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<document id="E4703613A7FF97210DE03C99A5457E22" ID-DOI="10.11646/zootaxa.5415.1.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="10692701" ID-ZooBank="8160D5A0-9A3E-4E65-868C-BEEB5CEFA254" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1708602603503" checkinUser="plazi" docAuthor="Tan, Ming Kai, Muhammad, Amira Aqilah, Gorochov, Andrei V. &amp; Abdullah, Nurul Ashikin" docDate="2024" docId="5B3A87E1FFBE5D5848A2FDAF28D8F9AC" docLanguage="en" docName="zootaxa.5415.1.5.pdf" docOrigin="Zootaxa 5415 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5415.1.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cercoteratura reflexa Tan &amp; Muhammad &amp; Gorochov &amp; Abdullah 2024, sp. n." docType="treatment" docVersion="2" lastPageNumber="134" masterDocId="A703FF99FFAE5D494835FF8C2938FFD9" masterDocTitle="An account on some katydids (Orthoptera: Tettigonioidea) from Panti Forest Reserve in Johor, Peninsular Malaysia" masterLastPageNumber="143" masterPageNumber="117" pageNumber="133" updateTime="1708603241094" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="7311720C4D37C5C4BD044F981E702CDB">An account on some katydids (Orthoptera: Tettigonioidea) from Panti Forest Reserve in Johor, Peninsular Malaysia</mods:title>
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<mods:namePart id="8A785CA017E4B62CA9E5CE25DD489173">Tan, Ming Kai</mods:namePart>
<mods:affiliation id="01E6D5DCA16D04D82949B356772167CD">Block 207 A, Woodleigh Link, Singapore 361207, Republic of Singapore</mods:affiliation>
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<mods:namePart id="D283281698A7EEC14EB8A7DB2AF8EF03">Muhammad, Amira Aqilah</mods:namePart>
<mods:affiliation id="A93BD2DB32005164C9F804D05916DD3C">Institute of Biological Sciences, Faculty of Science, Universiti Malaya, Kuala Lumpur 50603, Malaysia</mods:affiliation>
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<mods:namePart id="B9E9C14A76CC07C8384B15A020EEF8D6">Gorochov, Andrei V.</mods:namePart>
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<mods:nameIdentifier id="11C3D328E4790F9F41D535108D84B31E" type="email">gorochovandrei3@gmail.com</mods:nameIdentifier>
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<mods:namePart id="E039EE005DB74814C1BA7E968813D8DE">Abdullah, Nurul Ashikin</mods:namePart>
<mods:affiliation id="7263E6E42C45B56E086A9D31775E92C0">Institute of Biological Sciences, Faculty of Science, Universiti Malaya, Kuala Lumpur 50603, Malaysia</mods:affiliation>
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<mods:title id="3153003D6F86FE6662D5B810CB0DE92A">Zootaxa</mods:title>
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<mods:date id="73532EE8C4381096B8B736DA34C8397E">2024</mods:date>
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<treatment id="5B3A87E1FFBE5D5848A2FDAF28D8F9AC" ID-DOI="http://doi.org/10.5281/zenodo.10692713" ID-Zenodo-Dep="10692713" LSID="urn:lsid:plazi:treatment:5B3A87E1FFBE5D5848A2FDAF28D8F9AC" httpUri="http://treatment.plazi.org/id/5B3A87E1FFBE5D5848A2FDAF28D8F9AC" lastPageId="17" lastPageNumber="134" pageId="16" pageNumber="133">
<subSubSection id="9B89657CFFBE5D5948A2FDAF2BFEFDE4" box="[151,710,546,573]" pageId="16" pageNumber="133" type="nomenclature">
<paragraph id="D32C36F7FFBE5D5948A2FDAF2BFEFDE4" blockId="16.[151,710,546,609]" box="[151,710,546,573]" pageId="16" pageNumber="133">
<heading id="8864819BFFBE5D5948A2FDAF2BFEFDE4" bold="true" box="[151,710,546,573]" fontSize="11" level="1" pageId="16" pageNumber="133" reason="1">
<emphasis id="E1E7EAE5FFBE5D5948A2FDAF2BFEFDE4" bold="true" box="[151,710,546,573]" pageId="16" pageNumber="133">
<taxonomicName id="14934D74FFBE5D5948A2FDAF2B4FFDE5" authority="Tan &amp; Gorochov" authorityName="Tan &amp; Muhammad &amp; Gorochov &amp; Abdullah" authorityYear="2024" box="[151,631,546,573]" class="Insecta" family="Tettigoniidae" genus="Cercoteratura" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Orthoptera" pageId="16" pageNumber="133" phylum="Arthropoda" rank="species" species="reflexa" status="sp. nov.">
<emphasis id="E1E7EAE5FFBE5D5948A2FDAF28A5FDE4" bold="true" box="[151,413,546,573]" italics="true" pageId="16" pageNumber="133">Cercoteratura reflexa</emphasis>
Tan &amp; Gorochov
</taxonomicName>
,
<taxonomicNameLabel id="FAD4579EFFBE5D594AB6FDAF2BFEFDE4" box="[643,710,547,573]" pageId="16" pageNumber="133" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="9B89657CFFBE5D5948A2FDCB2A79FD28" pageId="16" pageNumber="133" type="description">
<paragraph id="D32C36F7FFBE5D5948A2FDCB2888FDB8" blockId="16.[151,710,546,609]" box="[151,432,583,609]" pageId="16" pageNumber="133">
(
<figureCitation id="4BA82A72FFBE5D5948AAFDCB29CEFDB8" box="[159,246,583,609]" captionStart="FIGURE 18" captionStartId="16.[151,250,1520,1544]" captionTargetBox="[254,1333,777,1496]" captionTargetId="figure-245@16.[254,1333,777,1496]" captionTargetPageId="16" captionText="FIGURE 18. Cercoteratura reflexa sp. n. ♁ adult in its natural environment." figureDoi="http://doi.org/10.5281/zenodo.10692753" httpUri="https://zenodo.org/record/10692753/files/figure.png" pageId="16" pageNumber="133">Figs 18</figureCitation>
