treatments-xml/data/F0/02/87/F00287A30E5BFFD2A6B2FC3AC838FB30.xml

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<mods:title id="5B535585E1D8202AFB0C5EF137DBE6C1">Identity of Pagurus watasei (Crustacea: Decapoda: Anomura: Paguroidea)</mods:title>
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<emphasis id="4ADFEAA70E5BFFD6A6B2FC3AC844FC76" bold="true" box="[941,1139,914,938]" italics="true" pageId="0" pageNumber="39">Dardanus aspersus</emphasis>
(
<bibRefCitation id="1C3A4B440E5BFFD6A160FC3AC92FFC75" author="Berthold, A. A." box="[1151,1304,914,938]" pageId="0" pageNumber="39" pagination="37 - 48" refId="ref3481" refString="Berthold, A. A. 1845. Ueber verschiedene neue oder seltene Reptilien aus Neu-Granada und Crustaceen aus China. Nachrichten von der Koniglichen Gesellschaft der Wissenschaften zu Gottingen 1845: 37 - 48." type="journal article" year="1845">Berthold, 1845</bibRefCitation>
)
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(
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Figs 
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, 
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<emphasis id="4ADFEAA70E5BFFD6A62BFC5CCFD5FBD4" box="[820,994,1012,1035]" italics="true" pageId="0" pageNumber="39">Pagurus aspersus</emphasis>
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<bibRefCitation id="1C3A4B440E5BFFD6A6F5FC5CC89DFBD4" author="Berthold, A. A." box="[1002,1194,1011,1036]" pageId="0" pageNumber="39" pagination="37 - 48" refId="ref3481" refString="Berthold, A. A. 1845. Ueber verschiedene neue oder seltene Reptilien aus Neu-Granada und Crustaceen aus China. Nachrichten von der Koniglichen Gesellschaft der Wissenschaften zu Gottingen 1845: 37 - 48." type="journal article" year="1845">Berthold, 1845: 45</bibRefCitation>
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(
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locality: 
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); 1846: 21, pl. 2, fig. 1.
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<taxonomicName id="BFAB4D360E5BFFD6A62BFB9DCFD5FB92" authorityName="Fabricius" authorityYear="1775" box="[820,994,1077,1101]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="diogenes">
<emphasis id="4ADFEAA70E5BFFD6A62BFB9DCFD5FB92" box="[820,994,1077,1101]" italics="true" pageId="0" pageNumber="39">Pagurus diogenes</emphasis>
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.—De Haan 1849: 208; 
<bibRefCitation id="1C3A4B440E5BFFD6A1C8FB9DC97BFB92" author="Terao, A." box="[1239,1356,1077,1101]" pageId="0" pageNumber="39" pagination="54 - 62" refId="ref4453" refString="Terao, A. 1932. Crustacea. Pls. 54 - 62. In: Ishikawa, C., Okamura, K., Tanaka, S., Maruyama, H., Terao, A., Higure, T., and Senou, H. (Eds.) Illustrations of Japanese Aquatic Plants and Animals, Volume 2. Fisheries Society of Japan, Tokyo. [In Japanese and English]" type="book chapter" year="1932">Terao 1932</bibRefCitation>
, pl. 57, fig. 2. [not 
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<emphasis id="4ADFEAA70E5BFFD6A6DFFBFEC856FBB2" box="[960,1121,1109,1133]" italics="true" pageId="0" pageNumber="39">Cancer diogenes</emphasis>
Linnaeus, 1758
</taxonomicName>
(= 
<taxonomicName id="BFAB4D360E5BFFD6A037FBFDC863FB51" ID-CoL="77dc9e1d-664e-43bb-8a33-fb85a46c5aa2" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Malacostraca" family="Diogenidae" genus="Petrochirus" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="diogenes">
<emphasis id="4ADFEAA70E5BFFD6A037FBFDCF95FB51" italics="true" pageId="0" pageNumber="39">Petrochirus diogenes</emphasis>
(Linnaeus, 1758)
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]
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<taxonomicName id="BFAB4D360E5BFFD6A62BFB3FC8AEFB71" authority="Terao, 1913: 380" authorityName="Terao" authorityPageNumber="380" authorityYear="1913" box="[820,1177,1174,1198]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="watasei">
<emphasis id="4ADFEAA70E5BFFD6A62BFB3FCFECFB71" box="[820,987,1175,1198]" italics="true" pageId="0" pageNumber="39">Pagurus watasei</emphasis>
<bibRefCitation id="1C3A4B440E5BFFD6A6F8FB3FC8AEFB71" author="Terao, A." box="[999,1177,1174,1198]" pageId="0" pageNumber="39" pagination="355 - 391" refId="ref4418" refString="Terao, A. 1913. A catalogue of hermit-crabs found in Japan (Paguridea excluding Lithodidae), with descriptions of four new species. Annotationes Zoologicae Japonenses 8: 355 - 391." type="journal article" year="1913">Terao, 1913: 380</bibRefCitation>
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, text-fig. 3. 
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locality: 
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, 
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(now located in 
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, 
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, 
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, southern 
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<taxonomicName id="BFAB4D360E5BFFD6A62BFB50CFC3FACF" authorityName="Gee" authorityYear="1935" box="[820,1012,1272,1296]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="diogenes">
<emphasis id="4ADFEAA70E5BFFD6A62BFB50CFC3FACF" box="[820,1012,1272,1296]" italics="true" pageId="0" pageNumber="39">Dardanus diogenes</emphasis>
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.— 
<treatmentCitation id="F90A10A40E5BFFD6A10FFB50C891FACF" author="Gee, N. G." box="[1040,1190,1272,1296]" page="159" pageId="0" pageNumber="39" year="1925">
<bibRefCitation id="1C3A4B440E5BFFD6A10FFB50C891FACF" author="Gee, N. G." box="[1040,1190,1272,1296]" pageId="0" pageNumber="39" pagination="156 - 166" refId="ref3719" refString="Gee, N. G. 1925. Tentative list of Chinese decapod Crustacea. Including those represented in the collections of the United States National Museum (marked with an *) with localities from which collected. The Lingnaam Agricultural Review 3: 156 - 166." type="journal article" year="1925">Gee 1925: 159</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="F90A10A40E5BFFD6A1ACFB50C95AFACF" author="Miyake, S." box="[1203,1389,1272,1296]" page="644" pageId="0" pageNumber="39" year="1965">
<bibRefCitation id="1C3A4B440E5BFFD6A1ACFB50C95AFACF" author="Miyake, S." box="[1203,1389,1272,1296]" pageId="0" pageNumber="39" pagination="630 - 652" refId="ref4215" refString="Miyake, S. 1965. Anomura. Pp. 630 - 652. In: Okada, K., Uchida, S. and Uchida, T. (Eds) New Illustrated Encyclopedia of the Fauna of Japan, Volume 2. Hokuryukan, Tokyo. [In Japanese]" type="book chapter" year="1965">Miyake 1965: 644</bibRefCitation>
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, fig. 1080.
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<treatmentCitationGroup id="58BB119B0E5BFFD6A62BFA91C844FA0C" pageId="0" pageNumber="39">
<taxonomicName id="BFAB4D360E5BFFD6A62BFA91CFC2FA8E" ID-CoL="3126f6b1-71c2-4c4c-910c-6875b407b0bc" baseAuthorityName="Berthold" baseAuthorityYear="1845" box="[820,1013,1337,1361]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="aspersus">
<emphasis id="4ADFEAA70E5BFFD6A62BFA91CFC2FA8E" box="[820,1013,1337,1361]" italics="true" pageId="0" pageNumber="39">Dardanus aspersus</emphasis>
</taxonomicName>
.— 
<treatmentCitation id="F90A10A40E5BFFD6A10DFA91C909FA8E" author="Holthuis, L. B. &amp; Sakai, T." box="[1042,1342,1337,1361]" page="96" pageId="0" pageNumber="39" year="1970">
<bibRefCitation id="1C3A4B440E5BFFD6A10DFA91C909FA8E" author="Holthuis, L. B. &amp; Sakai, T." box="[1042,1342,1337,1361]" pageId="0" pageNumber="39" refId="ref3844" refString="Holthuis, L. B. and Sakai, T. 1970. Ph. F. Von Siebold and Fauna Japonica. - A History of Early Japanese Zoology. Academic Press of Japan, Tokyo, xvi + 323 pp., 38 pls, 1 map. [In English and Japanese]" type="book" year="1970">Holthuis and Sakai 1970: 96</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="F90A10A40E5BFFD6A052FA91CF91FAAE" author="Miyake, S." page="64" pageId="0" pageNumber="39" year="1978">
<bibRefCitation id="1C3A4B440E5BFFD6A052FA91CF91FAAE" author="Miyake, S." pageId="0" pageNumber="39" refId="ref4268" refString="Miyake, S. 1978. The Crustacean Anomura of Sagami Bay. Biological Laboratory, Imperial Household, Tokyo, 200 (English) + 161 (Japanese) pp., 4 pls." type="book" year="1978">Miyake 1978: 64</bibRefCitation>
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, text-fig. 23, pl. 1, fig. 2; 1982: 109, pl. 37, fig. 2; 1998: 109, pl. 37, fig. 2; 
<treatmentCitation id="F90A10A40E5BFFD6A153FAD2C9A4FA4D" author="Yamaguchi, T. &amp; Baba, K." box="[1100,1427,1402,1426]" page="275" pageId="0" pageNumber="39" year="1993">
<bibRefCitation id="1C3A4B440E5BFFD6A153FAD2C9A4FA4D" author="Yamaguchi, T. &amp; Baba, K." box="[1100,1427,1402,1426]" pageId="0" pageNumber="39" pagination="145 - 570" refId="ref4576" refString="Yamaguchi, T. and Baba, K. 1993. Crustacean specimens collected in Japan by Ph. F. von Siebold and H. Burger and held by the Nationaal Natuurhistorisch Museum in Leiden and other museums. Pp. 145 - 570. In: Yamaguchi, T. (Ed.) Ph. F. Von Siebold and Natural History of Japan. Crustacea. Carcinological Society of Japan, Tokyo. [In English and Japanese]" type="book chapter" year="1993">
<collectingRegion id="BA6FF8570E5BFFD6A153FAD2C88BFA4D" box="[1100,1212,1402,1426]" country="Japan" name="Yamaguchi" pageId="0" pageNumber="39">Yamaguchi</collectingRegion>
and Baba 1993: 275
</bibRefCitation>
</treatmentCitation>
, fig. 74; 
<treatmentCitation id="F90A10A40E5BFFD6A682FA32C864FA6C" author="Asakura, A." box="[925,1107,1434,1459]" page="34" pageId="0" pageNumber="39" year="2006">
<bibRefCitation id="1C3A4B440E5BFFD6A682FA32C864FA6C" author="Asakura, A." box="[925,1107,1434,1459]" pageId="0" pageNumber="39" pagination="23 - 103" refId="ref3428" refString="Asakura, A. 2006. Shallow water hermit crabs of the families Pylochelidae, Diogenidae and Paguridae (Crustacea: Decapoda: Anomura) from the Sea of Japan, with a description of a new species of Diogenes. Bulletin of the Toyama Science Museum 29: 23 - 103." type="journal article" year="2006">Asakura 2006: 34</bibRefCitation>
</treatmentCitation>
, figs. 27, 28; 
<treatmentCitation id="F90A10A40E5BFFD6A1FFFA32CF93FA0C" author="McLaughlin, P. A. &amp; Rahayu, D. L. &amp; Komai, T. &amp; Chan, T. - Y." page="96" pageId="0" pageNumber="39" year="2007">
<bibRefCitation id="1C3A4B440E5BFFD6A1FFFA32CF93FA0C" author="McLaughlin, P. A. &amp; Rahayu, D. L. &amp; Komai, T. &amp; Chan, T. - Y." pageId="0" pageNumber="39" refId="ref4142" refString="McLaughlin, P. A., Rahayu, D. L., Komai, T. and Chan, T. - Y. 2007. A Catalog of the Hermit Crabs (Paguroidea) of Taiwan. National Taiwan Ocean University, Keelung, 367 pp." type="book" year="2007">
McLaughlin 
<emphasis id="4ADFEAA70E5BFFD6A07BFA33C9AFFA6D" box="[1380,1432,1434,1458]" italics="true" pageId="0" pageNumber="39">et al.</emphasis>
2007: 96
</bibRefCitation>
</treatmentCitation>
, 2 unnumbered figs.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="30B1653E0E5BFFD6A651FA54C8B2F927" pageId="0" pageNumber="39" type="materials_examined">
<paragraph id="781436B50E5BFFD6A651FA54C8B2F927" blockId="0.[820,1432,1532,2077]" pageId="0" pageNumber="39">
<emphasis id="4ADFEAA70E5BFFD6A651FA54C817F9CB" bold="true" box="[846,1056,1532,1556]" pageId="0" pageNumber="39">Material examined</emphasis>
. 
