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<mods:title id="C570252A02E1A684A46E11F7C7E2C30E">Taxonomic revision of the endemic Bornean genera Anexodus Pascoe and Pantilema Aurivillius (Coleoptera, Cerambycidae, Lamiinae)</mods:title>
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<mods:namePart id="5BBBEA6774FFB904CC0215BF349D8DA2">Gabris, Radim</mods:namePart>
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<mods:namePart id="E700C47BA434FB30E973CC114F65BF23">Wahab, Rodzay Abdul</mods:namePart>
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<paragraph id="31070394E2419947598903A943A8FCB9" pageId="1" pageNumber="30">
<taxonomicName id="36538A1DBF3D6D20E336DC5B6767EA3C" ID-CoL="66R4W" authority="Sudre, 1997" authorityName="Sudre" authorityYear="1997" class="Insecta" family="Cerambycidae" genus="Anexodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anexodus sarawakensis" order="Coleoptera" pageId="1" pageNumber="30" phylum="Arthropoda" rank="species" species="sarawakensis">Anexodus sarawakensis Sudre, 1997</taxonomicName>
Figs 5-6, 13-14, 18, 23-24, 29, 34
</paragraph>
</subSubSection>
<subSubSection id="02BCF17F7323A900E7267595D3603AF4" pageId="2" pageNumber="31" type="reference_group">
<paragraph id="53DA8AB9ABB52A18A848535C4A3A3DB9" pageId="2" pageNumber="31">
<taxonomicName id="49E47630CA425B2FC72FA4A134FCB9BF" class="Insecta" family="Cerambycidae" genus="Anexodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anexodus sarawakensis" order="Coleoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="sarawakensis">
<pageBreakToken id="53053F188470064E5D2457316E51FF54" pageId="2" pageNumber="31" start="start">Anexodus</pageBreakToken>
sarawakensis
</taxonomicName>
Sudre, 1997: 253.
</paragraph>
</subSubSection>
<subSubSection id="93A1F59ACF3090C21456870F0A3C3390" pageId="2" pageNumber="31" type="type material">
<paragraph id="F3714B6C7250E75B8F106A994AA51379" pageId="2" pageNumber="31">Type material examined.</paragraph>
<paragraph id="6FBFD4351A6971E1E7D93DF3FE83DA9D" pageId="2" pageNumber="31">
Holotype, male, &quot;E. MALAYSIA: Sarawak / confl. Sun Oyan and / Mujong riv., E. Kapit / 50m, 18.V.1994, # 5 /
<normalizedToken id="32BAB9C364EEA69CC7453B2CCA769781" originalValue="Löbl">Loebl</normalizedToken>
&amp; Burckhardt // Holotype //
<taxonomicName id="3B35CDC426FE87CDCAEA6CE3A70B3F7B" class="Insecta" family="Cerambycidae" genus="Anexodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anexodus" order="Coleoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="genus">Anexodus</taxonomicName>
/
<taxonomicName id="7C6C8B4D6E530C1B99B4988F7B524569" lsidName="sarawakensis" pageId="2" pageNumber="31" rank="species" species="sarawakensis">sarawakensis</taxonomicName>
Nov sp. / J. Sudre det. 1996&quot; (MHNG). Paratype, female, &quot;E. MALAYSIA: Sarawak / confl. Sun Oyan and / Mujong riv., E. Kapit / 50m, 18.V.1994, # 5 /
<normalizedToken id="734FE027E277E0EA559F33326069F020" originalValue="Löbl">Loebl</normalizedToken>
&amp; Burckhardt //
<normalizedToken id="0195FBD9AE163D7FD69B9FE54B73FA7D" originalValue="comparé">compare</normalizedToken>
zu type /
<normalizedToken id="1C29942FE3418862AABDDCAB5047D8D7" originalValue="d’">d'</normalizedToken>
<taxonomicName id="66A0B24B1A0E4185FD3789FA15FC230C" lsidName="A. aquilus" pageId="2" pageNumber="31" rank="species" species="aquilus">A. aquilus</taxonomicName>
per / J. Sudre 1996 // Paratype //
<taxonomicName id="961DC88AC24984119A1B4D706730496D" class="Insecta" family="Cerambycidae" genus="Anexodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anexodus" order="Coleoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="genus">Anexodus</taxonomicName>
(♀) /
<taxonomicName id="1CEF80DD74E8AB67A2DAF0E49617664C" lsidName="sarawakensis" pageId="2" pageNumber="31" rank="species" species="sarawakensis">sarawakensis</taxonomicName>
sp. n / J. Sudre det. 1996&quot; (MHNG).