,
<figureCitation id="4BA82A72FFBE5D594937FDCB286DFDB8" box="[258,341,583,609]" captionStart="FIGURE 19" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[195,1408,181,1915]" captionTargetId="figure-19@19.[179,1408,181,1919]" captionTargetPageId="19" captionText="FIGURE 19. Cercoteratura reflexa sp. n. (A–C) and Platycercotura caricauda gen. et sp. n. (D–F) ♁ adult holotypes: body and tegmina (A, D) in lateral views, head and pronotum in lateral (B, E) and dorsal (C, F) views. Scale bars: 5 mm (A, D), 2 mm (all except A, D)." figureDoi="http://doi.org/10.5281/zenodo.10692755" httpUri="https://zenodo.org/record/10692755/files/figure.png" pageId="16" pageNumber="133">19A–C</figureCitation>
,
<figureCitation id="4BA82A72FFBE5D594954FDCB28B8FDB8" box="[353,384,583,609]" captionStart="FIGURE 20" captionStartId="20.[151,250,1757,1781]" captionTargetBox="[151,1436,181,1725]" captionTargetId="figure-25@20.[151,1436,181,1733]" captionTargetPageId="20" captionText="FIGURE 20. Cercoteratura reflexa sp. n. ♁ adult holotype: abdominal apex in dorsal (A), posterior (B), lateral (C, D) and ventral (E) views (A–C dry pinned; D, E in ethanol with genitalia intact), cercus in lateral view (F), genitalia in ventral (G), dorsal (H) and lateral (I) views. Scale bars: 1 mm (all except G–I), 0.5 mm (G–I)." figureDoi="http://doi.org/10.5281/zenodo.10692757" httpUri="https://zenodo.org/record/10692757/files/figure.png" pageId="16" pageNumber="133">20</figureCitation>
,
<figureCitation id="4BA82A72FFBE5D5949BEFDCB2890FDB8" box="[395,424,583,609]" captionStart="FIGURE 21" captionStartId="21.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1920]" captionTargetId="figure-19@21.[214,1373,181,1920]" captionTargetPageId="21" captionText="FIGURE 21. Cercoteratura reflexa sp. n. ♁ adult holotype, freshly moulted." figureDoi="http://doi.org/10.5281/zenodo.10692769" httpUri="https://zenodo.org/record/10692769/files/figure.png" pageId="16" pageNumber="133">21</figureCitation>
)
</paragraph>
<paragraph id="D32C36F7FFBE5D5948A2FD022A79FD28" blockId="16.[151,1437,654,753]" pageId="16" pageNumber="133">
<emphasis id="E1E7EAE5FFBE5D5948A2FD0228A3FD71" bold="true" box="[151,411,654,680]" pageId="16" pageNumber="133">Specimens examined.</emphasis>
<materialsCitation id="63FB3CAAFFBE5D594993FD032B96FD28" collectingDate="2023-08-17" collectionCode="PFR, PANTI" collectorName="M. K. Tan &amp; Amira A. Muhammad &amp; N. Ashikin Abdullah" country="Malaysia" elevation="6" latitude="1.80951" location="Panti Barat" longLatPrecision="1" longitude="103.85115" pageId="16" pageNumber="133" specimenCount="1" stateProvince="Johor" typeStatus="holotype">
PENINSULAR
<collectingCountry id="AB847667FFBE5D594A6AFD032BCDFD70" box="[607,757,655,681]" name="Malaysia" pageId="16" pageNumber="133">MALAYSIA</collectingCountry>
:
<collectingRegion id="1157F815FFBE5D594B36FD032AB1FD70" box="[771,905,655,681]" country="Malaysia" name="Johor" pageId="16" pageNumber="133">Johor State</collectingRegion>
:
<collectionCode id="B582AE32FFBE5D594BADFD032AF2FD70" box="[920,970,655,681]" pageId="16" pageNumber="133">PFR</collectionCode>
<emphasis id="E1E7EAE5FFBE5D594BE0FD032AE7FD70" bold="true" box="[981,991,655,681]" pageId="16" pageNumber="133">•</emphasis>
♁ (
<typeStatus id="0C288855FFBE5D594C27FD032D42FD70" box="[1042,1146,655,681]" pageId="16" pageNumber="133" type="holotype">holotype</typeStatus>
);
<location id="D64C602CFFBE5D594CA7FD032C2EFD70" LSID="urn:lsid:plazi:treatment:5B3A87E1FFBE5D5848A2FDAF28D8F9AC:D64C602CFFBE5D594CA7FD032C2EFD70" box="[1170,1302,655,681]" country="Malaysia" latitude="1.80951" longLatPrecision="1" longitude="103.85115" name="Panti Barat" pageId="16" pageNumber="133" stateProvince="Johor">Panti Barat</location>
;
<geoCoordinate id="B6A75030FFBE5D594D13FD022CA0FD70" box="[1318,1432,654,681]" degrees="1.80951" direction="north" orientation="latitude" pageId="16" pageNumber="133" precision="1" value="1.80951">N1.80951</geoCoordinate>
,
<geoCoordinate id="B6A75030FFBE5D5948A2FD3E2819FD14" box="[151,289,690,717]" degrees="103.85115" direction="east" orientation="longitude" pageId="16" pageNumber="133" precision="1" value="103.85115">E103.85115</geoCoordinate>
, 76.6±
<quantity id="146B9B12FFBE5D594958FD3E288BFD14" box="[365,435,690,717]" metricMagnitude="0" metricUnit="m" metricValue="5.9" pageId="16" pageNumber="133" unit="m" value="5.9">
<elevation id="58BED1C4FFBE5D594958FD3E288BFD14" box="[365,435,690,717]" metricMagnitude="0" metricUnit="m" metricValue="5.9" pageId="16" pageNumber="133" unit="m" value="5.9">5.9 m</elevation>
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.a.s.l.;
<date id="A72D1037FFBE5D5949C0FD3F2B91FD14" box="[501,681,691,717]" pageId="16" pageNumber="133" value="2023-08-17">
<collectingDate id="B769E9DFFFBE5D5949C0FD3F2B91FD14" box="[501,681,691,717]" pageId="16" pageNumber="133" value="2023-08-17">17 August 2023</collectingDate>
</date>
, 19h51; moulting under the foliage of a tree; coll.
