species known in the genus. The characters useful to distinguish it from other
<taxonomicName id="F7E5756E0E03EFE2CA27659DD64294D1" lsidName="" pageId="22" pageNumber="67" rank="subfamily" subfamily="Asthenopodinae">Asthenopodinae</taxonomicName>
are listed in the generic diagnosis.
</paragraph>
<paragraph id="6A36D62FC479C69DF965B383055FAD25" pageId="22" pageNumber="67">
Male imago. See generic section above and original description in
<bibRefCitation id="9DFAA3E94E474A83BE01B4EA4C8F30F7" author="Dominguez, E" journalOrPublisher="Anales del Museo de Historia Natural de Valparaiso" pageId="51" pageNumber="96" pagination="21 - 26" title="Asthenopusgilliesi sp. n. y su importancia en la taxonomia de la subfamilia Asthenopodinae (Ephemeroptera: Polymitarcyidae)." volume="19" year="1988">
<normalizedToken id="ED8D8A835EBDAB204CB24AAF75EC44C5" originalValue="Domínguez">Dominguez</normalizedToken>
(1988)
</bibRefCitation>
.
</paragraph>
<paragraph id="387FCF93BD41EA9CCB37FCBE5C970AFD" lastPageId="23" lastPageNumber="68" pageId="22" pageNumber="67">
Female subimago. Length (mm): body, 6.5; FW, 8.9; HW, 3.3; cerci, 1.5. General coloration yellowish white with gray markings. Head cream extensively shaded gray dorsally, the shading is uniform anteriorly but in the form of a fine netted pattern posteriorly to lateral ocelli, occiput with a pair of submedian pale anterior spots and a pair of submedian dark posterior spots; venter of head whitish. Antennae yellowish white shaded with gray on scape and pedicel. Thorax cream. Anterior ring of prothorax very thin, less than 1/4 the dorsal length of posterior ring; ratio width/length: 2.6; pronotum shaded blackish on median area except pale medial line, presternum paler, shaded gray before coxa. Mesonotum shaded very diffusely with gray, darker on longitudinal carinae and between posteroscutal protuberances; mesosternum and pleurae paler, shaded gray on anterior corner of katepisternum. Metanotum shaded gray on posterior half, except on a pale median triangular mark, shaded darker posteriorly to this pale mark; metasternum whitish. Legs whitish except coxae yellowish shaded gray and apex
<pageBreakToken id="7B6BECC315B00422F2EA9DFDEE78B612" pageId="23" pageNumber="68" start="start">of</pageBreakToken>
hind trochanter pointed and yellowish orange. Wings (Figs 8
<normalizedToken id="4FCA1C7840B0110DC79656B85D8C488D" originalValue="HI">H-I</normalizedToken>
). Membrane of fore and hind wings slightly shaded with light brown, veins translucent shaded with light brown, markedly on larger veins (C, Sc, Rs); marginal intercalaries relatively long (Fig. 8H), MP1 fused with MP2 at base; 2 crossveins present between M and R stems basally to Rs fork. Abdomen whitish, shaded with gray dorsally and darkening posteriorly, shading on terga interrupted on medial paler line, this line is wide on terga 1-4, narrows posteriorly on 5-7, thin on 8, and widens posteriorly on 9-10; sterna whitish translucent except gill sclerites yellowish white; sternum 8 with anteromedian pair of relatively large sockets (Fig. 8J). Cerci whitish, about 0.2 the length of FW.
</paragraph>
<paragraph id="E1F24EBAFEDAD5033DB42069BDB1CFB6" pageId="23" pageNumber="68">Eggs. See generic description.</paragraph>
</subSubSection>
<subSubSection id="A27DF32D1120D4AE37D30A1EDB38F248" pageId="23" pageNumber="68" type="distribution">
<paragraph id="340FF9E52F1712EB843E3C7DF5ECC3C6" pageId="23" pageNumber="68">Distribution.</paragraph>
<paragraph id="2C5486388E8C49D5AAF6A16BDD778630" pageId="23" pageNumber="68">This species presents a wide geographic range that spans from the Amazon River in the North to the Uruguay River in the South, also extending towards the West in Bolivian Chiquitania.</paragraph>
</subSubSection>
<subSubSection id="0B6E274C56878FD85789E5E6C4A5B676" pageId="23" pageNumber="68" type="discussion">
<paragraph id="CA70243AF738D6D27CC63EE9083186F7" pageId="23" pageNumber="68">Discussion.</paragraph>
<paragraph id="418D48FAA0E0A53912A975A67F03429F" pageId="23" pageNumber="68">
<taxonomicName id="BAFD112A7EA7B6646378B81998E1E58F" genus="Priasthenopus" lsidName="Priasthenopus gilliesi" pageId="23" pageNumber="68" rank="species" species="gilliesi">Priasthenopus gilliesi</taxonomicName>
male imagos were adequately described by
<bibRefCitation id="EE2976916A66E46C21C4E218192DEBE0" author="Dominguez, E" journalOrPublisher="Anales del Museo de Historia Natural de Valparaiso" pageId="51" pageNumber="96" pagination="21 - 26" title="Asthenopusgilliesi sp. n. y su importancia en la taxonomia de la subfamilia Asthenopodinae (Ephemeroptera: Polymitarcyidae)." volume="19" year="1988">
<normalizedToken id="EB6AFCD62D0A33CEA116E4DAF4472698" originalValue="Domínguez">Dominguez</normalizedToken>
(1988)
</bibRefCitation>
, females and eggs are described here for the first time. There are no morphological differences between the male imagos examined from the different localities, except for the penes of the Colombian males are slightly stouter, and those from the Bolivian males are slightly slender than the penes of the Uruguayan types. In the forewings, vein MP1 is basally free except on Bolivian males where this vein tends to fuse with MP2, although not completely.
</paragraph>
</subSubSection>
</treatment>
</document>