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<document id="C56401DB07D88E1D9177CF105A2C2374" ID-CLB-Dataset="7126" ID-DOI="10.12782/sd.19.1.043" ID-GBIF-Dataset="238dd2d3-f38f-4e11-a2be-87c16ff21dd8" ID-ISSN="2189-7301" ID-Zenodo-Dep="4584938" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1614946654621" checkinUser="felipe" docAuthor="Dwibadra, Dhian, Takaku, Gen, Ôhara, Masahiro &amp; Ueda, Akira" docDate="2014" docId="0C62878DFFE9215BFF30FCEEFA94FC17" docLanguage="en" docName="SpeciesDiversity.19.1.43-57.pdf" docOrigin="Species Diversity 19 (1)" docSource="http://dx.doi.org/10.12782/sd.19.1.043" docStyle="DocumentStyle:27FECD2C3CD153DA405029F2D33C9514.7:SpeciesDiversity.2012-.journal_article" docStyleId="27FECD2C3CD153DA405029F2D33C9514" docStyleName="SpeciesDiversity.2012-.journal_article" docStyleVersion="7" docTitle="Macrocheles riparius Dwibadra and Takaku 2014, sp. nov." docType="treatment" docUuid="7E4E0752-0096-4467-A57A-BDD8E9CAAB4D" docUuidSource="ZooBank" docVersion="6" lastPageNumber="50" masterDocId="F05BFFF5FFEE215CFFF5FFE1FF89FFDD" masterDocTitle="Mites of the Family Macrochelidae (Acari: Gamasida) from Sungai Wain, East Kalimantan, Indonesia" masterLastPageNumber="57" masterPageNumber="43" pageNumber="50" updateTime="1698932732925" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="7056411A3A7DDB44FFAF5444011BE19F">
<mods:title id="E3E3687B4A0C046398C562D69D226EDD">Mites of the Family Macrochelidae (Acari: Gamasida) from Sungai Wain, East Kalimantan, Indonesia</mods:title>
</mods:titleInfo>
<mods:name id="704D746189DDFE0DC3AD2F1CE82D4C0E" type="personal">
<mods:role id="E178C7B744F3946FEC204E7329275590">
<mods:roleTerm id="AB35EE6BB7322DB7867A569271376A81">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1A9DA0F6AD7CAF8968394DF1F1C1D9A7">Dwibadra, Dhian</mods:namePart>
<mods:affiliation id="CD330FCFDBC836EB426A2A767230754F">Systematic Entomology Laboratory, Graduate School of Agriculture, Hokkaido University, Sapporo 060 - 8589, Japan E-mail: dwibadra _ yk @ yahoo. com &amp; Corresponding author</mods:affiliation>
<mods:nameIdentifier id="AAE77013563E72C801B069D8CA63CD55" type="email">dwibadra_yk@yahoo.com</mods:nameIdentifier>
</mods:name>
<mods:name id="773CE8DE63C4A13EE2C5193E639702D0" type="personal">
<mods:role id="8D0EAF149180286D3DAAF4174721E50A">
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</mods:role>
<mods:namePart id="2CAC069BA5D7C4F4798E4FC564CE6EB7">Takaku, Gen</mods:namePart>
<mods:affiliation id="3B57D728C5BB3FBCBFD9EB35EDA50F89">Biological Laboratory, Hokkaido University of Education Sapporo, 5 - 3 - 1 Ainosato, Kita-ku, Sapporo 002 - 8502, Japan</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="59BA3F92A3700DA9FF2205D21A09EA3B">Ôhara, Masahiro</mods:namePart>
<mods:affiliation id="7A8BC19F54B1EB38634BD43D4C0370D9">The Hokkaido University Museum, N 10 W 8, Kita-ku, Sapporo 060 - 0810, Japan</mods:affiliation>
</mods:name>
<mods:name id="A9D0A3530811D91ED452A524016DDDD8" type="personal">
<mods:role id="EF25CE08E27BFFE43980122689D1937E">
<mods:roleTerm id="1967F3BD31A5D09C7053B2BF15E13EBC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="25B074702B8B065A7766131C9E1F7F1F">Ueda, Akira</mods:namePart>
<mods:affiliation id="22B38D910B929943BBF0BD566B0926E5">Kyushu Research Center, Forestry and Forest Products Research Institute (FFPRI), 4 - 11 - 16 Kurokami, Kumamoto 860 - 0862, Japan</mods:affiliation>
</mods:name>
<mods:typeOfResource id="019EF0C14E433748C175C474B7709655">text</mods:typeOfResource>
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<mods:titleInfo id="3B5446AFF507F058BD28500EB0256E94">
