430 lines
56 KiB
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430 lines
56 KiB
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<document id="327D74C1F311C27BC68CF353A3BF1782" ID-DOI="10.11646/zootaxa.5123.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="6399827" ID-ZooBank="036F6B4D-CDCC-4CD7-A914-9A1D8C7A097A" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="karina" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,operationResults,plazi" IM.taxonomicNames_approvedBy="karina" IM.treatments_approvedBy="karina" checkinTime="1648673534065" checkinUser="plazi" docAuthor="Harzhauser, Mathias, Landau, Bernard & Janssen, Ronald" docDate="2022" docId="039487D1FF95FFB1FFBAF9296DE2FA01" docLanguage="en" docName="zootaxa.5123.1.1.pdf" docOrigin="Zootaxa 5123 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5123.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.18:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="18" docTitle="Clavatula sorini Harzhauser & Landau & Janssen 2022, nov. sp." docType="treatment" docVersion="4" lastPageNumber="26" masterDocId="FFADFFA9FF82FFA8FF2DFFAB6872FFC0" masterDocTitle="The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera" masterLastPageNumber="172" masterPageNumber="1" pageNumber="24" updateTime="1709132455857" updateUser="karina">
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<mods:title id="83B1559C0F7C6BEFC62F95BC56E6047A">The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera</mods:title>
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<mods:namePart id="83772CB23EC7AE7579FF53182E82B4BF">Harzhauser, Mathias</mods:namePart>
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<mods:namePart id="385FE4278A29E1F6319EF382C89A1968">Landau, Bernard</mods:namePart>
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<heading id="D0CA81ABFF95FFBFFFBAF92969CDF95D" bold="true" box="[151,447,1666,1693]" fontSize="11" level="2" pageId="23" pageNumber="24" reason="3">
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<taxonomicName id="4C3D4D44FF95FFBFFFBAF9296928F95D" ID-CoL="8YLQB" authority="Harzhauser & Landau & Janssen, 2022" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[151,346,1666,1693]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="sorini" status="sp. nov.">
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<emphasis id="B949EAD5FF95FFBFFFBAF9296928F95D" bold="true" box="[151,346,1666,1693]" italics="true" pageId="23" pageNumber="24">Clavatula sorini</emphasis>
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<paragraph id="8B8236C7FF95FFBFFFBAF90C6998F901" blockId="23.[151,490,1666,1735]" box="[151,490,1703,1735]" pageId="23" pageNumber="24">
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<figureCitation id="13062A42FF95FFBFFFBAF90C688FF901" box="[151,253,1703,1729]" captionStart="FIGURE 11" captionStartId="24.[151,250,1801,1825]" captionTargetBox="[164,1423,273,1778]" captionTargetId="figure-56@24.[164,1423,273,1778]" captionTargetPageId="24" captionText="FIGURE 11 A1–A3. Clavatula sorini nov. sp., holotype, NHMW 1865/0001/0224, Lăpugiu de Sus (Romania). B1–B3. Clavatula sorini nov. sp., paratype, NHMW 1874/0025/0046, Lăpugiu de Sus (Romania).C1–C3. ‘Clavatula’ camillae (Hoernes &Auinger, 1891), lectotype, NHMW 1949/0005/0042, Lăpugiu de Sus (Romania).D1–D3. ‘Clavatula’ camillae (Hoernes &Auinger, 1891), paralectotype, NHMW 1949/0005/0042b, Lăpugiu de Sus (Romania), E1–E2. ‘Clavatula’ atatuerki nov. sp., holotype,. NHMW 1847/0058/1216, Seyithasan (Turkey). F1–F2. ‘Clavatula’ atatuerki nov. sp., paratype, NHMW 1847/0058/1217, Seyithasan (Turkey). G1–G2. ‘Clavatula’ santhai Kovács & Vicián, 2021, holotype, HNHM, PAL 2021.20.1, Letkés (Hungary)." figureDoi="http://doi.org/10.5281/zenodo.6399855" httpUri="https://zenodo.org/record/6399855/files/figure.png" pageId="23" pageNumber="24">Figs 11A</figureCitation>
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<figureCitation id="13062A42FF95FFBFFFD3F912695AF901" box="[254,296,1703,1735]" captionStart="FIGURE 1" captionStartId="6.[151,250,1903,1927]" captionTargetBox="[171,1417,176,1880]" captionTargetId="figure-26@6.[171,1417,176,1880]" captionTargetPageId="6" captionText="FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E.-Sopr B: Eisenstadt–Sopron Basin)." figureDoi="http://doi.org/10.5281/zenodo.6399829" httpUri="https://zenodo.org/record/6399829/files/figure.png" pageId="23" pageNumber="24">
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–A
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</figureCitation>
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<figureCitation id="13062A42FF95FFBFFE05F9126942F907" box="[296,304,1721,1735]" captionStart="FIGURE 3" captionStartId="12.[151,250,1619,1643]" captionTargetBox="[151,1436,181,1595]" captionTargetId="figure-26@12.[151,1436,181,1595]" captionTargetPageId="12" captionText="FIGURE 3. Early teleoconch whorls of extant and fossil Clavatulidae species. A. Clavatula regia (Röding, 1798), NHMW 1683 6/28, West Africa. B. Clavatula rubrifasciata (Reeve, 1845), NHMW 10203, Gorée (Senegal). C. ‘Clavatula’ diadema (Kiener, 1839), NHMW 1884/0010/0023, Gorée (Senegal). D. ‘Clavatula’ mystica (Reeve, 1843), MNHN-IM-2013-59897, Est de l'ile de Gorée (Senegal). E. ‘Clavatula’ bimarginata (Lamarck, 1822), MNHN-IM-2013-43369, Dakar (Senegal). F. ‘Clavatula’ pyramidata (Kiener, 1839), MNHN-IM-2009-24982, Baie de Pointe-Noire (Republique du Congo). G. Clavatula sorini nov. sp., paratype, NHMW 1874/0025/0046, Lăpugiu de Sus (Romania). H. ‘Clavatula’ romana (Defrance, 1826), NHMW 1999/0098/0004, Modra-Kráľová (Slovakia). I. ‘Clavatula’ interrupta (Brocchi, 1814), NHMW 1861/0033/0420, Buituri (Romania). J. ‘Clavatula’ hirmetzli Kovács & Vicián, 2021, NHMW 1868/0001/0069, Baden-Sooss (Austria). K. ‘Clavatula’ irisae nov. sp., NHMW 2021/0126/0001, holotype, Gainfarn (Austria). L. ‘Clavatula’ sophiae (Hoernes & Auinger, 1891), lectotype, NHMW 2021/0127/0001, Enzesfeld (Austria). M. ‘Clavatula’ veronicae (Hoernes &Auinger, 1891), NHMW 1855/0043/0044b, Lăpugiu de Sus (Romania). N. ‘Clavatula’ santhai Kovács & Vicián, 2021, holotype, HNHM, PAL 2021.20.1, Letkés (Hungary).O. ‘Clavatula’ camillae (Hoernes &Auinger, 1891), lectotype, NHMW 1949/0005/0042, Lăpugiu de Sus (Romania). P. Megaclavatula amaliae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0049, Lăpugiu de Sus (Romania). Q. Megaclavatula grunerti nov. sp., paratype, NHMW 2021/0127/0003, Enzesfeld (Austria). R. Megaclavatula laevigata (Eichwald, 1830), NHMW 1853/0038/0035a, Korytnica (Poland). S. Megaclavatula pilleri nov. sp., holotype, NHMW 1949/0005/0051a, Baden-Sooss (Austria). T. Megaclavatula polonica (Pusch, 1837), NHMW 1862/0001/0324, Bad Vöslau (Austria).U. Granulatocincta angelae (Hoernes &Auinger, 1891), paralectotype, NHMW 1949/0005/0099, Möllersdorf (Austria). V. Granulatocincta granulatocincta (Münster in Goldfuss, 1841), NHMW 1846/0028/0009, Enzesfeld (Austria). W. Granulatocincta nataliae (Hoernes & Auinger, 1891), NHMW 1855/0035/0021, holotype, Belgrade/Rakovica (Serbia). X. Granulatocincta schreibersi (Hörnes, 1854), paralectotype, NHMW1997z0178/1133, Bad Vöslau (Austria).Y. Granulatocincta callim nov. sp., paratype, NHMW 2021/0128/0001, Bad Sauerbrunn (Austria). Z. Granulatocincta contorta (Švagrovský, 1958), holotype, SNMB Z 3076, Kuzmice (Slovakia), Aa. Granulatocincta theoderichi nov. sp., paratype, NHMW 2016/0026/0320b, Ritzing (Austria). Ab. Olegia doderleini (Hörnes, 1854), paralectotype, NHMW 1858/0015/0147, Wiesen (Austria). Ac. Olegia mandici nov. sp., paratype, NHMW 1872/0031/0008a, Bekića vrelo, Čremušnica at Donji Kirin (Croatia). Ae. Olegia dorotheae (Hoernes & Auinger, 1891), paralectotype, NHMW 1949/0005/0040, Kleinebersdorf (Austria). Scale bar is 5 mm for all specimens except for Granulatocincta contorta, Olegia doderleini and O. mandici." figureDoi="http://doi.org/10.5281/zenodo.6399835" httpUri="https://zenodo.org/record/6399835/files/figure.png" pageId="23" pageNumber="24">
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<subScript id="17B93482FF95FFBFFE05F9126942F907" attach="left" box="[296,304,1721,1735]" fontSize="6" pageId="23" pageNumber="24">3</subScript>
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, B
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<figureCitation id="13062A42FF95FFBFFE7CF912690BF901" box="[337,377,1703,1735]" captionStart="FIGURE 1" captionStartId="6.[151,250,1903,1927]" captionTargetBox="[171,1417,176,1880]" captionTargetId="figure-26@6.[171,1417,176,1880]" captionTargetPageId="6" captionText="FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E.-Sopr B: Eisenstadt–Sopron Basin)." figureDoi="http://doi.org/10.5281/zenodo.6399829" httpUri="https://zenodo.org/record/6399829/files/figure.png" pageId="23" pageNumber="24">
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–B
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</figureCitation>
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<figureCitation id="13062A42FF95FFBFFE57F91269C6F901" box="[378,436,1703,1735]" captionStart="FIGURE 3" captionStartId="12.[151,250,1619,1643]" captionTargetBox="[151,1436,181,1595]" captionTargetId="figure-26@12.[151,1436,181,1595]" captionTargetPageId="12" captionText="FIGURE 3. Early teleoconch whorls of extant and fossil Clavatulidae species. A. Clavatula regia (Röding, 1798), NHMW 1683 6/28, West Africa. B. Clavatula rubrifasciata (Reeve, 1845), NHMW 10203, Gorée (Senegal). C. ‘Clavatula’ diadema (Kiener, 1839), NHMW 1884/0010/0023, Gorée (Senegal). D. ‘Clavatula’ mystica (Reeve, 1843), MNHN-IM-2013-59897, Est de l'ile de Gorée (Senegal). E. ‘Clavatula’ bimarginata (Lamarck, 1822), MNHN-IM-2013-43369, Dakar (Senegal). F. ‘Clavatula’ pyramidata (Kiener, 1839), MNHN-IM-2009-24982, Baie de Pointe-Noire (Republique du Congo). G. Clavatula sorini nov. sp., paratype, NHMW 1874/0025/0046, Lăpugiu de Sus (Romania). H. ‘Clavatula’ romana (Defrance, 1826), NHMW 1999/0098/0004, Modra-Kráľová (Slovakia). I. ‘Clavatula’ interrupta (Brocchi, 1814), NHMW 1861/0033/0420, Buituri (Romania). J. ‘Clavatula’ hirmetzli Kovács & Vicián, 2021, NHMW 1868/0001/0069, Baden-Sooss (Austria). K. ‘Clavatula’ irisae nov. sp., NHMW 2021/0126/0001, holotype, Gainfarn (Austria). L. ‘Clavatula’ sophiae (Hoernes & Auinger, 1891), lectotype, NHMW 2021/0127/0001, Enzesfeld (Austria). M. ‘Clavatula’ veronicae (Hoernes &Auinger, 1891), NHMW 1855/0043/0044b, Lăpugiu de Sus (Romania). N. ‘Clavatula’ santhai Kovács & Vicián, 2021, holotype, HNHM, PAL 2021.20.1, Letkés (Hungary).O. ‘Clavatula’ camillae (Hoernes &Auinger, 1891), lectotype, NHMW 1949/0005/0042, Lăpugiu de Sus (Romania). P. Megaclavatula amaliae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0049, Lăpugiu de Sus (Romania). Q. Megaclavatula grunerti nov. sp., paratype, NHMW 2021/0127/0003, Enzesfeld (Austria). R. Megaclavatula laevigata (Eichwald, 1830), NHMW 1853/0038/0035a, Korytnica (Poland). S. Megaclavatula pilleri nov. sp., holotype, NHMW 1949/0005/0051a, Baden-Sooss (Austria). T. Megaclavatula polonica (Pusch, 1837), NHMW 1862/0001/0324, Bad Vöslau (Austria).U. Granulatocincta angelae (Hoernes &Auinger, 1891), paralectotype, NHMW 1949/0005/0099, Möllersdorf (Austria). V. Granulatocincta granulatocincta (Münster in Goldfuss, 1841), NHMW 1846/0028/0009, Enzesfeld (Austria). W. Granulatocincta nataliae (Hoernes & Auinger, 1891), NHMW 1855/0035/0021, holotype, Belgrade/Rakovica (Serbia). X. Granulatocincta schreibersi (Hörnes, 1854), paralectotype, NHMW1997z0178/1133, Bad Vöslau (Austria).Y. Granulatocincta callim nov. sp., paratype, NHMW 2021/0128/0001, Bad Sauerbrunn (Austria). Z. Granulatocincta contorta (Švagrovský, 1958), holotype, SNMB Z 3076, Kuzmice (Slovakia), Aa. Granulatocincta theoderichi nov. sp., paratype, NHMW 2016/0026/0320b, Ritzing (Austria). Ab. Olegia doderleini (Hörnes, 1854), paralectotype, NHMW 1858/0015/0147, Wiesen (Austria). Ac. Olegia mandici nov. sp., paratype, NHMW 1872/0031/0008a, Bekića vrelo, Čremušnica at Donji Kirin (Croatia). Ae. Olegia dorotheae (Hoernes & Auinger, 1891), paralectotype, NHMW 1949/0005/0040, Kleinebersdorf (Austria). Scale bar is 5 mm for all specimens except for Granulatocincta contorta, Olegia doderleini and O. mandici." figureDoi="http://doi.org/10.5281/zenodo.6399835" httpUri="https://zenodo.org/record/6399835/files/figure.png" pageId="23" pageNumber="24">
