treatments-xml/data/0C/07/87/0C0787D1FF90837EFE8DFAC9FA75E860.xml
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<document id="3A7998CE5C490AD677A8EDFFED3FED3C" ID-DOI="10.5252/zoosystema2020v42a10" ID-GBIF-Dataset="60ca36d6-4c1f-4050-88e2-38e8b28f3f5c" ID-ISSN="1638-9387" ID-Zenodo-Dep="3743146" ID-ZooBank="urn:lsid:zoobank.org:pub:21B9187A-D055-4946-8922-53FD10A159CE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1586267528490" checkinUser="felipe" docAuthor="Curler, Gregory" docDate="2020" docId="0C0787D1FF90837EFE8DFAC9FA75E860" docLanguage="en" docName="zoosystema.42.10.139-149.pdf.imf" docOrigin="Zoosystema 42 (10)" docStyle="DocumentStyle:5CC576F744449C1860B4E72ABD884631.7:Zoosystema.2020-.journal_article" docStyleId="5CC576F744449C1860B4E72ABD884631" docStyleName="Zoosystema.2020-.journal_article" docStyleVersion="7" docTitle="Polletomyia Curler 2020, n. gen." docType="treatment" docUuid="3869FC8A-73CD-4F9D-BBA1-F00A5BA77F4C" docUuidSource="ZooBank" docVersion="5" lastPageNumber="141" masterDocId="F03EFFA9FF94837AFFB6FFDFFF66EE0F" masterDocTitle="Descriptions of two new genera of Maruinini (Diptera, Psychodidae, Psychodinae) from the Mitaraka range of French Guiana" masterLastPageNumber="149" masterPageNumber="139" pageNumber="141" updateTime="1698736126426" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="915870D91E9316CEB331792EE71F3D1D">Descriptions of two new genera of Maruinini (Diptera, Psychodidae, Psychodinae) from the Mitaraka range of French Guiana</mods:title>
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<mods:namePart id="E66F083D9A08EE0C8DDB8C745432C456">Curler, Gregory</mods:namePart>
<mods:affiliation id="17922CA6AAFD9276C462A4CEE85F3F6D">Mississippi Entomological Museum, Mississippi State University, Old Highway 12, P. O. Drawer 9775, Mississippi State, MS 39762 - 9775 (USA) Collaborator, Smithsonian Institution, National Museum of Natural History; 10 &amp; Constitution NW, Washington, DC 20560 - 0169 (USA) gcurler @ gmail. com</mods:affiliation>
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<mods:title id="1D7D5C7673D4DB4477ECE9B150F2C070">Zoosystema</mods:title>
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<treatment id="0C0787D1FF90837EFE8DFAC9FA75E860" ID-DOI="http://doi.org/10.5281/zenodo.3809872" ID-GBIF-Taxon="163536911" ID-Zenodo-Dep="3809872" LSID="urn:lsid:zoobank.org:act:3869FC8A-73CD-4F9D-BBA1-F00A5BA77F4C" httpUri="http://treatment.plazi.org/id/0C0787D1FF90837EFE8DFAC9FA75E860" lastPageNumber="141" pageId="4" pageNumber="141">
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<paragraph id="841136C7FF90837EFE8DFAC9FD37EB3F" blockId="4.[315,593,1301,1328]" box="[315,593,1301,1328]" pageId="4" pageNumber="141">
<heading id="DF5981ABFF90837EFE8DFAC9FD37EB3F" box="[315,593,1301,1328]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="141" reason="2">
Genus
<taxonomicName id="43AE4D44FF90837EFE3EFACAFE99EB20" ID-CoL="8H3MK" authority="Curler, 2020" authorityName="Curler" authorityYear="2020" box="[392,511,1301,1327]" class="Insecta" family="Psychodidae" genus="Polletomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B6DAEAD5FF90837EFE3EFACAFE99EB20" box="[392,511,1301,1327]" italics="true" pageId="4" pageNumber="141">Polletomyia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADE957AEFF90837EFDB0FAC9FD37EB3F" box="[518,593,1302,1328]" pageId="4" pageNumber="141" rank="genus">n. gen.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="CCB4654CFF90837EFF2DFA85FD96EB63" box="[155,752,1370,1388]" pageId="4" pageNumber="141" type="description">
<paragraph id="841136C7FF90837EFF2DFA85FD96EB63" blockId="4.[155,752,1370,1388]" box="[155,752,1370,1388]" pageId="4" pageNumber="141">
<uri id="F03F3AC5FF90837EFF2DFA85FD96EB63" box="[155,752,1370,1388]" pageId="4" pageNumber="141">
urn:lsid:zoobank.org:act:
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<subSubSection id="CCB4654CFF90837EFF32FA53FD1AEBAB" box="[132,636,1420,1445]" pageId="4" pageNumber="141" type="type_taxon">