<mods:title id="81C3E69E6F2CEA68470BEBD5F5E6D534">Species Diversity</mods:title>
</mods:titleInfo>
<mods:part id="0B8C1B4D8D84CF636C51FF41033DE78D">
<mods:date id="E4F04407BF43100F2AB0CE186F1C9915">2014</mods:date>
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<mods:number id="CC1429F0C7FB664C1819AFE28CF921C6">2014-05-25</mods:number>
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<mods:number id="3B4B465A6E8118AD68CBDA83075C17B4">1</mods:number>
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<mods:start id="93AAED8264CCCA35C39C9769653DD23D">43</mods:start>
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<mods:identifier id="BB15692195546D564118BFD21A7F749E" type="DOI">10.12782/sd.19.1.043</mods:identifier>
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<treatment id="0C62878DFFE9215BFF30FCEEFA94FC17" ID-DOI="http://doi.org/10.5281/zenodo.4584932" ID-GBIF-Taxon="181762132" ID-Zenodo-Dep="4584932" LSID="urn:lsid:zoobank.org:act:7E4E0752-0096-4467-A57A-BDD8E9CAAB4D" httpUri="http://treatment.plazi.org/id/0C62878DFFE9215BFF30FCEEFA94FC17" lastPageNumber="50" pageId="7" pageNumber="50">
<subSubSection id="CCD16510FFE9215BFF30FCEEFE7BFC94" pageId="7" pageNumber="50" type="nomenclature">
<paragraph id="8474369BFFE9215BFF30FCEEFE7BFC94" blockId="7.[197,727,783,841]" pageId="7" pageNumber="50">
<heading id="DF3C81F7FFE9215BFF30FCEEFE7BFC94" centered="true" fontSize="10" level="2" pageId="7" pageNumber="50" reason="2">
<taxonomicName id="43CB4D18FFE9215BFF30FCEEFD0CFCF5" authority="Dwibadra and Takaku" authorityName="Dwibadra and Takaku" authorityYear="2014" box="[197,645,783,808]" class="Arachnida" family="Macrochelidae" genus="Macrocheles" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="50" phylum="Arthropoda" rank="species" species="riparius" status="sp. nov.">
<emphasis id="B6BFEA89FFE9215BFF30FCEEFE17FCF5" bold="true" box="[197,414,783,808]" italics="true" pageId="7" pageNumber="50">Macrocheles riparius</emphasis>
Dwibadra and Takaku
</taxonomicName>
<taxonomicNameLabel id="AD8C57F2FFE9215BFD7FFCF1FD5EFCFA" box="[650,727,784,807]" pageId="7" pageNumber="50" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="1CF02A1EFFE9215BFE45FCD1FE63FC94" box="[432,490,816,841]" captionStart="Fig" captionStartId="8.[156,187,1889,1911]" captionTargetBox="[222,1368,209,1849]" captionTargetId="figure-7@8.[218,1370,207,1851]" captionTargetPageId="8" captionText="Fig. 2. Macrocheles riparius Dwibadra and Takaku, sp. nov., female, holotype (MZB. Acar. 7566). A, dorsal shield (paratype, MZB. Acar. 7565.2); B, ventral shield; C, ventral view of gnathosoma (paratype, MZB. Acar. 7565.2); D, chelicera; E, tectum; F, sacculus foeminus (paratype, MZB. Acar. 8205). Scale bars: 50µm." figureDoi="http://doi.org/10.5281/zenodo.4584944" httpUri="https://zenodo.org/record/4584944/files/figure.png" pageId="7" pageNumber="50">Fig. 2</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CCD16510FFE9215BFF40FC93FD73FB53" pageId="7" pageNumber="50" type="materials_examined">
<paragraph id="8474369BFFE9215BFF40FC93FD73FB53" blockId="7.[156,768,881,2077]" pageId="7" pageNumber="50">
<emphasis id="B6BFEA89FFE9215BFF40FC93FEB5FC54" bold="true" box="[181,316,881,906]" pageId="7" pageNumber="50">Type series.</emphasis>
<materialsCitation id="34A33CC6FFE9215BFEA3FC90FEB9FC36" ID-GBIF-Occurrence="3095730301" collectingDate="2006-12-20" collectionCode="MZB" collectorName="A. Ueda &amp; D. Dwibadra" location="Balikpapan" municipality="Hutan Lindung Sungai Wain" pageId="7" pageNumber="50" specimenCount="1" stateProvince="Kalimantan Timur" typeStatus="holotype">