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, 3G
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</figureCitation>
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,
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<figureCitation id="13062A42FF95FFBFFEEDF90C69A2F902" box="[448,464,1703,1730]" captionStart="FIGURE 5" captionStartId="15.[151,250,1999,2023]" captionTargetBox="[151,1436,631,1976]" captionTargetId="figure-214@15.[151,1436,631,1976]" captionTargetPageId="15" captionText="FIGURE 5. Anal sinus morphology of Paratethyan Clavatulidae." figureDoi="http://doi.org/10.5281/zenodo.6399839" httpUri="https://zenodo.org/record/6399839/files/figure.png" pageId="23" pageNumber="24">5</figureCitation>
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,
|
||
<figureCitation id="13062A42FF95FFBFFEF1F90C6998F901" box="[476,490,1703,1729]" captionStart="FIGURE 6" captionStartId="16.[151,250,1938,1962]" captionTargetBox="[151,1436,345,1915]" captionTargetId="figure-106@16.[151,1436,345,1915]" captionTargetPageId="16" captionText="FIGURE 6. Apertures of Paratethyan Clavatulidae (C. = Clavatula, ‘C.’ = ‘Clavatula’, G. = Granulatocincta, M. = Megaclavaltula, O. = Olegia)." figureDoi="http://doi.org/10.5281/zenodo.6399841" httpUri="https://zenodo.org/record/6399841/files/figure.png" pageId="23" pageNumber="24">6</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C327654CFF95FFBFFFBAF940697BF8E2" pageId="23" pageNumber="24" type="reference_group">
|
||
<paragraph id="8B8236C7FF95FFBFFFBAF940697BF8E2" blockId="23.[151,1437,1771,1826]" pageId="23" pageNumber="24">
|
||
<treatmentCitationGroup id="AB2D11E9FF95FFBFFFBAF940697BF8E2" pageId="23" pageNumber="24">
|
||
<taxonomicName id="4C3D4D44FF95FFBFFFBAF9406A65F8C4" authority="Grat." authorityName="Grateloup" authorityYear="1832" box="[151,535,1771,1796]" class="Gastropoda" family="Turridae" genus="Pleurotoma" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="spinosa" subGenus="Clavatula">
|
||
<emphasis id="B949EAD5FF95FFBFFFBAF940697CF8C3" box="[151,270,1771,1795]" italics="true" pageId="23" pageNumber="24">Pleurotoma</emphasis>
|
||
(
|
||
<emphasis id="B949EAD5FF95FFBFFE30F94069F3F8C3" box="[285,385,1771,1795]" italics="true" pageId="23" pageNumber="24">Clavatula</emphasis>
|
||
)
|
||
<emphasis id="B949EAD5FF95FFBFFEBDF94769AFF8C3" box="[400,477,1772,1795]" italics="true" pageId="23" pageNumber="24">spinosa</emphasis>
|
||
Grat.
|
||
</taxonomicName>
|
||
—
|
||
<treatmentCitation id="0A9C10D6FF95FFBFFD1DF9476B1BF8C4" author="Hoernes, R. & Auinger, M." box="[560,873,1771,1796]" page="342" pageId="23" pageNumber="24" year="1891">
|
||
<bibRefCitation id="EFAC4B36FF95FFBFFD1DF9476B1BF8C4" author="Hoernes, R. & Auinger, M." box="[560,873,1771,1796]" pageId="23" pageNumber="24" pagination="283 - 382" refId="ref135318" refString="Hoernes, R. & Auinger, M. (1891) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der kaiserlich-koniglichen Geologischen Reichsanstalt, 12, 283 - 382, pls 37 - 50. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf]" type="journal article" year="1891">Hoernes & Auinger 1891: 342</bibRefCitation>
|
||
</treatmentCitation>
|
||
, pl. 46, figs 22a–b [
|
||
<emphasis id="B949EAD5FF95FFBFFB1BF9476C2EF8C3" box="[1078,1116,1772,1795]" italics="true" pageId="23" pageNumber="24">non</emphasis>
|
||
‘
|
||
<taxonomicName id="4C3D4D44FF95FFBFFB46F940688DF8E2" ID-CoL="7NZRZ" authority="Grateloup, 1832" authorityName="Lamarck" authorityYear="1801" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="24" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF95FFBFFB46F9406CBDF8C3" box="[1131,1231,1771,1795]" italics="true" pageId="23" pageNumber="24">Clavatula</emphasis>
|
||
’
|
||
<emphasis id="B949EAD5FF95FFBFFBF1F9476D5BF8C3" box="[1244,1321,1772,1795]" italics="true" pageId="23" pageNumber="24">spinosa</emphasis>
|
||
<bibRefCitation id="EFAC4B36FF95FFBFFA1CF940688DF8E2" author="Grateloup, J. P. S. de" pageId="23" pageNumber="24" pagination="314 - 344" refId="ref133310" refString="Grateloup, J. P. S. de (1832) Tableau des coquilles Fossiles qu'on rencontre dans les terrains calcaires tertiaries (faluns) des environs de Dax, dans le departement des Landes. Actes de la Societe Linneenne de Bordeaux, 5, 314 - 344. [https: // www. biodiversitylibrary. org / item / 102890 # page / 325 / mode / 1 up]" type="journal article" year="1832">Grateloup, 1832</bibRefCitation>
|
||
</taxonomicName>
|
||
].
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C327654CFF95FFB0FFBAF8FB6B27FF72" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24" type="materials_examined">
|
||
<paragraph id="8B8236C7FF95FFBFFFBAF8FB6B9AF872" blockId="23.[151,1436,1872,2042]" pageId="23" pageNumber="24">
|
||
<emphasis id="B949EAD5FF95FFBFFFBAF8FB693AF8AA" bold="true" box="[151,328,1872,1898]" pageId="23" pageNumber="24">Type material.</emphasis>
|
||
<materialsCitation id="3B553C9AFF95FFBFFE7EF8FB6B80F855" collectionCode="MD" collectorName="de Sus" country="Romania" location="Lapugiu de Sus" pageId="23" pageNumber="24" specimenCode="NHMW 1865" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus id="54868865FF95FFBFFE7EF8FB69B2F8AA" box="[339,448,1872,1898]" pageId="23" pageNumber="24" type="holotype">Holotype</typeStatus>
|
||
:
|
||
<specimenCode id="DB9B9EBCFF95FFBFFEE0F8FA6A18F8AB" box="[461,618,1872,1899]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="23" pageNumber="24" type="Museum">NHMW 1865</specimenCode>
|
||
/0001/0224,
|
||