<materialsCitation id="C8C33CE80E5BFFD6A120FA55C972F98A" ID-GBIF-Occurrence="3097977304" collectingDate="1896-04-20" collectorName="Mitsukuri &amp; Hara" location="Satsuma" municipality="Komenotsu" pageId="0" pageNumber="39" specimenCount="1" typeStatus="holotype">
<collectingMunicipality id="9870ACCF0E5BFFD6A120FA55C881F9CB" box="[1087,1206,1533,1556]" pageId="0" pageNumber="39">Komenotsu</collectingMunicipality>
, 
<location id="7D74606E0E5BFFD6A1D3FA54C914F9CB" LSID="urn:lsid:plazi:treatment:F00287A30E5BFFD2A6B2FC3AC838FB30:7D74606E0E5BFFD6A1D3FA54C914F9CB" box="[1228,1315,1532,1556]" municipality="Komenotsu" name="Satsuma" pageId="0" pageNumber="39">Satsuma</location>
, 
<date id="0C1510750E5BFFD6A026FA54CF50F9EA" pageId="0" pageNumber="39" value="1896-04-20">
<collectingDate id="1C51E99D0E5BFFD6A026FA54CF50F9EA" pageId="0" pageNumber="39" value="1896-04-20">20 April 1896</collectingDate>
</date>
, collected by 
<collectorName id="D55E53630E5BFFD6A6E4F9B5C857F9EA" box="[1019,1120,1565,1589]" pageId="0" pageNumber="39">Mitsukuri</collectorName>
and 
<collectorName id="D55E53630E5BFFD6A185F9B5C8F9F9EB" box="[1178,1230,1565,1588]" pageId="0" pageNumber="39">Hara</collectorName>
, 
<typeStatus id="A71088170E5BFFD6A1C3F9B5C902F9EA" box="[1244,1333,1565,1589]" pageId="0" pageNumber="39" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="BFAB4D360E5BFFD6A042F9B5CF9FF98A" authorityName="Terao" authorityYear="1913" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="watasei">
<emphasis id="4ADFEAA70E5BFFD6A042F9B5CF9FF98A" italics="true" pageId="0" pageNumber="39">Pagurus watasei</emphasis>
</taxonomicName>
, female (cl 
<quantity id="BF539B500E5BFFD6A135F995C8B2F98A" box="[1066,1157,1597,1621]" metricMagnitude="-2" metricUnit="m" metricValue="1.67" pageId="0" pageNumber="39" unit="mm" value="16.7">16.7 mm</quantity>
), UMUTZ-Crs-33
</materialsCitation>
. 
<materialsCitation id="C8C33CE80E5BFFD6A04DF996C918F949" ID-GBIF-Occurrence="3097977303" collectorName="Higher Normal School" location="No" pageId="0" pageNumber="39" specimenCount="1" specimenCount-male="1" stateProvince="Tokyo" typeStatus="holotype">
<location id="7D74606E0E5BFFD6A04DF996C946F98A" LSID="urn:lsid:plazi:treatment:F00287A30E5BFFD2A6B2FC3AC838FB30:7D74606E0E5BFFD6A04DF996C946F98A" box="[1362,1393,1598,1621]" name="No" pageId="0" pageNumber="39" stateProvince="Tokyo">No</location>
locality and date records, collected by 
<collectingRegion id="BA6FF8570E5BFFD6A1ADF9F6C8C6F9A9" box="[1202,1265,1630,1654]" country="Japan" name="Tokyo" pageId="0" pageNumber="39">Tokyo</collectingRegion>
<collectorName id="D55E53630E5BFFD6A1E6F9F6CF4CF949" pageId="0" pageNumber="39">Higher Normal School</collectorName>
, 
<specimenCount id="6EADFD3C0E5BFFD6A696F9D6CFE7F949" box="[905,976,1662,1686]" pageId="0" pageNumber="39" type="male">1 male</specimenCount>
(cl 14.0 mm), UMUTZ-Crs-c110
</materialsCitation>
. 
<collectingRegion id="BA6FF8570E5BFFD6A024F9D6CFA9F968" country="Japan" name="Nagasaki" pageId="0" pageNumber="39">Nagasaki Prefecture</collectingRegion>
, 1912, collected by 
<collectorName id="D55E53630E5BFFD6A17DF937C886F968" box="[1122,1201,1695,1719]" pageId="0" pageNumber="39">Kaneko</collectorName>
, 1 ovigerous female (cl 
<quantity id="BF539B500E5BFFD6A62BF917CFB8F908" box="[820,911,1727,1751]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" pageId="0" pageNumber="39" unit="mm" value="13.5">13.5 mm</quantity>
), UMUTZ-Crs-c112. 
<materialsCitation id="C8C33CE80E5BFFD6A16CF917C8B6F927" ID-GBIF-Occurrence="3097977301" location="Nagasaki Prefecture" pageId="0" pageNumber="39" specimenCount="1" specimenCount-male="1" stateProvince="Nagasaki" typeStatus="holotype">
<collectingRegion id="BA6FF8570E5BFFD6A16CF917C974F908" box="[1139,1347,1727,1751]" country="Japan" name="Nagasaki" pageId="0" pageNumber="39">Nagasaki Prefecture</collectingRegion>
, 
<specimenCount id="6EADFD3C0E5BFFD6A04FF917C9AFF908" box="[1360,1432,1727,1751]" pageId="0" pageNumber="39" type="male">1 male</specimenCount>
(cl 
<quantity id="BF539B500E5BFFD6A64BF948CF99F927" box="[852,942,1760,1784]" metricMagnitude="-2" metricUnit="m" metricValue="1.3599999999999999" pageId="0" pageNumber="39" unit="mm" value="13.6">13.6 mm</quantity>
), UMUTZ-Crs-c195
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="30B1653E0E5BFFD7A651F8A8CF54F9CB" lastPageId="1" lastPageNumber="40" pageId="0" pageNumber="39" type="description">
<paragraph id="781436B50E5BFFD6A651F8A8CF58F864" blockId="0.[820,1432,1532,2077]" pageId="0" pageNumber="39">
<emphasis id="4ADFEAA70E5BFFD6A651F8A8C90DF8C7" bold="true" box="[846,1338,1792,1816]" pageId="0" pageNumber="39">
Redescription of 
<typeStatus id="A71088170E5BFFD6A116F8A8C851F8C7" box="[1033,1126,1792,1816]" pageId="0" pageNumber="39" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="BFAB4D360E5BFFD6A192F8A9C90DF8C7" authorityName="Terao" authorityYear="1913" box="[1165,1338,1792,1816]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="watasei">
<emphasis id="4ADFEAA70E5BFFD6A192F8A9C90DF8C7" bold="true" box="[1165,1338,1792,1816]" italics="true" pageId="0" pageNumber="39">Pagurus watasei</emphasis>
</taxonomicName>
</emphasis>
. Shield (
<figureCitation id="E0902A300E5BFFD6A623F889CF90F8E6" box="[828,935,1825,1849]" captionStart="Fig" captionStartId="2.[156,187,1884,1906]" captionTargetBox="[170,1417,204,1862]" captionTargetId="figure-68@2.[170,1417,204,1862]" captionTargetPageId="2" captionText="Fig. 1. Dardanus aspersus (Berthold, 1845), holotype of Pagurus watasei Terao, 1913, female (cl 16.7mm), UMUTZ-Crs-33. A, entire animal, dorsal view; B, shield and cephalic appendages, dorsal view; C, left cheliped, outer view; D, right cheliped, outer view." figureDoi="http://doi.org/10.5281/zenodo.4584843" httpUri="https://zenodo.org/record/4584843/files/figure.png" pageId="0" pageNumber="39">Fig. 1A, B</figureCitation>
) approximately as long as broad; dorsal surface with tufts of stiff setae laterally and broad, U-shaped suture on posteromedian part; anterolateral margins terraced. Rostrum broadly triangular, nearly reaching tips of blunt lateral lobes.
</paragraph>
<paragraph id="781436B50E5BFFD7A651F86BCD74FF20" blockId="0.[820,1432,1532,2077]" lastBlockId="1.[156,768,198,2077]" lastPageId="1" lastPageNumber="40" pageId="0" pageNumber="39">
Ocular peduncles (
<figureCitation id="E0902A300E5BFFD6A10BF86BC868F803" box="[1044,1119,1987,2012]" captionStart="Fig" captionStartId="2.[156,187,1884,1906]" captionTargetBox="[170,1417,204,1862]" captionTargetId="figure-68@2.[170,1417,204,1862]" captionTargetPageId="2" captionText="Fig. 1. Dardanus aspersus (Berthold, 1845), holotype of Pagurus watasei Terao, 1913, female (cl 16.7mm), UMUTZ-Crs-33. A, entire animal, dorsal view; B, shield and cephalic appendages, dorsal view; C, left cheliped, outer view; D, right cheliped, outer view." figureDoi="http://doi.org/10.5281/zenodo.4584843" httpUri="https://zenodo.org/record/4584843/files/figure.png" pageId="0" pageNumber="39">Fig. 1B</figureCitation>
) slightly depressed dorsoventrally, stout, 0.6 of shield length; corneas somewhat dilated, broader than bases of peduncles, 0.4 of peduncular length; ocular acicles subquadrate, each with 3 or 4 small spines on anterior margin.
</paragraph>
<paragraph id="781436B50E5AFFD7A5AAFEAFCD52FE9F" blockId="1.[156,768,198,2077]" pageId="1" pageNumber="40">
Antennular peduncles (
<figureCitation id="E0902A300E5AFFD7A4BAFEAFCDD9FEFF" box="[421,494,263,288]" captionStart="Fig" captionStartId="2.[156,187,1884,1906]" captionTargetBox="[170,1417,204,1862]" captionTargetId="figure-68@2.[170,1417,204,1862]" captionTargetPageId="2" captionText="Fig. 1. Dardanus aspersus (Berthold, 1845), holotype of Pagurus watasei Terao, 1913, female (cl 16.7mm), UMUTZ-Crs-33. A, entire animal, dorsal view; B, shield and cephalic appendages, dorsal view; C, left cheliped, outer view; D, right cheliped, outer view." figureDoi="http://doi.org/10.5281/zenodo.4584843" httpUri="https://zenodo.org/record/4584843/files/figure.png" pageId="1" pageNumber="40">Fig. 1B</figureCitation>
) slightly overreaching distal corneal margins.
</paragraph>
<paragraph id="781436B50E5AFFD7A5AAFEE0CE6CFE3D" blockId="1.[156,768,198,2077]" pageId="1" pageNumber="40">
Antennal peduncles (
<figureCitation id="E0902A300E5AFFD7A490FEE0CDEEFEBE" box="[399,473,328,353]" captionStart="Fig" captionStartId="2.[156,187,1884,1906]" captionTargetBox="[170,1417,204,1862]" captionTargetId="figure-68@2.[170,1417,204,1862]" captionTargetPageId="2" captionText="Fig. 1. Dardanus aspersus (Berthold, 1845), holotype of Pagurus watasei Terao, 1913, female (cl 16.7mm), UMUTZ-Crs-33. A, entire animal, dorsal view; B, shield and cephalic appendages, dorsal view; C, left cheliped, outer view; D, right cheliped, outer view." figureDoi="http://doi.org/10.5281/zenodo.4584843" httpUri="https://zenodo.org/record/4584843/files/figure.png" pageId="1" pageNumber="40">Fig. 1B</figureCitation>
) slender, overreaching bases but not reaching distal margins of corneas; antennal acicle slender, overreaching base of fifth peduncular segment, terminating in small bifid spine, mesial margin with row of small spines partially obscured by stiff setae.