<typeStatus id="5B708839FFE9215BFEA3FC90FE3DFC54" box="[342,436,881,905]" pageId="7" pageNumber="50" type="holotype">Holotype</typeStatus>
(
<collectionCode id="E2DAAE5EFFE9215BFE3BFC93FD8EFC57" box="[462,519,882,906]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="7" pageNumber="50" type="Museum">MZB</collectionCode>
. Acar. 7566): female,
<collectingMunicipality id="6410ACE1FFE9215BFF69FC73FE5DFC77" box="[156,468,914,938]" pageId="7" pageNumber="50">Hutan Lindung Sungai Wain</collectingMunicipality>
,
<location id="81146040FFE9215BFE11FC73FDD1FC77" LSID="urn:lsid:plazi:treatment:0C62878DFFE9215BFF30FCEEFA94FC17:81146040FFE9215BFE11FC73FDD1FC77" box="[484,600,914,938]" municipality="Hutan Lindung Sungai Wain" name="Balikpapan" pageId="7" pageNumber="50" stateProvince="Kalimantan Timur">Balikpapan</location>
,
<collectingRegion id="460FF879FFE9215BFD9DFC73FF37FC17" country="Indonesia" name="Kalimantan Timur" pageId="7" pageNumber="50">East Kalimantan</collectingRegion>
,
<date id="F075105BFFE9215BFF3EFC52FE19FC17" box="[203,400,946,971]" pageId="7" pageNumber="50" value="2006-12-20">
<collectingDate id="E031E9B3FFE9215BFF3EFC52FE19FC17" box="[203,400,946,971]" pageId="7" pageNumber="50" value="2006-12-20">20 December 2006</collectingDate>
</date>
,
<emphasis id="B6BFEA89FFE9215BFE69FC52FE3BFC17" box="[412,434,947,970]" italics="true" pageId="7" pageNumber="50">ex</emphasis>
residue in vial,
<collectorName id="293E534DFFE9215BFDA9FC52FD39FC17" box="[604,688,946,971]" pageId="7" pageNumber="50">A. Ueda</collectorName>
and
<collectorName id="293E534DFFE9215BFD1DFC52FE88FC36" pageId="7" pageNumber="50">D. Dwibadra</collectorName>
leg.
</materialsCitation>
<materialsCitation id="34A33CC6FFE9215BFECCFC35FD9EFBD6" ID-GBIF-Occurrence="3095730310" collectingDate="2006-12-17" collectionCode="SWPF" pageId="7" pageNumber="50" specimenCount="5" specimenCount-female="5" stateProvince="Kalimantan Timur" typeStatus="paratype">
<typeStatus id="5B708839FFE9215BFECCFC35FE14FC31" box="[313,413,980,1004]" pageId="7" pageNumber="50" type="paratype">Paratypes</typeStatus>
:
<emphasis id="B6BFEA89FFE9215BFE59FC32FE66FC36" bold="true" box="[428,495,979,1003]" pageId="7" pageNumber="50">
<collectionCode id="E2DAAE5EFFE9215BFE59FC32FE66FC36" box="[428,495,979,1003]" pageId="7" pageNumber="50">SWPF</collectionCode>
</emphasis>
:
<specimenCount id="92CDFD12FFE9215BFE0AFC32FDECFC36" box="[511,613,979,1003]" pageId="7" pageNumber="50" type="female">5 females</specimenCount>
,
<date id="F075105BFFE9215BFD86FC32FF46FBD6" pageId="7" pageNumber="50" value="2006-12-17">
<collectingDate id="E031E9B3FFE9215BFD86FC32FF46FBD6" pageId="7" pageNumber="50" value="2006-12-17">17 December 2006</collectingDate>
</date>
,
<emphasis id="B6BFEA89FFE9215BFF2DFC15FE70FBD6" box="[216,505,1012,1036]" italics="true" pageId="7" pageNumber="50">
ex
<taxonomicName id="43CB4D18FFE9215BFF00FC15FE70FBD6" authorityName="Harold" authorityYear="1877" box="[245,505,1012,1036]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="50" phylum="Arthropoda" rank="species" species="semicupreus">Onthophagus semicupreus</taxonomicName>
</emphasis>
,
<taxonomicName id="43CB4D18FFE9215BFDF0FC15FD9EFBD6" authorityName="Boucomont" authorityYear="1914" box="[517,535,1012,1035]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="50" phylum="Arthropoda" rank="species" species="waterstradti">
<emphasis id="B6BFEA89FFE9215BFDF0FC15FD9EFBD6" box="[517,535,1012,1035]" italics="true" pageId="7" pageNumber="50">O</emphasis>
</taxonomicName>
</materialsCitation>
.