<location id="8EE2601CFF95FFBFFDD6F8FB6BC2F8AA" LSID="urn:lsid:plazi:treatment:039487D1FF95FFB1FFBAF9296DE2FA01:8EE2601CFF95FFBFFDD6F8FB6BC2F8AA" box="[763,944,1872,1898]" country="Romania" name="Lapugiu de Sus" pageId="23" pageNumber="24">
|
||
Lăpugiu
|
||
<collectorName id="26C85311FF95FFBFFC4EF8FB6BC2F8AA" box="[867,944,1872,1898]" pageId="23" pageNumber="24">de Sus</collectorName>
|
||
</location>
|
||
(
|
||
<collectingCountry id="F32A7657FF95FFBFFCEEF8FB6C5CF8AB" box="[963,1070,1872,1899]" name="Romania" pageId="23" pageNumber="24">Romania</collectingCountry>
|
||
), SL:
|
||
<quantity id="4CC59B22FF95FFBFFB5AF8FB6C92F8AB" box="[1143,1248,1872,1899]" metricMagnitude="-2" metricUnit="m" metricValue="2.98" pageId="23" pageNumber="24" unit="mm" value="29.8">29.8 mm</quantity>
|
||
,
|
||
<collectionCode id="ED2CAE02FF95FFBFFBC2F8FA6D52F8AB" box="[1263,1312,1873,1899]" country="Germany" name="Museum Donaueschingen" pageId="23" pageNumber="24">MD</collectionCode>
|
||
:
|
||
<quantity id="4CC59B22FF95FFBFFA03F8FB6DEAF8AB" box="[1326,1432,1872,1899]" metricMagnitude="-2" metricUnit="m" metricValue="1.56" pageId="23" pageNumber="24" unit="mm" value="15.6">15.6 mm</quantity>
|
||
, illustrated in
|
||
<bibRefCitation id="EFAC4B36FF95FFBFFE1FF8DE6A11F84E" author="Hoernes, R. & Auinger, M." box="[306,611,1908,1935]" pageId="23" pageNumber="24" pagination="283 - 382" refId="ref135318" refString="Hoernes, R. & Auinger, M. (1891) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der kaiserlich-koniglichen Geologischen Reichsanstalt, 12, 283 - 382, pls 37 - 50. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf]" type="journal article" year="1891">Hoernes & Auinger (1891</bibRefCitation>
|
||
, pl. 46, figs 22a–b); figs 11A
|
||
<subScript id="17B93482FF95FFBFFC92F82C6BB5F855" attach="left" box="[959,967,1927,1941]" fontSize="6" pageId="23" pageNumber="24">1</subScript>
|
||
–A
|
||
<subScript id="17B93482FF95FFBFFCC7F82C6B80F855" attach="left" box="[1002,1010,1927,1941]" fontSize="6" pageId="23" pageNumber="24">3</subScript>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B553C9AFF95FFBFFB29F8DE6A1AF873" collectionCode="NHMW" collectorName="de Sus" country="Romania" location="Lapugiu de Sus" pageId="23" pageNumber="24" specimenCode="NHMW 1874" specimenCount="1" typeStatus="paratype">
|
||
<typeStatus id="54868865FF95FFBFFB29F8DE6C18F84F" box="[1028,1130,1909,1935]" pageId="23" pageNumber="24" type="paratype">Paratype</typeStatus>
|
||
:
|
||
<specimenCode id="DB9B9EBCFF95FFBFFB55F8DE6D67F84F" box="[1144,1301,1908,1935]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="23" pageNumber="24" type="Museum">NHMW 1874</specimenCode>
|
||
/0025/0046,
|
||
<location id="8EE2601CFF95FFBFFFBAF8336937F872" LSID="urn:lsid:plazi:treatment:039487D1FF95FFB1FFBAF9296DE2FA01:8EE2601CFF95FFBFFFBAF8336937F872" box="[151,325,1944,1970]" country="Romania" name="Lapugiu de Sus" pageId="23" pageNumber="24">
|
||
Lăpugiu
|
||
<collectorName id="26C85311FF95FFBFFFD1F8336937F872" box="[252,325,1944,1970]" pageId="23" pageNumber="24">de Sus</collectorName>
|
||
</location>
|
||
(
|
||
<collectingCountry id="F32A7657FF95FFBFFE79F83369CDF873" box="[340,447,1944,1971]" name="Romania" pageId="23" pageNumber="24">Romania</collectingCountry>
|
||
), SL:
|
||
<quantity id="4CC59B22FF95FFBFFD2FF8336A1AF873" box="[514,616,1944,1971]" metricMagnitude="-2" metricUnit="m" metricValue="3.09" pageId="23" pageNumber="24" unit="mm" value="30.9">30.9 mm</quantity>
|
||
</materialsCitation>
|
||
,
|
||
<collectionCode id="ED2CAE02FF95FFBFFD5EF8326AD6F873" box="[627,676,1945,1971]" country="Germany" name="Museum Donaueschingen" pageId="23" pageNumber="24">MD</collectionCode>
|
||
:
|
||
<quantity id="4CC59B22FF95FFBFFD82F8336B67F873" box="[687,789,1944,1971]" metricMagnitude="-2" metricUnit="m" metricValue="1.47" pageId="23" pageNumber="24" unit="mm" value="14.7">14.7 mm</quantity>
|
||
, figs 11B
|
||
<subScript id="17B93482FF95FFBFFC52F8006BF5F879" attach="left" box="[895,903,1963,1977]" fontSize="6" pageId="23" pageNumber="24">1</subScript>
|
||
–B
|
||
<subScript id="17B93482FF95FFBFFC85F8006BC2F879" attach="left" box="[936,944,1963,1977]" fontSize="6" pageId="23" pageNumber="24">3</subScript>
|
||
, 3G.
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FF95FFBFFFEAF8176B46F816" blockId="23.[151,1436,1872,2042]" box="[199,820,1980,2007]" pageId="23" pageNumber="24">
|
||
<materialsCitation id="3B553C9AFF95FFBFFFEAF8176B42F816" box="[199,816,1980,2007]" collectorName="de Sus" country="Romania" location="Lapugiu de Sus" pageId="23" pageNumber="24" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="B949EAD5FF95FFBFFFEAF8176914F816" bold="true" box="[199,358,1980,2006]" pageId="23" pageNumber="24">
|
||
<typeStatus id="54868865FF95FFBFFFEAF8176970F816" box="[199,258,1980,2006]" pageId="23" pageNumber="24">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<location id="8EE2601CFF95FFBFFE41F8176A68F816" LSID="urn:lsid:plazi:treatment:039487D1FF95FFB1FFBAF9296DE2FA01:8EE2601CFF95FFBFFE41F8176A68F816" box="[364,538,1980,2006]" country="Romania" name="Lapugiu de Sus" pageId="23" pageNumber="24">
|
||
Lăpugiu
|
||
<collectorName id="26C85311FF95FFBFFEFCF8176A68F816" box="[465,538,1980,2006]" pageId="23" pageNumber="24">de Sus</collectorName>
|
||
</location>
|
||
(
|
||
<collectingCountry id="F32A7657FF95FFBFFD04F8176AE6F817" box="[553,660,1980,2007]" name="Romania" pageId="23" pageNumber="24">Romania</collectingCountry>
|
||
), Făget Basin
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FF95FFBFFFEAF84B6A8EF83A" blockId="23.[151,1436,1872,2042]" box="[199,764,2016,2042]" pageId="23" pageNumber="24">
|
||
<emphasis id="B949EAD5FF95FFBFFFEAF84B691DF83A" bold="true" box="[199,367,2016,2042]" pageId="23" pageNumber="24">
|
||
<typeStatus id="54868865FF95FFBFFFEAF84B6970F83A" box="[199,258,2016,2042]" pageId="23" pageNumber="24">Type</typeStatus>
|
||
stratum.
|
||
</emphasis>
|
||
Silt and clay of the Dej Formation.
