plazi/treatments-xml
2
0
Fork 0
Code Issues Pull requests Releases Activity
treatments-xml/data/F8/95/E5/F895E5960025550FA394CC1243A1DE43.xml

481 lines
58 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document ID-CLB-Dataset="129873" ID-DOI="http://dx.doi.org/10.3897/zookeys.1119.85400" ID-GBIF-Dataset="418bd521-e4b9-4ee6-981f-a6f471eab287" ID-PMC="PMC9848625" ID-Pensoft-Pub="1313-2970-1119-1" ID-Pensoft-UUID="F158A7C5261D52B69288A62F3C777CAF" ID-PubMed="36762355" ID-ZooBank="A3BE91C6B79344E1A886A803BF104D8B" ModsDocID="1313-2970-1119-1" checkinTime="1661532673136" checkinUser="pensoft" docAuthor="Jirapatrasilp, Parin, Sutcharit, Chirasak &amp; Panha, Somsak" docDate="2022" docId="F895E5960025550FA394CC1243A1DE43" docLanguage="en" docName="ZooKeys 1119: 1-115" docOrigin="ZooKeys 1119" docPubDate="2022-08-25" docSource="http://dx.doi.org/10.3897/zookeys.1119.85400" docTitle="Pupina aureola Stoliczka 1872" docType="treatment" docVersion="6" id="F158A7C5261D52B69288A62F3C777CAF" lastPageNumber="1" masterDocId="F158A7C5261D52B69288A62F3C777CAF" masterDocTitle="Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia" masterLastPageNumber="115" masterPageNumber="1" pageNumber="1" updateTime="1732923707409" updateUser="ExternalLinkService">
<mods:mods id="BDD6CC9920F0670D2D6B38C7A4E61C30" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="BAF9A2D3FDC28F4BC76FCC3FA5648535">
<mods:title id="DBB1A0DA93FA6B7AA47831C33806049E">Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia</mods:title>
</mods:titleInfo>
<mods:name id="FC6C08F4A5566B1648C7E992D1A3C950" type="personal">
<mods:role id="F7C0FA71574EA3ECD4CE2AB7AB584E67">
<mods:roleTerm id="8C90D32CD44019F8BBFFC357A983C6B5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="540C0EA93807807BB6B6AD9BAC19479B">Jirapatrasilp, Parin</mods:namePart>
<mods:nameIdentifier id="74DFE74ABF2FA18CE72C23C31B4E23A2" type="ORCID">https://orcid.org/0000-0002-5591-6724</mods:nameIdentifier>
<mods:affiliation id="60134C60017F31CD7C98F05428CCC192">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
</mods:name>
<mods:name id="2B6E961FB99F35FE9AF4B1814AF460F3" type="personal">
<mods:role id="0C5BBEA93CFD488FF0FDC922ECE0A922">
<mods:roleTerm id="0DAACDC9504EFD82316BBCF32B21947C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="59A7930876355B289782DCEDBA734548">Sutcharit, Chirasak</mods:namePart>
<mods:affiliation id="595AC02C2F06DE8EBA601FD7C2E39651">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
<mods:nameIdentifier id="78F6F45EE19CCAB735C348858F1FC563" type="email">jirasak4@yahoo.com</mods:nameIdentifier>
</mods:name>
<mods:name id="98C84E824EBE91DA1A1F904E6CE86A25" type="personal">
<mods:role id="1124AB7081930058A93945B600E307EF">
<mods:roleTerm id="B727CB21290C711F9FAD084D9D49EDA6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7F4B3613275FBBA22F0A8D4BE01DC2BD">Panha, Somsak</mods:namePart>
<mods:affiliation id="3226E254EDE8BBA2D76E2269FAF614D7">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand &amp; Academy of Science, The Royal Society of Thailand, Bangkok 10300, Thailand</mods:affiliation>
<mods:nameIdentifier id="26CF4606ED034B7482CC76EF8B9375B7" type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="1DBEBB5FF8CE497B9F478D10D9A53D77">text</mods:typeOfResource>
<mods:relatedItem id="2628D85D04DACA00794517F6A39CD129" type="host">
<mods:titleInfo id="EF2CA2AE0228E6A7C132F5226164626A">
<mods:title id="055356A4FC18B51DFC10E20609517885">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="4428CE19CA11CED6994308EDF0B53822">
<mods:date id="3305729ACD127BD98B4582A756D84AB3">2022</mods:date>
<mods:detail id="1B470C57396D03068C27748B86467B0F" type="pubDate">
<mods:number id="6588AD915E60DA52AA0D04B63DC198E6">2022-08-25</mods:number>
</mods:detail>
<mods:detail id="32997095365371DB530EF8AD3F34F913" type="volume">
<mods:number id="3450BC784BD61CD7DB2674BD787BF4EA">1119</mods:number>
</mods:detail>
<mods:extent id="81734B29A8CD2BBF6214FCC036EA1EFB" unit="page">
<mods:start id="7F4239811645A3401268892B3BD58005">1</mods:start>
<mods:end id="027600726CEA1BC3EE133D7C558F12F8">115</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="78B45473171E3D9DC0310F79559BE34F">
<mods:url id="42C38ED62F24FE4656F278D4DA0CA51E">http://dx.doi.org/10.3897/zookeys.1119.85400</mods:url>
</mods:location>
<mods:classification id="6717729AD4E13B16DBCD1F2565455A98">journal article</mods:classification>
<mods:identifier id="6B66D4E7701E8E62FB65384AACA29EED" type="DOI">http://dx.doi.org/10.3897/zookeys.1119.85400</mods:identifier>
<mods:identifier id="7364EF5A9560441A91CE53A0DBB47ACA" type="Pensoft-Pub">1313-2970-1119-1</mods:identifier>
<mods:identifier id="B9F1E8AB2CAD2317BB1B1AABC7163AD2" type="ZooBank">A3BE91C6B79344E1A886A803BF104D8B</mods:identifier>
<mods:identifier id="192C6E714ADB58D37B598E16AF0D9CD6" type="Pensoft-UUID">F158A7C5261D52B69288A62F3C777CAF</mods:identifier>
</mods:mods>
<treatment id="F895E5960025550FA394CC1243A1DE43" ID-GBIF-Taxon="199839000" LSID="urn:lsid:plazi:treatment:F895E5960025550FA394CC1243A1DE43" httpUri="http://treatment.plazi.org/id/F895E5960025550FA394CC1243A1DE43" lastPageNumber="1" pageId="0" pageNumber="1" scope_class="Gastropoda" scope_family="Pupinidae" scope_phylum="Mollusca" scope_subClass="Caenogastropoda" scope_subFamily="Pupininae" scope_superFamily="Cyclophoroidea">
<subSubSection id="5203601766402F522A79EE2483D9A3E4" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="B26DC3B319D71DFE47ADD877B3ED264D" pageId="0" pageNumber="1">
<taxonomicName id="34EC0968D688358E08A22792DB718711" ID-CoL="7SWXF" LSID="F895E596-0025-550F-A394-CC1243A1DE43" authority="Stoliczka, 1872" authorityName="Stoliczka" authorityYear="1872" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina aureola" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="aureola">Pupina aureola Stoliczka, 1872</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="368CA242B40E30312F830FB0EDC3B9F0" pageId="0" pageNumber="1" type="description">
<paragraph id="7567E9DC5B861CFA0A9C1007F80D5462" pageId="0" pageNumber="1">
<figureCitation id="B2369A7E4A792D9A50A46FBCCC399C78" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Umbilical, columellar and parietal views of A Pupinella mansuyi, specimen CUMZ 12148 from Pha Chu, Nan B Pupina artata from the Pupina artata species group, specimen CUMZ 12003 from Ban Ping Khong, Chiang Mai C Pupina godwinausteni sp. nov. from the Pupina arula species group, holotype CUMZ 12090 / 1 D Pupina aureola from the Pupina aureola species group, specimen CUMZ 12130 from Sra Morakot, Krabi." figureDoi="10.3897/zookeys.1119.85400.figure10" httpUri="https://binary.pensoft.net/fig/735371" pageId="0" pageNumber="1">Figs 10D</figureCitation>
<figureCitation id="E9D3BA0927A6D703DD3707EB9F38E000" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Opercula of three mainland Southeast Asian Pupina species groups A Pupina bensoni sp. nov. from the Pupina artata species group, specimen CUMZ 12047 B Pupina siamensis from the Pupina arula species group, specimen CUMZ 12067, and C Pupina aureola from the Pupina aureola species group, specimen CUMZ 12116. All not to scale." figureDoi="10.3897/zookeys.1119.85400.figure19" httpUri="https://binary.pensoft.net/fig/735380" pageId="0" pageNumber="1">, 19C</figureCitation>