<materialsCitation id="34A33CC6FFE9215BFDD6FC15FEC6FBB0" ID-GBIF-Occurrence="3095730325" collectingDate="2006-12-20" collectingDateMax="2008-12-12" collectingDateMin="2006-12-20" collectionCode="MZB" collectorName="A. Ueda &amp; D. Dwibadra" location="Balikpapan" municipality="Hutan Lindung Sungai Wain" pageId="7" pageNumber="50" specimenCount="3" specimenCount-female="3" stateProvince="Kalimantan Timur" typeStatus="paratype">
<emphasis id="B6BFEA89FFE9215BFDD6FC15FD13FBD1" box="[547,666,1012,1036]" italics="true" pageId="7" pageNumber="50">waterstradti</emphasis>
, and residue in vials;
<specimenCount id="92CDFD12FFE9215BFEEEFBF5FEF5FBF1" box="[283,380,1044,1068]" pageId="7" pageNumber="50" type="female">2 females</specimenCount>
,
<date id="F075105BFFE9215BFE70FBF5FDCFFBF1" box="[389,582,1044,1068]" pageId="7" pageNumber="50" value="2006-12-20">
<collectingDate id="E031E9B3FFE9215BFE70FBF5FDCFFBF1" box="[389,582,1044,1068]" pageId="7" pageNumber="50" value="2006-12-20">20 December 2006</collectingDate>
</date>
,
<emphasis id="B6BFEA89FFE9215BFDBAFBF4FDECFBF1" box="[591,613,1045,1068]" italics="true" pageId="7" pageNumber="50">ex</emphasis>
residue in vial;
<specimenCount id="92CDFD12FFE9215BFF69FBD4FF7FFB90" box="[156,246,1077,1101]" pageId="7" pageNumber="50" type="female">1 female</specimenCount>
,
<date id="F075105BFFE9215BFEF7FBD4FE41FB90" box="[258,456,1077,1101]" pageId="7" pageNumber="50" value="2008-12-12">
<collectingDate id="E031E9B3FFE9215BFEF7FBD4FE41FB90" box="[258,456,1077,1101]" pageId="7" pageNumber="50" value="2008-12-12">12 December 2008</collectingDate>
</date>
,
<emphasis id="B6BFEA89FFE9215BFE21FBD4FE63FB91" box="[468,490,1077,1100]" italics="true" pageId="7" pageNumber="50">ex</emphasis>
residue in vial, other data same as holotype
</materialsCitation>
.
<materialsCitation id="34A33CC6FFE9215BFEACFBB4FE65FB50" ID-GBIF-Occurrence="3095730321" collectingDate="2006-12-21" location="Balikpapan" pageId="7" pageNumber="50" specimenCount="2" specimenCount-female="2" stateProvince="Kalimantan Timur" typeStatus="paratype">
<emphasis id="B6BFEA89FFE9215BFEACFBB4FE29FBB0" bold="true" box="[345,416,1109,1133]" pageId="7" pageNumber="50">Km 24</emphasis>
:
<specimenCount id="92CDFD12FFE9215BFE5BFBB4FD98FBB0" box="[430,529,1109,1133]" pageId="7" pageNumber="50" type="female">2 females</specimenCount>
,
<location id="81146040FFE9215BFDEEFBB4FD06FBB0" LSID="urn:lsid:plazi:treatment:0C62878DFFE9215BFF30FCEEFA94FC17:81146040FFE9215BFDEEFBB4FD06FBB0" box="[539,655,1109,1133]" name="Balikpapan" pageId="7" pageNumber="50" stateProvince="Kalimantan Timur">Balikpapan</location>
,
<collectingRegion id="460FF879FFE9215BFD6EFBB7FF65FB50" country="Indonesia" name="Kalimantan Timur" pageId="7" pageNumber="50">East Kalimantan</collectingRegion>
,
<date id="F075105BFFE9215BFF00FB97FE3CFB53" box="[245,437,1142,1166]" pageId="7" pageNumber="50" value="2006-12-21">
<collectingDate id="E031E9B3FFE9215BFF00FB97FE3CFB53" box="[245,437,1142,1166]" pageId="7" pageNumber="50" value="2006-12-21">21 December 2006</collectingDate>
</date>
, ex
<taxonomicName id="43CB4D18FFE9215BFE2FFB97FE65FB50" authorityName="Vollenhoven" authorityYear="1864" box="[474,492,1142,1165]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="50" phylum="Arthropoda" rank="species" species="schwaneri">
<emphasis id="B6BFEA89FFE9215BFE2FFB97FE65FB50" box="[474,492,1142,1165]" italics="true" pageId="7" pageNumber="50">O</emphasis>
</taxonomicName>
</materialsCitation>
.