<paragraph id="841136C7FF90837EFF32FA53FD1AEBAB" blockId="4.[132,636,1420,1445]" box="[132,636,1420,1445]" pageId="4" pageNumber="141">
<typeStatus id="5B158865FF90837EFF32FA53FFD3EBAE" box="[132,181,1420,1444]" pageId="4" pageNumber="141">TYPE</typeStatus>
SPECIES. —
<taxonomicName id="43AE4D44FF90837EFE98FA53FE96EBAB" ID-CoL="8H2QC" authority="Curler, 2020" authorityName="Curler" authorityYear="2020" box="[302,496,1420,1444]" class="Insecta" family="Psychodidae" genus="Polletomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="species" species="subulata" status="n. gen., n. sp.">
<emphasis id="B6DAEAD5FF90837EFE98FA53FE96EBAB" box="[302,496,1420,1444]" italics="true" pageId="4" pageNumber="141">Polletomyia subulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADE957AEFF90837EFE41FA52FD1AEBAB" box="[503,636,1421,1445]" pageId="4" pageNumber="141" rank="species">n. gen., n. sp.</taxonomicNameLabel>
</paragraph>
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<subSubSection id="CCB4654CFF90837EFF32FA64FF8AEBE1" pageId="4" pageNumber="141" type="description">
<paragraph id="841136C7FF90837EFF32FA64FF8AEBE1" blockId="4.[132,776,1467,1518]" pageId="4" pageNumber="141">
ILLUSTRATIONS. — Head:
<figureCitation id="1C952A42FF90837EFE31FA63FEA9EBDB" box="[391,463,1468,1492]" captionStart="FIG" captionStartId="5.[132,143,1940,1958]" captionTargetBox="[160,1421,243,1885]" captionTargetId="figure@5.[132,1455,214,1920]" captionTargetPageId="5" captionText="FIG. 1. — Polletomyia subulata n. gen., n. sp.: A, male head, frontal view; B, wing; C, male antennal flagellomeres 3-4, dorsal view; D, male antennal flagellomeres 12-14, dorsal view; E, male terminalia,epandrium removed, dorsal view; F, male terminalia,epandrium removed, aedeagal structure expanded,dorsal view; G, male terminalia, epandrium removed, aedeagal structure partly expanded, ventral view. Abbreviations: see Material and methods. Scale bars: A, 0.25 mm; B, 0.5 mm; C-G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3743148" httpUri="https://zenodo.org/record/3743148/files/figure.png" pageId="4" pageNumber="141">Fig. 1A</figureCitation>
; Wing:
<figureCitation id="1C952A42FF90837EFD97FA63FD0FEBDB" box="[545,617,1468,1492]" captionStart="FIG" captionStartId="5.[132,143,1940,1958]" captionTargetBox="[160,1421,243,1885]" captionTargetId="figure@5.[132,1455,214,1920]" captionTargetPageId="5" captionText="FIG. 1. — Polletomyia subulata n. gen., n. sp.: A, male head, frontal view; B, wing; C, male antennal flagellomeres 3-4, dorsal view; D, male antennal flagellomeres 12-14, dorsal view; E, male terminalia,epandrium removed, dorsal view; F, male terminalia,epandrium removed, aedeagal structure expanded,dorsal view; G, male terminalia, epandrium removed, aedeagal structure partly expanded, ventral view. Abbreviations: see Material and methods. Scale bars: A, 0.25 mm; B, 0.5 mm; C-G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3743148" httpUri="https://zenodo.org/record/3743148/files/figure.png" pageId="4" pageNumber="141">Fig. 1B</figureCitation>
; Male genitalia:
<figureCitation id="1C952A42FF90837EFF32FA09FFDEEBE1" box="[132,184,1494,1518]" captionStart="FIG" captionStartId="5.[132,143,1940,1958]" captionTargetBox="[160,1421,243,1885]" captionTargetId="figure@5.[132,1455,214,1920]" captionTargetPageId="5" captionText="FIG. 1. — Polletomyia subulata n. gen., n. sp.: A, male head, frontal view; B, wing; C, male antennal flagellomeres 3-4, dorsal view; D, male antennal flagellomeres 12-14, dorsal view; E, male terminalia,epandrium removed, dorsal view; F, male terminalia,epandrium removed, aedeagal structure expanded,dorsal view; G, male terminalia, epandrium removed, aedeagal structure partly expanded, ventral view. Abbreviations: see Material and methods. Scale bars: A, 0.25 mm; B, 0.5 mm; C-G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3743148" httpUri="https://zenodo.org/record/3743148/files/figure.png" pageId="4" pageNumber="141">Fig. 1</figureCitation>
E-G.