<figureCitation id="FDCF16DE592952CD58BAC671B9D3F5C9" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Live specimens of A Pupina siamensis, specimen CUMZ 12071 from Tham Khao Loi Temple, Rayong B-D Pupina bilabiata sp. nov. B paratype CUMZ 12073 / 2 from Banpot Pisai Temple, Chumphon and specimens C CUMZ 12082 from Pha Jor Cave, Nong Bua Lam Phu and D CUMZ 12087 from Ban Yai, Surat Thani E, F Pupina godwinausteni sp. nov.: paratypes E CUMZ 12090 / 26 and F CUMZ 12091 from Khao Wong Cave, Uthai Thani G, H Pupina aureola: specimens G CUMZ 12117 from Lod Cave, Nakhon Sri Thammarat and H CUMZ 12121 from Tham Thong Panara Temple, Nakhon Sri Thammarat, showing its microhabitat in rotten log. All not to scale." figureDoi="10.3897/zookeys.1119.85400.figure32" httpUri="https://binary.pensoft.net/fig/735393" pageId="0" pageNumber="1">, 32G, H</figureCitation>
<figureCitation id="8A39020C366FA7F1EEF98AB73179542D" captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. A Pupina vescoi, specimen SMF 109956 / 1 from Cochin China B, C Pupina exclamationis B syntype MNHN-IM- 2000 - 35840 from Tonkin and C specimen NHMUK 1901.12.23.205 &quot; forma minor &quot; from Than-Moi, Tonkin D Pupina perakensis, lectotype SMF 109969 / 1 from Bukit Pondong, Perak E Pupina excisa, lectotype SMF 110778 / 1 from Kelantan F Pupina aureola, possible syntype NHMUK 1988.12.4.101 from Pinang. Photo: P. Maestrati, MNHN (B)." figureDoi="10.3897/zookeys.1119.85400.figure34" httpUri="https://binary.pensoft.net/fig/735395" pageId="0" pageNumber="1">, 34F</figureCitation>
<figureCitation id="9A35A8C341DF437B4DE1E7B0FB7C5B8F" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Shells of Pupina aureola species group from mainland Southeast Asia A-F Pupina aureola A possible syntype NHMUK 1988.12.4.101 and specimens B CUMZ 12124 C CUMZ 12126 D CUMZ 12130 E CUMZ 12133, and F CUMZ 12121 G, H Pupina stoliczkai sp. nov. G holotype CUMZ 12147 / 1 and H paratype NHMUK 20210336 I, J Pupina dorri dorri I paralectotype MNHN-IM- 2000 - 35836 and J specimen NHMUK ex. A. J. Piele Colln. Acc. No. 2242 K, L Pupina dorri isanensis spp. nov. K holotype CUMZ 12140 / 1 and L specimen CUMZ 12137 M Pupina brachysoma, lectotype MNHN-IM- 2000 - 9652. Photo: P. Maestrati, MNHN (I, M)" figureDoi="10.3897/zookeys.1119.85400.figure36" httpUri="https://binary.pensoft.net/fig/735397" pageId="0" pageNumber="1">, 36A-F</figureCitation>
<figureCitation id="454F518C120EC7CE0E43EA45DA4990DE" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. A-E Pupina aureola: specimens A CUMZ 12121 from Tham Thong Panara Temple, Nakhon Sri Thammarat B CUMZ 12124 from Talot Cave, Nakhon Sri Thammarat C CUMZ 12126 from Khao Huai Haeng Temple, Trang D CUMZ 12130 from Sra Morakot, Krabi, and E CUMZ 12133 from Khantiphol Cave, Satun F, G Pupina paviei: paralectotypes F MNHN-IM- 2000 - 35837 from Chaine de l'Elephant, Kampot, Cambodge and G RBINS 525404 from Kampot et forets de la chaine de l'Elephant, Cambodge et Kamchay. Photo: P. Maestrati, MNHN (F), F. Trus, RBINS (G)." figureDoi="10.3897/zookeys.1119.85400.figure38" httpUri="https://binary.pensoft.net/fig/735399" pageId="0" pageNumber="1">, 38A-E</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="DD0DF38A8EFD81EC3F9A4EE3DCE5CC24" pageId="0" pageNumber="1" type="reference_group">
<paragraph id="269FF20A86B2674A542C217C1EA5A3B5" pageId="0" pageNumber="1">
<taxonomicName id="19811ABDFBB88CF83C2E1B9588460EE3" authorityName="Stoliczka" authorityYear="1872" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina aureola" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="aureola">Pupina aureola</taxonomicName>
Stoliczka, 1872: 267, pl. 10, figs 11, 12. Type locality: Penang [Penang State, Malaysia].
<bibRefCitation id="97928D71EF27CADD76781CC1F70EDEAF" DOI="https://doi.org/10.5962/bhl.title.11957" author="Nevill, G" journalOrPublisher="Indian Museum, Calcutta" pageId="0" pageNumber="1" refId="B104" refString="Nevill, G, 1878. Hand List of Mollusca in the Indian Museum, Calcutta. Part 1. Gastropoda. Pulmonata and Prosobranchia-Neurobranchia. Indian Museum, Calcutta, DOI: https://doi.org/10.5962/bhl.title.11957" title="Hand List of Mollusca in the Indian Museum, Calcutta. Part 1. Gastropoda. Pulmonata and Prosobranchia-Neurobranchia." url="https://doi.org/10.5962/bhl.title.11957" year="1878">Nevill 1878</bibRefCitation>
: 299. de Morgan 1885: 414, Poulo Pinang, mont Tchora,
<normalizedToken id="76B531CC40B7E59344C1C21EA8887300" originalValue="près">pres</normalizedToken>
<normalizedToken id="00C98EB9CE2AAF8B485D8EF426034711" originalValue="d’Ipoh">d'Ipoh</normalizedToken>
(Kinta), [Perak State, Malaysia].
<bibRefCitation id="633AF5E2361C6415490BA1FA0794417C" DOI="https://doi.org/10.1111/j.1096-3642.1891.tb01757.x" author="von Moellendorff, OF" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" pagination="330 - 348" refId="B191" refString="von Moellendorff, OF, 1891. On the land and freshwater shells of Perak. Proceedings of the Zoological Society of London 59 (3): 330 - 348, DOI: https://doi.org/10.1111/j.1096-3642.1891.tb01757.x" title="On the land and freshwater shells of Perak." url="https://doi.org/10.1111/j.1096-3642.1891.tb01757.x" volume="59" year="1891">
von
<normalizedToken id="F478A372D532DF3F82EFE37CA9BBCF93" originalValue="Möllendorff">Moellendorff</normalizedToken>
1891
</bibRefCitation>
: 345.
<bibRefCitation id="6A6BB224DD57753B749B414E094C2C61" author="Sykes, ER" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" pagination="194 - 199" refId="B160" refString="Sykes, ER, 1903. On the land operculate mollusks collected during the &quot;Skeat Expedition&quot; to the Malay peninsula in 1899-1900. Proceedings of the Zoological Society of London 1903: 194 - 199" title="On the land operculate mollusks collected during the &quot; Skeat Expedition &quot; to the Malay peninsula in 1899 - 1900." volume="1903" year="1903">Sykes 1903</bibRefCitation>
: 197, Jalor [Yala Province, Thailand].
<bibRefCitation id="15845C7EAFF115E4B7C2FED20F96BCEE" author="van Benthem Jutting, WSS" journalOrPublisher="Bulletin of the Raffles Museum" pageId="0" pageNumber="1" pagination="50 - 77" refId="B180" refString="van Benthem Jutting, WSS, 1949. On a collection of non-marine Mollusca from Malaya in the Raffles Museum, Singapore, with an appendix on cave shells. Bulletin of the Raffles Museum 19: 50 - 77" title="On a collection of non-marine Mollusca from Malaya in the Raffles Museum, Singapore, with an appendix on cave shells." volume="19" year="1949">van Benthem Jutting 1949</bibRefCitation>
: 57, Gunong Pulai, Johore [Johor State, Malaysia].
<bibRefCitation id="E2DB432ECFD123B1D127A79DC5E9F1E6" author="van Benthem Jutting, WSS" journalOrPublisher="Basteria" pageId="0" pageNumber="1" pagination="10 - 20" refId="B181" refString="van Benthem Jutting, WSS, 1960. Some notes on land and freshwater Mollusca of Malaya. Basteria 24: 10 - 20" title="Some notes on land and freshwater Mollusca of Malaya." volume="24" year="1960">van Benthem Jutting 1960</bibRefCitation>
: 13, limestone hill near kampong Tebing Tinggi, N. of Kangar, Perlis [Malaysia].