</paragraph>
<paragraph id="781436B50E5AFFD7A5AAFE43CD49FDFB" blockId="1.[156,768,198,2077]" pageId="1" pageNumber="40">Carpus of third maxilliped with 2 tiny spines on distal half of ventral margin.</paragraph>
<paragraph id="781436B50E5AFFD7A5AAFD84CE96FC15" blockId="1.[156,768,198,2077]" pageId="1" pageNumber="40">
Left cheliped (
<figureCitation id="E0902A300E5AFFD7A451FD84CDAAFD9B" box="[334,413,556,580]" captionStart="Fig" captionStartId="2.[156,187,1884,1906]" captionTargetBox="[170,1417,204,1862]" captionTargetId="figure-68@2.[170,1417,204,1862]" captionTargetPageId="2" captionText="Fig. 1. Dardanus aspersus (Berthold, 1845), holotype of Pagurus watasei Terao, 1913, female (cl 16.7mm), UMUTZ-Crs-33. A, entire animal, dorsal view; B, shield and cephalic appendages, dorsal view; C, left cheliped, outer view; D, right cheliped, outer view." figureDoi="http://doi.org/10.5281/zenodo.4584843" httpUri="https://zenodo.org/record/4584843/files/figure.png" pageId="1" pageNumber="40">Fig. 1C</figureCitation>
) massive, appreciably larger than right. Chela broadly subovate in general outline, 1.4 times longer than wide; outer surface convex, covered with numerous short tubercles each bearing sharp corneous spinule and tuft of short, stiff setae; fingers without hiatus, each with row of blunt teeth and terminating in small, plate-like claw. Dactylus slightly longer than palm. Palm with row of small corneous-tipped spines on upper margin and row of blunt spines on lower margin. Carpus with sparse small spines on outer surface; upper margin with row of moderately large, sharp, corneous-tipped spines increasing in size distally. Merus with row of small spines on ventromesial and on ventrolateral margin, ventromesial spines pronounced.
</paragraph>
<paragraph id="781436B50E5AFFD7A5AAFC7BCEEFFA2C" blockId="1.[156,768,198,2077]" pageId="1" pageNumber="40">
Right cheliped (
<figureCitation id="E0902A300E5AFFD7A443FC7BCD9CFC34" box="[348,427,979,1003]" captionStart="Fig" captionStartId="2.[156,187,1884,1906]" captionTargetBox="[170,1417,204,1862]" captionTargetId="figure-68@2.[170,1417,204,1862]" captionTargetPageId="2" captionText="Fig. 1. Dardanus aspersus (Berthold, 1845), holotype of Pagurus watasei Terao, 1913, female (cl 16.7mm), UMUTZ-Crs-33. A, entire animal, dorsal view; B, shield and cephalic appendages, dorsal view; C, left cheliped, outer view; D, right cheliped, outer view." figureDoi="http://doi.org/10.5281/zenodo.4584843" httpUri="https://zenodo.org/record/4584843/files/figure.png" pageId="1" pageNumber="40">Fig. 1D</figureCitation>
) short. Chela elongate subtriangular in general outline, 1.9 times longer than wide; fingers without hiatus, each with row of blunt teeth and terminating in large, plate-like claw. Dactylus about 1.3 times as long as palm; outer surface with small tubercles near upper margin, each tubercle bearing corneous spinule and tuft of stiff setae. Palm slightly convex on outer face, covered with short, transverse ridges each bearing tuft of short, stiff setae, ridges near upper and lower margins and on fixed finger with 1 or 2 corneous spinules each; upper margin with row of sharp corneous-tipped spines; lower margin nearly straight, rounded. Carpus with small corneous-tipped spines on upper side and short, transverse ridges on lower side of outer surface; upper margin with row of moderately large, sharp corneous-tipped spines. Merus also with row of small corneous-tipped spines on ventromesial margin; ventrolateral margin distally with 2 small corneous-tipped spines.
</paragraph>
<paragraph id="781436B50E5AFFD7A5AAFA54C886FD39" blockId="1.[156,768,198,2077]" lastBlockId="1.[820,1433,198,2076]" pageId="1" pageNumber="40">
Ambulatory legs (
<figureCitation id="E0902A300E5AFFD7A471FA54CDEEF9CB" box="[366,473,1532,1556]" captionStart="Fig" captionStartId="3.[156,187,1884,1906]" captionTargetBox="[171,1416,204,1861]" captionTargetId="figure-4@3.[171,1416,204,1861]" captionTargetPageId="3" captionText="Fig. 2. Dardanus aspersus (Berthold, 1845), holotype of Pagurus watasei Terao, 1913, female (cl 16.7mm), UMUTZ-Crs-33. A, left second and third pereopods, lateral view; B, right second and third pereopods, lateral view; C, left fourth pereopod, lateral view; D, sixth pleonal tergite and telson, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4584847" httpUri="https://zenodo.org/record/4584847/files/figure.png" pageId="1" pageNumber="40">Fig. 2A, B</figureCitation>
) relatively short; second pereopods generally similar on right and left sides but third pereopods noticeably dissimilar to each other (dactylus, propodus, and carpus of left third pereopod stouter and with stronger armature than those of right); most spines corneous-tipped and most tubercles bearing slender corneous spinule; tufts of short and long stiff setae present on spines, tubercles, and ridges, particularly numerous on left third pereopod. Dactyli 1.3 (right second pereopod)–1.6 (left third) times longer than propodi, lengths 7.2–7.3 (right and left second), 8.4 (right third), and 4.9 (left third) times respective proximal widths; dorsal margins each with row of small spines; lateral and mesial surfaces flattened, each with shallow longitudinal sulcus on midline, on left third pereopod, lateral surface also with scattered rounded and low tubercles. Propodi 2.8 (right second and third pereopods), 3.0 (left second), and 2.0 (left third) times respective proximal widths, somewhat tapering distally and convex ventrally; dorsal, lateral, and ventral surfaces of right second, left second, and right third pereopods with occasionally granulate, short, transverse ridges (dorsal ridges each with 1 or 2 corneous spinules), but on left third pereopod, dorsal surface with irregular, longitudinal double row of small spines, lateral surface with 4 longitudinal rows of tubercles, ventral margin with row of blunt spines; lateral surface of left third pereopod flattened, but those of right second, left second, and right third slightly convex. Carpi each with dorsal row of small spines, but on right third pereopod, these reduced in size; lateral surfaces each with median row of short transverse ridges, distal ridges denticulate on left third pereopod. Meri unarmed on dorsal margins; lateral surfaces with short, sparse transverse ridges; ventral margins with some tiny spines or protuberances, these pronounced on left third pereopod, but obsolete on right third.
</paragraph>
<paragraph id="781436B50E5AFFD7A651FD47CF4DFC15" blockId="1.[820,1433,198,2076]" pageId="1" pageNumber="40">
Fourth pereopods (
<figureCitation id="E0902A300E5AFFD7A10CFD47C856FCD8" box="[1043,1121,751,775]" captionStart="Fig" captionStartId="3.[156,187,1884,1906]" captionTargetBox="[171,1416,204,1861]" captionTargetId="figure-4@3.[171,1416,204,1861]" captionTargetPageId="3" captionText="Fig. 2. Dardanus aspersus (Berthold, 1845), holotype of Pagurus watasei Terao, 1913, female (cl 16.7mm), UMUTZ-Crs-33. A, left second and third pereopods, lateral view; B, right second and third pereopods, lateral view; C, left fourth pereopod, lateral view; D, sixth pleonal tergite and telson, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4584847" httpUri="https://zenodo.org/record/4584847/files/figure.png" pageId="1" pageNumber="40">Fig. 2C</figureCitation>
) semichelate. Dactyli distinctly longer than palm, each with dorsal row of small corneous-tipped spines; preungual process absent. Propodi each with small, corneous-tipped spine near dorsodistal end; rasp well developed, consisting of numerous small, subovate scales. Carpi each with dorsal row of sharp, corneous-tipped spines.
</paragraph>
<paragraph id="781436B50E5AFFD7A651FC7BC8F2FBD4" blockId="1.[820,1433,198,2076]" pageId="1" pageNumber="40">Sixth thoracic sternite with anterior lobe narrow, rounded, and with tuft of stiff setae anteriorly.</paragraph>
<paragraph id="781436B50E5AFFD7A651FBBCC86DFA6D" blockId="1.[820,1433,198,2076]" pageId="1" pageNumber="40">
Second to fifth pleomeres (
<figureCitation id="E0902A300E5AFFD7A16CFBBCC8F2FBF3" box="[1139,1221,1044,1068]" captionStart="Fig" captionStartId="2.[156,187,1884,1906]" captionTargetBox="[170,1417,204,1862]" captionTargetId="figure-68@2.[170,1417,204,1862]" captionTargetPageId="2" captionText="Fig. 1. Dardanus aspersus (Berthold, 1845), holotype of Pagurus watasei Terao, 1913, female (cl 16.7mm), UMUTZ-Crs-33. A, entire animal, dorsal view; B, shield and cephalic appendages, dorsal view; C, left cheliped, outer view; D, right cheliped, outer view." figureDoi="http://doi.org/10.5281/zenodo.4584843" httpUri="https://zenodo.org/record/4584843/files/figure.png" pageId="1" pageNumber="40">Fig. 1A</figureCitation>
) with unpaired left pleopods; second to fourth pleopods well developed, triramous; fifth pleopod short, biramous, with endopod greatly reduced in size; short, subtriangular plate present between fourth and fifth pleopods. Sixth pleonal tergite (
<figureCitation id="E0902A300E5AFFD7A007FB3EC952FB71" box="[1304,1381,1174,1198]" captionStart="Fig" captionStartId="3.[156,187,1884,1906]" captionTargetBox="[171,1416,204,1861]" captionTargetId="figure-4@3.[171,1416,204,1861]" captionTargetPageId="3" captionText="Fig. 2. Dardanus aspersus (Berthold, 1845), holotype of Pagurus watasei Terao, 1913, female (cl 16.7mm), UMUTZ-Crs-33. A, left second and third pereopods, lateral view; B, right second and third pereopods, lateral view; C, left fourth pereopod, lateral view; D, sixth pleonal tergite and telson, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4584847" httpUri="https://zenodo.org/record/4584847/files/figure.png" pageId="1" pageNumber="40">Fig. 2D</figureCitation>
) unarmed, widened posteriorly, approximately as long as wide, divided into 2 parts by distinct transverse groove crossing at nearly half length; lateral indentations narrow; posterior part with deep median groove. Telson (
<figureCitation id="E0902A300E5AFFD7A1CCFAB0C913FAEF" box="[1235,1316,1304,1328]" captionStart="Fig" captionStartId="3.[156,187,1884,1906]" captionTargetBox="[171,1416,204,1861]" captionTargetId="figure-4@3.[171,1416,204,1861]" captionTargetPageId="3" captionText="Fig. 2. Dardanus aspersus (Berthold, 1845), holotype of Pagurus watasei Terao, 1913, female (cl 16.7mm), UMUTZ-Crs-33. A, left second and third pereopods, lateral view; B, right second and third pereopods, lateral view; C, left fourth pereopod, lateral view; D, sixth pleonal tergite and telson, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4584847" httpUri="https://zenodo.org/record/4584847/files/figure.png" pageId="1" pageNumber="40">Fig. 2D</figureCitation>
) with posterior lobes broadly subtriangular, unequal (left lobe larger), separated by narrow median cleft; terminal margins strongly oblique, each with 6 or 7 small corneous-tipped spines extending onto lateral margins.
</paragraph>
<paragraph id="781436B50E5AFFD7A651FA12CF54F9CB" blockId="1.[820,1433,198,2076]" pageId="1" pageNumber="40">
<emphasis id="4ADFEAA70E5AFFD7A651FA12CFF5FA0D" bold="true" box="[846,962,1466,1490]" pageId="1" pageNumber="40">Coloration</emphasis>
. See 
<bibRefCitation id="1C3A4B440E5AFFD7A111FA13C8FEFA0D" author="Miyake, S." box="[1038,1225,1466,1491]" pageId="1" pageNumber="40" refId="ref4268" refString="Miyake, S. 1978. The Crustacean Anomura of Sagami Bay. Biological Laboratory, Imperial Household, Tokyo, 200 (English) + 161 (Japanese) pp., 4 pls." type="book" year="1978">Miyake (1978: 65</bibRefCitation>
, pl. 1, fig. 2; 1998: pl. 37, fig. 2) and 
<bibRefCitation id="1C3A4B440E5AFFD7A6F4FA73C93FFA2C" author="McLaughlin, P. A. &amp; Rahayu, D. L. &amp; Komai, T. &amp; Chan, T. - Y." box="[1003,1288,1499,1523]" pageId="1" pageNumber="40" refId="ref4142" refString="McLaughlin, P. A., Rahayu, D. L., Komai, T. and Chan, T. - Y. 2007. A Catalog of the Hermit Crabs (Paguroidea) of Taiwan. National Taiwan Ocean University, Keelung, 367 pp." type="book" year="2007">
McLaughlin 
<emphasis id="4ADFEAA70E5AFFD7A171FA74C896FA2C" box="[1134,1185,1499,1523]" italics="true" pageId="1" pageNumber="40">et al.</emphasis>
(2007: 97
</bibRefCitation>
, unnumbered fig.).