</paragraph>
</subSubSection>
<subSubSection id="CCB4654CFF90837EFF32F9DAFD9BE85C" pageId="4" pageNumber="141" type="etymology">
<paragraph id="841136C7FF90837EFF32F9DAFD9BE85C" blockId="4.[132,775,1541,1619]" pageId="4" pageNumber="141">
ETYMOLOGY. — This genus is named in honor of Dr Marc Pollet, who collected the
<typeStatus id="5B158865FF90837EFE98F9FEFE31E836" box="[302,343,1569,1593]" pageId="4" pageNumber="141">type</typeStatus>
specimens, forwarded them to the author and provided supporting information during the course of this study.
</paragraph>
</subSubSection>
<subSubSection id="CCB4654CFF90837EFF32F9B5FDE6E959" pageId="4" pageNumber="141" type="diagnosis">
<paragraph id="841136C7FF90837EFF32F9B5FDE6E959" blockId="4.[132,776,1642,1878]" pageId="4" pageNumber="141">
DIAGNOSIS. — The following combination of characters is diagnostic for the genus: male head: antennal flagellomeres fusiform, with paired, unbranched ascoids on f4 f7 (
<figureCitation id="1C952A42FF90837EFE4BF940FD2FE8B8" box="[509,585,1695,1719]" captionStart="FIG" captionStartId="5.[132,143,1940,1958]" captionTargetBox="[160,1421,243,1885]" captionTargetId="figure@5.[132,1455,214,1920]" captionTargetPageId="5" captionText="FIG. 1. — Polletomyia subulata n. gen., n. sp.: A, male head, frontal view; B, wing; C, male antennal flagellomeres 3-4, dorsal view; D, male antennal flagellomeres 12-14, dorsal view; E, male terminalia,epandrium removed, dorsal view; F, male terminalia,epandrium removed, aedeagal structure expanded,dorsal view; G, male terminalia, epandrium removed, aedeagal structure partly expanded, ventral view. Abbreviations: see Material and methods. Scale bars: A, 0.25 mm; B, 0.5 mm; C-G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3743148" httpUri="https://zenodo.org/record/3743148/files/figure.png" pageId="4" pageNumber="141">Fig. 1C</figureCitation>
), single ascoids on f8 f11; male terminalia: bilaterally symmetrical; aedeagus comprised of four movable, blade-like sclerites surrounded by membranous sheath; outer aedeagal sclerites sinuous, spreading laterally when extended posteriorly (
<figureCitation id="1C952A42FF90837EFEEAF8D6FEC4E92E" box="[348,418,1801,1825]" captionStart="FIG" captionStartId="5.[132,143,1940,1958]" captionTargetBox="[160,1421,243,1885]" captionTargetId="figure@5.[132,1455,214,1920]" captionTargetPageId="5" captionText="FIG. 1. — Polletomyia subulata n. gen., n. sp.: A, male head, frontal view; B, wing; C, male antennal flagellomeres 3-4, dorsal view; D, male antennal flagellomeres 12-14, dorsal view; E, male terminalia,epandrium removed, dorsal view; F, male terminalia,epandrium removed, aedeagal structure expanded,dorsal view; G, male terminalia, epandrium removed, aedeagal structure partly expanded, ventral view. Abbreviations: see Material and methods. Scale bars: A, 0.25 mm; B, 0.5 mm; C-G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3743148" httpUri="https://zenodo.org/record/3743148/files/figure.png" pageId="4" pageNumber="141">Fig. 1F</figureCitation>
); gonostyli with stout, conical setae inserted apically (
<figureCitation id="1C952A42FF90837EFE85F8FBFE0EE933" box="[307,360,1828,1852]" captionStart="FIG" captionStartId="5.[132,143,1940,1958]" captionTargetBox="[160,1421,243,1885]" captionTargetId="figure@5.[132,1455,214,1920]" captionTargetPageId="5" captionText="FIG. 1. — Polletomyia subulata n. gen., n. sp.: A, male head, frontal view; B, wing; C, male antennal flagellomeres 3-4, dorsal view; D, male antennal flagellomeres 12-14, dorsal view; E, male terminalia,epandrium removed, dorsal view; F, male terminalia,epandrium removed, aedeagal structure expanded,dorsal view; G, male terminalia, epandrium removed, aedeagal structure partly expanded, ventral view. Abbreviations: see Material and methods. Scale bars: A, 0.25 mm; B, 0.5 mm; C-G, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3743148" httpUri="https://zenodo.org/record/3743148/files/figure.png" pageId="4" pageNumber="141">Fig. 1</figureCitation>