<bibRefCitation id="16BA1263E4BC5D1B46120E92FBE32FE9" author="Chan, S-Y" journalOrPublisher="The Papustyla" pageId="0" pageNumber="1" pagination="4 - 5" refId="B17" refString="Chan, S-Y, 1998a. A brief collecting trip to Perak, West Malaysia, part one. The Papustyla 12: 4 - 5" title="A brief collecting trip to Perak, West Malaysia, part one." volume="12" year="1998 a">Chan 1998a</bibRefCitation>
: 4, Ipoh, Perak.
<bibRefCitation id="BD5D0A2D825F959FABA1629127724CD5" author="Maassen, WJM" journalOrPublisher="Proceedings of the United States National Museum" pageId="0" pageNumber="1" refId="B87" refString="Maassen, WJM, 2001. A preliminary checklist of the non-marine molluscs of West-Malaysia, A handlist. De Kreukel (Supplement): 1-155." title="A preliminary checklist of the non-marine molluscs of West-Malaysia, A handlist. De Kreukel (Supplement): 1 - 155." year="2001">Maassen 2001</bibRefCitation>
: 40, 41.
<bibRefCitation id="B482A154F86ABA8EC5B85C2AF6CBCB40" author="Bavay, A" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" refId="B8" refString="BEDO (2017) Land Snails: Checklist of Molluscan Biodiversity in Thailand.BEDO, Bangkok, 300 pp. [in Thai]" year="1904">BEDO 2017</bibRefCitation>
: 88.
<bibRefCitation id="E733E25456F045FD580AF113D706F9AC" author="Sutcharit, C" journalOrPublisher="Zino Publishing, Bangkok" pageId="0" pageNumber="1" refId="B156" refString="Sutcharit, C, Tongkerd, P, Panha, S, 2018. Land snails: The invaluable bio-resources for the Kingdom of Thailand. Zino Publishing, Bangkok" title="Land snails: The invaluable bio-resources for the Kingdom of Thailand." year="2018">Sutcharit et al. 2018</bibRefCitation>
: fig. 5-13f.
</paragraph>
<paragraph id="C3D383DFB0B32B96F6D5F405479BBB5F" pageId="0" pageNumber="1">
<taxonomicName id="399FBAA28C0C41577E2687CAEE24DAF1" authorityName="Stoliczka" authorityYear="1872" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina (Tylotoechus) aureola" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="aureola" subGenus="Tylotoechus">Pupina (Tylotoechus) aureola</taxonomicName>
-
<bibRefCitation id="7752CD5AA36B3BB78C585AF63BF396A7" author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="0" pageNumber="1" refId="B76" refString="Kobelt, W, 1902. Cyclophoridae. Das Tierreich. R. Friedlaender und Sohn, Berlin" title="Cyclophoridae. Das Tierreich. R." year="1902">Kobelt 1902</bibRefCitation>
: 307.
<bibRefCitation id="40B2456F0095681C6D4E8894078395F9" author="Laidlaw, FF" journalOrPublisher="Journal of the Malaysian Branch of the Royal Asiatic Society" pageId="0" pageNumber="1" pagination="25 - 37" refId="B82" refString="Laidlaw, FF, 1928. A list of the land and fresh-water mollusca of the Malay Peninsula with notes. Journal of the Malaysian Branch of the Royal Asiatic Society 6: 25 - 37" title="A list of the land and fresh-water mollusca of the Malay Peninsula with notes." volume="6" year="1928">Laidlaw 1928</bibRefCitation>
: 34.
<bibRefCitation id="AF71ABA346F75F3562FD2FC4882D4A1A" author="Hemmen, J" journalOrPublisher="Schriften zur Malakozoologie" pageId="0" pageNumber="1" pagination="35 - 70" refId="B66" refString="Hemmen, J, Hemmen, C, 2001. Aktualisierte Liste der terrestrischen Gastropoden Thailands. Schriften zur Malakozoologie 18: 35 - 70" title="Aktualisierte Liste der terrestrischen Gastropoden Thailands." volume="18" year="2001">Hemmen and Hemmen 2001</bibRefCitation>
: 39.
</paragraph>
<paragraph id="18B7988B09D2308041C730D2E7FD2745" pageId="0" pageNumber="1">
<taxonomicName id="82DB93D01E019F824E514C4B18D02305" authorityName="Mollendorff" authorityYear="1891" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina arula subsp. perakensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="arula" subSpecies="perakensis">Pupina arula perakensis</taxonomicName>
[non
<normalizedToken id="4DEFC4824261C7B877CB270DD72B06E4" originalValue="Möllendorff">Moellendorff</normalizedToken>
]-
<bibRefCitation id="3D55571753D07D2EC22A0F4622B90696" DOI="https://doi.org/10.3897/zookeys.682.12999" author="Foon, JK" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 94" refId="B48" refString="Foon, JK, Clements, GR, Liew, T-S, 2017. Diversity and biogeography of land snails (Mollusca, Gastropoda) in the limestone hills of Perak, Peninsular Malaysia. ZooKeys 682: 1 - 94, DOI: https://doi.org/10.3897/zookeys.682.12999" title="Diversity and biogeography of land snails (Mollusca, Gastropoda) in the limestone hills of Perak, Peninsular Malaysia." url="https://doi.org/10.3897/zookeys.682.12999" volume="682" year="2017">Foon et al. 2017</bibRefCitation>
: 40, fig. 15c, Ipoh, Perak.
</paragraph>
<paragraph id="F079C3A5B771E71E2649C94EA255345B" pageId="0" pageNumber="1">
<taxonomicName id="ACCF5D1165795F661A5D29B30810018D" authorityName="Vignard" authorityYear="1829" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Pupina</taxonomicName>
sp.-
<bibRefCitation id="8211E8CCC267C2262F1C08BC975F84C5" author="Sutcharit, C" journalOrPublisher="Zino Publishing, Bangkok" pageId="0" pageNumber="1" refId="B156" refString="Sutcharit, C, Tongkerd, P, Panha, S, 2018. Land snails: The invaluable bio-resources for the Kingdom of Thailand. Zino Publishing, Bangkok" title="Land snails: The invaluable bio-resources for the Kingdom of Thailand." year="2018">Sutcharit et al. 2018</bibRefCitation>
: fig. 5-11b.
</paragraph>
</subSubSection>
<subSubSection id="6CCDF49D3FAD7A5C8951730E81F17618" pageId="0" pageNumber="1" type="type material">
<paragraph id="72357822F0AB462CBA4EC1EB05C95410" pageId="0" pageNumber="1">Type material examined.</paragraph>
<paragraph id="9D627DF0ECF627BC24649F6EEA05E4B2" pageId="0" pageNumber="1">
<emphasis id="910D95057CE7E90C7D962DEF7AD6D03D" bold="true" italics="true" pageId="0" pageNumber="1">Possible syntype</emphasis>
NHMUK 1988.12.4.101 (Figs
<figureCitation id="313E3154E7005AB17839FAC53041EA7F" captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. A Pupina vescoi, specimen SMF 109956 / 1 from Cochin China B, C Pupina exclamationis B syntype MNHN-IM- 2000 - 35840 from Tonkin and C specimen NHMUK 1901.12.23.205 &quot; forma minor &quot; from Than-Moi, Tonkin D Pupina perakensis, lectotype SMF 109969 / 1 from Bukit Pondong, Perak E Pupina excisa, lectotype SMF 110778 / 1 from Kelantan F Pupina aureola, possible syntype NHMUK 1988.12.4.101 from Pinang. Photo: P. Maestrati, MNHN (B)." figureDoi="10.3897/zookeys.1119.85400.figure34" httpUri="https://binary.pensoft.net/fig/735395" pageId="0" pageNumber="1">34F</figureCitation>
,
<figureCitation id="0A600CA928319B6B81CBC8C02C4A62E0" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Shells of Pupina aureola species group from mainland Southeast Asia A-F Pupina aureola A possible syntype NHMUK 1988.12.4.101 and specimens B CUMZ 12124 C CUMZ 12126 D CUMZ 12130 E CUMZ 12133, and F CUMZ 12121 G, H Pupina stoliczkai sp. nov. G holotype CUMZ 12147 / 1 and H paratype NHMUK 20210336 I, J Pupina dorri dorri I paralectotype MNHN-IM- 2000 - 35836 and J specimen NHMUK ex. A. J. Piele Colln. Acc. No. 2242 K, L Pupina dorri isanensis spp. nov. K holotype CUMZ 12140 / 1 and L specimen CUMZ 12137 M Pupina brachysoma, lectotype MNHN-IM- 2000 - 9652. Photo: P. Maestrati, MNHN (I, M)" figureDoi="10.3897/zookeys.1119.85400.figure36" httpUri="https://binary.pensoft.net/fig/735397" pageId="0" pageNumber="1">36A</figureCitation>
) from Pinang.