<collectorName id="7E665321FFBE5D594CE2FD3F2C73FD14" box="[1239,1355,691,717]" pageId="16" pageNumber="133">M. K. Tan</collectorName>
,
<collectorName id="7E665321FFBE5D594D60FD3F2878FD28" pageId="16" pageNumber="133">Amira A. Muhammad</collectorName>
&amp;
<collectorName id="7E665321FFBE5D594950FD5B2B77FD28" box="[357,591,727,753]" pageId="16" pageNumber="133">N. Ashikin Abdullah</collectorName>
;
<collectionCode id="B582AE32FFBE5D594A6FFD5B2B96FD28" box="[602,686,727,753]" pageId="16" pageNumber="133">PANTI</collectionCode>
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.23.16 (ZRC)
</paragraph>
</subSubSection>
<caption id="87EC667FFFBE5D5948A2FA7C2A9EF9D1" ID-DOI="http://doi.org/10.5281/zenodo.10692753" ID-Zenodo-Dep="10692753" box="[151,934,1520,1546]" httpUri="https://zenodo.org/record/10692753/files/figure.png" pageId="16" pageNumber="133" startId="16.[151,250,1520,1544]" targetBox="[254,1333,777,1496]" targetPageId="16" targetType="figure">
<paragraph id="D32C36F7FFBE5D5948A2FA7C2A9EF9D1" blockId="16.[151,934,1520,1546]" box="[151,934,1520,1546]" pageId="16" pageNumber="133">
<emphasis id="E1E7EAE5FFBE5D5948A2FA7C2818F9D1" bold="true" box="[151,288,1520,1544]" pageId="16" pageNumber="133">FIGURE 18.</emphasis>
<taxonomicName id="14934D74FFBE5D594913FA7C2B39F9D1" authority="Tan &amp; Muhammad &amp; Gorochov &amp; Abdullah, 2024" authorityName="Tan &amp; Muhammad &amp; Gorochov &amp; Abdullah" authorityYear="2024" box="[294,513,1520,1544]" class="Insecta" family="Tettigoniidae" genus="Cercoteratura" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Orthoptera" pageId="16" pageNumber="133" phylum="Arthropoda" rank="species" species="reflexa" status="sp. nov.">
<emphasis id="E1E7EAE5FFBE5D594913FA7C2B39F9D1" box="[294,513,1520,1544]" italics="true" pageId="16" pageNumber="133">Cercoteratura reflexa</emphasis>
</taxonomicName>
<emphasis id="E1E7EAE5FFBE5D594A3DFA7C2B79F9D1" bold="true" box="[520,577,1520,1544]" pageId="16" pageNumber="133">
<taxonomicNameLabel id="FAD4579EFFBE5D594A3DFA7C2B79F9D1" box="[520,577,1520,1544]" pageId="16" pageNumber="133" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
♁ adult in its natural environment.
</paragraph>
</caption>
<subSubSection id="9B89657CFFBE5D5948F2F9D62B21F965" pageId="16" pageNumber="133" type="etymology">
<paragraph id="D32C36F7FFBE5D5948F2F9D62B21F965" blockId="16.[151,1437,1626,1977]" pageId="16" pageNumber="133">
<emphasis id="E1E7EAE5FFBE5D5948F2F9D62876F9AD" bold="true" box="[199,334,1626,1652]" pageId="16" pageNumber="133">Etymology.</emphasis>
The species name refers to the diagnostic long lamellar lobes of the epiproct appearing to bend backwards (anteriorly) being curved ventrally and having a tooth-like ventral lobe pointing ventro-anteriorly at the apex; reflexa = bent back in Latin.
</paragraph>
</subSubSection>
<subSubSection id="9B89657CFFBE5D5848F2F94A2B5AFE98" lastPageId="17" lastPageNumber="134" pageId="16" pageNumber="133" type="diagnosis">
<paragraph id="D32C36F7FFBE5D5948F2F94A2A41F861" blockId="16.[151,1437,1626,1977]" pageId="16" pageNumber="133">
<emphasis id="E1E7EAE5FFBE5D5948F2F94A287AF939" bold="true" box="[199,322,1734,1760]" pageId="16" pageNumber="133">Diagnosis.</emphasis>
The new species is characterised most by the epiproct (a pair of rather large and long lamellar lobes with its apex having an obtuse, stout dorsal lobe pointing dorso-posteriorly and a tooth-like ventral lobe pointing ventro-anteriorly) and the phallic complex (with yoke-shaped epiphallus producing into two lateral stout and rounded lobes at the apex with dentated margins); as well as the combination of the shapes of the male last abdominal tergite (with small triangular postero-median lobe), male cercus (S-shaped in profile view; with the basal part thickened and having a tongue-shaped inner lobe; almost lamellar rest part; at the apex with a thin and arcuate apical spine and a smaller stout lobule dorsad of the apical spine).