E-G); epandrial claspers each with single, setiform tenaculum inserted dorsoapically (
<figureCitation id="1C952A42FF90837EFD9DF8E1FD14E959" box="[555,626,1854,1878]" captionStart="FIG" captionStartId="6.[132,143,1987,2005]" captionTargetBox="[163,1420,231,1895]" captionTargetId="figure@6.[132,1455,216,1902]" captionTargetPageId="6" captionText="FIG. 2. — Polletomyia subulata n. gen., n. sp.: A, male terminalia, lateral view; B, male epandrium, proctiger and epandrial claspers, dorsal view; C, female terminalia, external structure, ventral view; D, female terminalia, internal structure, ventral view. Abbreviations: see Material and methods. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3743150" httpUri="https://zenodo.org/record/3743150/files/figure.png" pageId="4" pageNumber="141">Fig. 2B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CCB4654CFF90837EFF32F8B2FE91E9AF" pageId="4" pageNumber="141" type="distribution">
<paragraph id="841136C7FF90837EFF32F8B2FE91E9AF" blockId="4.[132,775,1901,1953]" pageId="4" pageNumber="141">
DISTRIBUTION. —
<taxonomicName id="43AE4D44FF90837EFE8CF8B1FEC0E989" authority="Curler, 2020" authorityName="Curler" authorityYear="2020" box="[314,422,1902,1926]" class="Insecta" family="Psychodidae" genus="Polletomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B6DAEAD5FF90837EFE8CF8B1FEC0E989" box="[314,422,1902,1926]" italics="true" pageId="4" pageNumber="141">Polletomyia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADE957AEFF90837EFE1CF8B1FE8AE989" box="[426,492,1902,1926]" pageId="4" pageNumber="141" rank="genus">n. gen.</taxonomicNameLabel>
is currently known only from the Mitaraka range of
<collectingCountry id="FCB97657FF90837EFEE8F857FE95E9AF" box="[350,499,1928,1952]" name="French Guiana" pageId="4" pageNumber="141">French Guiana</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="CCB4654CFF90837EFF32F868FD9AE9E5" pageId="4" pageNumber="141" type="biology_ecology">
<paragraph id="841136C7FF90837EFF32F868FD9AE9E5" blockId="4.[132,775,1975,2026]" pageId="4" pageNumber="141">HABITAT. — Little information regarding specific habitat is available; however, all specimens were collected in lowland rainforest.</paragraph>
</subSubSection>
<subSubSection id="CCB4654CFF90837EFC9BFF06FB6FE800" pageId="4" pageNumber="141" type="discussion">
<paragraph id="841136C7FF90837EFC9BFF06FCF7EEFE" blockId="4.[813,1456,217,1551]" box="[813,913,217,243]" pageId="4" pageNumber="141">REMARKS</paragraph>
<paragraph id="841136C7FF90837EFC9BFF26FBFAEC7D" blockId="4.[813,1456,217,1551]" pageId="4" pageNumber="141">
The relatively small size of
<taxonomicName id="43AE4D44FF90837EFBF0FF26FBDBEF1C" authority="Curler, 2020" authorityName="Curler" authorityYear="2020" box="[1094,1213,249,275]" class="Insecta" family="Psychodidae" genus="Polletomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B6DAEAD5FF90837EFBF0FF26FBDBEF1C" box="[1094,1213,249,275]" italics="true" pageId="4" pageNumber="141">Polletomyia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADE957AEFF90837EFB75FF25FA68EF1B" box="[1219,1294,250,276]" pageId="4" pageNumber="141" rank="genus">n. gen.</taxonomicNameLabel>
, as well as their lanceolate wings and dark coloration make it possible to differentiate them in a sample comprised of many psychodid genera and species. These superficial characters also make it possible to associate males and females, as the latter are otherwise lacking in characters that readily differentiate them from other psychodine females. Indeed, if further species of
<taxonomicName id="43AE4D44FF90837EFC9BFE07FCC2EFFD" authority="Curler, 2020" authorityName="Curler" authorityYear="2020" box="[813,932,472,498]" class="Insecta" family="Psychodidae" genus="Polletomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B6DAEAD5FF90837EFC9BFE07FCC2EFFD" box="[813,932,472,498]" italics="true" pageId="4" pageNumber="141">Polletomyia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADE957AEFF90837EFC1CFE06FC95EFFC" box="[938,1011,473,499]" pageId="4" pageNumber="141" rank="genus">n. gen.</taxonomicNameLabel>