</paragraph>
<caption id="285E015B485F5A178345BBD5A1DCBFD7" pageId="2" pageNumber="31">
<paragraph id="9ED2430377C3864553B6EE94B6944F2A" pageId="2" pageNumber="31">
Figures 5-8. Dorsal habitus of
<taxonomicName id="8711161CB9BD7B6CB0B84044C00C4C96" class="Insecta" family="Cerambycidae" genus="Anexodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anexodus" order="Coleoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="genus">Anexodus</taxonomicName>
spp.: 5
<taxonomicName id="4CC93D65B800ADB6C8E2533222ACB36E" class="Insecta" family="Cerambycidae" genus="Anexodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anexodus sarawakensis" order="Coleoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="sarawakensis">Anexodus sarawakensis</taxonomicName>
Sudre, holotype male 6
<taxonomicName id="360360138E85B1EDFC0994A37A4A4153" class="Insecta" family="Cerambycidae" genus="Anexodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anexodus sarawakensis" order="Coleoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="sarawakensis">Anexodus sarawakensis</taxonomicName>
Sudre, paratype female 7
<taxonomicName id="1A8A524755B5A1A8F55CF19B27DEDD33" class="Insecta" family="Cerambycidae" genus="Anexodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anexodus tufi" order="Coleoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="tufi">Anexodus tufi</taxonomicName>
sp. n., holotype male 8
<taxonomicName id="A7D1C6F5E411E5D56B035284C642B8E7" class="Insecta" family="Cerambycidae" genus="Anexodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anexodus tufi" order="Coleoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="tufi">Anexodus tufi</taxonomicName>
sp. n., paratype female. Scale bar 5 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="1BE252A2B3CEEEBA77B710326C176926" pageId="2" pageNumber="31" type="differential diagnosis">
<paragraph id="08D17611CB047BB1F73A08076907E94E" pageId="2" pageNumber="31">Differential diagnosis.</paragraph>
<paragraph id="24266BE2C152F978F6F63240572EFA60" pageId="2" pageNumber="31">
<taxonomicName id="E2390D887A39ADAA5B794B52FF6F53E8" class="Insecta" family="Cerambycidae" genus="Anexodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anexodus sarawakensis" order="Coleoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="sarawakensis">Anexodus sarawakensis</taxonomicName>
is similar to
<taxonomicName id="FD348CEF321516317975BFA1F4F012D5" lsidName="A. syptakovae" pageId="2" pageNumber="31" rank="species" species="syptakovae">A. syptakovae</taxonomicName>
sp. n. in having uniformly colored habitus and labrum with one row of distinct punctures with setae (Fig. 18). For more details see the differential diagnosis under the latter species.
</paragraph>
</subSubSection>
<subSubSection id="4F36FED1DBCB21D106003DE61D4DA4AF" pageId="2" pageNumber="31" type="description">
<paragraph id="99A26117AA4CB1758DA8C5FD381EABF8" pageId="2" pageNumber="31">Description.</paragraph>
<paragraph id="5AA13E151370D5D8BE912C347B03959D" pageId="2" pageNumber="31">Holotype (male). BL 9.2 mm, BW 3.1 mm. Body uniformly brown, mouthparts lighter. Body densely clothed with very short brown pubescence. Head about as wide as anterior margin of pronotum. Labrum transverse, with one row of punctures bearing long setae (Fig. 18). Eyes less emarginate at antennal insertions (Figs 13-14). Antennae 0.8 times as long as body; scape gradually slightly widened towards apex, thickest at apical part, apex simple (Figs 23-24); the relative ratio of antennomere lengths 6.3: 2.5: 1.0: 1.3: 1.1: 1.0: 1.0: 0.9: 0.8: 1.0: 2.0.</paragraph>
<paragraph id="166772C9768B113B17C537AE642788C1" pageId="2" pageNumber="31">Prothorax as long as wide, laterally with one moderately distinct tubercle; pronotal disc with a pair of distinct tubercles near middle and one median at second half; pronotal tubercles punctate. Prosternum in front of coxae 0.9 times shorter than diameter of coxal cavity. Scutellum transverse, two times as wide as long. Elytra elongate, 1.6 times as long as wide at widest part, 1.8 times as long as pronotum, widest near middle; each elytron with three rows of distinct tubercles (Figs 5-6), sparsely covered with large deep punctures arranged in slightly irregular rows. Legs long, slender; relative lengths of metatarsomeres 1.0: 0.7: 1.1: 2.0.</paragraph>
<caption id="1AB0B45F9321F439C2FA83AFA6E505FA" pageId="2" pageNumber="31">
<paragraph id="417A20742967D8E5955992B1A8694064" pageId="2" pageNumber="31">
Figures 9-16. Lateral habitus of