</paragraph>
</subSubSection>
<subSubSection id="3E6EF5476910F2FDA458D49F5B37164C" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="C6AFAD51EF0698EF69D0765F94ADDB25" pageId="0" pageNumber="1">Other material examined.</paragraph>
<paragraph id="BE83294D8D7B9895B881DE9524089A8F" pageId="0" pageNumber="1">
<materialsCitation id="4060368F439431981283A86FF87039FF" collectingDate="1998-04-04" location="Phra Kayang Cave" specimenCount="6" stateProvince="Kra Buri District, Ranong Province">
CUMZ 12112 (2 shells and
<specimenCount id="452C9298D875339615FE6EF098439722" type="generic">6 specimens</specimenCount>
in ethanol) from
<location id="820F10D269C8682B14CDE1E8FB5A0A28" LSID="urn:lsid:plazi:treatment:F895E5960025550FA394CC1243A1DE43:820F10D269C8682B14CDE1E8FB5A0A28" name="Phra Kayang Cave" stateProvince="Kra Buri District, Ranong Province">Phra Kayang Cave</location>
,
<collectingRegion id="CC692531407C7B3A42A398DF369D6370" isEnumeration="true">Kra Buri District, Ranong Province</collectingRegion>
,
<collectingDate id="896A457513A497489683D20CE4336226" value="1998-04-04">4 Apr. 1998</collectingDate>
</materialsCitation>
.
<materialsCitation id="8FF641CD3DCEAA96EC629F40B2F2A3D1" collectingDate="2007-03-04" collectorName="Na Mueang Waterfall" location="Ko Samui District, Surat Thani Province" specimenCount="3" stateProvince="Ko Samui District, Surat Thani Province">
CUMZ 12113 (
<specimenCount id="48DC994639D1CA1E3D36CE2FCCC15097" type="generic">3 specimens</specimenCount>
in ethanol) from
<collectorName id="8C81F6D6E19FB81FCF2A7EF63A6E952C">Na Mueang Waterfall</collectorName>
,
<collectingRegion id="A60D5360A0EC2F4DD94A627D02ADD026" isEnumeration="true">Ko Samui District, Surat Thani Province</collectingRegion>
,
<collectingDate id="02CA843CF59B11393EA308367D6F069E" value="2007-03-04">4 Mar. 2007</collectingDate>
</materialsCitation>
.
<materialsCitation id="39F5D343262AF68FC942708ABA109555" collectingDate="2015-12-03" collectorName="Na Mueang Waterfall" location="Ko Samui District, Surat Thani Province" specimenCount="5" stateProvince="Ko Samui District, Surat Thani Province">
CUMZ 12114 (
<specimenCount id="A309A2DDDA11A5E436F63AA6D4441FFC" type="generic">5 specimens</specimenCount>
in ethanol) from
<collectorName id="CEBB7DCDF0554D8C9943D258DF960882">Na Mueang Waterfall</collectorName>
,
<collectingRegion id="BAE4C46CD80058ACC2704FA0C852B4F7" isEnumeration="true">Ko Samui District, Surat Thani Province</collectingRegion>
,
<collectingDate id="5E85AA02BFE733B0FDD8FF0B47F204FC" value="2015-12-03">3 Dec. 2015</collectingDate>
</materialsCitation>
.
<materialsCitation id="E707DB520956A0C04A138ECB265D888D" collectingDate="2016-12-06" collectorName="Pra Puttabhat Sri Suratth Temple" location="Kanchanadit District, Surat Thani Province" specimenCount="4" stateProvince="Kanchanadit District, Surat Thani Province">
CUMZ 12115 (
<specimenCount id="D84D695825AA989F752D3728FA5F96AB" type="generic">4 specimens</specimenCount>
in ethanol) from
<collectorName id="34B8450466A62A1A62950C0E37B451AA">Pra Puttabhat Sri Suratth Temple</collectorName>
,
<collectingRegion id="03034CE1389A6FDE6E8B8A6D535B7893" isEnumeration="true">Kanchanadit District, Surat Thani Province</collectingRegion>
,
<collectingDate id="11DBB478EF4CCFABA933610DDBB97AF9" value="2016-12-06">6 Dec. 2016</collectingDate>
</materialsCitation>
.
<materialsCitation id="E30BCDC05C9A14367B742C242E8EE9B2" collectingDate="2017-07-04" collectorName="Fig., Khiri Rat Phatthana Temple" location="Wiang Sa District, Surat Thani Province" specimenCount="7" stateProvince="Wiang Sa District, Surat Thani Province">
CUMZ 12116 (
<specimenCount id="5ECE39B9C6751438F257A8A92073D204" type="generic">7 specimens</specimenCount>
in ethanol;
<collectorName id="9D6F861A566EB1105FF2EC7FA69EB363">Fig.</collectorName>
<figureCitation id="6987A5FDD015FF89E86D5B39894731E9" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Opercula of three mainland Southeast Asian Pupina species groups A Pupina bensoni sp. nov. from the Pupina artata species group, specimen CUMZ 12047 B Pupina siamensis from the Pupina arula species group, specimen CUMZ 12067, and C Pupina aureola from the Pupina aureola species group, specimen CUMZ 12116. All not to scale." figureDoi="10.3897/zookeys.1119.85400.figure19" httpUri="https://binary.pensoft.net/fig/735380" pageId="0" pageNumber="1">19C</figureCitation>
) from
<collectorName id="6EE80F26548E867AF04AEA66EAA93382">Khiri Rat Phatthana Temple</collectorName>
,
<collectingRegion id="38D0491FE825ABF31B7BB053E89F1710" isEnumeration="true">Wiang Sa District, Surat Thani Province</collectingRegion>
,
<collectingDate id="2B4E6E7B2E565CE20D703F7A35CD17C2" value="2017-07-04">4 July 2017</collectingDate>
</materialsCitation>
.
<materialsCitation id="D66B46AD3BB4550A95049DB8489B4A1A" collectingDate="2017-03-11" collectorName="Fig." location="Lod Cave" specimenCount="42" stateProvince="Nopphitam District, Nakhon Si Thammarat District">
CUMZ 12117 (4 shells and
<specimenCount id="CE04C00062F7ED6C48822FAEEA0C454F" type="generic">42 specimens</specimenCount>
in ethanol;
<collectorName id="5CE0F8CAC7DFA5EB32376F3442ED0FE9">Fig.</collectorName>
<figureCitation id="66FD2477C94929CDAA8EF4193151E347" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Live specimens of A Pupina siamensis, specimen CUMZ 12071 from Tham Khao Loi Temple, Rayong B-D Pupina bilabiata sp. nov. B paratype CUMZ 12073 / 2 from Banpot Pisai Temple, Chumphon and specimens C CUMZ 12082 from Pha Jor Cave, Nong Bua Lam Phu and D CUMZ 12087 from Ban Yai, Surat Thani E, F Pupina godwinausteni sp. nov.: paratypes E CUMZ 12090 / 26 and F CUMZ 12091 from Khao Wong Cave, Uthai Thani G, H Pupina aureola: specimens G CUMZ 12117 from Lod Cave, Nakhon Sri Thammarat and H CUMZ 12121 from Tham Thong Panara Temple, Nakhon Sri Thammarat, showing its microhabitat in rotten log. All not to scale." figureDoi="10.3897/zookeys.1119.85400.figure32" httpUri="https://binary.pensoft.net/fig/735393" pageId="0" pageNumber="1">32G</figureCitation>
) from
<location id="ECB345616F6A2E67869DB84ABD3ECE2D" LSID="urn:lsid:plazi:treatment:F895E5960025550FA394CC1243A1DE43:ECB345616F6A2E67869DB84ABD3ECE2D" name="Lod Cave" stateProvince="Nopphitam District, Nakhon Si Thammarat District">Lod Cave</location>
,
<collectingRegion id="990FC89BBCA26B51685B5EE28A7E6134" isEnumeration="true">Nopphitam District, Nakhon Si Thammarat District</collectingRegion>
,
<collectingDate id="4F466A65939DB5D52767FB74B2B6E90E" value="2017-03-11">11 Mar. 2017</collectingDate>
</materialsCitation>
.