are discovered, it may become difficult or impossible to differentiate females within the genus. Despite the relative ease of differentiating them in a bulk sample, accurate diagnosis of this genus is dependent on examination of slide-mounted male terminalia.
</paragraph>
<paragraph id="841136C7FF90837EFCF5FDA7FCB2EA3F" blockId="4.[813,1456,217,1551]" pageId="4" pageNumber="141">
Currently, the following character states are sufficient for distinguishing this genus from all other
<taxonomicName id="43AE4D44FF90837EFB41FD47FA08ECBD" authorityName="Enderlein" authorityYear="1937" box="[1271,1390,664,690]" class="Insecta" family="Psychodidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="tribe" tribe="Maruinini">Maruinini</taxonomicName>
: male terminalia bilaterally symmetrical; gonostyli each with a conical, spiniform seta inserted apically; tenacula setiform. Nonetheless, additional characters are included above in the diagnosis because they are also unique to the genus, thereby making the diagnosis more robust. Some other genera of
<taxonomicName id="43AE4D44FF90837EFC9BFC88FCF9ED7E" authorityName="Enderlein" authorityYear="1937" box="[813,927,855,881]" class="Insecta" family="Psychodidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="tribe" tribe="Maruinini">Maruinini</taxonomicName>
also have bilaterally symmetrical male terminalia in some or all constituent species (e.g.
<taxonomicName id="43AE4D44FF90837EFB7BFCA8FA76ED9E" authorityName="Enderlein" authorityYear="1937" box="[1229,1296,887,913]" class="Insecta" family="Psychodidae" genus="Alepia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="B6DAEAD5FF90837EFB7BFCA8FA76ED9E" box="[1229,1296,887,913]" italics="true" pageId="4" pageNumber="141">Alepia</emphasis>
</taxonomicName>
,
<taxonomicName id="43AE4D44FF90837EFAABFCA8FC2CEDBF" authorityName="Wagner" authorityYear="1981" class="Insecta" family="Psychodidae" genus="Caenobrunettia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="B6DAEAD5FF90837EFAABFCA8FC2CEDBF" italics="true" pageId="4" pageNumber="141">Caenobrunettia</emphasis>
</taxonomicName>
,
<taxonomicName id="43AE4D44FF90837EFCECFC48FB3FEDBD" authority="Muller, 1895" authorityName="Muller" authorityYear="1895" box="[858,1113,919,946]" class="Insecta" family="Psychodidae" genus="Maruina" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="B6DAEAD5FF90837EFCECFC48FCDDEDBF" box="[858,955,919,944]" italics="true" pageId="4" pageNumber="141">Maruina</emphasis>
Müller, 1895
</taxonomicName>
, and
<taxonomicName id="43AE4D44FF90837EFB2EFC48FA70EDBF" authority="Curler, 2020" authorityName="Curler" authorityYear="2020" box="[1176,1302,919,944]" class="Insecta" family="Psychodidae" genus="Myiomystax" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B6DAEAD5FF90837EFB2EFC48FA70EDBF" box="[1176,1302,919,944]" italics="true" pageId="4" pageNumber="141">Myiomystax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADE957AEFF90837EFA96FC48FA17EDBE" box="[1312,1393,919,945]" pageId="4" pageNumber="141" rank="genus">n. gen.</taxonomicNameLabel>
). Yet, each of these groups has a unique build to their aedeagus that is distinct from
<taxonomicName id="43AE4D44FF90837EFBA7FC09FBEDEDFF" authority="Curler, 2020" authorityName="Curler" authorityYear="2020" box="[1041,1163,982,1008]" class="Insecta" family="Psychodidae" genus="Polletomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B6DAEAD5FF90837EFBA7FC09FBEDEDFF" box="[1041,1163,982,1008]" italics="true" pageId="4" pageNumber="141">Polletomyia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADE957AEFF90837EFB25FC08FB86EDFE" box="[1171,1248,983,1009]" pageId="4" pageNumber="141" rank="genus">n. gen.</taxonomicNameLabel>
and none of them have conspicuous, spiniform setae inserted at the apices of their gonostyli.