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FF9AFFB0FFEAFF3D6B27FF72" blockId="24.[151,1437,150,249]" box="[199,853,150,178]" pageId="24" pageNumber="25">
|
||
<emphasis id="B949EAD5FF9AFFB0FFEAFF3D688FFF70" bold="true" box="[199,253,150,176]" pageId="24" pageNumber="25">Age.</emphasis>
|
||
Middle Miocene, early/middle Badenian (Langhian).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C327654CFF9AFFB0FFEAFF106BD6FF39" pageId="24" pageNumber="25" type="etymology">
|
||
<paragraph id="8B8236C7FF9AFFB0FFEAFF106BD6FF39" blockId="24.[151,1437,150,249]" pageId="24" pageNumber="25">
|
||
<emphasis id="B949EAD5FF9AFFB0FFEAFF10693CFF15" bold="true" box="[199,334,187,213]" pageId="24" pageNumber="25">Etymology.</emphasis>
|
||
In honor of Sorin Filipescu (Babeş-Bolyai University,
|
||
<collectingRegion id="49F9F825FF9AFFB0FCCEFF106C64FF15" box="[995,1046,187,213]" country="Romania" name="Cluj" pageId="24" pageNumber="25">Cluj</collectingRegion>
|
||
,
|
||
<collectingCountry id="F32A7657FF9AFFB0FB0AFF106CE3FF16" box="[1063,1169,187,214]" name="Romania" pageId="24" pageNumber="25">Romania</collectingCountry>
|
||
), in recognition of his contributions to the Neogene stratigraphy of the Transylvanian Basin.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF42664FFF9AFFB0FFBAF8A26D43F816" ID-DOI="http://doi.org/10.5281/zenodo.6399855" ID-Zenodo-Dep="6399855" httpUri="https://zenodo.org/record/6399855/files/figure.png" pageId="24" pageNumber="25" startId="24.[151,250,1801,1825]" targetBox="[164,1423,273,1778]" targetPageId="24">
|
||
<paragraph id="8B8236C7FF9AFFB0FFBAF8A26D43F816" blockId="24.[151,1437,1801,2011]" pageId="24" pageNumber="25">
|
||
<emphasis id="B949EAD5FF9AFFB0FFBAF8A26913F8E2" bold="true" box="[151,353,1801,1831]" pageId="24" pageNumber="25">
|
||
FIGURE 11 A
|
||
<subScript id="17B93482FF9AFFB0FE00F8B26946F8E7" attach="left" box="[301,308,1817,1831]" fontSize="6" pageId="24" pageNumber="25">1</subScript>
|
||
–A
|
||
<subScript id="17B93482FF9AFFB0FE79F8B26929F8E7" attach="left" box="[340,347,1817,1831]" fontSize="6" pageId="24" pageNumber="25">3</subScript>
|
||
.
|
||
</emphasis>
|
||
<taxonomicName id="4C3D4D44FF9AFFB0FE4AF8A16A79F8E1" authority="Harzhauser & Landau & Janssen, 2022" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[359,523,1802,1826]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="sorini" status="sp. nov.">
|
||
<emphasis id="B949EAD5FF9AFFB0FE4AF8A16A79F8E1" box="[359,523,1802,1826]" italics="true" pageId="24" pageNumber="25">Clavatula sorini</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57AEFF9AFFB0FD3DF8A16A2DF8E1" box="[528,607,1802,1825]" pageId="24" pageNumber="25" rank="species">nov. sp.</taxonomicNameLabel>
|
||
, holotype, NHMW 1865/0001/0224, Lăpugiu de Sus (Romania).
|
||
<emphasis id="B949EAD5FF9AFFB0FBC2F8A16D41F8E2" bold="true" box="[1263,1331,1802,1831]" pageId="24" pageNumber="25">
|
||
B
|
||
<subScript id="17B93482FF9AFFB0FA2DF8B26D75F8E7" attach="left" box="[1280,1287,1817,1831]" fontSize="6" pageId="24" pageNumber="25">1</subScript>
|
||
–B
|
||
<subScript id="17B93482FF9AFFB0FA08F8B26D5EF8E7" attach="left" box="[1317,1324,1817,1831]" fontSize="6" pageId="24" pageNumber="25">3</subScript>
|
||
.
|
||
</emphasis>
|
||
<taxonomicName id="4C3D4D44FF9AFFB0FA15F8A168A0F885" authority="Harzhauser & Landau & Janssen, 2022" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="sorini" status="sp. nov.">
|
||
<emphasis id="B949EAD5FF9AFFB0FA15F8A168A0F885" italics="true" pageId="24" pageNumber="25">Clavatula sorini</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A57AEFF9AFFB0FFFAF8856957F885" box="[215,293,1838,1861]" pageId="24" pageNumber="25" rank="species">nov. sp.</taxonomicNameLabel>
|
||
, paratype, NHMW 1874/0025/0046, Lăpugiu de Sus (Romania).
|
||
<emphasis id="B949EAD5FF9AFFB0FC81F8866B80F886" bold="true" box="[940,1010,1837,1867]" pageId="24" pageNumber="25">
|
||
C
|
||
<subScript id="17B93482FF9AFFB0FC92F8966BB4F88B" attach="right" box="[959,966,1853,1867]" fontSize="6" pageId="24" pageNumber="25">1</subScript>
|
||
–C
|
||
<subScript id="17B93482FF9AFFB0FCC8F8966B9EF88B" attach="left" box="[997,1004,1853,1867]" fontSize="6" pageId="24" pageNumber="25">3</subScript>
|
||
.
|
||
</emphasis>
|
||
‘
|
||
<taxonomicName id="4C3D4D44FF9AFFB0FB2DF8856C16F886" authorityName="Lamarck" authorityYear="1801" box="[1024,1124,1838,1862]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="25" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF9AFFB0FB2DF8856C16F886" box="[1024,1124,1838,1862]" italics="true" pageId="24" pageNumber="25">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<taxonomicName id="4C3D4D44FF9AFFB0FB5DF88568A7F8AA" authority="(Hoernes & Auinger, 1891)" baseAuthorityName="Hoernes & Auinger" baseAuthorityYear="1891" class="Gastropoda" family="Turridae" genus="Pleurotoma" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="camillae">
|
||
<emphasis id="B949EAD5FF9AFFB0FB5DF8856CB5F886" box="[1136,1223,1838,1862]" italics="true" pageId="24" pageNumber="25">camillae</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF9AFFB0FBFEF88568BFF8AA" author="Hoernes, R. & Auinger, M." pageId="24" pageNumber="25" pagination="283 - 382" refId="ref135318" refString="Hoernes, R. & Auinger, M. (1891) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der kaiserlich-koniglichen Geologischen Reichsanstalt, 12, 283 - 382, pls 37 - 50. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf]" type="journal article" year="1891">Hoernes & Auinger, 1891</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, lectotype, NHMW 1949/0005/0042, Lăpugiu de Sus (Romania).
|
||
<emphasis id="B949EAD5FF9AFFB0FC49F8F96BD8F8AA" bold="true" box="[868,938,1874,1903]" pageId="24" pageNumber="25">
|
||
D
|
||
<subScript id="17B93482FF9AFFB0FC5AF8CA6B0CF8AF" attach="right" box="[887,894,1889,1903]" fontSize="6" pageId="24" pageNumber="25">1</subScript>
|
||
–D
|
||
<subScript id="17B93482FF9AFFB0FCB0F8CA6BD6F8AF" attach="left" box="[925,932,1889,1903]" fontSize="6" pageId="24" pageNumber="25">3</subScript>
|
||
.
|
||
</emphasis>
|
||
‘
|
||
<taxonomicName id="4C3D4D44FF9AFFB0FC95F8F96C6EF8AA" authorityName="Lamarck" authorityYear="1801" box="[952,1052,1874,1898]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="25" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF9AFFB0FC95F8F96C6EF8AA" box="[952,1052,1874,1898]" italics="true" pageId="24" pageNumber="25">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<taxonomicName id="4C3D4D44FF9AFFB0FB04F8F96DEAF8AA" authority="(Hoernes & Auinger, 1891)" baseAuthorityName="Hoernes & Auinger" baseAuthorityYear="1891" box="[1065,1432,1874,1898]" class="Gastropoda" family="Turridae" genus="Pleurotoma" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="camillae">
|
||
<emphasis id="B949EAD5FF9AFFB0FB04F8F96CF2F8AA" box="[1065,1152,1874,1898]" italics="true" pageId="24" pageNumber="25">camillae</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF9AFFB0FBA1F8F96DE3F8AA" author="Hoernes, R. & Auinger, M." box="[1164,1425,1874,1898]" pageId="24" pageNumber="25" pagination="283 - 382" refId="ref135318" refString="Hoernes, R. & Auinger, M. (1891) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der kaiserlich-koniglichen Geologischen Reichsanstalt, 12, 283 - 382, pls 37 - 50. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf]" type="journal article" year="1891">Hoernes & Auinger, 1891</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, paralectotype, NHMW 1949/0005/0042b, Lăpugiu de Sus (Romania),
|
||
<emphasis id="B949EAD5FF9AFFB0FC70F8DD6BD2F84E" bold="true" box="[861,928,1910,1939]" pageId="24" pageNumber="25">
|
||
E
|
||
<subScript id="17B93482FF9AFFB0FC43F82E6B07F853" attach="left" box="[878,885,1925,1939]" fontSize="6" pageId="24" pageNumber="25">1</subScript>
|
||
–E
|
||
<subScript id="17B93482FF9AFFB0FCBEF82E6BE8F853" attach="left" box="[915,922,1925,1939]" fontSize="6" pageId="24" pageNumber="25">2</subScript>
|
||
.