<materialsCitation id="8F9FEE96489A61744E272150EEA8940A" collectingDate="2017-03-11" location="Kaeo Surakan Cave" specimenCount="1" stateProvince="Lan Saka District, Nakhon Si Thammarat Province">
CUMZ 12118 (1 shell) from
<location id="1E7B4ABB898CCF2FADFDA386E4058BEA" LSID="urn:lsid:plazi:treatment:F895E5960025550FA394CC1243A1DE43:1E7B4ABB898CCF2FADFDA386E4058BEA" name="Kaeo Surakan Cave" stateProvince="Lan Saka District, Nakhon Si Thammarat Province">Kaeo Surakan Cave</location>
,
<collectingRegion id="903A6BE011AC68B3950D75CE91081F00" isEnumeration="true">Lan Saka District, Nakhon Si Thammarat Province</collectingRegion>
,
<collectingDate id="AE874239E1B9C076B7AE442E4868F9EB" value="2017-03-11">11 Mar. 2017</collectingDate>
</materialsCitation>
.
<materialsCitation id="C7B845DF51ADC57728D182DAAADE7460" collectingDate="2003-04-04" collectorName="Tham Thong Panara Temple" location="Tham Phannara District, Nakhon Sri Thammarat Province" specimenCount="6" stateProvince="Tham Phannara District, Nakhon Sri Thammarat Province">
CUMZ 12119 (
<specimenCount id="A63E914CE02C2A65BF6A3525D7A80AA3" type="generic">6 specimens</specimenCount>
in ethanol) from
<collectorName id="E8ABC41CECB5B77487532E071F504FC8">Tham Thong Panara Temple</collectorName>
,
<collectingRegion id="42E0E338835A4BB0B9083458C5978437" isEnumeration="true">Tham Phannara District, Nakhon Sri Thammarat Province</collectingRegion>
,
<collectingDate id="597EDE152564663089FF3FAC5AF45020" value="2003-04-04">4 Apr. 2003</collectingDate>
</materialsCitation>
.
<materialsCitation id="723DDC952A39C235CC327F8E88A1293C" collectingDate="2006-10-11" collectorName="Tham Thong Panara Temple" location="Tham Phannara District, Nakhon Sri Thammarat Province" specimenCount="1" stateProvince="Tham Phannara District, Nakhon Sri Thammarat Province">
CUMZ 12120 (36 shells and
<specimenCount id="EECEE0AABE72CC1846332C31A70B93CC" type="generic">1 specimen</specimenCount>
in ethanol) from
<collectorName id="E095AE68F1E16932B895B4355FDC3DAA">Tham Thong Panara Temple</collectorName>
,
<collectingRegion id="8D78D3056C3AED40D5E3E7A59D451592" isEnumeration="true">Tham Phannara District, Nakhon Sri Thammarat Province</collectingRegion>
,
<collectingDate id="F89D5D9855A2144C824653EC0247CF47" value="2006-10-11">11 Oct. 2006</collectingDate>
</materialsCitation>
.
<materialsCitation id="35D59F5B102AC81F2A63ECCCACCBE6F5" collectingDate="2012-06-11" collectorName="Figs" location="Tham Thong Panara Temple" specimenCount="100" stateProvince="Tham Phannara District, Nakhon Sri Thammarat Province">
CUMZ 12121 (&gt;
<specimenCount id="0F332736F146D75A86B06AC34B920E4C" type="generic">100 specimens</specimenCount>
in ethanol;
<collectorName id="7FA19CB94803D5B9B23D125E15EDBA9E">Figs</collectorName>
<figureCitation id="F006355920222ACBBD0FA50B273DC636" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Live specimens of A Pupina siamensis, specimen CUMZ 12071 from Tham Khao Loi Temple, Rayong B-D Pupina bilabiata sp. nov. B paratype CUMZ 12073 / 2 from Banpot Pisai Temple, Chumphon and specimens C CUMZ 12082 from Pha Jor Cave, Nong Bua Lam Phu and D CUMZ 12087 from Ban Yai, Surat Thani E, F Pupina godwinausteni sp. nov.: paratypes E CUMZ 12090 / 26 and F CUMZ 12091 from Khao Wong Cave, Uthai Thani G, H Pupina aureola: specimens G CUMZ 12117 from Lod Cave, Nakhon Sri Thammarat and H CUMZ 12121 from Tham Thong Panara Temple, Nakhon Sri Thammarat, showing its microhabitat in rotten log. All not to scale." figureDoi="10.3897/zookeys.1119.85400.figure32" httpUri="https://binary.pensoft.net/fig/735393" pageId="0" pageNumber="1">32H</figureCitation>
,
<figureCitation id="735A686A8FA938B4F0E06C298B95B48A" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Shells of Pupina aureola species group from mainland Southeast Asia A-F Pupina aureola A possible syntype NHMUK 1988.12.4.101 and specimens B CUMZ 12124 C CUMZ 12126 D CUMZ 12130 E CUMZ 12133, and F CUMZ 12121 G, H Pupina stoliczkai sp. nov. G holotype CUMZ 12147 / 1 and H paratype NHMUK 20210336 I, J Pupina dorri dorri I paralectotype MNHN-IM- 2000 - 35836 and J specimen NHMUK ex. A. J. Piele Colln. Acc. No. 2242 K, L Pupina dorri isanensis spp. nov. K holotype CUMZ 12140 / 1 and L specimen CUMZ 12137 M Pupina brachysoma, lectotype MNHN-IM- 2000 - 9652. Photo: P. Maestrati, MNHN (I, M)" figureDoi="10.3897/zookeys.1119.85400.figure36" httpUri="https://binary.pensoft.net/fig/735397" pageId="0" pageNumber="1">36F</figureCitation>
,
<figureCitation id="BE7A69F849616E0C76381AB191A24101" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. A-E Pupina aureola: specimens A CUMZ 12121 from Tham Thong Panara Temple, Nakhon Sri Thammarat B CUMZ 12124 from Talot Cave, Nakhon Sri Thammarat C CUMZ 12126 from Khao Huai Haeng Temple, Trang D CUMZ 12130 from Sra Morakot, Krabi, and E CUMZ 12133 from Khantiphol Cave, Satun F, G Pupina paviei: paralectotypes F MNHN-IM- 2000 - 35837 from Chaine de l'Elephant, Kampot, Cambodge and G RBINS 525404 from Kampot et forets de la chaine de l'Elephant, Cambodge et Kamchay. Photo: P. Maestrati, MNHN (F), F. Trus, RBINS (G)." figureDoi="10.3897/zookeys.1119.85400.figure38" httpUri="https://binary.pensoft.net/fig/735399" pageId="0" pageNumber="1">38A</figureCitation>
) from
<location id="F22207C5A6FCB89A905B9D770931BF73" LSID="urn:lsid:plazi:treatment:F895E5960025550FA394CC1243A1DE43:F22207C5A6FCB89A905B9D770931BF73" name="Tham Thong Panara Temple" stateProvince="Tham Phannara District, Nakhon Sri Thammarat Province">Tham Thong Panara Temple</location>
,
<collectingRegion id="B348456C4FC011881B0243E164F00DB7" isEnumeration="true">Tham Phannara District, Nakhon Sri Thammarat Province</collectingRegion>
,
<collectingDate id="FF3E653788EC5BF322682E913AB58AFB" value="2012-06-11">11 June 2012</collectingDate>
</materialsCitation>
.
<materialsCitation id="20694EB39165E97FB5699909A1D90F17" collectingDate="2014-01-15" collectorName="Tham Thong Panara Temple" location="Tham Phannara District, Nakhon Sri Thammarat Province" specimenCount="15" stateProvince="Tham Phannara District, Nakhon Sri Thammarat Province">
CUMZ 12122 (
<specimenCount id="D49F2C8800F6C0F86142A306F4304FCA" type="generic">15 specimens</specimenCount>
in ethanol) from
<collectorName id="420E36D1DFDF3C0F552D8BD4620AD317">Tham Thong Panara Temple</collectorName>
,
<collectingRegion id="19746FF0FA5AAFFD8F10D02D8E15B19F" isEnumeration="true">Tham Phannara District, Nakhon Sri Thammarat Province</collectingRegion>
,
<collectingDate id="E3CAFCAAE9B12F5CC52B995645311BC9" value="2014-01-15">15 Jan. 2014</collectingDate>
</materialsCitation>
.