</paragraph>
</subSubSection>
<subSubSection id="30B1653E0E5AFFD7A651F9B4CF43F927" pageId="1" pageNumber="40" type="distribution">
<paragraph id="781436B50E5AFFD7A651F9B4C8A3F949" blockId="1.[820,1433,198,2076]" pageId="1" pageNumber="40">
<emphasis id="4ADFEAA70E5AFFD7A651F9B4CFE4F9EB" bold="true" box="[846,979,1564,1588]" pageId="1" pageNumber="40">Distribution</emphasis>
. Boso Peninsula and 
<collectingRegion id="BA6FF8570E5AFFD7A1D3F9B5C92BF9EA" box="[1228,1308,1565,1589]" country="Japan" name="Toyama" pageId="1" pageNumber="40">Toyama</collectingRegion>
Bay southward on both coasts of Honshu and Kyushu in 
<collectingCountry id="00BC76250E5AFFD7A036F995C953F98B" box="[1321,1380,1597,1620]" name="Japan" pageId="1" pageNumber="40">Japan</collectingCountry>
, 
<collectingCountry id="00BC76250E5AFFD7A06FF995CF57F9AA" name="Taiwan" pageId="1" pageNumber="40">Taiwan</collectingCountry>
, East and South 
<collectingCountry id="00BC76250E5AFFD7A104F9F5C86EF9AA" box="[1051,1113,1629,1653]" name="China" pageId="1" pageNumber="40">China</collectingCountry>
Seas, Andaman Sea; 
<quantity id="BF539B500E5AFFD7A05DF9F5C9AFF9AA" box="[1346,1432,1629,1653]" metricMagnitude="1" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="1" pageNumber="40" unit="m" value="35.0" valueMax="50.0" valueMin="20.0">20–50 m</quantity>
(
<bibRefCitation id="1C3A4B440E5AFFD7A624F9D6CFC6F949" author="McLaughlin, P. A." box="[827,1009,1662,1686]" pageId="1" pageNumber="40" pagination="385 - 460" refId="ref3930" refString="McLaughlin, P. A. 2002. A review of the hermit crab (Decapoda: Anomura: Paguridea) fauna of southern Thailand, with particular emphasis on the Andaman Sea, and descriptions of three new species. In: Bruce, N., Berggren, M. and Bussawarit, S. (Eds) Proceedings of the International Workshop on the Crustacea in the Andaman Sea, Phuket Marine Biological Center 29 November- 20 December, 1998. Phuket Marine Biological Center, Special Publication 23: 385 - 460." type="journal article" year="2002">McLaughlin 2002</bibRefCitation>
; 
<bibRefCitation id="1C3A4B440E5AFFD7A6E4F9D6C8BEF949" author="Asakura, A." box="[1019,1161,1662,1686]" pageId="1" pageNumber="40" pagination="23 - 103" refId="ref3428" refString="Asakura, A. 2006. Shallow water hermit crabs of the families Pylochelidae, Diogenidae and Paguridae (Crustacea: Decapoda: Anomura) from the Sea of Japan, with a description of a new species of Diogenes. Bulletin of the Toyama Science Museum 29: 23 - 103." type="journal article" year="2006">Asakura 2006</bibRefCitation>
).
</paragraph>
<paragraph id="781436B50E5AFFD7A651F936CF43F927" blockId="1.[820,1433,198,2076]" pageId="1" pageNumber="40">
<emphasis id="4ADFEAA70E5AFFD7A651F936CFA9F969" bold="true" box="[846,926,1694,1718]" pageId="1" pageNumber="40">Habitat</emphasis>
. Gastropod shells, often encrusted with sea anemones, on sandy-mud substrates (
<bibRefCitation id="1C3A4B440E5AFFD7A1CCF917CF5EF927" author="McLaughlin, P. A. &amp; Rahayu, D. L. &amp; Komai, T. &amp; Chan, T. - Y." pageId="1" pageNumber="40" refId="ref4142" refString="McLaughlin, P. A., Rahayu, D. L., Komai, T. and Chan, T. - Y. 2007. A Catalog of the Hermit Crabs (Paguroidea) of Taiwan. National Taiwan Ocean University, Keelung, 367 pp." type="book" year="2007">
McLaughlin 
<emphasis id="4ADFEAA70E5AFFD7A040F968C9AEF908" box="[1375,1433,1727,1751]" italics="true" pageId="1" pageNumber="40">et al.</emphasis>
2007
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="30B1653E0E5AFFD2A651F8A8C838FB30" lastPageId="4" lastPageNumber="43" pageId="1" pageNumber="40" type="discussion">
<paragraph id="781436B50E5AFFD2A651F8A8CD20FE5E" blockId="1.[820,1433,198,2076]" lastBlockId="4.[156,771,198,2077]" lastPageId="4" lastPageNumber="43" pageId="1" pageNumber="40">
<emphasis id="4ADFEAA70E5AFFD7A651F8A8CF9CF8C7" bold="true" box="[846,939,1792,1816]" pageId="1" pageNumber="40">Remarks</emphasis>
. The 
<typeStatus id="A71088170E5AFFD7A6E6F8A8C865F8C7" box="[1017,1106,1792,1816]" pageId="1" pageNumber="40" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="BFAB4D360E5AFFD7A166F8A9C92AF8C7" authorityName="Terao" authorityYear="1913" box="[1145,1309,1793,1816]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="40" phylum="Arthropoda" rank="species" species="watasei">
<emphasis id="4ADFEAA70E5AFFD7A166F8A9C92AF8C7" box="[1145,1309,1793,1816]" italics="true" pageId="1" pageNumber="40">Pagurus watasei</emphasis>
</taxonomicName>
agrees well with the previous descriptions of 
<taxonomicName id="BFAB4D360E5AFFD7A181F888C969F8E7" baseAuthorityName="Berthold" baseAuthorityYear="1845" box="[1182,1374,1824,1848]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="40" phylum="Arthropoda" rank="species" species="aspersus">
<emphasis id="4ADFEAA70E5AFFD7A181F888C969F8E7" box="[1182,1374,1824,1848]" italics="true" pageId="1" pageNumber="40">Dardanus aspersus</emphasis>
</taxonomicName>
(
<emphasis id="4ADFEAA70E5AFFD7A06EF889C9A4F8E7" box="[1393,1427,1825,1848]" italics="true" pageId="1" pageNumber="40">e.g.</emphasis>
, 
<bibRefCitation id="1C3A4B440E5AFFD7A62BF8E9CF8AF886" author="Miyake, S." box="[820,957,1857,1881]" pageId="1" pageNumber="40" refId="ref4268" refString="Miyake, S. 1978. The Crustacean Anomura of Sagami Bay. Biological Laboratory, Imperial Household, Tokyo, 200 (English) + 161 (Japanese) pp., 4 pls." type="book" year="1978">Miyake 1978</bibRefCitation>
, 
<bibRefCitation id="1C3A4B440E5AFFD7A6D3F8E9CFC8F886" author="Miyake, S." box="[972,1023,1857,1881]" pageId="1" pageNumber="40" refId="ref4345" refString="Miyake, S. 1998. Japanese Crustacean Decapods and Stomatopods in Color. I. Macrura, Anomura, and Stomatopoda. Third printing. Hoikusha Publishing Company, Osaka, vii + 261 pp. [In Japanese]" type="book" year="1998">1998</bibRefCitation>
; 
<bibRefCitation id="1C3A4B440E5AFFD7A112F8E9C930F886" author="McLaughlin, P. A. &amp; Rahayu, D. L. &amp; Komai, T. &amp; Chan, T. - Y." box="[1037,1287,1857,1881]" pageId="1" pageNumber="40" refId="ref4142" refString="McLaughlin, P. A., Rahayu, D. L., Komai, T. and Chan, T. - Y. 2007. A Catalog of the Hermit Crabs (Paguroidea) of Taiwan. National Taiwan Ocean University, Keelung, 367 pp." type="book" year="2007">
McLaughlin 
<emphasis id="4ADFEAA70E5AFFD7A18BF8EAC8FDF886" box="[1172,1226,1857,1881]" italics="true" pageId="1" pageNumber="40">et al.</emphasis>
2007
</bibRefCitation>
) in every diagnostic aspect, particularly in the stout ocular peduncles with dilated corneas, the markedly unequal chelipeds with the left chela covered with numerous tubercles each bearing a corneous spinule, and the flattened lateral surfaces of the dactylus and propodus of the left third pereopod. 
<bibRefCitation id="1C3A4B440E5AFFD7A041F84CCF41F7C3" author="Terao, A." pageId="1" pageNumber="40" pagination="355 - 391" refId="ref4418" refString="Terao, A. 1913. A catalogue of hermit-crabs found in Japan (Paguridea excluding Lithodidae), with descriptions of four new species. Annotationes Zoologicae Japonenses 8: 355 - 391." type="journal article" year="1913">Terao (1913)</bibRefCitation>
described the coloration of his material of 
<taxonomicName id="BFAB4D360E5AFFD4A058F7ADCCD1F804" authorityName="Terao" authorityYear="1913" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" lastPageId="2" lastPageNumber="41" order="Decapoda" pageId="1" pageNumber="40" phylum="Arthropoda" rank="species" species="watasei">
<emphasis id="4ADFEAA70E5AFFD4A058F7ADCCD1F804" italics="true" lastPageId="2" lastPageNumber="41" pageId="1" pageNumber="40">Pagurus watasei</emphasis>
</taxonomicName>
preserved in alcohol and formalin, and this is also consistent with that reported for 
<taxonomicName id="BFAB4D360E59FFD4A71BF84CCEF2F823" baseAuthorityName="Berthold" baseAuthorityYear="1845" box="[516,709,2020,2044]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="species" species="aspersus">
<emphasis id="4ADFEAA70E59FFD4A71BF84CCEF2F823" box="[516,709,2020,2044]" italics="true" pageId="2" pageNumber="41">Dardanus aspersus</emphasis>
</taxonomicName>
(
<emphasis id="4ADFEAA70E59FFD4A7C6F84DCECDF823" box="[729,762,2021,2044]" italics="true" pageId="2" pageNumber="41">e.g.</emphasis>
, 
<bibRefCitation id="1C3A4B440E59FFD4A583F7ADCD60F7C2" author="Miyake, S." box="[156,343,2053,2077]" pageId="2" pageNumber="41" refId="ref4345" refString="Miyake, S. 1998. Japanese Crustacean Decapods and Stomatopods in Color. I. Macrura, Anomura, and Stomatopoda. Third printing. Hoikusha Publishing Company, Osaka, vii + 261 pp. [In Japanese]" type="book" year="1998">Miyake, 1998: 111</bibRefCitation>
, pl. 38, fig. 2; 
<bibRefCitation id="1C3A4B440E59FFD4A4F7F7ADCECCF7C2" author="McLaughlin, P. A. &amp; Rahayu, D. L. &amp; Komai, T. &amp; Chan, T. - Y." box="[488,763,2053,2077]" pageId="2" pageNumber="41" refId="ref4142" refString="McLaughlin, P. A., Rahayu, D. L., Komai, T. and Chan, T. - Y. 2007. A Catalog of the Hermit Crabs (Paguroidea) of Taiwan. National Taiwan Ocean University, Keelung, 367 pp." type="book" year="2007">
McLaughlin 
<emphasis id="4ADFEAA70E59FFD4A774F7AECEAAF7C2" box="[619,669,2053,2077]" italics="true" pageId="2" pageNumber="41">et al.</emphasis>
2007: 96
</bibRefCitation>
, unnumbered fig.) in the presence of numerous red spots on the carapace, pleonal tergites, and pereopods, darker coloration on the dactyli of the ambulatory legs than on their proximal segments, and a broad purple band on the median part of each ocular peduncle; the characteristic color pattern of the ocular peduncle is still preserved in the 
<typeStatus id="A71088170E5FFFD2A796FEAFCED5FEC0" box="[649,738,263,287]" pageId="4" pageNumber="43" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="BFAB4D360E5FFFD2A583FE81CD09FE9F" authorityName="Terao" authorityYear="1913" box="[156,318,297,320]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="watasei">
<emphasis id="4ADFEAA70E5FFFD2A583FE81CD09FE9F" box="[156,318,297,320]" italics="true" pageId="4" pageNumber="43">Pagurus watasei</emphasis>
</taxonomicName>
. Thus, this taxon is considered to be identical with 
<taxonomicName id="BFAB4D360E5FFFD2A5EEFEE0CD9AFEBF" baseAuthorityName="Berthold" baseAuthorityYear="1845" box="[241,429,328,352]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="aspersus">
<emphasis id="4ADFEAA70E5FFFD2A5EEFEE0CD9AFEBF" box="[241,429,328,352]" italics="true" pageId="4" pageNumber="43">Dardanus aspersus</emphasis>
</taxonomicName>
and is placed in the synonymy of that species.