</paragraph>
<paragraph id="D32C36F7FFBF5D5848F2FF1B2B5AFE98" blockId="17.[151,1437,151,1653]" pageId="17" pageNumber="134">
The new species is most similar to
<taxonomicName id="14934D74FFBF5D584A61FF1B2D1AFF68" authority="Gorochov, 2019" authorityName="Gorochov" authorityYear="2019" box="[596,1058,151,177]" class="Insecta" family="Tettigoniidae" genus="Cercoteratura" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Orthoptera" pageId="17" pageNumber="134" phylum="Arthropoda" rank="species" species="variegata">
<emphasis id="E1E7EAE5FFBF5D584A61FF1B2A5DFF69" box="[596,869,151,177]" italics="true" pageId="17" pageNumber="134">Cercoteratura variegata</emphasis>
<bibRefCitation id="B7024B06FFBF5D584B58FF1B2D1AFF68" author="Gorochov, A. V." box="[877,1058,151,177]" pageId="17" pageNumber="134" pagination="1 - 24" refId="ref8484" refString="Gorochov, A. V. (2019) Taxonomy of the katydids (Orthoptera: Tettigoniidae) from East Asia and adjacent islands. Communication 12. Far Eastern Entomologist, 379, 1 - 24. https: // doi. org / 10.25221 / fee. 379.1" type="journal article" year="2019">Gorochov, 2019</bibRefCitation>
</taxonomicName>
from
<collectingRegion id="1157F815FFBF5D584C5DFF1B2DFDFF68" box="[1128,1221,151,177]" country="Indonesia" name="Sumatera Utara" pageId="17" pageNumber="134">Sumatra</collectingRegion>
and Borneo by the shape of the last abdominal tergite with a small postero-median lobe, of the epiproct consisting of a pair of rather large and long lamellar lobes and of the male cercus S-shaped with a thin and arcuate apical spine. It differs by the epiproct forming two apical lobes (absent in
<taxonomicName id="14934D74FFBF5D584ABEFE8F2AA4FEC5" authorityName="Gorochov" authorityYear="2019" box="[651,924,259,285]" class="Insecta" family="Tettigoniidae" genus="Cercoteratura" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Orthoptera" pageId="17" pageNumber="134" phylum="Arthropoda" rank="species" species="variegata">
<emphasis id="E1E7EAE5FFBF5D584ABEFE8F2AA4FEC5" box="[651,924,259,285]" italics="true" pageId="17" pageNumber="134">Cercoteratura variegata</emphasis>
</taxonomicName>
) and the cercus having a tongue-shaped inner basal lobe (instead of having four lobes).
</paragraph>
</subSubSection>
<subSubSection id="9B89657CFFBF5D5848F2FEC72B44FE50" pageId="17" pageNumber="134" type="discussion">
<paragraph id="D32C36F7FFBF5D5848F2FEC72B44FE50" blockId="17.[151,1437,151,1653]" pageId="17" pageNumber="134">
<emphasis id="E1E7EAE5FFBF5D5848F2FEC72803FEBC" bold="true" box="[199,315,331,357]" pageId="17" pageNumber="134">Remarks.</emphasis>
This represents the first record of
<taxonomicName id="14934D74FFBF5D584A8FFEC72A61FEBC" authorityName="Gorochov" authorityYear="2019" box="[698,857,331,357]" class="Insecta" family="Tettigoniidae" genus="Cercoteratura" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Orthoptera" pageId="17" pageNumber="134" phylum="Arthropoda" rank="genus">
<emphasis id="E1E7EAE5FFBF5D584A8FFEC72A61FEBC" box="[698,857,331,357]" italics="true" pageId="17" pageNumber="134">Cercoteratura</emphasis>
</taxonomicName>
in Malay Peninsula, having previously known from Thai-Peninsula,
<collectingRegion id="1157F815FFBF5D58497AFEE32897FE50" box="[335,431,367,393]" country="Indonesia" name="Sumatera Utara" pageId="17" pageNumber="134">Sumatra</collectingRegion>
, Borneo and
<collectingRegion id="1157F815FFBF5D584A76FEE32B40FE50" box="[579,632,367,393]" country="Indonesia" name="Jawa Timur" pageId="17" pageNumber="134">Java</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="9B89657CFFBF5D5848F2FE1E2859FA18" pageId="17" pageNumber="134" type="description">
<paragraph id="D32C36F7FFBF5D5848F2FE1E2B8CFBC8" blockId="17.[151,1437,151,1653]" pageId="17" pageNumber="134">
<emphasis id="E1E7EAE5FFBF5D5848F2FE1E2861FE75" bold="true" box="[199,345,402,428]" pageId="17" pageNumber="134">Description.</emphasis>
Habitus typical of congeners; generally green with dark-brown patterns on head, pronotum, tegmina and legs when alive (
<figureCitation id="4BA82A72FFBF5D5849DFFE3B2B7AFE08" box="[490,578,439,465]" captionStart="FIGURE 18" captionStartId="16.[151,250,1520,1544]" captionTargetBox="[254,1333,777,1496]" captionTargetId="figure-245@16.[254,1333,777,1496]" captionTargetPageId="16" captionText="FIGURE 18. Cercoteratura reflexa sp. n. ♁ adult in its natural environment." figureDoi="http://doi.org/10.5281/zenodo.10692753" httpUri="https://zenodo.org/record/10692753/files/figure.png" pageId="17" pageNumber="134">Figs 18</figureCitation>
,