<materialsCitation id="86D0EA14223859375928955FD82F8733" collectingDate="2017-07-04" collectorName="Tham Thong Panara Temple" location="Tham Phannara District, Nakhon Sri Thammarat Province" specimenCount="1" stateProvince="Tham Phannara District, Nakhon Sri Thammarat Province">
CUMZ 12123 (12 shells) from
<collectorName id="FD08C519F7899007DA35CDF717E8681A">Tham Thong Panara Temple</collectorName>
,
<collectingRegion id="46B49891ADC41CBF50FA28BD9E24CBCC" isEnumeration="true">Tham Phannara District, Nakhon Sri Thammarat Province</collectingRegion>
,
<collectingDate id="C51DDFE9D3B3A7D23BBF61DB6F609CBD" value="2017-07-04">4 July 2017</collectingDate>
</materialsCitation>
.
<materialsCitation id="4521D4807DA8FF9C49251688F5E197BA" collectingDate="2017-07-05" collectorName="Figs" country="Thailand" location="Talot Cave" specimenCount="1" stateProvince="Thung Song District, Nakhon Sri Thammarat Province">
CUMZ 12124 (3 shells and
<specimenCount id="8AA7C8A25C7DC65529B27802DFF1AAA6" type="generic">1 specimen</specimenCount>
in ethanol;
<collectorName id="2A8872EBCFE543709E3E87AD03B0A307">Figs</collectorName>
<figureCitation id="299CD3C993AC2B4FBEE8574FDE4E5FE8" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Shells of Pupina aureola species group from mainland Southeast Asia A-F Pupina aureola A possible syntype NHMUK 1988.12.4.101 and specimens B CUMZ 12124 C CUMZ 12126 D CUMZ 12130 E CUMZ 12133, and F CUMZ 12121 G, H Pupina stoliczkai sp. nov. G holotype CUMZ 12147 / 1 and H paratype NHMUK 20210336 I, J Pupina dorri dorri I paralectotype MNHN-IM- 2000 - 35836 and J specimen NHMUK ex. A. J. Piele Colln. Acc. No. 2242 K, L Pupina dorri isanensis spp. nov. K holotype CUMZ 12140 / 1 and L specimen CUMZ 12137 M Pupina brachysoma, lectotype MNHN-IM- 2000 - 9652. Photo: P. Maestrati, MNHN (I, M)" figureDoi="10.3897/zookeys.1119.85400.figure36" httpUri="https://binary.pensoft.net/fig/735397" pageId="0" pageNumber="1">36B</figureCitation>
,
<figureCitation id="E8C6FB6B8C26E10D868C88EC67248C22" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. A-E Pupina aureola: specimens A CUMZ 12121 from Tham Thong Panara Temple, Nakhon Sri Thammarat B CUMZ 12124 from Talot Cave, Nakhon Sri Thammarat C CUMZ 12126 from Khao Huai Haeng Temple, Trang D CUMZ 12130 from Sra Morakot, Krabi, and E CUMZ 12133 from Khantiphol Cave, Satun F, G Pupina paviei: paralectotypes F MNHN-IM- 2000 - 35837 from Chaine de l'Elephant, Kampot, Cambodge and G RBINS 525404 from Kampot et forets de la chaine de l'Elephant, Cambodge et Kamchay. Photo: P. Maestrati, MNHN (F), F. Trus, RBINS (G)." figureDoi="10.3897/zookeys.1119.85400.figure38" httpUri="https://binary.pensoft.net/fig/735399" pageId="0" pageNumber="1">38B</figureCitation>
) from
<location id="8401E5DA6353DD6B47E252250844E15D" LSID="urn:lsid:plazi:treatment:F895E5960025550FA394CC1243A1DE43:8401E5DA6353DD6B47E252250844E15D" country="Thailand" name="Talot Cave" stateProvince="Thung Song District, Nakhon Sri Thammarat Province">Talot Cave</location>
,
<collectingRegion id="EDC722DA0583C126C1F628EC85C5A788" isEnumeration="true">Thung Song District, Nakhon Sri Thammarat Province</collectingRegion>
,
<collectingCountry id="95DBC6165D04E85C34277DE3F1347670" name="Thailand">Thailand</collectingCountry>
,
<collectingDate id="2A5A20666B0D36ACD4E31BAD96F511A9" value="2017-07-05">5 July 2017</collectingDate>
</materialsCitation>
.
<materialsCitation id="4245BC9077C2ADB8F22B6460D4AADD2F" collectingDate="2015-08-06" country="Thailand" location="Mueang" municipality="Nam Phut Cave" specimenCount="1" stateProvince="Phang Nga District">
CUMZ 12125 (1 shell and
<specimenCount id="6B0AECC1811FAA45EE98DFF6353E9C85" type="generic">1 specimen</specimenCount>
in ethanol) from
<collectingMunicipality id="E110B898A8876FFAB423A2B996295D2C">Nam Phut Cave</collectingMunicipality>
,
<location id="665637C4BD0D6A7E4D6B2FBF1E9CB364" LSID="urn:lsid:plazi:treatment:F895E5960025550FA394CC1243A1DE43:665637C4BD0D6A7E4D6B2FBF1E9CB364" country="Thailand" municipality="Nam Phut Cave" name="Mueang" stateProvince="Phang Nga District">Mueang</location>
<collectingRegion id="A2264F0CEFA3ACD437BB71076346989D" country="Thailand" name="Phangnga">Phang Nga District</collectingRegion>
,
<collectingRegion id="3418599883F8B1BF8F1AC3AE7ED39176" country="Thailand" name="Phangnga">Phang Nga Province</collectingRegion>
,
<collectingDate id="37B838838CE3A16615A3CB8FFEED1895" value="2015-08-06">6 Aug. 2015</collectingDate>
</materialsCitation>
.
<materialsCitation id="2E69DE1D21127D7E892B5F3777FA0063" collectingDate="2006-10-06" collectorName="Figs" country="Thailand" location="Huai Yot District" specimenCount="1" stateProvince="Trang Province">
CUMZ 12126 (9 shells;
<collectorName id="9D0A13128922FE234280A7F985515D60">Figs</collectorName>
<figureCitation id="B9D394342D3210E9BA0471595B715D34" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Shells of Pupina aureola species group from mainland Southeast Asia A-F Pupina aureola A possible syntype NHMUK 1988.12.4.101 and specimens B CUMZ 12124 C CUMZ 12126 D CUMZ 12130 E CUMZ 12133, and F CUMZ 12121 G, H Pupina stoliczkai sp. nov. G holotype CUMZ 12147 / 1 and H paratype NHMUK 20210336 I, J Pupina dorri dorri I paralectotype MNHN-IM- 2000 - 35836 and J specimen NHMUK ex. A. J. Piele Colln. Acc. No. 2242 K, L Pupina dorri isanensis spp. nov. K holotype CUMZ 12140 / 1 and L specimen CUMZ 12137 M Pupina brachysoma, lectotype MNHN-IM- 2000 - 9652. Photo: P. Maestrati, MNHN (I, M)" figureDoi="10.3897/zookeys.1119.85400.figure36" httpUri="https://binary.pensoft.net/fig/735397" pageId="0" pageNumber="1">36C</figureCitation>
,
<figureCitation id="A45DCF559553E24308BD175834A6B530" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. A-E Pupina aureola: specimens A CUMZ 12121 from Tham Thong Panara Temple, Nakhon Sri Thammarat B CUMZ 12124 from Talot Cave, Nakhon Sri Thammarat C CUMZ 12126 from Khao Huai Haeng Temple, Trang D CUMZ 12130 from Sra Morakot, Krabi, and E CUMZ 12133 from Khantiphol Cave, Satun F, G Pupina paviei: paralectotypes F MNHN-IM- 2000 - 35837 from Chaine de l'Elephant, Kampot, Cambodge and G RBINS 525404 from Kampot et forets de la chaine de l'Elephant, Cambodge et Kamchay. Photo: P. Maestrati, MNHN (F), F. Trus, RBINS (G)." figureDoi="10.3897/zookeys.1119.85400.figure38" httpUri="https://binary.pensoft.net/fig/735399" pageId="0" pageNumber="1">38C</figureCitation>
) from Khao Huai Haeng Temple,
<location id="119318DDEAAC67C8D1AE6F8E3946EAD4" LSID="urn:lsid:plazi:treatment:F895E5960025550FA394CC1243A1DE43:119318DDEAAC67C8D1AE6F8E3946EAD4" country="Thailand" name="Huai Yot District" stateProvince="Trang Province">Huai Yot District</location>
,
<collectingRegion id="90F60DEB650D1FB38739155328F8C708" country="Thailand" name="Trang">Trang Province</collectingRegion>
,
<collectingDate id="C7F53F5D31162F68951F9B4D78B6F2ED" value="2006-10-06">6 Oct. 2006</collectingDate>
</materialsCitation>
.