</paragraph>
<caption id="2CD4663D0E59FFD4A583F8F4C8DDF851" ID-DOI="http://doi.org/10.5281/zenodo.4584843" ID-Zenodo-Dep="4584843" httpUri="https://zenodo.org/record/4584843/files/figure.png" pageId="2" pageNumber="41" startId="2.[156,187,1884,1906]" targetBox="[170,1417,204,1862]" targetPageId="2">
<paragraph id="781436B50E59FFD4A583F8F4C8DDF851" blockId="2.[156,1431,1884,1934]" pageId="2" pageNumber="41">
Fig. 1. 
<taxonomicName id="BFAB4D360E59FFD4A5EFF8F4CE00F8AD" authority="(Berthold, 1845)" baseAuthorityName="Berthold" baseAuthorityYear="1845" box="[240,567,1884,1906]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="species" species="aspersus">
<emphasis id="4ADFEAA70E59FFD4A5EFF8F4CDAFF8AD" box="[240,408,1884,1906]" italics="true" pageId="2" pageNumber="41">Dardanus aspersus</emphasis>
(
<bibRefCitation id="1C3A4B440E59FFD4A4BAF8F4CE07F8AD" author="Berthold, A. A." box="[421,560,1884,1906]" pageId="2" pageNumber="41" pagination="37 - 48" refId="ref3481" refString="Berthold, A. A. 1845. Ueber verschiedene neue oder seltene Reptilien aus Neu-Granada und Crustaceen aus China. Nachrichten von der Koniglichen Gesellschaft der Wissenschaften zu Gottingen 1845: 37 - 48." type="journal article" year="1845">Berthold, 1845</bibRefCitation>
)
</taxonomicName>
, holotype of 
<taxonomicName id="BFAB4D360E59FFD4A7B0F8F5CF86F8AE" authority="Terao, 1913" authorityName="Terao" authorityYear="1913" box="[687,945,1884,1906]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="species" species="watasei">
<emphasis id="4ADFEAA70E59FFD4A7B0F8F5CF76F8AD" box="[687,833,1885,1906]" italics="true" pageId="2" pageNumber="41">Pagurus watasei</emphasis>
<bibRefCitation id="1C3A4B440E59FFD4A659F8F5CF86F8AE" author="Terao, A." box="[838,945,1884,1906]" pageId="2" pageNumber="41" pagination="355 - 391" refId="ref4418" refString="Terao, A. 1913. A catalogue of hermit-crabs found in Japan (Paguridea excluding Lithodidae), with descriptions of four new species. Annotationes Zoologicae Japonenses 8: 355 - 391." type="journal article" year="1913">Terao, 1913</bibRefCitation>
</taxonomicName>
, female (cl 16.7mm), UMUTZ-Crs-33. A, entire animal, dorsal view; B, shield and cephalic appendages, dorsal view; C, left cheliped, outer view; D, right cheliped, outer view.
</paragraph>
</caption>
<caption id="2CD4663D0E58FFD5A583F8F4CDA2F875" ID-DOI="http://doi.org/10.5281/zenodo.4584847" ID-Zenodo-Dep="4584847" httpUri="https://zenodo.org/record/4584847/files/figure.png" pageId="3" pageNumber="42" startId="3.[156,187,1884,1906]" targetBox="[171,1416,204,1861]" targetPageId="3">
<paragraph id="781436B50E58FFD5A583F8F4CDA2F875" blockId="3.[156,1432,1883,1962]" pageId="3" pageNumber="42">
Fig. 2. 
<taxonomicName id="BFAB4D360E58FFD5A5EFF8F4CE00F8AE" authority="(Berthold, 1845)" baseAuthorityName="Berthold" baseAuthorityYear="1845" box="[240,567,1883,1906]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="species" species="aspersus">
<emphasis id="4ADFEAA70E58FFD5A5EFF8F4CDAFF8AE" box="[240,408,1884,1905]" italics="true" pageId="3" pageNumber="42">Dardanus aspersus</emphasis>
(
<bibRefCitation id="1C3A4B440E58FFD5A4BBF8F4CE18F8AE" author="Berthold, A. A." box="[420,559,1883,1906]" pageId="3" pageNumber="42" pagination="37 - 48" refId="ref3481" refString="Berthold, A. A. 1845. Ueber verschiedene neue oder seltene Reptilien aus Neu-Granada und Crustaceen aus China. Nachrichten von der Koniglichen Gesellschaft der Wissenschaften zu Gottingen 1845: 37 - 48." type="journal article" year="1845">Berthold, 1845</bibRefCitation>
)
</taxonomicName>
, holotype of 
<taxonomicName id="BFAB4D360E58FFD5A7B1F8F4CF87F8AE" authority="Terao, 1913" authorityName="Terao" authorityYear="1913" box="[686,944,1884,1905]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="42" phylum="Arthropoda" rank="species" species="watasei">
<emphasis id="4ADFEAA70E58FFD5A7B1F8F4CF08F8AE" box="[686,831,1884,1905]" italics="true" pageId="3" pageNumber="42">Pagurus watasei</emphasis>
<bibRefCitation id="1C3A4B440E58FFD5A65AF8F4CF87F8AE" author="Terao, A." box="[837,944,1884,1905]" pageId="3" pageNumber="42" pagination="355 - 391" refId="ref4418" refString="Terao, A. 1913. A catalogue of hermit-crabs found in Japan (Paguridea excluding Lithodidae), with descriptions of four new species. Annotationes Zoologicae Japonenses 8: 355 - 391." type="journal article" year="1913">Terao, 1913</bibRefCitation>
</taxonomicName>
, female (cl 16.7 mm), UMUTZ-Crs-33. A, left second and third pereopods, lateral view; B, right second and third pereopods, lateral view; C, left fourth pereopod, lateral view; D, sixth pleonal tergite and telson, dorsal view.
</paragraph>
</caption>
<paragraph id="781436B50E5FFFD2A5AAFE22CD72FD19" blockId="4.[156,771,198,2077]" pageId="4" pageNumber="43">
In addition to the 
<typeStatus id="A71088170E5FFFD2A472FE21CDF1FE7E" box="[365,454,393,417]" pageId="4" pageNumber="43" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="BFAB4D360E5FFFD2A4F8FE22CE70FE7E" authorityName="Terao" authorityYear="1913" box="[487,583,394,417]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="watasei">
<emphasis id="4ADFEAA70E5FFFD2A4F8FE22CE70FE7E" box="[487,583,394,417]" italics="true" pageId="4" pageNumber="43">P. watasei</emphasis>
</taxonomicName>
, three other specimens identified with this taxon by Terao were examined, although one of them (UMUTZ-Crs-c112) was not included in 
<bibRefCitation id="1C3A4B440E5FFFD2A5A9FE43CD75FDDC" author="Terao, A." box="[182,322,491,515]" pageId="4" pageNumber="43" pagination="355 - 391" refId="ref4418" refString="Terao, A. 1913. A catalogue of hermit-crabs found in Japan (Paguridea excluding Lithodidae), with descriptions of four new species. Annotationes Zoologicae Japonenses 8: 355 - 391." type="journal article" year="1913">Terao’s (1913)</bibRefCitation>
description. One male (UMUTZ-Crs-c195) is now in poor condition (dry and partially fragmental), but the other two specimens are still in good condition and preserved in ethanol. As in the 
<typeStatus id="A71088170E5FFFD2A4DBFDE5CE28FDBA" box="[452,543,589,613]" pageId="4" pageNumber="43" type="holotype">holotype</typeStatus>
, these two specimens display the diagnostic characters of 
<taxonomicName id="BFAB4D360E5FFFD2A70BFDC5CEE4FD5A" baseAuthorityName="Berthold" baseAuthorityYear="1845" box="[532,723,621,645]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="aspersus">
<emphasis id="4ADFEAA70E5FFFD2A70BFDC5CEE4FD5A" box="[532,723,621,645]" italics="true" pageId="4" pageNumber="43">Dardanus aspersus</emphasis>
</taxonomicName>
, including the broad purple band on the median part of each ocular peduncle.
</paragraph>
<paragraph id="781436B50E5FFFD2A5AAFD67CCEBF845" blockId="4.[156,771,198,2077]" pageId="4" pageNumber="43">
De Haan (1849) reported 
<taxonomicName id="BFAB4D360E5FFFD2A4C4FD78CCFAFCD8" authority="Fabricius, 1775" authorityName="Fabricius" authorityYear="1775" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="diogenes">
<emphasis id="4ADFEAA70E5FFFD2A4C4FD78CEB8FD38" box="[475,655,719,743]" italics="true" pageId="4" pageNumber="43">Pagurus diogenes</emphasis>
Fabricius, 1775
</taxonomicName>
[as Fabricius, 1798; = 
<taxonomicName id="BFAB4D360E5FFFD2A4A2FD47CCE0FCF7" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Malacostraca" family="Diogenidae" genus="Petrochirus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="diogenes">
<emphasis id="4ADFEAA70E5FFFD2A4A2FD47CEBDFCD8" box="[445,650,751,775]" italics="true" pageId="4" pageNumber="43">Petrochirus diogenes</emphasis>
(Linnaeus, 1758)
</taxonomicName>
; cf. 
<bibRefCitation id="1C3A4B440E5FFFD2A412FCB8CE44FCF7" author="McLaughlin, P. A. &amp; Holthuis, L. B." box="[269,627,784,808]" pageId="4" pageNumber="43" pagination="257 - 273" refId="ref4027" refString="McLaughlin, P. A. and Holthuis, L. B. 2001. In pursuit of J. F. W. Herbst's species of Diogenes (Anomura: Paguridea: Diogenidae). Journal of Crustacean Biology 21: 257 - 273." type="journal article" year="2001">McLaughlin and Holthuis (2001)</bibRefCitation>
] from 
<collectingCountry id="00BC76250E5FFFD2A7D8FCB8CEC8FCF8" box="[711,767,784,807]" name="Japan" pageId="4" pageNumber="43">Japan</collectingCountry>
and included 
<taxonomicName id="BFAB4D360E5FFFD2A437FC99CDE4FC97" authorityName="Berthold" authorityYear="1845" box="[296,467,817,840]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="aspersus">
<emphasis id="4ADFEAA70E5FFFD2A437FC99CDE4FC97" box="[296,467,817,840]" italics="true" pageId="4" pageNumber="43">Pagurus aspersus</emphasis>
</taxonomicName>
in the synonymy of that species. As was discussed at some length by 
<bibRefCitation id="1C3A4B440E5FFFD2A74BFCF9CD72FC55" author="McLaughlin, P. A. &amp; Holthuis, L. B." pageId="4" pageNumber="43" pagination="257 - 273" refId="ref4027" refString="McLaughlin, P. A. and Holthuis, L. B. 2001. In pursuit of J. F. W. Herbst's species of Diogenes (Anomura: Paguridea: Diogenidae). Journal of Crustacean Biology 21: 257 - 273." type="journal article" year="2001">McLaughlin and Holthuis (2001)</bibRefCitation>
, De Haan’s (1849) misinterpretation led numerous subsequent authors to identify 
<taxonomicName id="BFAB4D360E5FFFD2A749FC3ACC8DFC15" baseAuthorityName="Berthold" baseAuthorityYear="1845" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="aspersus">
<emphasis id="4ADFEAA70E5FFFD2A749FC3ACC8DFC15" italics="true" pageId="4" pageNumber="43">Dardanus aspersus</emphasis>
</taxonomicName>
with 
<taxonomicName id="BFAB4D360E5FFFD2A5E7FC1BCD92FC15" authorityName="Fabricius" authorityYear="1775" box="[248,421,946,970]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="diogenes">
<emphasis id="4ADFEAA70E5FFFD2A5E7FC1BCD92FC15" box="[248,421,946,970]" italics="true" pageId="4" pageNumber="43">Pagurus diogenes</emphasis>
</taxonomicName>
or 
<taxonomicName id="BFAB4D360E5FFFD2A4D4FC1ACEBDFC15" authorityName="Gee" authorityYear="1935" box="[459,650,946,970]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="diogenes">
<emphasis id="4ADFEAA70E5FFFD2A4D4FC1ACEBDFC15" box="[459,650,946,970]" italics="true" pageId="4" pageNumber="43">Dardanus diogenes</emphasis>
</taxonomicName>
. De Haan’s material is unquestionably referred to 
<taxonomicName id="BFAB4D360E5FFFD2A739FC7BCED4FC34" baseAuthorityName="Berthold" baseAuthorityYear="1845" box="[550,739,979,1003]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="aspersus">
<emphasis id="4ADFEAA70E5FFFD2A739FC7BCED4FC34" box="[550,739,979,1003]" italics="true" pageId="4" pageNumber="43">Dardanus aspersus</emphasis>
</taxonomicName>
as shown by 
<bibRefCitation id="1C3A4B440E5FFFD2A415FC5CCE69FBD4" author="Yamaguchi, T. &amp; Baba, K." box="[266,606,1011,1036]" pageId="4" pageNumber="43" pagination="145 - 570" refId="ref4576" refString="Yamaguchi, T. and Baba, K. 1993. Crustacean specimens collected in Japan by Ph. F. von Siebold and H. Burger and held by the Nationaal Natuurhistorisch Museum in Leiden and other museums. Pp. 145 - 570. In: Yamaguchi, T. (Ed.) Ph. F. Von Siebold and Natural History of Japan. Crustacea. Carcinological Society of Japan, Tokyo. [In English and Japanese]" type="book chapter" year="1993">
<collectingRegion id="BA6FF8570E5FFFD2A415FC5CCD4DFBD3" box="[266,378,1012,1036]" country="Japan" name="Yamaguchi" pageId="4" pageNumber="43">Yamaguchi</collectingRegion>
and Baba (1993: 275
</bibRefCitation>
, fig. 74). 