<figureCitation id="4BA82A72FFBF5D584A7AFE3B2BBBFE08" box="[591,643,439,465]" captionStart="FIGURE 19" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[195,1408,181,1915]" captionTargetId="figure-19@19.[179,1408,181,1919]" captionTargetPageId="19" captionText="FIGURE 19. Cercoteratura reflexa sp. n. (A–C) and Platycercotura caricauda gen. et sp. n. (D–F) ♁ adult holotypes: body and tegmina (A, D) in lateral views, head and pronotum in lateral (B, E) and dorsal (C, F) views. Scale bars: 5 mm (A, D), 2 mm (all except A, D)." figureDoi="http://doi.org/10.5281/zenodo.10692755" httpUri="https://zenodo.org/record/10692755/files/figure.png" pageId="17" pageNumber="134">19A</figureCitation>
). Head generally pale green to green, dorsum with four dark-coloured longitudinal bands converging at frontal rostrum; antennal scapus and pedicel dark coloured (
<figureCitation id="4BA82A72FFBF5D584C9EFE572C6AFE2C" box="[1195,1362,475,501]" captionStart="FIGURE 19" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[195,1408,181,1915]" captionTargetId="figure-19@19.[179,1408,181,1919]" captionTargetPageId="19" captionText="FIGURE 19. Cercoteratura reflexa sp. n. (A–C) and Platycercotura caricauda gen. et sp. n. (D–F) ♁ adult holotypes: body and tegmina (A, D) in lateral views, head and pronotum in lateral (B, E) and dorsal (C, F) views. Scale bars: 5 mm (A, D), 2 mm (all except A, D)." figureDoi="http://doi.org/10.5281/zenodo.10692755" httpUri="https://zenodo.org/record/10692755/files/figure.png" pageId="17" pageNumber="134">Figs 19B, 19C</figureCitation>
). Eyes globular (
<figureCitation id="4BA82A72FFBF5D584932FE732849FDC0" box="[263,369,511,537]" captionStart="FIGURE 19" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[195,1408,181,1915]" captionTargetId="figure-19@19.[179,1408,181,1919]" captionTargetPageId="19" captionText="FIGURE 19. Cercoteratura reflexa sp. n. (A–C) and Platycercotura caricauda gen. et sp. n. (D–F) ♁ adult holotypes: body and tegmina (A, D) in lateral views, head and pronotum in lateral (B, E) and dorsal (C, F) views. Scale bars: 5 mm (A, D), 2 mm (all except A, D)." figureDoi="http://doi.org/10.5281/zenodo.10692755" httpUri="https://zenodo.org/record/10692755/files/figure.png" pageId="17" pageNumber="134">Fig. 19B</figureCitation>
). Frontal rostrum dark coloured, conical, stout, with rounded apex of upper tubercle having median sulcus feebly present (
<figureCitation id="4BA82A72FFBF5D5849D3FDAF2B74FDE4" box="[486,588,547,573]" captionStart="FIGURE 19" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[195,1408,181,1915]" captionTargetId="figure-19@19.[179,1408,181,1919]" captionTargetPageId="19" captionText="FIGURE 19. Cercoteratura reflexa sp. n. (A–C) and Platycercotura caricauda gen. et sp. n. (D–F) ♁ adult holotypes: body and tegmina (A, D) in lateral views, head and pronotum in lateral (B, E) and dorsal (C, F) views. Scale bars: 5 mm (A, D), 2 mm (all except A, D)." figureDoi="http://doi.org/10.5281/zenodo.10692755" httpUri="https://zenodo.org/record/10692755/files/figure.png" pageId="17" pageNumber="134">Fig. 19C</figureCitation>
). Antennal segments with apical parts dark (
<figureCitation id="4BA82A72FFBF5D584C00FDAF2DA2FDE4" box="[1077,1178,547,573]" captionStart="FIGURE 19" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[195,1408,181,1915]" captionTargetId="figure-19@19.[179,1408,181,1919]" captionTargetPageId="19" captionText="FIGURE 19. Cercoteratura reflexa sp. n. (A–C) and Platycercotura caricauda gen. et sp. n. (D–F) ♁ adult holotypes: body and tegmina (A, D) in lateral views, head and pronotum in lateral (B, E) and dorsal (C, F) views. Scale bars: 5 mm (A, D), 2 mm (all except A, D)." figureDoi="http://doi.org/10.5281/zenodo.10692755" httpUri="https://zenodo.org/record/10692755/files/figure.png" pageId="17" pageNumber="134">Fig. 19B</figureCitation>
). Mouthparts generally light coloured (
<figureCitation id="4BA82A72FFBF5D58497DFDCB2888FDB8" box="[328,432,583,609]" captionStart="FIGURE 19" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[195,1408,181,1915]" captionTargetId="figure-19@19.[179,1408,181,1919]" captionTargetPageId="19" captionText="FIGURE 19. Cercoteratura reflexa sp. n. (A–C) and Platycercotura caricauda gen. et sp. n. (D–F) ♁ adult holotypes: body and tegmina (A, D) in lateral views, head and pronotum in lateral (B, E) and dorsal (C, F) views. Scale bars: 5 mm (A, D), 2 mm (all except A, D)." figureDoi="http://doi.org/10.5281/zenodo.10692755" httpUri="https://zenodo.org/record/10692755/files/figure.png" pageId="17" pageNumber="134">Fig. 19B</figureCitation>
). Apical segment of maxillary palpi long and slender, widened apically and blackish at apex; fourth slightly shorter than third segment in length (