<materialsCitation id="F1590804F9F37BA3C1FDFA93D8491C93" collectingDate="2006-10-06" collectorName="Ban Khao Poon" country="Thailand" location="Huai Yot District" specimenCount="5" stateProvince="Trang Province">
CUMZ 12127 (
<specimenCount id="6DD2FCDC048EC436EE32074AAB45E305" type="generic">5 specimens</specimenCount>
in ethanol) from
<collectorName id="A440D6CC22807C66A4608CBE6F9CD3E4">Ban Khao Poon</collectorName>
,
<location id="DCE0361EC8520005C9FFFFAE4736AD0E" LSID="urn:lsid:plazi:treatment:F895E5960025550FA394CC1243A1DE43:DCE0361EC8520005C9FFFFAE4736AD0E" country="Thailand" name="Huai Yot District" stateProvince="Trang Province">Huai Yot District</location>
,
<collectingRegion id="891D1A0A76BE1B24D90BBB7FFF8D5F3C" country="Thailand" name="Trang">Trang Province</collectingRegion>
,
<collectingDate id="576CBF783376FE377DE28DC976432558" value="2006-10-06">6 Oct. 2006</collectingDate>
</materialsCitation>
.
<materialsCitation id="22F7ECB37B5ECE3031D4AD211C10F494" collectingDate="1999-08-06" collectorName="Botanical Garden" country="Thailand" location="Yan Ta Khao District" specimenCount="1" stateProvince="Trang">
CUMZ 12128 (1 shell) from
<collectingRegion id="3BB1DFB680111A26E807FD8719A0E4D1" country="Thailand" name="Trang">Trang</collectingRegion>
<collectorName id="CC5F5BFBE4EF82589E94EAF52196C670">Botanical Garden</collectorName>
,
<location id="22762CA77470A4E168F8FFAF2096F1F3" LSID="urn:lsid:plazi:treatment:F895E5960025550FA394CC1243A1DE43:22762CA77470A4E168F8FFAF2096F1F3" country="Thailand" name="Yan Ta Khao District" stateProvince="Trang">Yan Ta Khao District</location>
,
<collectingRegion id="E07156C6F41D62B2932E91ED96328A11" country="Thailand" name="Trang">Trang Province</collectingRegion>
,
<collectingDate id="930CCEF30245871A30B3CAEE559F2757" value="1999-08-06">6 Aug. 1999</collectingDate>
</materialsCitation>
.
<materialsCitation id="354BA86C6A5828F4C2733DF9BA1C5A5E" collectingDate="2017-07-08" country="Thailand" location="Na Yong District" municipality="Khao Pu Chao Bureau of Monks" specimenCount="4" stateProvince="Trang Province">
CUMZ 12129 (
<specimenCount id="B04447A69D9C4DE01CBB2F1F982A24E5" type="generic">4 specimens</specimenCount>
in ethanol) from
<collectingMunicipality id="5CCAF99028798611B254D74F0E0D6DD0">Khao Pu Chao Bureau of Monks</collectingMunicipality>
,
<location id="337ED40940C45624F23EAAE58810BE66" LSID="urn:lsid:plazi:treatment:F895E5960025550FA394CC1243A1DE43:337ED40940C45624F23EAAE58810BE66" country="Thailand" municipality="Khao Pu Chao Bureau of Monks" name="Na Yong District" stateProvince="Trang Province">Na Yong District</location>
,
<collectingRegion id="0C8DA73ED1BD99B9E14913CD1E4184CC" country="Thailand" name="Trang">Trang Province</collectingRegion>
,
<collectingDate id="AAA66C2EDF14FF6865F283BC1987DE06" value="2017-07-08">8 July 2017</collectingDate>
</materialsCitation>
.
<materialsCitation id="ACE814744246877D649D877B0C2176AD" collectingDate="2009-01-15" collectorName="Figs" country="Thailand" location="Khlong Thom District" municipality="Sra Morakot" specimenCount="1" stateProvince="Krabi Province">
CUMZ 12130 (8 shells;
<collectorName id="CB5151969D2E5E4DD752A6D50F7397AC">Figs</collectorName>
<figureCitation id="65C6C0B7160FBD96E3B53FE3C662E7B3" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Umbilical, columellar and parietal views of A Pupinella mansuyi, specimen CUMZ 12148 from Pha Chu, Nan B Pupina artata from the Pupina artata species group, specimen CUMZ 12003 from Ban Ping Khong, Chiang Mai C Pupina godwinausteni sp. nov. from the Pupina arula species group, holotype CUMZ 12090 / 1 D Pupina aureola from the Pupina aureola species group, specimen CUMZ 12130 from Sra Morakot, Krabi." figureDoi="10.3897/zookeys.1119.85400.figure10" httpUri="https://binary.pensoft.net/fig/735371" pageId="0" pageNumber="1">10D</figureCitation>
,
<figureCitation id="70D6338BBF6FFFC28F471EBA09D57D3E" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Shells of Pupina aureola species group from mainland Southeast Asia A-F Pupina aureola A possible syntype NHMUK 1988.12.4.101 and specimens B CUMZ 12124 C CUMZ 12126 D CUMZ 12130 E CUMZ 12133, and F CUMZ 12121 G, H Pupina stoliczkai sp. nov. G holotype CUMZ 12147 / 1 and H paratype NHMUK 20210336 I, J Pupina dorri dorri I paralectotype MNHN-IM- 2000 - 35836 and J specimen NHMUK ex. A. J. Piele Colln. Acc. No. 2242 K, L Pupina dorri isanensis spp. nov. K holotype CUMZ 12140 / 1 and L specimen CUMZ 12137 M Pupina brachysoma, lectotype MNHN-IM- 2000 - 9652. Photo: P. Maestrati, MNHN (I, M)" figureDoi="10.3897/zookeys.1119.85400.figure36" httpUri="https://binary.pensoft.net/fig/735397" pageId="0" pageNumber="1">36D</figureCitation>
,
<figureCitation id="1DAFC5D0F45DFCCDB62416986B6280AA" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. A-E Pupina aureola: specimens A CUMZ 12121 from Tham Thong Panara Temple, Nakhon Sri Thammarat B CUMZ 12124 from Talot Cave, Nakhon Sri Thammarat C CUMZ 12126 from Khao Huai Haeng Temple, Trang D CUMZ 12130 from Sra Morakot, Krabi, and E CUMZ 12133 from Khantiphol Cave, Satun F, G Pupina paviei: paralectotypes F MNHN-IM- 2000 - 35837 from Chaine de l'Elephant, Kampot, Cambodge and G RBINS 525404 from Kampot et forets de la chaine de l'Elephant, Cambodge et Kamchay. Photo: P. Maestrati, MNHN (F), F. Trus, RBINS (G)." figureDoi="10.3897/zookeys.1119.85400.figure38" httpUri="https://binary.pensoft.net/fig/735399" pageId="0" pageNumber="1">38D</figureCitation>
) from
<collectingMunicipality id="FF3071D5AAB71707EB2DE66F3A1FB5BF">Sra Morakot</collectingMunicipality>
,
<location id="A59C92B6842316F7FDF3DF52B24C19E6" LSID="urn:lsid:plazi:treatment:F895E5960025550FA394CC1243A1DE43:A59C92B6842316F7FDF3DF52B24C19E6" country="Thailand" municipality="Sra Morakot" name="Khlong Thom District" stateProvince="Krabi Province">Khlong Thom District</location>
,
<collectingRegion id="281ED97149DE52AE586A9DE246852A2C" country="Thailand" name="Krabi">Krabi Province</collectingRegion>
,
<collectingDate id="3B4FCE223642630FF3D99EACCE0DE7B5" value="2009-01-15">15 Jan. 2009</collectingDate>
</materialsCitation>
.