<bibRefCitation id="1C3A4B440E5FFFD2A7D9FC5CCCE9FBF3" author="Terao, A." pageId="4" pageNumber="43" pagination="355 - 391" refId="ref4418" refString="Terao, A. 1913. A catalogue of hermit-crabs found in Japan (Paguridea excluding Lithodidae), with descriptions of four new species. Annotationes Zoologicae Japonenses 8: 355 - 391." type="journal article" year="1913">Terao (1913)</bibRefCitation>
listed 
<taxonomicName id="BFAB4D360E5FFFD2A439FBBDCDE4FBF3" authorityName="Fabricius" authorityYear="1775" box="[294,467,1044,1068]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="diogenes">
<emphasis id="4ADFEAA70E5FFFD2A439FBBDCDE4FBF3" box="[294,467,1044,1068]" italics="true" pageId="4" pageNumber="43">Pagurus diogenes</emphasis>
</taxonomicName>
in the catalogue of Japanese hermit crabs, although there was no material referred to that taxon available for his examination at that time, and he remarked that 
<taxonomicName id="BFAB4D360E5FFFD2A428FBDECDADFB52" authorityName="Terao" authorityYear="1913" box="[311,410,1142,1165]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="watasei">
<emphasis id="4ADFEAA70E5FFFD2A428FBDECDADFB52" box="[311,410,1142,1165]" italics="true" pageId="4" pageNumber="43">P. watasei</emphasis>
</taxonomicName>
was closely similar to 
<taxonomicName id="BFAB4D360E5FFFD2A793FBDECECEFB51" authorityName="Fabricius" authorityYear="1775" box="[652,761,1142,1166]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="diogenes">
<emphasis id="4ADFEAA70E5FFFD2A793FBDECECEFB51" box="[652,761,1142,1166]" italics="true" pageId="4" pageNumber="43">P. diogenes</emphasis>
</taxonomicName>
, but distinguishable by the relatively longer left chela. 
<bibRefCitation id="1C3A4B440E5FFFD2A7D9FB3FCCEFFB10" author="Terao, A." pageId="4" pageNumber="43" pagination="54 - 62" refId="ref4453" refString="Terao, A. 1932. Crustacea. Pls. 54 - 62. In: Ishikawa, C., Okamura, K., Tanaka, S., Maruyama, H., Terao, A., Higure, T., and Senou, H. (Eds.) Illustrations of Japanese Aquatic Plants and Animals, Volume 2. Fisheries Society of Japan, Tokyo. [In Japanese and English]" type="book chapter" year="1932">Terao (1932</bibRefCitation>
: pl. 57, fig. 2) subsequently provided a color illustration of 
<taxonomicName id="BFAB4D360E5FFFD2A5F4FB70CD6EFB30" authorityName="Fabricius" authorityYear="1775" box="[235,345,1239,1263]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="diogenes">
<emphasis id="4ADFEAA70E5FFFD2A5F4FB70CD6EFB30" box="[235,345,1239,1263]" italics="true" pageId="4" pageNumber="43">P. diogenes</emphasis>
</taxonomicName>
with a short description, both of which agree well with 
<taxonomicName id="BFAB4D360E5FFFD2A426FB50CDC2FACF" baseAuthorityName="Berthold" baseAuthorityYear="1845" box="[313,501,1272,1296]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="aspersus">
<emphasis id="4ADFEAA70E5FFFD2A426FB50CDC2FACF" box="[313,501,1272,1296]" italics="true" pageId="4" pageNumber="43">Dardanus aspersus</emphasis>
</taxonomicName>
, furthermore, the illustration shows the left chela as being much larger than that of the 
<typeStatus id="A71088170E5FFFD2A5DAFA91CD29FA8E" box="[197,286,1337,1361]" pageId="4" pageNumber="43" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="BFAB4D360E5FFFD2A45BFA92CD9FFA8E" authorityName="Terao" authorityYear="1913" box="[324,424,1338,1361]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="watasei">
<emphasis id="4ADFEAA70E5FFFD2A45BFA92CD9FFA8E" box="[324,424,1338,1361]" italics="true" pageId="4" pageNumber="43">P. watasei</emphasis>
</taxonomicName>
. In the second description of 
<taxonomicName id="BFAB4D360E5FFFD2A7F7FA92CDACFAAE" authority=", Berthold (1846" authorityName=", Berthold" authorityYear="1846" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="aspersus">
<emphasis id="4ADFEAA70E5FFFD2A7F7FA92CCD8FAAE" italics="true" pageId="4" pageNumber="43">D. aspersus</emphasis>
, 
<bibRefCitation id="1C3A4B440E5FFFD2A5E2FAF1CDACFAAE" author="Berthold, A. A." box="[253,411,1369,1393]" pageId="4" pageNumber="43" pagination="3 - 32" refId="ref3517" refString="Berthold, A. A. 1846. Uber verschiedene neue oder seltene Reptilien aus Neu-Granada und Crustaceen aus China. Abhandlungen der Koniglichen Gesellschaft der Wissenschaften zu Gottingen (Physicalischen Classe) 3: 3 - 32, pls 1 - 3." type="journal article" year="1846">Berthold (1846</bibRefCitation>
</taxonomicName>
: pl. 2, fig. 1) had also given an illustration showing a similarly proportioned large left chela. These facts suggest that 
<bibRefCitation id="1C3A4B440E5FFFD2A4B7FA33CE07FA6C" author="Terao, A." box="[424,560,1435,1459]" pageId="4" pageNumber="43" pagination="355 - 391" refId="ref4418" refString="Terao, A. 1913. A catalogue of hermit-crabs found in Japan (Paguridea excluding Lithodidae), with descriptions of four new species. Annotationes Zoologicae Japonenses 8: 355 - 391." type="journal article" year="1913">Terao (1913)</bibRefCitation>
misinterpreted the comparative sizes of the left chelae of 
<taxonomicName id="BFAB4D360E5FFFD2A705FA14CE8BFA0C" authorityName="Terao" authorityYear="1913" box="[538,700,1468,1491]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="watasei">
<emphasis id="4ADFEAA70E5FFFD2A705FA14CE8BFA0C" box="[538,700,1468,1491]" italics="true" pageId="4" pageNumber="43">Pagurus watasei</emphasis>
</taxonomicName>
and 
<taxonomicName id="BFAB4D360E5FFFD2A7F0FA14CCD8FA2C" authorityName=", Berthold" authorityYear="1846" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="aspersus">
<emphasis id="4ADFEAA70E5FFFD2A7F0FA14CCD8FA2C" italics="true" pageId="4" pageNumber="43">P. aspersus</emphasis>
</taxonomicName>
(as 
<taxonomicName id="BFAB4D360E5FFFD2A405FA74CDB2FA2C" authorityName="Fabricius" authorityYear="1775" box="[282,389,1499,1523]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="diogenes">
<emphasis id="4ADFEAA70E5FFFD2A405FA74CDB2FA2C" box="[282,389,1499,1523]" italics="true" pageId="4" pageNumber="43">P. diogenes</emphasis>
</taxonomicName>
). Nevertheless, such a difference can often be attributed to intraspecific variations; chela size generally varies to some degree in hermit crab species, particularly between males and females (for 
<taxonomicName id="BFAB4D360E5FFFD2A702F995CEEDF98A" authorityName=", Berthold" authorityYear="1846" box="[541,730,1597,1621]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="aspersus">
<emphasis id="4ADFEAA70E5FFFD2A702F995CEEDF98A" box="[541,730,1597,1621]" italics="true" pageId="4" pageNumber="43">Dardanus aspersus</emphasis>
</taxonomicName>
, cf. 
<bibRefCitation id="1C3A4B440E5FFFD2A583F9F6CD16F9A9" author="Miyake, S." box="[156,289,1630,1654]" pageId="4" pageNumber="43" refId="ref4268" refString="Miyake, S. 1978. The Crustacean Anomura of Sagami Bay. Biological Laboratory, Imperial Household, Tokyo, 200 (English) + 161 (Japanese) pp., 4 pls." type="book" year="1978">Miyake 1978</bibRefCitation>
: pl. 1, fig. 2; 1998: 109, pl. 37, fig. 2; 
<bibRefCitation id="1C3A4B440E5FFFD2A789F9F6CD04F949" author="McLaughlin, P. A. &amp; Rahayu, D. L. &amp; Komai, T. &amp; Chan, T. - Y." pageId="4" pageNumber="43" refId="ref4142" refString="McLaughlin, P. A., Rahayu, D. L., Komai, T. and Chan, T. - Y. 2007. A Catalog of the Hermit Crabs (Paguroidea) of Taiwan. National Taiwan Ocean University, Keelung, 367 pp." type="book" year="2007">
McLaughlin 
<emphasis id="4ADFEAA70E5FFFD2A5DEF9D7CCC0F949" box="[193,247,1662,1686]" italics="true" pageId="4" pageNumber="43">et al.</emphasis>
2007
</bibRefCitation>
; 96, unnumbered fig.). Furthermore, 
<taxonomicName id="BFAB4D360E5FFFD2A7DCF9D7CD0FF968" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Malacostraca" family="Diogenidae" genus="Petrochirus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="diogenes">
<emphasis id="4ADFEAA70E5FFFD2A7DCF9D7CD0FF968" italics="true" pageId="4" pageNumber="43">Petrochirus diogenes</emphasis>
</taxonomicName>
is immediately distinguished from 
<bibRefCitation id="1C3A4B440E5FFFD2A7A4F937CCE9F908" author="Terao, A." pageId="4" pageNumber="43" pagination="355 - 391" refId="ref4418" refString="Terao, A. 1913. A catalogue of hermit-crabs found in Japan (Paguridea excluding Lithodidae), with descriptions of four new species. Annotationes Zoologicae Japonenses 8: 355 - 391." type="journal article" year="1913">Terao’s (1913)</bibRefCitation>
taxon by the much longer and more slender ocular peduncles (cf. 
<bibRefCitation id="1C3A4B440E5FFFD2A42CF948CDFBF927" author="Williams, A. B." box="[307,460,1760,1784]" pageId="4" pageNumber="43" refId="ref4532" refString="Williams, A. B. 1984. Shrimps, Lobsters, and Crabs of the Atlantic Coast of the Eastern United States, Maine to Florida. Smithsonian Institution Press, Washington, D. C., xviii + 550 pp." type="book" year="1984">Williams 1984</bibRefCitation>
). It seems unlikely that 
<bibRefCitation id="1C3A4B440E5FFFD2A7D9F948CCE9F8C7" author="Terao, A." pageId="4" pageNumber="43" pagination="355 - 391" refId="ref4418" refString="Terao, A. 1913. A catalogue of hermit-crabs found in Japan (Paguridea excluding Lithodidae), with descriptions of four new species. Annotationes Zoologicae Japonenses 8: 355 - 391." type="journal article" year="1913">Terao (1913)</bibRefCitation>
overlooked this distinction, because he correctly remarked that 
<taxonomicName id="BFAB4D360E5FFFD2A401F88ACDF7F8E7" authorityName="Terao" authorityYear="1913" box="[286,448,1825,1849]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="watasei">
<emphasis id="4ADFEAA70E5FFFD2A401F88ACDF7F8E7" box="[286,448,1825,1849]" italics="true" pageId="4" pageNumber="43">Pagurus watasei</emphasis>
</taxonomicName>
differed from 
<taxonomicName id="BFAB4D360E5FFFD2A74AF889CD64F886" authority="(Herbst, 1804)" baseAuthorityName="Herbst" baseAuthorityYear="1804" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="megistos">
<emphasis id="4ADFEAA70E5FFFD2A74AF889CC8FF886" italics="true" pageId="4" pageNumber="43">Dardanus megistos</emphasis>
(Herbst, 1804)
</taxonomicName>
(as 
<taxonomicName id="BFAB4D360E5FFFD2A461F8EACDF8F886" box="[382,463,1858,1881]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="4ADFEAA70E5FFFD2A461F8EACDF8F886" box="[382,463,1858,1881]" italics="true" pageId="4" pageNumber="43">Pagurus</emphasis>
</taxonomicName>
) in having shorter ocular peduncles (for 
<taxonomicName id="BFAB4D360E5FFFD2A402F8CBCDA7F8A5" baseAuthorityName="Herbst" baseAuthorityYear="1804" box="[285,400,1891,1914]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="megistos">
<emphasis id="4ADFEAA70E5FFFD2A402F8CBCDA7F8A5" box="[285,400,1891,1914]" italics="true" pageId="4" pageNumber="43">D. megistos</emphasis>
</taxonomicName>
, cf. 