<emphasis id="B6BFEA89FFE9215BFE0DFB97FDD2FB53" box="[504,603,1142,1166]" italics="true" pageId="7" pageNumber="50">schwaneri</emphasis>
, residue in vial.
</paragraph>
</subSubSection>
<subSubSection id="CCD16510FFE9215BFF40FB77FB2BFDB9" pageId="7" pageNumber="50" type="description">
<paragraph id="8474369BFFE9215BFF40FB77FDEDFB32" blockId="7.[156,768,881,2077]" pageId="7" pageNumber="50">
<emphasis id="B6BFEA89FFE9215BFF40FB77FEB0FB73" bold="true" box="[181,313,1174,1198]" pageId="7" pageNumber="50">Description.</emphasis>
<emphasis id="B6BFEA89FFE9215BFEA6FB77FE13FB73" box="[339,410,1174,1198]" italics="true" pageId="7" pageNumber="50">Female</emphasis>
. Length of dorsal shield 731.5 (680770), width at level of coxae II 467.5 (410510) (
<emphasis id="B6BFEA89FFE9215BFF51FB39FF38FB32" box="[164,177,1240,1263]" italics="true" pageId="7" pageNumber="50">n</emphasis>
=10). Specimens yellowish brown in color.
</paragraph>
<paragraph id="8474369BFFE9215BFF40FB19FDA5FA4F" blockId="7.[156,768,881,2077]" pageId="7" pageNumber="50">
Dorsum (
<figureCitation id="1CF02A1EFFE9215BFEE3FB19FEECFACD" box="[278,357,1272,1296]" captionStart="Fig" captionStartId="8.[156,187,1889,1911]" captionTargetBox="[222,1368,209,1849]" captionTargetId="figure-7@8.[218,1370,207,1851]" captionTargetPageId="8" captionText="Fig. 2. Macrocheles riparius Dwibadra and Takaku, sp. nov., female, holotype (MZB. Acar. 7566). A, dorsal shield (paratype, MZB. Acar. 7565.2); B, ventral shield; C, ventral view of gnathosoma (paratype, MZB. Acar. 7565.2); D, chelicera; E, tectum; F, sacculus foeminus (paratype, MZB. Acar. 8205). Scale bars: 50µm." figureDoi="http://doi.org/10.5281/zenodo.4584944" httpUri="https://zenodo.org/record/4584944/files/figure.png" pageId="7" pageNumber="50">Fig. 2A</figureCitation>
). Dorsal shield with distinct punctations and reticulation; lateral margin of shield smooth; shield bearing 28 pairs of dorsal setae and 22 pairs of pores; seta j1 and Z5 pilose distally; J5 entirely pilose, r2 simple but sometimes slightly pilose; other setae simple.
</paragraph>
<paragraph id="8474369BFFE9215BFF40FA7AFE17F96B" blockId="7.[156,768,881,2077]" pageId="7" pageNumber="50">
Venter (
<figureCitation id="1CF02A1EFFE9215BFEFFFA7BFED3FA6E" box="[266,346,1434,1459]" captionStart="Fig" captionStartId="8.[156,187,1889,1911]" captionTargetBox="[222,1368,209,1849]" captionTargetId="figure-7@8.[218,1370,207,1851]" captionTargetPageId="8" captionText="Fig. 2. Macrocheles riparius Dwibadra and Takaku, sp. nov., female, holotype (MZB. Acar. 7566). A, dorsal shield (paratype, MZB. Acar. 7565.2); B, ventral shield; C, ventral view of gnathosoma (paratype, MZB. Acar. 7565.2); D, chelicera; E, tectum; F, sacculus foeminus (paratype, MZB. Acar. 8205). Scale bars: 50µm." figureDoi="http://doi.org/10.5281/zenodo.4584944" httpUri="https://zenodo.org/record/4584944/files/figure.png" pageId="7" pageNumber="50">Fig. 2B</figureCitation>
). Sternal shield wider than long; length 144 (130150), width at level of coxae II 154.5 (150160) (
<emphasis id="B6BFEA89FFE9215BFF51FA3DFF38FA2E" box="[164,177,1500,1523]" italics="true" pageId="7" pageNumber="50">n</emphasis>
=10); shield with 3 pairs of simple setae and 2 pairs of pores; l. ang. with punctation, disjunct from l. arc.; l. o. a. punctate; two l. arc. present as irregular punctate lines; l. m. t. punctate; l. o. p. present as paired transverse lines, with punctations, not connected to l. m. t.; two a. p. l. present. Metasternal shields oval and free; each shield with 1 simple seta and an anterior pore.