<figureCitation id="4BA82A72FFBF5D584B12FDE72AB7FD5C" box="[807,911,619,645]" captionStart="FIGURE 19" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[195,1408,181,1915]" captionTargetId="figure-19@19.[179,1408,181,1919]" captionTargetPageId="19" captionText="FIGURE 19. Cercoteratura reflexa sp. n. (A–C) and Platycercotura caricauda gen. et sp. n. (D–F) ♁ adult holotypes: body and tegmina (A, D) in lateral views, head and pronotum in lateral (B, E) and dorsal (C, F) views. Scale bars: 5 mm (A, D), 2 mm (all except A, D)." figureDoi="http://doi.org/10.5281/zenodo.10692755" httpUri="https://zenodo.org/record/10692755/files/figure.png" pageId="17" pageNumber="134">Fig. 19B</figureCitation>
). Pronotal disc covering most part of tegminal mirror, anterior margin slightly convex, anterior half green with broad median brown band, posterior half brown with two black spots, posterior margin subobtusely rounded (
<figureCitation id="4BA82A72FFBF5D584B08FD3F2A9BFD14" box="[829,931,691,717]" captionStart="FIGURE 19" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[195,1408,181,1915]" captionTargetId="figure-19@19.[179,1408,181,1919]" captionTargetPageId="19" captionText="FIGURE 19. Cercoteratura reflexa sp. n. (A–C) and Platycercotura caricauda gen. et sp. n. (D–F) ♁ adult holotypes: body and tegmina (A, D) in lateral views, head and pronotum in lateral (B, E) and dorsal (C, F) views. Scale bars: 5 mm (A, D), 2 mm (all except A, D)." figureDoi="http://doi.org/10.5281/zenodo.10692755" httpUri="https://zenodo.org/record/10692755/files/figure.png" pageId="17" pageNumber="134">Fig. 19C</figureCitation>
). Ventral margin of pronotal lateral lobe short and rounded (
<figureCitation id="4BA82A72FFBF5D584902FD5B28A7FD28" box="[311,415,727,753]" captionStart="FIGURE 19" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[195,1408,181,1915]" captionTargetId="figure-19@19.[179,1408,181,1919]" captionTargetPageId="19" captionText="FIGURE 19. Cercoteratura reflexa sp. n. (A–C) and Platycercotura caricauda gen. et sp. n. (D–F) ♁ adult holotypes: body and tegmina (A, D) in lateral views, head and pronotum in lateral (B, E) and dorsal (C, F) views. Scale bars: 5 mm (A, D), 2 mm (all except A, D)." figureDoi="http://doi.org/10.5281/zenodo.10692755" httpUri="https://zenodo.org/record/10692755/files/figure.png" pageId="17" pageNumber="134">Fig. 19B</figureCitation>
). Thoracic auditory spiracle (= thoracic foramen) large and bean-shaped; not covered by pronotal lateral lobe (
<figureCitation id="4BA82A72FFBF5D5849BEFD7728CAFCCC" box="[395,498,763,789]" captionStart="FIGURE 19" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[195,1408,181,1915]" captionTargetId="figure-19@19.[179,1408,181,1919]" captionTargetPageId="19" captionText="FIGURE 19. Cercoteratura reflexa sp. n. (A–C) and Platycercotura caricauda gen. et sp. n. (D–F) ♁ adult holotypes: body and tegmina (A, D) in lateral views, head and pronotum in lateral (B, E) and dorsal (C, F) views. Scale bars: 5 mm (A, D), 2 mm (all except A, D)." figureDoi="http://doi.org/10.5281/zenodo.10692755" httpUri="https://zenodo.org/record/10692755/files/figure.png" pageId="17" pageNumber="134">Fig. 19B</figureCitation>
). Tegmen rather narrow, with oblique apex, reaching basal part of hind tibiae; with numerous irregular black or dark spots (
<figureCitation id="4BA82A72FFBF5D584A64FC932B80FCE0" box="[593,696,799,825]" captionStart="FIGURE 19" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[195,1408,181,1915]" captionTargetId="figure-19@19.[179,1408,181,1919]" captionTargetPageId="19" captionText="FIGURE 19. Cercoteratura reflexa sp. n. (A–C) and Platycercotura caricauda gen. et sp. n. (D–F) ♁ adult holotypes: body and tegmina (A, D) in lateral views, head and pronotum in lateral (B, E) and dorsal (C, F) views. Scale bars: 5 mm (A, D), 2 mm (all except A, D)." figureDoi="http://doi.org/10.5281/zenodo.10692755" httpUri="https://zenodo.org/record/10692755/files/figure.png" pageId="17" pageNumber="134">Fig. 19A</figureCitation>
). Hind wing surpassing tegminal apices (
<figureCitation id="4BA82A72FFBF5D584C4BFC932DDDFCE0" box="[1150,1253,799,825]" captionStart="FIGURE 19" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[195,1408,181,1915]" captionTargetId="figure-19@19.[179,1408,181,1919]" captionTargetPageId="19" captionText="FIGURE 19. Cercoteratura reflexa sp. n. (A–C) and Platycercotura caricauda gen. et sp. n. (D–F) ♁ adult holotypes: body and tegmina (A, D) in lateral views, head and pronotum in lateral (B, E) and dorsal (C, F) views. Scale bars: 5 mm (A, D), 2 mm (all except A, D)." figureDoi="http://doi.org/10.5281/zenodo.10692755" httpUri="https://zenodo.org/record/10692755/files/figure.png" pageId="17" pageNumber="134">Fig. 19A</figureCitation>