<bibRefCitation id="1C3A4B440E5FFFD2A4A4F8CACE77F8A5" author="Miyake, S." box="[443,576,1890,1914]" pageId="4" pageNumber="43" refId="ref4345" refString="Miyake, S. 1998. Japanese Crustacean Decapods and Stomatopods in Color. I. Macrura, Anomura, and Stomatopoda. Third printing. Hoikusha Publishing Company, Osaka, vii + 261 pp. [In Japanese]" type="book" year="1998">Miyake 1998</bibRefCitation>
; 
<bibRefCitation id="1C3A4B440E5FFFD2A754F8CACCE6F845" author="McLaughlin, P. A. &amp; Rahayu, D. L. &amp; Komai, T. &amp; Chan, T. - Y." pageId="4" pageNumber="43" refId="ref4142" refString="McLaughlin, P. A., Rahayu, D. L., Komai, T. and Chan, T. - Y. 2007. A Catalog of the Hermit Crabs (Paguroidea) of Taiwan. National Taiwan Ocean University, Keelung, 367 pp." type="book" year="2007">
McLaughlin 
<emphasis id="4ADFEAA70E5FFFD2A7D1F8CBCF37F8A5" box="[718,768,1890,1914]" italics="true" pageId="4" pageNumber="43">et al.</emphasis>
2007
</bibRefCitation>
).
</paragraph>
<paragraph id="781436B50E5FFFD2A5AAF80BC877FCD8" blockId="4.[156,771,198,2077]" lastBlockId="4.[820,1433,198,1263]" pageId="4" pageNumber="43">
<bibRefCitation id="1C3A4B440E5FFFD2A5AAF80BCD7BF864" author="Miyake, S." box="[181,332,1955,1979]" pageId="4" pageNumber="43" pagination="127 - 140" refId="ref4192" refString="Miyake, S. 1951. List of Anomura from Kishu. Nankiseibutsu 2: 127 - 140. [In Japanese]" type="journal article" year="1951">Miyake (1951)</bibRefCitation>
gave an annotated list of anomurans recorded from Kii Peninsula, western 
<collectingCountry id="00BC76250E5FFFD2A70FF86CCE7CF804" box="[528,587,1988,2011]" name="Japan" pageId="4" pageNumber="43">Japan</collectingCountry>
. His list included 
<taxonomicName id="BFAB4D360E5FFFD2A583F84DCD76F823" authorityName="Terao" authorityYear="1913" box="[156,321,2021,2044]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="watasei">
<emphasis id="4ADFEAA70E5FFFD2A583F84DCD76F823" box="[156,321,2021,2044]" italics="true" pageId="4" pageNumber="43">Pagurus watasei</emphasis>
</taxonomicName>
based on the published records by 
<bibRefCitation id="1C3A4B440E5FFFD2A7D9F84CCCE9F7C2" author="Terao, A." pageId="4" pageNumber="43" pagination="355 - 391" refId="ref4418" refString="Terao, A. 1913. A catalogue of hermit-crabs found in Japan (Paguridea excluding Lithodidae), with descriptions of four new species. Annotationes Zoologicae Japonenses 8: 355 - 391." type="journal article" year="1913">Terao (1913)</bibRefCitation>
and 
<bibRefCitation id="1C3A4B440E5FFFD2A40CF7ACCD97F7C2" author="Shiino, S. M." box="[275,416,2052,2077]" pageId="4" pageNumber="43" pagination="249 - 300" refId="ref4386" refString="Shiino, S. M. 1933. Bopyrids from Tanabe Bay. Memoirs of the College of Science, Kyoto Imperial University, Series Biology 8: 249 - 300." type="journal article" year="1933">Shiino (1933)</bibRefCitation>
; the latter author only reported 
<taxonomicName id="BFAB4D360E5FFFD2A7F0F7ADCF49FF01" authorityName="Terao" authorityYear="1913" box="[751,894,199,2076]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="watasei">
<emphasis id="4ADFEAA70E5FFFD2A7F0F7ADCF49FF01" box="[751,894,199,2076]" italics="true" pageId="4" pageNumber="43">P. watasei</emphasis>
</taxonomicName>
as a host of his new parasitic isopod 
<taxonomicName id="BFAB4D360E5FFFD2A009FF6FC814FF20" authority="Shiino, 1933" authorityName="Shiino" authorityYear="1933" class="Malacostraca" family="Bopyridae" genus="Pagurion" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="4ADFEAA70E5FFFD2A009FF6FCFA7FF20" italics="true" pageId="4" pageNumber="43">Pagurion tuberculata</emphasis>
<bibRefCitation id="1C3A4B440E5FFFD2A682FF4FC814FF20" author="Shiino, S. M." box="[925,1059,231,255]" pageId="4" pageNumber="43" pagination="249 - 300" refId="ref4386" refString="Shiino, S. M. 1933. Bopyrids from Tanabe Bay. Memoirs of the College of Science, Kyoto Imperial University, Series Biology 8: 249 - 300." type="journal article" year="1933">Shiino, 1933</bibRefCitation>
</taxonomicName>
. Nevertheless, 
<bibRefCitation id="1C3A4B440E5FFFD2A1DAFF4FC969FF20" author="Miyake, S." box="[1221,1374,231,255]" pageId="4" pageNumber="43" refId="ref4268" refString="Miyake, S. 1978. The Crustacean Anomura of Sagami Bay. Biological Laboratory, Imperial Household, Tokyo, 200 (English) + 161 (Japanese) pp., 4 pls." type="book" year="1978">Miyake (1978)</bibRefCitation>
later suggested that 
<taxonomicName id="BFAB4D360E5FFFD2A6D2FEAFC8B5FEC0" baseAuthorityName="Terao" baseAuthorityYear="1913" box="[973,1154,263,287]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="watasei">
<emphasis id="4ADFEAA70E5FFFD2A6D2FEAFC8B5FEC0" box="[973,1154,263,287]" italics="true" pageId="4" pageNumber="43">Dardanus watasei</emphasis>
</taxonomicName>
might be identical with 
<taxonomicName id="BFAB4D360E5FFFD2A09EFEA0CFACFE9F" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1848" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="scutellatus">
<emphasis id="4ADFEAA70E5FFFD2A09EFEA0CFACFE9F" italics="true" pageId="4" pageNumber="43">D. scutellatus</emphasis>
</taxonomicName>
based on his personal investigation, while in the same publication he questionably placed 
<taxonomicName id="BFAB4D360E5FFFD2A1CCFEE1C90DFEBF" baseAuthorityName="Terao" baseAuthorityYear="1913" box="[1235,1338,329,352]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="watasei">
<emphasis id="4ADFEAA70E5FFFD2A1CCFEE1C90DFEBF" box="[1235,1338,329,352]" italics="true" pageId="4" pageNumber="43">D. watasei</emphasis>
</taxonomicName>
under in the synonymy of 
<taxonomicName id="BFAB4D360E5FFFD2A6F6FEC1C858FE5E" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1848" box="[1001,1135,361,385]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="scutellatus">
<emphasis id="4ADFEAA70E5FFFD2A6F6FEC1C858FE5E" box="[1001,1135,361,385]" italics="true" pageId="4" pageNumber="43">D. scutellatus</emphasis>
</taxonomicName>
in a list of Japanese anomurans. 
<bibRefCitation id="1C3A4B440E5FFFD2A672FE21CFCDFE7E" author="Miyake, S." box="[877,1018,393,417]" pageId="4" pageNumber="43" refId="ref4304" refString="Miyake, S. 1982. Japanese Crustacean Decapods and Stomatopods in Color. I. Macrura, Anomura, and Stomatopoda. First edition. Hoikusha Publishing Company, Osaka, vii + 261 pp. [In Japanese]" type="book" year="1982">Miyake (1982</bibRefCitation>
, 
<bibRefCitation id="1C3A4B440E5FFFD2A11AFE21C800FE7E" author="Miyake, S." box="[1029,1079,393,417]" pageId="4" pageNumber="43" refId="ref4345" refString="Miyake, S. 1998. Japanese Crustacean Decapods and Stomatopods in Color. I. Macrura, Anomura, and Stomatopoda. Third printing. Hoikusha Publishing Company, Osaka, vii + 261 pp. [In Japanese]" type="book" year="1998">1998</bibRefCitation>
) did the same, and 
<bibRefCitation id="1C3A4B440E5FFFD2A01CFE21CFA0FE1D" author="McLaughlin, P. A. &amp; Komai, T. &amp; Lemaitre, R. &amp; Rahayu, D. L." pageId="4" pageNumber="43" pagination="5 - 107" refId="ref4073" refString="McLaughlin, P. A., Komai, T., Lemaitre, R. and Rahayu, D. L. 2010. Annotated checklist of anomuran decapod crustaceans of the world (exclusive of the Kiwaoidea and families Chirostylidae and Galatheidae of the Galatheoidea) Part I-Lithodoidea, Lomisoidea and Paguroidea. The Raffles Bulletin of Zoology (Supplement 23): 5 - 107." type="journal article" year="2010">
McLaughlin 
<emphasis id="4ADFEAA70E5FFFD2A099FE22CF79FE1D" italics="true" pageId="4" pageNumber="43">et al.</emphasis>
(2010)
</bibRefCitation>
also followed this synonymy in their checklist of the hermit crab species of the world. However, as has been shown above, 
<taxonomicName id="BFAB4D360E5FFFD2A6D8FE44C807FDDC" baseAuthorityName="Terao" baseAuthorityYear="1913" box="[967,1072,492,515]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="watasei">
<emphasis id="4ADFEAA70E5FFFD2A6D8FE44C807FDDC" box="[967,1072,492,515]" italics="true" pageId="4" pageNumber="43">D. watasei</emphasis>
</taxonomicName>
is actually a junior synonym of 
<taxonomicName id="BFAB4D360E5FFFD2A09EFE44CFB0FDFC" authorityName=", Berthold" authorityYear="1846" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="aspersus">
<emphasis id="4ADFEAA70E5FFFD2A09EFE44CFB0FDFC" italics="true" pageId="4" pageNumber="43">D. aspersus</emphasis>
</taxonomicName>
. 
<taxonomicName id="BFAB4D360E5FFFD2A68AFDA3C850FDFC" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1848" box="[917,1127,523,547]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="scutellatus">
<emphasis id="4ADFEAA70E5FFFD2A68AFDA3C850FDFC" box="[917,1127,523,547]" italics="true" pageId="4" pageNumber="43">Dardanus scutellatus</emphasis>
</taxonomicName>
is readily distinguished from 
<taxonomicName id="BFAB4D360E5FFFD2A62BFD85CF9FFD9B" authorityName=", Berthold" authorityYear="1846" box="[820,936,557,580]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="aspersus">
<emphasis id="4ADFEAA70E5FFFD2A62BFD85CF9FFD9B" box="[820,936,557,580]" italics="true" pageId="4" pageNumber="43">D. aspersus</emphasis>
</taxonomicName>
by the comparatively slender ocular peduncles that are longer than the antennular peduncles, the non-dilated corneas, the lack of distinct corneous-tipped spines or tubercles on the outer face of the left chela, and the dorsally concave lateral surfaces of the dactylus and propodus of the left third pereopod (cf. 