</paragraph>
<paragraph id="8474369BFFE9215BFF40F95EFD59F92A" blockId="7.[156,768,881,2077]" pageId="7" pageNumber="50">
Width of epigynial shield 184.5 (160200) (
<emphasis id="B6BFEA89FFE9215BFD86F921FD09F90A" box="[627,640,1728,1751]" italics="true" pageId="7" pageNumber="50">n</emphasis>
=10); shield ornamented with punctuations and pair of simple setae.
</paragraph>
<paragraph id="8474369BFFE9215BFF40F8E1FF73F867" blockId="7.[156,768,881,2077]" pageId="7" pageNumber="50">
Ventrianal shield longer than wide; length 260 (235280), width 221 (195245) (
<emphasis id="B6BFEA89FFE9215BFE7FF8C3FE1EF8E4" box="[394,407,1826,1849]" italics="true" pageId="7" pageNumber="50">n</emphasis>
=10); shield ornamented with reticulation and punctations; 3 pairs of preanal setae, 1 pair of paranal setae, and 1 postanal seta present; postanal seta pilose, other setae simple; cribrum located posterior to postanal seta.
</paragraph>
<paragraph id="8474369BFFE9215BFF40F822FC0EFEC2" blockId="7.[156,768,881,2077]" lastBlockId="7.[820,1432,198,970]" pageId="7" pageNumber="50">
Gnathosoma (
<figureCitation id="1CF02A1EFFE9215BFEBCF822FE15F801" box="[329,412,1987,2012]" captionStart="Fig" captionStartId="8.[156,187,1889,1911]" captionTargetBox="[222,1368,209,1849]" captionTargetId="figure-7@8.[218,1370,207,1851]" captionTargetPageId="8" captionText="Fig. 2. Macrocheles riparius Dwibadra and Takaku, sp. nov., female, holotype (MZB. Acar. 7566). A, dorsal shield (paratype, MZB. Acar. 7565.2); B, ventral shield; C, ventral view of gnathosoma (paratype, MZB. Acar. 7565.2); D, chelicera; E, tectum; F, sacculus foeminus (paratype, MZB. Acar. 8205). Scale bars: 50µm." figureDoi="http://doi.org/10.5281/zenodo.4584944" httpUri="https://zenodo.org/record/4584944/files/figure.png" pageId="7" pageNumber="50">Fig. 2C</figureCitation>
). Well developed and sclerotized, differing from that of
<taxonomicName id="43CB4D18FFE9215BFE79F805FDD8F821" authorityName="Dwibadra and Takaku" authorityYear="2014" box="[396,593,2020,2044]" class="Arachnida" family="Macrochelidae" genus="Macrocheles" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="50" phylum="Arthropoda" rank="species" species="dayaci">
<emphasis id="B6BFEA89FFE9215BFE79F805FDD8F821" box="[396,593,2020,2044]" italics="true" pageId="7" pageNumber="50">Macrocheles dayaci</emphasis>
</taxonomicName>
only in median process of epistome (
<figureCitation id="1CF02A1EFFE9215BFE8DF7E5FE4DF7C0" box="[376,452,2052,2077]" captionStart="Fig" captionStartId="8.[156,187,1889,1911]" captionTargetBox="[222,1368,209,1849]" captionTargetId="figure-7@8.[218,1370,207,1851]" captionTargetPageId="8" captionText="Fig. 2. Macrocheles riparius Dwibadra and Takaku, sp. nov., female, holotype (MZB. Acar. 7566). A, dorsal shield (paratype, MZB. Acar. 7565.2); B, ventral shield; C, ventral view of gnathosoma (paratype, MZB. Acar. 7565.2); D, chelicera; E, tectum; F, sacculus foeminus (paratype, MZB. Acar. 8205). Scale bars: 50µm." figureDoi="http://doi.org/10.5281/zenodo.4584944" httpUri="https://zenodo.org/record/4584944/files/figure.png" pageId="7" pageNumber="50">Fig. 2E</figureCitation>
) bearing many small spicules. Chelicera (