<taxonomicName id="2E169B37F548E60F0F0807D3A3561F58" class="Insecta" family="Cerambycidae" genus="Anexodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anexodus" order="Coleoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="genus">Anexodus</taxonomicName>
spp.: 9
<taxonomicName id="A2A14C63EB9AE9574C7C01C88A295C26" class="Insecta" family="Cerambycidae" genus="Anexodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anexodus aquilus" order="Coleoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="aquilus">Anexodus aquilus</taxonomicName>
Pascoe, holotype male 10
<taxonomicName id="94239F9E7D4E440100A298D16EE11FA6" class="Insecta" family="Cerambycidae" genus="Anexodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anexodus aquilus" order="Coleoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="aquilus">Anexodus aquilus</taxonomicName>
Pascoe, female (= holotype of
<taxonomicName id="BCF2EFF81894707D16DDDF50627A5F0E" class="Insecta" family="Cerambycidae" genus="Anexodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anexodus kuntzeni" order="Coleoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="kuntzeni">Anexodus kuntzeni</taxonomicName>
Kriesche) 11
<taxonomicName id="3F4F4730B889BBD1E974B8969B30A5CA" class="Insecta" family="Cerambycidae" genus="Anexodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anexodus aquilus" order="Coleoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="aquilus">Anexodus aquilus</taxonomicName>
Pascoe, large male from Trus Madi 12
<taxonomicName id="D6D3B5010D7EE993F8DCF23409015367" class="Insecta" family="Cerambycidae" genus="Anexodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anexodus syptakovae" order="Coleoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="syptakovae">Anexodus syptakovae</taxonomicName>
sp. n., holotype male 13
<taxonomicName id="7D25C27D3E04D69453CAD820EA7CD672" class="Insecta" family="Cerambycidae" genus="Anexodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anexodus sarawakensis" order="Coleoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="sarawakensis">Anexodus sarawakensis</taxonomicName>
Sudre, holotype male 14
<taxonomicName id="2849D1A637DC0813296F79AF89FC605C" class="Insecta" family="Cerambycidae" genus="Anexodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anexodus sarawakensis" order="Coleoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="sarawakensis">Anexodus sarawakensis</taxonomicName>
Sudre, paratype female 15
<taxonomicName id="759DF45293CB626DB8D68A45863AAD67" class="Insecta" family="Cerambycidae" genus="Anexodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anexodus tufi" order="Coleoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="tufi">Anexodus tufi</taxonomicName>
sp. n., holotype male 16
<taxonomicName id="5474054D4F508254106A08D54951BB8F" class="Insecta" family="Cerambycidae" genus="Anexodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anexodus tufi" order="Coleoptera" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="tufi">Anexodus tufi</taxonomicName>
sp. n., paratype female. Scale bar 5 mm.
</paragraph>
</caption>
<paragraph id="03580EC1DF41738E50971918911F8B80" pageId="2" pageNumber="31">Male genitalia with tegmen elongate, widest near middle, basally with short strut; parameres elongate, 2.9 times longer than wide, apically with long setae (Fig. 29). Penis subparallel-sided, apically subacute; dorsal struts diverged from about one third of penis length. Internal sac long, with paired small medial sclerites and distinct flagellar sclerites.</paragraph>
<paragraph id="CB0DEC70B385D4A7CE1D7A1667B33D87" pageId="2" pageNumber="31">
Female. Most characters same as for males. BL 9.9 mm, BW 3.5 mm. Body more convex dorsally. Antennae slightly shorter than in male, 0.7 times as long as body; length ratio of antennomeres
<normalizedToken id="3886335CA4E5C6DD24D8108D270C41DB" originalValue="I–III">I-III</normalizedToken>
: 6.5: 2.7: 1.0. Elytra 1.7 times as long as wide, 2.3 times as long as pronotum. Spermatheca only slightly sclerotized, slender, elongate; sclerotized part of spermathecal duct strongly coiled (Fig. 34).
</paragraph>
</subSubSection>
<subSubSection id="76C3F7BE0A39AA1FB669A3DC35FB2944" pageId="2" pageNumber="31" type="distribution">
<paragraph id="F2EA48E35E4CC02D649AB9962C074686" pageId="2" pageNumber="31">Distribution.</paragraph>
<paragraph id="EB5EE2D195D9136C642B7956C0380A35" pageId="2" pageNumber="31">Malaysia: Borneo (Sarawak: Kapit; Fig. 43).</paragraph>
</subSubSection>
</treatment>
</document>
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