). Legs generally pale green; fore and middle femora dark coloured towards apex and knees; hind femur with large dark-coloured patch at middle of external surface and with knee dark coloured; tarsi and apex of tibiae black (
<figureCitation id="4BA82A72FFBF5D584CFFFCEB2C0AFC58" box="[1226,1330,871,897]" captionStart="FIGURE 19" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[195,1408,181,1915]" captionTargetId="figure-19@19.[179,1408,181,1919]" captionTargetPageId="19" captionText="FIGURE 19. Cercoteratura reflexa sp. n. (A–C) and Platycercotura caricauda gen. et sp. n. (D–F) ♁ adult holotypes: body and tegmina (A, D) in lateral views, head and pronotum in lateral (B, E) and dorsal (C, F) views. Scale bars: 5 mm (A, D), 2 mm (all except A, D)." figureDoi="http://doi.org/10.5281/zenodo.10692755" httpUri="https://zenodo.org/record/10692755/files/figure.png" pageId="17" pageNumber="134">Fig. 19A</figureCitation>
). Femora without spine; fore and middle tibiae each with four outer and four inner subapical spines (spines black and tibia at base of spine dark coloured); inner and outer knees of hind femora with rounded lobes, inner knee with a small apical spine; hind tibiae ventrally and dorsally with numerous outer and inner dark-coloured spines as well as two ventral and one dorsal apical spurs on each side.
</paragraph>
<paragraph id="D32C36F7FFBF5D5848F2FB972D8CFA8C" blockId="17.[151,1437,151,1653]" pageId="17" pageNumber="134">
<emphasis id="E1E7EAE5FFBF5D5848F2FB972833FBEC" bold="true" box="[199,267,1051,1077]" pageId="17" pageNumber="134">Male.</emphasis>
Abdominal apex as in
<figureCitation id="4BA82A72FFBF5D584A2EFB972B9DFBEC" box="[539,677,1051,1077]" captionStart="FIGURE 20" captionStartId="20.[151,250,1757,1781]" captionTargetBox="[151,1436,181,1725]" captionTargetId="figure-25@20.[151,1436,181,1733]" captionTargetPageId="20" captionText="FIGURE 20. Cercoteratura reflexa sp. n. ♁ adult holotype: abdominal apex in dorsal (A), posterior (B), lateral (C, D) and ventral (E) views (A–C dry pinned; D, E in ethanol with genitalia intact), cercus in lateral view (F), genitalia in ventral (G), dorsal (H) and lateral (I) views. Scale bars: 1 mm (all except G–I), 0.5 mm (G–I)." figureDoi="http://doi.org/10.5281/zenodo.10692757" httpUri="https://zenodo.org/record/10692757/files/figure.png" pageId="17" pageNumber="134">Figs 20A–F</figureCitation>
. Last abdominal tergite with small triangular postero-median lobe (apex obtuse) located in depth of large postero-median notch. Epiproct consisting of a pair of rather large and long lamellar lobes, curved ventrally in profile, at the apex having obtuse, stout dorsal lobe pointing dorso-posteriorly and a tooth-like ventral lobe pointing ventro-anteriorly; ventral lobe distinctly slenderer and longer than dorsal lobe. Cercus S-shaped in profile view; with basal part thickened and having tongue-shaped inner lobe; thereafter almost lamellar; at apex with thin and arcuate apical spine and smaller stout lobule dorsad of apical spine. Subgenital plate with long styles. Phallic complex consists of yoke-shaped epiphallus (
<figureCitation id="4BA82A72FFBF5D584B96FB7F2D1EFAD4" box="[931,1062,1267,1293]" captionStart="FIGURE 20" captionStartId="20.[151,250,1757,1781]" captionTargetBox="[151,1436,181,1725]" captionTargetId="figure-25@20.[151,1436,181,1733]" captionTargetPageId="20" captionText="FIGURE 20. Cercoteratura reflexa sp. n. ♁ adult holotype: abdominal apex in dorsal (A), posterior (B), lateral (C, D) and ventral (E) views (A–C dry pinned; D, E in ethanol with genitalia intact), cercus in lateral view (F), genitalia in ventral (G), dorsal (H) and lateral (I) views. Scale bars: 1 mm (all except G–I), 0.5 mm (G–I)." figureDoi="http://doi.org/10.5281/zenodo.10692757" httpUri="https://zenodo.org/record/10692757/files/figure.png" pageId="17" pageNumber="134">Figs 20G–I</figureCitation>
), with small basal lobe (in profile view), anterior half slightly swollen, posterior half slightly narrowing, posterior apex emarginated and with two lateral stout and rounded lobes pointing dorsad (in profile view); lobes with dentated margins.
</paragraph>
<paragraph id="D32C36F7FFBF5D5848F2FAD3289AFAA0" blockId="17.[151,1437,151,1653]" box="[199,418,1375,1401]" pageId="17" pageNumber="134">
<emphasis id="E1E7EAE5FFBF5D5848F2FAD3281DFAA0" bold="true" box="[199,293,1375,1401]" pageId="17" pageNumber="134">Female.</emphasis>
Unknown.