<figureCitation id="1CF02A1EFFE9215BFC53FF27FC71FF03" box="[934,1016,198,222]" captionStart="Fig" captionStartId="8.[156,187,1889,1911]" captionTargetBox="[222,1368,209,1849]" captionTargetId="figure-7@8.[218,1370,207,1851]" captionTargetPageId="8" captionText="Fig. 2. Macrocheles riparius Dwibadra and Takaku, sp. nov., female, holotype (MZB. Acar. 7566). A, dorsal shield (paratype, MZB. Acar. 7565.2); B, ventral shield; C, ventral view of gnathosoma (paratype, MZB. Acar. 7565.2); D, chelicera; E, tectum; F, sacculus foeminus (paratype, MZB. Acar. 8205). Scale bars: 50µm." figureDoi="http://doi.org/10.5281/zenodo.4584944" httpUri="https://zenodo.org/record/4584944/files/figure.png" pageId="7" pageNumber="50">Fig. 2D</figureCitation>
) same as in
<taxonomicName id="43CB4D18FFE9215BFB73FF26FB64FF03" authorityName="Dwibadra and Takaku" authorityYear="2014" box="[1158,1261,198,222]" class="Arachnida" family="Macrochelidae" genus="Macrocheles" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="50" phylum="Arthropoda" rank="species" species="dayaci">
<emphasis id="B6BFEA89FFE9215BFB73FF26FB15FF03" box="[1158,1180,199,222]" italics="true" pageId="7" pageNumber="50">M</emphasis>
.
<emphasis id="B6BFEA89FFE9215BFB59FF27FB64FF03" box="[1196,1261,198,222]" italics="true" pageId="7" pageNumber="50">dayaci</emphasis>
</taxonomicName>
; length of fixed digit 217.5 (210220) and movable digit 69.8 (67.572.5) (
<emphasis id="B6BFEA89FFE9215BFCC8FEE9FCC3FEC2" box="[829,842,264,287]" italics="true" pageId="7" pageNumber="50">n</emphasis>
=10).
</paragraph>
<paragraph id="8474369BFFE9215BFCBBFEC9FB5AFE3F" blockId="7.[820,1432,198,970]" pageId="7" pageNumber="50">
Legs. Most leg segments with simple and pilose setae except for coxae IIV, trochanters IIII and tarsus I with only simple setae. Leg chaetotaxy typical for this genus. Genu IV with 6 pilose setae. Leg length (excluding ambulacrum,
<emphasis id="B6BFEA89FFE9215BFCC1FE4AFCC8FE1F" box="[820,833,427,450]" italics="true" pageId="7" pageNumber="50">n</emphasis>
=10); leg I, 638.5 (610665); leg II, 615 (585680); leg III, 573.5 (525625); leg IV, 847.5 (815930).
</paragraph>
<paragraph id="8474369BFFE9215BFCBBFE0AFCECFD99" blockId="7.[820,1432,198,970]" pageId="7" pageNumber="50">
Sacculus foemineus (
<figureCitation id="1CF02A1EFFE9215BFBC7FE0AFB0DFDDE" box="[1074,1156,491,515]" captionStart="Fig" captionStartId="8.[156,187,1889,1911]" captionTargetBox="[222,1368,209,1849]" captionTargetId="figure-7@8.[218,1370,207,1851]" captionTargetPageId="8" captionText="Fig. 2. Macrocheles riparius Dwibadra and Takaku, sp. nov., female, holotype (MZB. Acar. 7566). A, dorsal shield (paratype, MZB. Acar. 7565.2); B, ventral shield; C, ventral view of gnathosoma (paratype, MZB. Acar. 7565.2); D, chelicera; E, tectum; F, sacculus foeminus (paratype, MZB. Acar. 8205). Scale bars: 50µm." figureDoi="http://doi.org/10.5281/zenodo.4584944" httpUri="https://zenodo.org/record/4584944/files/figure.png" pageId="7" pageNumber="50">Fig. 2F</figureCitation>
). Pair of sacculi present; cornu distinct, small and rounded distally; spermatheca oval.
</paragraph>
<paragraph id="8474369BFFE9215BFCBBFDADFB2BFDB9" blockId="7.[820,1432,198,970]" box="[846,1186,588,613]" pageId="7" pageNumber="50">Male and other stages. Unknown.</paragraph>
</subSubSection>
<subSubSection id="CCD16510FFE9215BFCBBFD8CFBA9FD78" pageId="7" pageNumber="50" type="etymology">
<paragraph id="8474369BFFE9215BFCBBFD8CFBA9FD78" blockId="7.[820,1432,198,970]" pageId="7" pageNumber="50">
<emphasis id="B6BFEA89FFE9215BFCBBFD8CFC4CFD58" bold="true" box="[846,965,621,645]" pageId="7" pageNumber="50">Etymology.</emphasis>
This specific name alludes to its occurrence around the Wain River.