<materialsCitation id="E19772A881AAD233E133C0590BB6F088" collectingDate="2012-05-17" collectorName="Sra Morakot" country="Thailand" location="Khlong Thom District" specimenCount="2" stateProvince="Krabi Province">
CUMZ 12131 (
<specimenCount id="3E5B7763FECA61F949A692D4E1F5085E" type="generic">2 specimens</specimenCount>
in ethanol) from
<collectorName id="C8F02BB5AFE62D75F7F310AC35FD7802">Sra Morakot</collectorName>
,
<location id="27334F659AFD56B92979B7E4BB2ED61E" LSID="urn:lsid:plazi:treatment:F895E5960025550FA394CC1243A1DE43:27334F659AFD56B92979B7E4BB2ED61E" country="Thailand" name="Khlong Thom District" stateProvince="Krabi Province">Khlong Thom District</location>
,
<collectingRegion id="FDDAA13EB37D1AD8C7CCAD167E9877BC" country="Thailand" name="Krabi">Krabi Province</collectingRegion>
,
<collectingDate id="C3C94BD910A5A70D114354FD0B631026" value="2012-05-17">17 May 2012</collectingDate>
</materialsCitation>
.
<materialsCitation id="46903918CC134AB441FD9BAC4F214EDB" collectingDate="2008-04-07" collectorName="Toe Bu Cliff Viewpoint" country="Thailand" location="Mueang" specimenCount="15" stateProvince="Satun District">
CUMZ 12132 (
<specimenCount id="07C529E7B53A2CC00252B815635B4177" type="generic">15 specimens</specimenCount>
in ethanol) from
<collectorName id="3389F0E5572AB05A21A405E392A83C0A">Toe Bu Cliff Viewpoint</collectorName>
,
<location id="355E27B8F298F29C2C699FD81ABAA124" LSID="urn:lsid:plazi:treatment:F895E5960025550FA394CC1243A1DE43:355E27B8F298F29C2C699FD81ABAA124" country="Thailand" name="Mueang" stateProvince="Satun District">Mueang</location>
<collectingRegion id="3968C9CCE2ACA119A1E23525CB010C02" country="Thailand" name="Satun">Satun District</collectingRegion>
,
<collectingRegion id="57AC5903FE788F6589DB4915D638CDB8" country="Thailand" name="Satun">Satun Province</collectingRegion>
,
<collectingDate id="1A3CBAB2CA2FC7DB85CCA9D164329F9A" value="2008-04-07">7 Apr. 2008</collectingDate>
</materialsCitation>
.
<materialsCitation id="26AD49A7A35759A9CC1C190DDEA95085" collectingDate="2009-01-13" collectorName="Figs" country="Thailand" location="Thung Wa District" municipality="Khantiphol Cave" specimenCount="1" stateProvince="Satun Province">
CUMZ 12133 (7 shells;
<collectorName id="CA06C163CF2E067BB04B67C0A1909C58">Figs</collectorName>
36E, 38E) from
<collectingMunicipality id="F58CB114FF9BB76CE6BC1E3FD37CCA12">Khantiphol Cave</collectingMunicipality>
,
<location id="49E05BF3EA3C148A31465911288B5A40" LSID="urn:lsid:plazi:treatment:F895E5960025550FA394CC1243A1DE43:49E05BF3EA3C148A31465911288B5A40" country="Thailand" municipality="Khantiphol Cave" name="Thung Wa District" stateProvince="Satun Province">Thung Wa District</location>
,
<collectingRegion id="3488375C5FA80BA00A7D5CA300574624" country="Thailand" name="Satun">Satun Province</collectingRegion>
,
<collectingDate id="47A3A7680792261CB9662C0D103E9914" value="2009-01-13">13 Jan. 2009</collectingDate>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="FF040BEC670DEC33AFFEFA3E086F7857" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="46D5C4D85E68C22BA38B87A368EAD0C8" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="625D8171899CCF7CCCA09B88EC95DE46" pageId="0" pageNumber="1">Shell ovate to fusiform; last whorl ca. 70-75% of shell height. Apertural lip thickened to highly thickened but not expanded. Parietal tooth thickened, fin-shaped or tooth-like, always located next to but not covering posterior canal; columellar tooth fin-shaped, thickened, located next to anterior canal. Posterior canal slightly bulging outward.</paragraph>
</subSubSection>
<subSubSection id="EF54FBB5F06ED2BFA46DDE8B5885B60A" pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph id="6568F1402410EE7CA59EE2A3D2B67A10" pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph id="85D14DBCDAD80747D240DE940207578E" pageId="0" pageNumber="1">
<taxonomicName id="61332EDAC1B67061CE84668BF9E74D36" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. aureola" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="aureola">
<emphasis id="1DF44ECBFC164E563A97C48E3CC5AD7F" italics="true" pageId="0" pageNumber="1">P. aureola</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="A30C0608CE7AA177C7DE934490D3D5C2" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. stoliczkai" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="stoliczkai">
<emphasis id="592972832731B4C0F881795F96695BCD" italics="true" pageId="0" pageNumber="1">P. stoliczkai</emphasis>
</taxonomicName>
sp. nov. in shell shape and having both fin-shaped and highly thickened parietal and columellar teeth located next to their respective canals; the posterior canal slightly bulges outward. However,
<taxonomicName id="96E5F5D285576D1B6DF5D7E59C029AF4" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. aureola" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="aureola">
<emphasis id="B44E48A2A11D40E8F4CC14951DDBE389" italics="true" pageId="0" pageNumber="1">P. aureola</emphasis>
</taxonomicName>
does not have a furrow between inner and outer peristomes.
</paragraph>
</subSubSection>
<subSubSection id="9149C00BB65A4985A373E441CB843BD3" pageId="0" pageNumber="1" type="distribution">
<paragraph id="269F48A69F259CCCF7FDB9E3464C83B2" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="7982FFA9E160DD221340DFFD0D96AB40" pageId="0" pageNumber="1">
Malaysia and southern Thailand (
<bibRefCitation id="00D4CBDED65F35AB1BA7052068E888C7" author="Maassen, WJM" journalOrPublisher="Proceedings of the United States National Museum" pageId="0" pageNumber="1" refId="B87" refString="Maassen, WJM, 2001. A preliminary checklist of the non-marine molluscs of West-Malaysia, A handlist. De Kreukel (Supplement): 1-155." title="A preliminary checklist of the non-marine molluscs of West-Malaysia, A handlist. De Kreukel (Supplement): 1 - 155." year="2001">Maassen 2001</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C28F76E4F2485C9B62995B7CB306616D" pageId="0" pageNumber="1" type="remarks">
<paragraph id="182E977D6EC9316C94C0E2C4A91695AB" pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph id="F0FA29973EF81F7D6F6C41B8BC992805" pageId="0" pageNumber="1">
This species has high variation in shell shape from ovate to fusiform, and the parietal tooth varies from fin-shaped to tooth-like. Despite those shell variations, we assign these shell morphs to
<taxonomicName id="3F1EF810B4ADE73B342374FA276378A5" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. aureola" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="aureola">
<emphasis id="DCD426019C68555D33096E7C6ECEB061" italics="true" pageId="0" pageNumber="1">P. aureola</emphasis>
</taxonomicName>
due to the uniform position of a parietal tooth that is always located next to the posterior canal, and a columellar tooth that is always fin-shaped and not extending over the apertural lip.
</paragraph>
<paragraph id="900FA062112D0A88274CEC8196F22A12" pageId="0" pageNumber="1">
By comparing with the possible type specimen, the specimen of
<taxonomicName id="816C310D96F47F067DB585AA1FE507BC" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. arula subsp. perakensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="arula" subSpecies="perakensis">
<emphasis id="5B78999F5440E6454C53C69B3C2A8831" italics="true" pageId="0" pageNumber="1">P. arula perakensis</emphasis>
</taxonomicName>
figured in
<bibRefCitation id="4D7C02BBDB7FA9CDF019158A3B22C744" DOI="https://doi.org/10.3897/zookeys.682.12999" author="Foon, JK" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 94" refId="B48" refString="Foon, JK, Clements, GR, Liew, T-S, 2017. Diversity and biogeography of land snails (Mollusca, Gastropoda) in the limestone hills of Perak, Peninsular Malaysia. ZooKeys 682: 1 - 94, DOI: https://doi.org/10.3897/zookeys.682.12999" title="Diversity and biogeography of land snails (Mollusca, Gastropoda) in the limestone hills of Perak, Peninsular Malaysia." url="https://doi.org/10.3897/zookeys.682.12999" volume="682" year="2017">Foon et al. (2017</bibRefCitation>
: fig. 15c) from Gunung Datok Plot, Ipoh, Perak should belong to
<taxonomicName id="145D9E916FF9AE7124A28FE9F3CA4BE8" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. aureola" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="aureola">
<emphasis id="E2081D2EE266CE85DA6B6CC617D32846" italics="true" pageId="0" pageNumber="1">P. aureola</emphasis>
</taxonomicName>
(Foon, pers. comm.).
</paragraph>
</subSubSection>
</treatment>
</document>
Reference in a new issue View git blame Copy permalink