|
||
</emphasis>
|
||
‘
|
||
<taxonomicName id="4C3D4D44FF9AFFB0FC82F8DD6C61F84E" authorityName="Lamarck" authorityYear="1801" box="[943,1043,1910,1934]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="25" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF9AFFB0FC82F8DD6C61F84E" box="[943,1043,1910,1934]" italics="true" pageId="24" pageNumber="25">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<emphasis id="B949EAD5FF9AFFB0FB32F8DD6C08F84E" box="[1055,1146,1910,1934]" italics="true" pageId="24" pageNumber="25">atatuerki</emphasis>
|
||
nov. sp., holotype,. NHMW 1847/0058/1216, Seyithasan (Turkey).
|
||
<emphasis id="B949EAD5FF9AFFB0FD01F8316A1FF872" bold="true" box="[556,621,1946,1975]" pageId="24" pageNumber="25">
|
||
F
|
||
<subScript id="17B93482FF9AFFB0FD11F8026A31F877" attach="right" box="[572,579,1961,1975]" fontSize="6" pageId="24" pageNumber="25">1</subScript>
|
||
–F
|
||
<subScript id="17B93482FF9AFFB0FD72F8026A14F877" attach="left" box="[607,614,1961,1975]" fontSize="6" pageId="24" pageNumber="25">2</subScript>
|
||
.
|
||
</emphasis>
|
||
‘
|
||
<taxonomicName id="4C3D4D44FF9AFFB0FD53F8316A90F872" authorityName="Lamarck" authorityYear="1801" box="[638,738,1946,1970]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="25" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF9AFFB0FD53F8316A90F872" box="[638,738,1946,1970]" italics="true" pageId="24" pageNumber="25">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<emphasis id="B949EAD5FF9AFFB0FDDFF8316B3FF872" box="[754,845,1946,1970]" italics="true" pageId="24" pageNumber="25">atatuerki</emphasis>
|
||
nov. sp., paratype, NHMW 1847/0058/1217, Seyithasan (Turkey).
|
||
<emphasis id="B949EAD5FF9AFFB0FFD0F8166934F816" bold="true" box="[253,326,1981,2011]" pageId="24" pageNumber="25">
|
||
G
|
||
<subScript id="17B93482FF9AFFB0FE3CF866696AF81B" attach="left" box="[273,280,1997,2011]" fontSize="6" pageId="24" pageNumber="25">1</subScript>
|
||
–G
|
||
<subScript id="17B93482FF9AFFB0FE15F866694DF81B" attach="left" box="[312,319,1997,2011]" fontSize="6" pageId="24" pageNumber="25">2</subScript>
|
||
.
|
||
</emphasis>
|
||
‘
|
||
<taxonomicName id="4C3D4D44FF9AFFB0FE78F81569CBF816" authorityName="Lamarck" authorityYear="1801" box="[341,441,1982,2006]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="25" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B949EAD5FF9AFFB0FE78F81569CBF816" box="[341,441,1982,2006]" italics="true" pageId="24" pageNumber="25">Clavatula</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<taxonomicName id="4C3D4D44FF9AFFB0FEE5F8156B7AF816" authority="Kovacs & Vician, 2021" authorityName="Kovacs & Vician" authorityYear="2021" box="[456,776,1982,2006]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="santhai">
|
||
<emphasis id="B949EAD5FF9AFFB0FEE5F8156A61F816" box="[456,531,1982,2006]" italics="true" pageId="24" pageNumber="25">santhai</emphasis>
|
||
<bibRefCitation id="EFAC4B36FF9AFFB0FD34F8156B7AF816" author="Kovacs, Z. & Vician, Z." box="[537,776,1982,2006]" pageId="24" pageNumber="25" pagination="137 - 158" refId="ref137046" refString="Kovacs, Z. & Vician, Z. (2021) Middle Miocene Conoidea (Neogastropoda) assemblage of Letkes (Hungary), Part II. (Borsoniidae, Cochlespiridae, Clavatulidae, Turridae, Fusiturridae). Foldtani Kozlony, 151, 137 - 158. https: // doi. org / 10.23928 / foldt. kozl. 2021.151.2.137" type="journal article" year="2021">Kovács & Vicián, 2021</bibRefCitation>
|
||
</taxonomicName>
|
||
, holotype, HNHM, PAL 2021.20.1, Letkés (Hungary).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C327654CFF9BFFB1FFEAFF3D692EFF39" pageId="25" pageNumber="26" type="diagnosis">
|
||
<paragraph id="8B8236C7FF9BFFB1FFEAFF3D692EFF39" blockId="25.[151,1437,150,1474]" pageId="25" pageNumber="26">
|
||
<emphasis id="B949EAD5FF9BFFB1FFEAFF3D6930FF70" bold="true" box="[199,322,150,176]" pageId="25" pageNumber="26">Diagnosis.</emphasis>
|
||
Medium-sized, moderately broad biconic, with coronate spire, open spines on subsutural collar, last whorl with sharply angular shoulder, cancellate sculpture at mid-whorl, base with irregular cords, siphonal canal moderately short.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C327654CFF9BFFB1FFEAFEA969A8FCF9" pageId="25" pageNumber="26" type="description">
|
||
<paragraph id="8B8236C7FF9BFFB1FFEAFEA969A8FCF9" blockId="25.[151,1437,150,1474]" pageId="25" pageNumber="26">
|
||
<emphasis id="B949EAD5FF9BFFB1FFEAFEA9692BFEDC" bold="true" box="[199,345,258,284]" pageId="25" pageNumber="26">Description.</emphasis>
|
||
Shell medium-sized, moderately broad biconic, solid, with coronate conical spire, apical angle about 45–47°. Protoconch not preserved. Teleoconch of up to nine whorls (earliest whorls not preserved). Early teleoconch whorls with bipartite sculpture; weakly granulose subsutural cord, fine spiral threads in concave portion at mid-whorl, bifid beaded suprasutural cord in apposition to abapical suture. On the sixth spire whorl sculpture changes abruptly. Sharp open spines appear on subsutural collar. Spines slightly bent adapically, mid-whorl portion concave, weakly striate. Beads along abapical suture broaden into low triangular riblets. Suture narrowly impressed, weakly undulating. Last whorl 60% of total height; subsutural collar bearing 12–14 sharp spiny tubercles; subsutural ramp very broad, steep, concave, weakly spirally striate, with weak ribs on abapical half of ramp strengthening towards shoulder; shoulder sharp; ribs most strongly developed at mid-whorl, crossing three spiral cords resulting in cancellate mid-whorlsculpture. Base moderately well delimited by strengthened peribasal cord. Base strongly constricted, concave, bearing cords of irregular strength, two stronger just above the siphonal fasciole, beaded by axial ribs that weaken abapically. Siphonal fasciole poorly delimited, bearing numerous fine spiral cords. Aperture ovate, outer lip not thickened, smooth within. Anal sinus moderately wide, moderately deep, asymmetrically Ushaped. Siphonal canal moderately short, shallowly notched at tip. Columella strongly excavated in adapical third, straight below, weakly twisted, smooth. Columellar and parietal callus thickened, sharply delimited, forming moderately broad callus rim.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C327654CFF9BFFB1FFEAFCE96A73FACD" pageId="25" pageNumber="26" type="discussion">