</paragraph>
<paragraph id="841136C7FF90837EFCF5FBE9FB6FE800" blockId="4.[813,1456,217,1551]" pageId="4" pageNumber="141">
Aside from the male terminalia, some character states present in
<taxonomicName id="43AE4D44FF90837EFCCEFB89FC89EA7F" authority="Curler, 2020" authorityName="Curler" authorityYear="2020" box="[888,1007,1110,1136]" class="Insecta" family="Psychodidae" genus="Polletomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B6DAEAD5FF90837EFCCEFB89FC89EA7F" box="[888,1007,1110,1136]" italics="true" pageId="4" pageNumber="141">Polletomyia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADE957AEFF90837EFC42FB88FB5BEA7E" box="[1012,1085,1111,1137]" pageId="4" pageNumber="141" rank="genus">n. gen.</taxonomicNameLabel>
are found as well in other genera of
<taxonomicName id="43AE4D44FF90837EFC9BFBA9FCFBEA9F" authorityName="Enderlein" authorityYear="1937" box="[813,925,1142,1168]" class="Insecta" family="Psychodidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="tribe" tribe="Maruinini">Maruinini</taxonomicName>
, such as a contiguous eye bridge comprised of three rows of ommatidia. These states are also present in
<taxonomicName id="43AE4D44FF90837EFAE6FB48FAC9EABF" authorityName="Muller" authorityYear="1895" box="[1360,1455,1175,1200]" class="Insecta" family="Psychodidae" genus="Maruina" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="B6DAEAD5FF90837EFAE6FB48FAC9EABF" box="[1360,1455,1175,1200]" italics="true" pageId="4" pageNumber="141">Maruina</emphasis>
</taxonomicName>
(so far only in subgenus
<taxonomicName id="43AE4D44FF90837EFB9DFB69FA7FEADF" authority="Hogue, 1973" authorityName="Hogue" authorityYear="1973" box="[1067,1305,1206,1232]" class="Insecta" family="Psychodidae" genus="Aculcina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="B6DAEAD5FF90837EFB9DFB69FBE3EADF" box="[1067,1157,1206,1232]" italics="true" pageId="4" pageNumber="141">Aculcina</emphasis>
Hogue, 1973
</taxonomicName>
) and
<taxonomicName id="43AE4D44FF90837EFAE5FB69FCAEEAFF" authority="Satchell, 1955" authorityName="Satchell" authorityYear="1955" class="Insecta" family="Psychodidae" genus="Arisemus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="B6DAEAD5FF90837EFAE5FB69FAC9EADF" box="[1363,1455,1206,1232]" italics="true" pageId="4" pageNumber="141">Arisemus</emphasis>
<bibRefCitation id="E03F4B36FF90837EFC9BFB09FCAEEAFF" author="SATCHELL G. H." box="[813,968,1237,1264]" pageId="4" pageNumber="141" pagination="85 - 93" refId="ref7264" refString="SATCHELL G. H. 1955. - Two new subgenera of Psychodidae (Diptera) from Jamaica, with descriptions of five new species. Annals and Magazine of Natural History 12: 85 - 93. https: // doi. org / 10.1080 / 00222935508651831" type="journal article" year="1955">Satchell, 1955</bibRefCitation>
</taxonomicName>
and have been indicated by some authors as diagnostic for those genera (e.g.