<bibRefCitation id="1C3A4B440E5FFFD2A128FD66C8E8FD38" author="Buitendijk, A. M." box="[1079,1247,718,743]" pageId="4" pageNumber="43" pagination="251 - 280" refId="ref3561" refString="Buitendijk, A. M. 1937. Biological results of the Snellius Expedition. IV. The Paguridea of the Snellius Expedition. Temminckia 2: 251 - 280." type="journal article" year="1937">Buitendijk 1937</bibRefCitation>
; 
<bibRefCitation id="1C3A4B440E5FFFD2A1F1FD66CF50FCD8" author="Fize, A. &amp; Serene, R." pageId="4" pageNumber="43" pagination="1 - 228" refId="ref3691" refString="Fize, A. and Serene, R. 1955. Les Pagures du Vietnam. Institut Oceanographique Nhatrang Note 45: i-ix, 1 - 228." type="book chapter" year="1955">Fize and Serene 1955</bibRefCitation>
; 
<bibRefCitation id="1C3A4B440E5FFFD2A66EFD47C802FCD8" author="Haig, J. &amp; Ball, E. E." box="[881,1077,751,775]" pageId="4" pageNumber="43" pagination="151 - 196" refId="ref3768" refString="Haig, J. and Ball, E. E. 1988. Hermit crabs from northern Australian and eastern Indonesian waters (Crustacea Decapoda: Anomura: Paguroidea) collected during the 1975 Alpha Helix Expedition. Records of the Australian Museum 40: 151 - 196." type="journal article" year="1988">Haig and Ball 1988</bibRefCitation>
).
</paragraph>
<paragraph id="781436B50E5FFFD2A651FCB8C838FB30" blockId="4.[820,1433,198,1263]" pageId="4" pageNumber="43">
The original description of 
<taxonomicName id="BFAB4D360E5FFFD2A17FFCB8C8E7FCF8" authorityName=", Berthold" authorityYear="1846" box="[1120,1232,784,807]" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="aspersus">
<emphasis id="4ADFEAA70E5FFFD2A17FFCB8C8E7FCF8" box="[1120,1232,784,807]" italics="true" pageId="4" pageNumber="43">D. aspersus</emphasis>
</taxonomicName>
has been attributed frequently to 
<bibRefCitation id="1C3A4B440E5FFFD2A6D9FC98C859FC97" author="Berthold, A. A." box="[966,1134,816,840]" pageId="4" pageNumber="43" pagination="3 - 32" refId="ref3517" refString="Berthold, A. A. 1846. Uber verschiedene neue oder seltene Reptilien aus Neu-Granada und Crustaceen aus China. Abhandlungen der Koniglichen Gesellschaft der Wissenschaften zu Gottingen (Physicalischen Classe) 3: 3 - 32, pls 1 - 3." type="journal article" year="1846">Berthold (1846)</bibRefCitation>
, a work containing illustrations of his new taxa of crustaceans, reptiles, and amphibians (
<emphasis id="4ADFEAA70E5FFFD2A670FCDACFA7FC56" box="[879,912,882,905]" italics="true" pageId="4" pageNumber="43">e.g.</emphasis>
, 
<bibRefCitation id="1C3A4B440E5FFFD2A682FCD9C813FC56" author="Miyake, S." box="[925,1060,881,905]" pageId="4" pageNumber="43" refId="ref4268" refString="Miyake, S. 1978. The Crustacean Anomura of Sagami Bay. Biological Laboratory, Imperial Household, Tokyo, 200 (English) + 161 (Japanese) pp., 4 pls." type="book" year="1978">Miyake 1978</bibRefCitation>
; 
<bibRefCitation id="1C3A4B440E5FFFD2A12FFCD9C914FC56" author="McLaughlin, P. A. &amp; Rahayu, D. L. &amp; Komai, T. &amp; Chan, T. - Y." box="[1072,1315,881,905]" pageId="4" pageNumber="43" refId="ref4142" refString="McLaughlin, P. A., Rahayu, D. L., Komai, T. and Chan, T. - Y. 2007. A Catalog of the Hermit Crabs (Paguroidea) of Taiwan. National Taiwan Ocean University, Keelung, 367 pp." type="book" year="2007">
McLaughlin 
<emphasis id="4ADFEAA70E5FFFD2A1ABFCDAC8DFFC56" box="[1204,1256,881,905]" italics="true" pageId="4" pageNumber="43">et al.</emphasis>
2007
</bibRefCitation>
, 
<bibRefCitation id="1C3A4B440E5FFFD2A030FCD9C953FC56" author="McLaughlin, P. A. &amp; Komai, T. &amp; Lemaitre, R. &amp; Rahayu, D. L." box="[1327,1380,881,905]" pageId="4" pageNumber="43" pagination="5 - 107" refId="ref4073" refString="McLaughlin, P. A., Komai, T., Lemaitre, R. and Rahayu, D. L. 2010. Annotated checklist of anomuran decapod crustaceans of the world (exclusive of the Kiwaoidea and families Chirostylidae and Galatheidae of the Galatheoidea) Part I-Lithodoidea, Lomisoidea and Paguroidea. The Raffles Bulletin of Zoology (Supplement 23): 5 - 107." type="journal article" year="2010">2010</bibRefCitation>
), but the correct attribution is to 
<bibRefCitation id="1C3A4B440E5FFFD2A173FC3AC920FC75" author="Berthold, A. A." box="[1132,1303,914,938]" pageId="4" pageNumber="43" pagination="37 - 48" refId="ref3481" refString="Berthold, A. A. 1845. Ueber verschiedene neue oder seltene Reptilien aus Neu-Granada und Crustaceen aus China. Nachrichten von der Koniglichen Gesellschaft der Wissenschaften zu Gottingen 1845: 37 - 48." type="journal article" year="1845">Berthold (1845)</bibRefCitation>
. This paper had the same title as the 1846 work but comprised a shorter, unillustrated text. In addition to 
<taxonomicName id="BFAB4D360E5FFFD2A1A6FC7BCF41FBD4" authority=", Berthold (1845)" authorityName="Berthold" authorityYear="1845" class="Malacostraca" family="Diogenidae" genus="Dardanus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="aspersus">
<emphasis id="4ADFEAA70E5FFFD2A1A6FC7BC919FC35" box="[1209,1326,979,1002]" italics="true" pageId="4" pageNumber="43">D. aspersus</emphasis>
, 
<bibRefCitation id="1C3A4B440E5FFFD2A021FC7BCF41FBD4" author="Berthold, A. A." pageId="4" pageNumber="43" pagination="37 - 48" refId="ref3481" refString="Berthold, A. A. 1845. Ueber verschiedene neue oder seltene Reptilien aus Neu-Granada und Crustaceen aus China. Nachrichten von der Koniglichen Gesellschaft der Wissenschaften zu Gottingen 1845: 37 - 48." type="journal article" year="1845">Berthold (1845)</bibRefCitation>
</taxonomicName>
included descriptions of three other new crustacean taxa: one slipper lobster, 
<taxonomicName id="BFAB4D360E5FFFD2A156FBBCC9AFFBF3" authority="Berthold, 1845" authorityName="Berthold" authorityYear="1845" box="[1097,1432,1044,1068]" class="Malacostraca" family="Scyllaridae" genus="Scyllarus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="haanii">
<emphasis id="4ADFEAA70E5FFFD2A156FBBCC8D8FBF3" box="[1097,1263,1044,1068]" italics="true" pageId="4" pageNumber="43">Scyllarus haanii</emphasis>
<bibRefCitation id="1C3A4B440E5FFFD2A1E4FBBCC9AFFBF3" author="Berthold, A. A." box="[1275,1432,1044,1068]" pageId="4" pageNumber="43" pagination="37 - 48" refId="ref3481" refString="Berthold, A. A. 1845. Ueber verschiedene neue oder seltene Reptilien aus Neu-Granada und Crustaceen aus China. Nachrichten von der Koniglichen Gesellschaft der Wissenschaften zu Gottingen 1845: 37 - 48." type="journal article" year="1845">Berthold, 1845</bibRefCitation>
</taxonomicName>
[= 
<taxonomicName id="BFAB4D360E5FFFD2A64DFB9DC81FFB93" box="[850,1064,1076,1100]" class="Malacostraca" family="Scyllaridae" genus="Remiarctus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="bertholdii">
<emphasis id="4ADFEAA70E5FFFD2A64DFB9DC81FFB93" box="[850,1064,1076,1100]" italics="true" pageId="4" pageNumber="43">Remiarctus bertholdii</emphasis>
</taxonomicName>
(Paul’son, 1875)], and two stomatopods, 
<taxonomicName id="BFAB4D360E5FFFD2A66BFBFDC8A9FBB2" authority="Berthold, 1845" authorityName="Berthold" authorityYear="1845" box="[884,1182,1109,1133]" class="Malacostraca" family="Squillidae" genus="Squilla" kingdom="Animalia" order="Stomatopoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="affinis">
<emphasis id="4ADFEAA70E5FFFD2A66BFBFDCFC9FBB2" box="[884,1022,1109,1133]" italics="true" pageId="4" pageNumber="43">Squilla affinis</emphasis>
<bibRefCitation id="1C3A4B440E5FFFD2A119FBFDC8A9FBB2" author="Berthold, A. A." box="[1030,1182,1109,1133]" pageId="4" pageNumber="43" pagination="37 - 48" refId="ref3481" refString="Berthold, A. A. 1845. Ueber verschiedene neue oder seltene Reptilien aus Neu-Granada und Crustaceen aus China. Nachrichten von der Koniglichen Gesellschaft der Wissenschaften zu Gottingen 1845: 37 - 48." type="journal article" year="1845">Berthold, 1845</bibRefCitation>
</taxonomicName>
[= 
<taxonomicName id="BFAB4D360E5FFFD2A1DBFBFDCFD9FB52" authority="(De Haan, 1844)" baseAuthorityName="De Haan" baseAuthorityYear="1844" class="Malacostraca" family="Squillidae" genus="Oratosquilla" kingdom="Animalia" order="Stomatopoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="oratoria">
<emphasis id="4ADFEAA70E5FFFD2A1DBFBFDC9AFFBB2" box="[1220,1432,1109,1133]" italics="true" pageId="4" pageNumber="43">Oratosquilla oratoria</emphasis>
(De Haan, 1844)
</taxonomicName>
] and 
<taxonomicName id="BFAB4D360E5FFFD2A129FBDDCF52FB71" authority="Berthold, 1845" authorityName="Berthold" authorityYear="1845" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylus" kingdom="Animalia" order="Stomatopoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="edwardsii">
<emphasis id="4ADFEAA70E5FFFD2A129FBDDC91CFB52" box="[1078,1323,1141,1165]" italics="true" pageId="4" pageNumber="43">Gonodactylus edwardsii</emphasis>
<bibRefCitation id="1C3A4B440E5FFFD2A027FBDDCF52FB71" author="Berthold, A. A." pageId="4" pageNumber="43" pagination="37 - 48" refId="ref3481" refString="Berthold, A. A. 1845. Ueber verschiedene neue oder seltene Reptilien aus Neu-Granada und Crustaceen aus China. Nachrichten von der Koniglichen Gesellschaft der Wissenschaften zu Gottingen 1845: 37 - 48." type="journal article" year="1845">Berthold, 1845</bibRefCitation>
</taxonomicName>
[= 
<taxonomicName id="BFAB4D360E5FFFD2A694FB3EC970FB71" authority="(De Haan, 1844)" baseAuthorityName="De Haan" baseAuthorityYear="1844" box="[907,1351,1174,1198]" class="Malacostraca" family="Odontodactylidae" genus="Odontodactylus" kingdom="Animalia" order="Stomatopoda" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="4ADFEAA70E5FFFD2A694FB3EC8B9FB71" box="[907,1166,1174,1198]" italics="true" pageId="4" pageNumber="43">Odontodactylus japonicus</emphasis>
(De Haan, 1844)
</taxonomicName>
], which have already been referred to the year 1845 (
<emphasis id="4ADFEAA70E5FFFD2A013FB1FC91AFB11" box="[1292,1325,1207,1230]" italics="true" pageId="4" pageNumber="43">e.g.</emphasis>
, 
<bibRefCitation id="1C3A4B440E5FFFD2A024FB1ECF50FB30" author="Manning, R. B." pageId="4" pageNumber="43" pagination="1 - 339" refId="ref3900" refString="Manning, R. B. 1995. Stomatopod Crustacea of Vietnam: the legacy of Raoul Serene. Crustacean Research, Special Number 4: 1 - 339." type="book chapter" year="1995">Manning 1995</bibRefCitation>
; 
<bibRefCitation id="1C3A4B440E5FFFD2A66EFB7FC834FB30" author="Holthuis, L. B." box="[881,1027,1239,1263]" pageId="4" pageNumber="43" pagination="499 - 683" refId="ref3817" refString="Holthuis, L. B. 2002. The Indo-Pacific scyllarine lobsters (Crustacea, Decapoda, Scyllaridae). Zoosystema 24: 499 - 683." type="journal article" year="2002">Holthuis 2002</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>