|
||
<paragraph id="8B8236C7FF9BFFB1FFEAFCE96D79FC65" blockId="25.[151,1437,150,1474]" pageId="25" pageNumber="26">
|
||
<emphasis id="B949EAD5FF9BFFB1FFEAFCE9693EFC9C" bold="true" box="[199,332,834,860]" pageId="25" pageNumber="26">Discussion.</emphasis>
|
||
<taxonomicName id="4C3D4D44FF9BFFB1FE78FCE86A7FFC9C" authorityName="Harzhauser & Landau & Janssen" authorityYear="2022" box="[341,525,835,861]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="sorini">
|
||
<emphasis id="B949EAD5FF9BFFB1FE78FCE86A7FFC9C" box="[341,525,835,861]" italics="true" pageId="25" pageNumber="26">Clavatula sorini</emphasis>
|
||
</taxonomicName>
|
||
is morphologically very close to the extant
|
||
<taxonomicName id="4C3D4D44FF9BFFB1FB25FCE86D2BFC9D" authority="(Lamarck, 1822)" authorityName="" baseAuthorityName="Lamarck" baseAuthorityYear="1822" box="[1032,1369,835,861]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="muricata">
|
||
<emphasis id="B949EAD5FF9BFFB1FB25FCE86CFDFC9C" box="[1032,1167,835,861]" italics="true" pageId="25" pageNumber="26">C. muricata</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF9BFFB1FB8DFCE86D23FC9D" author="Lamarck, J. - B. M. de" box="[1184,1361,835,861]" pageId="25" pageNumber="26" refId="ref137485" refString="Lamarck, J. - B. M. de. (1822) Histoire naturelle des animaux sans vertebres. Tome Septieme. Published by the Author, Paris, 711 pp. [http: // www. biodiversitylibrary. org / item / 47433]" type="book" year="1822">Lamarck, 1822</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Both species are coronate with prominent, open spines and sharp shoulder.
|
||
<taxonomicName id="4C3D4D44FF9BFFB1FC97FCCC6CE5FC40" authorityName="" baseAuthorityName="Lamarck" baseAuthorityYear="1822" box="[954,1175,871,897]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="muricata">
|
||
<emphasis id="B949EAD5FF9BFFB1FC97FCCC6CE5FC40" box="[954,1175,871,897]" italics="true" pageId="25" pageNumber="26">Clavatula muricata</emphasis>
|
||
</taxonomicName>
|
||
, however differs in its more conical spire, the narrower anal sinus and the less prominent and raised spiral cords on the base.
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FF9BFFB1FFEAFC046A73FACD" blockId="25.[151,1437,150,1474]" pageId="25" pageNumber="26">
|
||
When describing this species,
|
||
<bibRefCitation id="EFAC4B36FF9BFFB1FD1FFC046B1DFC09" author="Hoernes, R. & Auinger, M." box="[562,879,943,969]" pageId="25" pageNumber="26" pagination="283 - 382" refId="ref135318" refString="Hoernes, R. & Auinger, M. (1891) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der kaiserlich-koniglichen Geologischen Reichsanstalt, 12, 283 - 382, pls 37 - 50. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf]" type="journal article" year="1891">Hoernes & Auinger (1891)</bibRefCitation>
|
||
referred to a species established by
|
||
<bibRefCitation id="EFAC4B36FF9BFFB1FA01FC0468A9FC2D" author="Grateloup, J. P. S. de" pageId="25" pageNumber="26" pagination="314 - 344" refId="ref133310" refString="Grateloup, J. P. S. de (1832) Tableau des coquilles Fossiles qu'on rencontre dans les terrains calcaires tertiaries (faluns) des environs de Dax, dans le departement des Landes. Actes de la Societe Linneenne de Bordeaux, 5, 314 - 344. [https: // www. biodiversitylibrary. org / item / 102890 # page / 325 / mode / 1 up]" type="journal article" year="1832">Grateloup (1832</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFAC4B36FF9BFFB1FFCAFC786952FC2E" author="Grateloup, J. P. S. de" box="[231,288,979,1006]" pageId="25" pageNumber="26" pagination="1 - 12" refId="ref133384" refString="Grateloup, J. P. S. de (1846) s. n. In: Conchyliologie fossile des terrains tertiaires du Bassin de l'Adour, (environs de Dax). Tome Ier. Univalves. Atlas. Th. Lafargue, Bordeaux, pls. 1 - 45 (1840) + pp. i - xx + 1 - 12 pp., pls. 46 - 48 (1846). [https: // archive. org / details / conchyliologief 00 gratgoog]" type="book chapter" year="1846">1846</bibRefCitation>
|
||
) from the Burdigalian of Dax in the Aquitaine Basin as
|
||
<taxonomicName id="4C3D4D44FF9BFFB1FCB5FC786C0AFC2C" authorityName="Grateloup" authorityYear="1832" box="[920,1144,979,1005]" class="Gastropoda" family="Turridae" genus="Pleurotoma" kingdom="Animalia" order="Neogastropoda" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="spinosa">
|
||
<emphasis id="B949EAD5FF9BFFB1FCB5FC786C0AFC2C" box="[920,1144,979,1005]" italics="true" pageId="25" pageNumber="26">Pleurotoma spinosa</emphasis>
|
||
</taxonomicName>
|
||
. The status of this French species is not fully clear and
|
||
<bibRefCitation id="EFAC4B36FF9BFFB1FECCFC5C6ADBFBD1" author="Peyrot, A." box="[481,681,1015,1041]" pageId="25" pageNumber="26" pagination="5 - 288" refId="ref141284" refString="Peyrot, A. (1931 & 1933) Conchologie neogenique de l'Aquitaine. Actes de la Societe Linneenne de Bordeaux, 83, 5 - 116 (1931), 84, 5 - 288 (1933). [https: // www. biodiversitylibrary. org / item / 185325 # page / 11 / mode / 1 up]" type="journal article" year="1931">Peyrot (1931: 18)</bibRefCitation>
|
||
treated it as synonym of
|
||
<taxonomicName id="4C3D4D44FF9BFFB1FCE5FC5C6D09FBD1" authority="(Lamarck, 1822)" baseAuthorityName="Lamarck" baseAuthorityYear="1822" box="[968,1403,1015,1041]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="asperulata">
|
||
<emphasis id="B949EAD5FF9BFFB1FCE5FC5C6CC5FBD1" box="[968,1207,1015,1041]" italics="true" pageId="25" pageNumber="26">Clavatula asperulata</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC4B36FF9BFFB1FBEBFC5C6D01FBD1" author="Lamarck, J. - B. M. de" box="[1222,1395,1015,1041]" pageId="25" pageNumber="26" refId="ref137485" refString="Lamarck, J. - B. M. de. (1822) Histoire naturelle des animaux sans vertebres. Tome Septieme. Published by the Author, Paris, 711 pp. [http: // www. biodiversitylibrary. org / item / 47433]" type="book" year="1822">Lamarck, 1822</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
[=
|
||