</paragraph>
<paragraph id="D32C36F7FFBF5D5848F2FA0F2859FA18" blockId="17.[151,1437,151,1653]" pageId="17" pageNumber="134">
<emphasis id="E1E7EAE5FFBF5D5848F2FA0F2B21FA45" bold="true" box="[199,537,1410,1440]" pageId="17" pageNumber="134">Measurements (1♁, in mm).</emphasis>
BL = 8.7 (dry-pinned), BWL = 21.7, PL = 3.8, TL = 16.6, HWT = 2.4, HFL = 11.2, HTL = 12.0.
</paragraph>
</subSubSection>
<subSubSection id="9B89657CFFBF5D5848F2FA472B18F9D0" pageId="17" pageNumber="134" type="biology_ecology">
<paragraph id="D32C36F7FFBF5D5848F2FA472B18F9D0" blockId="17.[151,1437,151,1653]" pageId="17" pageNumber="134">
<emphasis id="E1E7EAE5FFBF5D5848F2FA472814FA3C" bold="true" box="[199,300,1483,1509]" pageId="17" pageNumber="134">Ecology.</emphasis>
The specimen was collected when it had just moulted (
<figureCitation id="4BA82A72FFBF5D584B95FA472ACAFA3C" box="[928,1010,1483,1509]" captionStart="FIGURE 21" captionStartId="21.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1920]" captionTargetId="figure-19@21.[214,1373,181,1920]" captionTargetPageId="21" captionText="FIGURE 21. Cercoteratura reflexa sp. n. ♁ adult holotype, freshly moulted." figureDoi="http://doi.org/10.5281/zenodo.10692769" httpUri="https://zenodo.org/record/10692769/files/figure.png" pageId="17" pageNumber="134">Fig. 21</figureCitation>
) and the colouration differs distinctly from its final colouration (
<figureCitation id="4BA82A72FFBF5D58498AFA632B2BF9D0" box="[447,531,1519,1545]" captionStart="FIGURE 18" captionStartId="16.[151,250,1520,1544]" captionTargetBox="[254,1333,777,1496]" captionTargetId="figure-245@16.[254,1333,777,1496]" captionTargetPageId="16" captionText="FIGURE 18. Cercoteratura reflexa sp. n. ♁ adult in its natural environment." figureDoi="http://doi.org/10.5281/zenodo.10692753" httpUri="https://zenodo.org/record/10692753/files/figure.png" pageId="17" pageNumber="134">Fig. 18</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="9B89657CFFBF5D5848F2F99F2AB8F9F4" box="[199,896,1554,1581]" pageId="17" pageNumber="134" type="materials_examined">
<paragraph id="D32C36F7FFBF5D5848F2F99F2AB8F9F4" blockId="17.[151,1437,151,1653]" box="[199,896,1554,1581]" pageId="17" pageNumber="134">
<materialsCitation id="63FB3CAAFFBF5D5848F2F99F2AB8F9F4" box="[199,896,1554,1581]" collectionCode="PFR" country="Malaysia" location="Johor State" pageId="17" pageNumber="134" specimenCount="1" stateProvince="Johor" typeStatus="holotype">
<emphasis id="E1E7EAE5FFBF5D5848F2F99F285EF9F5" bold="true" box="[199,358,1554,1581]" pageId="17" pageNumber="134">
<typeStatus id="0C288855FFBF5D5848F2F99F283AF9F4" box="[199,258,1555,1581]" pageId="17" pageNumber="134">Type</typeStatus>
locality.
</emphasis>
PENINSULAR
<collectingCountry id="AB847667FFBF5D584A17F99F2B80F9F4" box="[546,696,1555,1581]" name="Malaysia" pageId="17" pageNumber="134">MALAYSIA</collectingCountry>
:
<collectingRegion id="1157F815FFBF5D584AF7F99F2A7BF9F4" box="[706,835,1555,1581]" country="Malaysia" name="Johor" pageId="17" pageNumber="134">Johor State</collectingRegion>
:
<collectionCode id="B582AE32FFBF5D584B7BF99F2AB8F9F4" box="[846,896,1555,1581]" pageId="17" pageNumber="134">PFR</collectionCode>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="9B89657CFFBF5D5848F2F9BA28D8F9AC" pageId="17" pageNumber="134" type="distribution">
<paragraph id="D32C36F7FFBF5D5848F2F9BA2ABBF988" blockId="17.[151,1437,151,1653]" box="[199,899,1590,1617]" pageId="17" pageNumber="134">
<emphasis id="E1E7EAE5FFBF5D5848F2F9BA2859F989" bold="true" box="[199,353,1590,1616]" pageId="17" pageNumber="134">Distribution.</emphasis>
PENINSULAR
<collectingCountry id="AB847667FFBF5D584A28F9BB2B95F988" box="[541,685,1591,1617]" name="Malaysia" pageId="17" pageNumber="134">MALAYSIA</collectingCountry>
(
<collectingRegion id="1157F815FFBF5D584A8EF9BB2A06F988" box="[699,830,1591,1617]" country="Malaysia" name="Johor" pageId="17" pageNumber="134">Johor State</collectingRegion>
: PFR)
</paragraph>
<paragraph id="D32C36F7FFBF5D5848F2F9D628D8F9AC" blockId="17.[151,1437,151,1653]" box="[199,480,1626,1653]" pageId="17" pageNumber="134">
<emphasis id="E1E7EAE5FFBF5D5848F2F9D6285BF9AC" bold="true" box="[199,355,1626,1653]" pageId="17" pageNumber="134">Calling song.</emphasis>
Unknown.
</paragraph>
</subSubSection>
</treatment>
</document>
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