<bibRefCitation id="E03F4B36FF90837EFB3DFB2AFAF4EB1F" author="CAMICO J. &amp; CORDEIRO D. &amp; CHAGAS C." box="[1163,1426,1269,1296]" pageId="4" pageNumber="141" pagination="296 - 316" refId="ref6793" refString="CAMICO J., CORDEIRO D. &amp; CHAGAS C. 2019. - Maruina Muller, 1895 (Diptera: Psychodidae): description of four new species from Brazil, with revised diagnoses for the genus and subgenera. Zootaxa 4652 (2): 296 - 316. https: // doi. org / 10.11646 / zootaxa. 4652.2.4" type="journal article" year="2019">
Camico
<emphasis id="B6DAEAD5FF90837EFB51FB29FA7DEB00" box="[1255,1307,1269,1295]" italics="true" pageId="4" pageNumber="141">et al.</emphasis>
2019:
<quantity id="43569B22FF90837EFADEFB29FAF4EB1F" box="[1384,1426,1270,1296]" metricMagnitude="0" metricUnit="m" metricValue="7.5691999999999995" pageId="4" pageNumber="141" unit="in" value="298.0">298</quantity>
</bibRefCitation>
in their diagnosis of
<emphasis id="B6DAEAD5FF90837EFC59FAC9FB69EB20" box="[1007,1039,1302,1327]" italics="true" pageId="4" pageNumber="141">M.</emphasis>
(
<taxonomicName id="43AE4D44FF90837EFBA9FACAFB1FEB20" authorityName="Hogue" authorityYear="1973" box="[1055,1145,1301,1327]" class="Insecta" family="Psychodidae" genus="Aculcina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="B6DAEAD5FF90837EFBA9FACAFB1FEB20" box="[1055,1145,1301,1327]" italics="true" pageId="4" pageNumber="141">Aculcina</emphasis>
</taxonomicName>
)). Despite this, the presence of these character states in multiple genera indicates that they are not diagnostic in and of themselves. Rather, it is necessary to determine whether there are other nuances that distinguish the eye bridge of each genus from others that have the same general appearance. In the case of
<taxonomicName id="43AE4D44FF90837EFB1BFA6AFA40EBC0" authority="Curler, 2020" authorityName="Curler" authorityYear="2020" box="[1197,1318,1461,1487]" class="Insecta" family="Psychodidae" genus="Polletomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B6DAEAD5FF90837EFB1BFA6AFA40EBC0" box="[1197,1318,1461,1487]" italics="true" pageId="4" pageNumber="141">Polletomyia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADE957AEFF90837EFA86FA6AFAE6EBC0" box="[1328,1408,1461,1487]" pageId="4" pageNumber="141" rank="genus">n. gen.</taxonomicNameLabel>
, the eye bridge is not particularly distinct, so it is omitted as a diagnostic character.
</paragraph>
</subSubSection>
<subSubSection id="CCB4654CFF90837EFC9BF9EBFA75E860" pageId="4" pageNumber="141" type="description">
<paragraph id="841136C7FF90837EFC9BF9EBFCDBE843" blockId="4.[813,1299,1588,1647]" box="[813,957,1588,1614]" pageId="4" pageNumber="141">DESCRIPTION</paragraph>
<paragraph id="841136C7FF90837EFC9BF98BFA75E860" blockId="4.[813,1299,1588,1647]" box="[813,1299,1620,1647]" pageId="4" pageNumber="141">
See below,
<taxonomicName id="43AE4D44FF90837EFC15F98BFB1FE861" authority="Curler, 2020" authorityName="Curler" authorityYear="2020" box="[931,1145,1620,1646]" class="Insecta" family="Psychodidae" genus="Polletomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="141" phylum="Arthropoda" rank="species" species="subulata" status="n. gen., n. sp.">
<emphasis id="B6DAEAD5FF90837EFC15F98BFB1FE861" box="[931,1145,1620,1646]" italics="true" pageId="4" pageNumber="141">Polletomyia subulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ADE957AEFF90837EFB36F98AFA75E860" box="[1152,1299,1621,1647]" pageId="4" pageNumber="141" rank="species">n. gen., n. sp.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>