</paragraph>
</subSubSection>
<subSubSection id="CCD16510FFE9215BFCBBFD4FFA94FC17" pageId="7" pageNumber="50" type="discussion">
<paragraph id="8474369BFFE9215BFCBBFD4FFA94FC17" blockId="7.[820,1432,198,970]" pageId="7" pageNumber="50">
<emphasis id="B6BFEA89FFE9215BFCBBFD4FFC38FD1B" bold="true" box="[846,945,686,710]" pageId="7" pageNumber="50">Remarks.</emphasis>
The sternal punctate ornamentation of the present species is similar in pattern to that of
<taxonomicName id="43CB4D18FFE9215BFAEAFD2FFB40FCDA" authority="Krantz and Whitaker, 1988" authorityName="Krantz and Whitaker" authorityYear="1988" class="Arachnida" family="Macrochelidae" genus="Macrocheles" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="50" phylum="Arthropoda" rank="species" species="adenostictus">
<emphasis id="B6BFEA89FFE9215BFAEAFD2FFC26FCDA" italics="true" pageId="7" pageNumber="50">Macrocheles adenostictus</emphasis>
<bibRefCitation id="E05A4B6AFFE9215BFC40FD11FB40FCDA" author="Krantz, G. W. &amp; Whitaker, J. O." box="[949,1225,751,775]" pageId="7" pageNumber="50" pagination="225 - 259" refId="ref13652" refString="Krantz, G. W. and Whitaker, J. O. 1988. Mites of the genus Macrocheles (Acari: Macrochelidae) associated with small mammals in North America. Acarologia 29: 225 - 259." type="journal article" year="1988">Krantz and Whitaker, 1988</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="E05A4B6AFFE9215BFB23FD0EFC06FCF5" author="Krantz, G. W. &amp; Whitaker, J. O." pageId="7" pageNumber="50" pagination="225 - 259" refId="ref13652" refString="Krantz, G. W. and Whitaker, J. O. 1988. Mites of the genus Macrocheles (Acari: Macrochelidae) associated with small mammals in North America. Acarologia 29: 225 - 259." type="journal article" year="1988">Krantz and Whitaker 1988</bibRefCitation>
) recorded from the
<collectingCountry id="FCDC760BFFE9215BFB93FCF1FB10FCF5" box="[1126,1177,784,808]" name="United States of America" pageId="7" pageNumber="50">USA</collectingCountry>
. However,
<taxonomicName id="43CB4D18FFE9215BFAF9FCF1FAF6FCFA" authorityName="Dwibadra and Takaku" authorityYear="2014" box="[1292,1407,784,807]" class="Arachnida" family="Macrochelidae" genus="Macrocheles" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="50" phylum="Arthropoda" rank="species" species="riparius">
<emphasis id="B6BFEA89FFE9215BFAF9FCF1FAABFCFA" box="[1292,1314,784,807]" italics="true" pageId="7" pageNumber="50">M</emphasis>
.
<emphasis id="B6BFEA89FFE9215BFAC5FCF1FAF6FCFA" box="[1328,1407,784,807]" italics="true" pageId="7" pageNumber="50">riparius</emphasis>
</taxonomicName>
is distinguishable by the following characters (corresponding conditions of
<taxonomicName id="43CB4D18FFE9215BFC31FCB0FBEBFCB4" authorityName="Krantz and Whitaker" authorityYear="1988" box="[964,1122,849,873]" class="Arachnida" family="Macrochelidae" genus="Macrocheles" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="50" phylum="Arthropoda" rank="species" species="adenostictus">
<emphasis id="B6BFEA89FFE9215BFC31FCB0FC53FCB5" box="[964,986,849,872]" italics="true" pageId="7" pageNumber="50">M</emphasis>
.
<emphasis id="B6BFEA89FFE9215BFC1DFCB0FBEBFCB4" box="[1000,1122,849,873]" italics="true" pageId="7" pageNumber="50">adenostictus</emphasis>
</taxonomicName>
in parentheses): 1) most dorsal setae, including setae of z-Z and s-S series, simple (all z-Z and s-S series setae strongly pilose distally); and 2) ophisthogastric setae simple (weakly pilose distally).
</paragraph>
</subSubSection>
</treatment>
</document>