<taxonomicName id="4C3D4D44FF9BFFB1FFBAFBB069B2FBF5" baseAuthorityName="Lamarck" baseAuthorityYear="1822" box="[151,448,1051,1077]" class="Gastropoda" family="Clavatulidae" genus="Megaclavatula" kingdom="Animalia" order="Neogastropoda" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="asperulata">
|
||
<emphasis id="B949EAD5FF9BFFB1FFBAFBB069B2FBF5" box="[151,448,1051,1077]" italics="true" pageId="25" pageNumber="26">Megaclavatula asperulata</emphasis>
|
||
</taxonomicName>
|
||
]. The illustrations of
|
||
<bibRefCitation id="EFAC4B36FF9BFFB1FD94FBB06B04FBF5" author="Grateloup, J. P. S. de" box="[697,886,1051,1077]" pageId="25" pageNumber="26" pagination="1 - 12" refId="ref133384" refString="Grateloup, J. P. S. de (1846) s. n. In: Conchyliologie fossile des terrains tertiaires du Bassin de l'Adour, (environs de Dax). Tome Ier. Univalves. Atlas. Th. Lafargue, Bordeaux, pls. 1 - 45 (1840) + pp. i - xx + 1 - 12 pp., pls. 46 - 48 (1846). [https: // archive. org / details / conchyliologief 00 gratgoog]" type="book chapter" year="1846">Grateloup (1846</bibRefCitation>
|
||
, pl. 19, fig. 24) and
|
||
<bibRefCitation id="EFAC4B36FF9BFFB1FB4FFBB06C85FBF5" author="Peyrot, A." box="[1122,1271,1051,1077]" pageId="25" pageNumber="26" pagination="5 - 288" refId="ref141284" refString="Peyrot, A. (1931 & 1933) Conchologie neogenique de l'Aquitaine. Actes de la Societe Linneenne de Bordeaux, 83, 5 - 116 (1931), 84, 5 - 288 (1933). [https: // www. biodiversitylibrary. org / item / 185325 # page / 11 / mode / 1 up]" type="journal article" year="1931">Peyrot (1931</bibRefCitation>
|
||
, pl. 5, figs 11, 15) differ clearly from the Paratethyan species in their slender outline, high and flat-sided early spire whorls, and the much more prominent and spiny nodes on the penultimate and last whorls. These features distinguish also the late Miocene species from Stazzano and S. Agata in
|
||
<collectingCountry id="F32A7657FF9BFFB1FDC4FB2C6B6FFB61" box="[745,797,1159,1185]" name="Italy" pageId="25" pageNumber="26">Italy</collectingCountry>
|
||
, which were erroneously identified as
|
||
<taxonomicName id="4C3D4D44FF9BFFB1FBFCFB2C6975FB05" authority="Grateloup" authorityName="Grateloup" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="spinosa">
|
||
<emphasis id="B949EAD5FF9BFFB1FBFCFB2C6DEFFB60" box="[1233,1437,1159,1185]" italics="true" pageId="25" pageNumber="26">Clavatula spinosa</emphasis>
|
||
Grateloup
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFAC4B36FF9BFFB1FE1FFB006993FB05" author="Bellardi, L" box="[306,481,1195,1221]" pageId="25" pageNumber="26" refId="ref129307" refString="Bellardi, L (1877) I molluschi dei terreni terziarii del Piemonte e della Liguria. Parte II. Gasteropoda (Pleurotomidae). Stamperia Reale, Torino, 342 pp. phttps: // www. biodiversitylibrary. org / item / 45375 # page / 5 / mode / 1 up [" type="book" year="1877">Bellardi (1877)</bibRefCitation>
|
||
. The Italian shells differ in their more slender shape, longer siphonal canal, narrower last whorl, only one stronger subperipheral spiral cord on the basis and more numerous but less spiny nodes on the shoulder (pers. obs. R. Janssen).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C327654CFF9BFFB1FFEAFABD6A39FA96" pageId="25" pageNumber="26" type="biology_ecology">
|
||
<paragraph id="8B8236C7FF9BFFB1FFEAFABD6A39FA96" blockId="25.[151,1437,150,1474]" pageId="25" pageNumber="26">
|
||
<emphasis id="B949EAD5FF9BFFB1FFEAFABD69DAFAF0" bold="true" box="[199,424,1302,1328]" pageId="25" pageNumber="26">Paleoenvironment.</emphasis>
|
||
Unknown, probably middle to outer neritic environments based on assemblages from Lăpugiu de Sus (
|
||
<collectingCountry id="F32A7657FF9BFFB1FE79FA9069CFFA96" box="[340,445,1339,1366]" name="Romania" pageId="25" pageNumber="26">Romania</collectingCountry>
|
||
) (own data).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C327654CFF9BFFB1FFEAFAF56DE2FA01" pageId="25" pageNumber="26" type="distribution">
|
||
<paragraph id="8B8236C7FF9BFFB1FFEAFAF569B9FA5D" blockId="25.[151,1437,150,1474]" pageId="25" pageNumber="26">
|
||
<emphasis id="B949EAD5FF9BFFB1FFEAFAF56A03FAB9" bold="true" box="[199,625,1374,1401]" pageId="25" pageNumber="26">Distribution in Central Paratethys.</emphasis>
|
||
Badenian (middle Miocene):
|
||
<emphasis id="B949EAD5FF9BFFB1FCE1FAF46C24FAB8" box="[972,1110,1375,1401]" italics="true" pageId="25" pageNumber="26">Făget Basin</emphasis>
|
||
: Lăpugiu de Sus (
|
||
<collectingCountry id="F32A7657FF9BFFB1FA01FAF46DE7FABA" box="[1324,1429,1375,1402]" name="Romania" pageId="25" pageNumber="26">Romania</collectingCountry>
|
||
) (
|
||
<bibRefCitation id="EFAC4B36FF9BFFB1FFB2FA2869CDFA5D" author="Hoernes, R. & Auinger, M." box="[159,447,1411,1437]" pageId="25" pageNumber="26" pagination="283 - 382" refId="ref135318" refString="Hoernes, R. & Auinger, M. (1891) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der kaiserlich-koniglichen Geologischen Reichsanstalt, 12, 283 - 382, pls 37 - 50. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf]" type="journal article" year="1891">Hoernes & Auinger 1891</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8236C7FF9BFFB1FFEAFA0D6DE2FA01" blockId="25.[151,1437,150,1474]" box="[199,1424,1446,1474]" pageId="25" pageNumber="26">
|
||
<emphasis id="B949EAD5FF9BFFB1FFEAFA0D6A73FA00" bold="true" box="[199,513,1446,1472]" pageId="25" pageNumber="26">Proto-Mediterranean Sea:</emphasis>
|
||
? Tortonian (late Miocene):
|
||
<emphasis id="B949EAD5FF9BFFB1FC6FFA0C6BDAFA00" box="[834,936,1447,1472]" italics="true" pageId="25" pageNumber="26">Po Basin</emphasis>
|
||
: S. Agata, Stazzano (
|
||
<collectingCountry id="F32A7657FF9BFFB1FBBAFA0C6CBFFA02" box="[1175,1229,1447,1474]" name="Italy" pageId="25" pageNumber="26">Italy</collectingCountry>
|
||
) (
|
||
<bibRefCitation id="EFAC4B36FF9BFFB1FBC9FA0C6DF1FA01" author="Bellardi, L" box="[1252,1411,1447,1473]" pageId="25" pageNumber="26" refId="ref129307" refString="Bellardi, L (1877) I molluschi dei terreni terziarii del Piemonte e della Liguria. Parte II. Gasteropoda (Pleurotomidae). Stamperia Reale, Torino, 342 pp. phttps: // www. biodiversitylibrary. org / item / 45375 # page / 5 / mode / 1 up [" type="book" year="1877">Bellardi 1877</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |