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<document id="BF246822D88C91BBCE6F1AA30F3E7F97" ID-CLB-Dataset="2419" ID-DOI="10.21248/contrib.entomol.66.1.13-111" ID-GBIF-Dataset="d2933fad-1cac-4a5a-a558-2633f8314541" ID-ISSN="0005-805X" ID-Zenodo-Dep="6421171" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630270776760" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2016" docId="766F7C36FFF0FFAAFF3675D4DC89FC02" docLanguage="en" docName="ContribEntomol.2016.66.1.13-111.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 66 (1)" docSource="https://www.contributions-to-entomology.org/article/view/1904" docStyle="DocumentStyle:05A48F5F6902F879680B6E64765AA512.2:ContribEntomol.2013-2016.journal_article" docStyleId="05A48F5F6902F879680B6E64765AA512" docStyleName="ContribEntomol.2013-2016.journal_article" docStyleVersion="2" docTitle="Drusilla bifida Assing 2016, spec. nov." docType="treatment" docVersion="9" lastPageNumber="78" masterDocId="8A56044EFFB1FFEBFF8B7527D878FF83" masterDocTitle="On some Lomechusini of the Palaearctic and Oriental regions (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="111" masterPageNumber="13" pageNumber="78" updateTime="1699235685730" updateUser="ExternalLinkService">
<mods:mods id="4A2B3426F784E41AE70F3E478531A46B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E26ECAE331D226D41A328F6E32AB2C0F">
<mods:title id="AB0AD1252C2C60B8DCC826F27F078128">On some Lomechusini of the Palaearctic and Oriental regions (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
</mods:titleInfo>
<mods:name id="FC11F59BAEDA1C472CED42B359BE36D1" type="personal">
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<mods:namePart id="F4922532ECFCD46EFDAC44FA3DBDEC8B">Assing, Volker</mods:namePart>
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<mods:typeOfResource id="4816A846EF9DD87714611257B14F5F54">text</mods:typeOfResource>
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<mods:title id="41A15D02FD317910CDB712964A8FC22D">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
</mods:titleInfo>
<mods:part id="CE643AE35D31FA958B996DBE2263E875">
<mods:date id="B52FF81C4B682D0B37A2F46AE9C3597C">2016</mods:date>
<mods:detail id="AB6F9C84A2C6F4EB1E20C6576BDFC1A2" type="pubDate">
<mods:number id="282B7378A984C61D0193B3BAB8506BCD">2016-12-20</mods:number>
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<mods:detail id="16A77AF0EAD553AF2916712133B5AF77" type="volume">
<mods:number id="5DA7C2996274107E6696F0A0A680BAEF">66</mods:number>
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<mods:number id="DBDAE4B16F2EF37F88EEAF17878BC7B7">1</mods:number>
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<mods:url id="6BAC57A7523E16B29DB1A7589AA32D87">https://www.contributions-to-entomology.org/article/view/1904</mods:url>
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<mods:classification id="A963DA2D1F37E4A825B5D6D66907817A">journal article</mods:classification>
<mods:identifier id="AEF96D416BF6A42B2F5639B93926274B" type="CLB-Dataset">2419</mods:identifier>
<mods:identifier id="1FD645178CBB03559EC089E1D2DAF48E" type="DOI">10.21248/contrib.entomol.66.1.13-111</mods:identifier>
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<treatment id="766F7C36FFF0FFAAFF3675D4DC89FC02" ID-DOI="http://doi.org/10.5281/zenodo.5903486" ID-GBIF-Taxon="192602800" ID-Zenodo-Dep="5903486" LSID="urn:lsid:plazi:treatment:766F7C36FFF0FFAAFF3675D4DC89FC02" httpUri="http://treatment.plazi.org/id/766F7C36FFF0FFAAFF3675D4DC89FC02" lastPageNumber="78" pageId="65" pageNumber="78">
<subSubSection id="B6DC9EABFFF0FFAAFF3675D4D9C2FE8D" box="[189,442,243,270]" pageId="65" pageNumber="78" type="nomenclature">
<paragraph id="FE79CD20FFF0FFAAFF3675D4D9C2FE8D" blockId="65.[189,442,243,303]" box="[189,442,243,270]" pageId="65" pageNumber="78">
<heading id="A5317A4CFFF0FFAAFF3675D4D9C2FE8D" box="[189,442,243,270]" centered="true" fontSize="11" level="2" pageId="65" pageNumber="78" reason="3">
<taxonomicName id="39C6B6A3FFF0FFAAFF3675D4D933FE8D" ID-CoL="37TDW" authority="Assing, 2016" authorityName="Assing" authorityYear="2016" box="[189,331,243,270]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="78" phylum="Arthropoda" rank="species" species="bifida" status="sp. nov.">
<emphasis id="CCB21132FFF0FFAAFF3675D4D933FE8D" box="[189,331,243,270]" italics="true" pageId="65" pageNumber="78">Drusilla bifida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D781AC49FFF0FFAAFEDA75D4D9C2FE8D" box="[337,442,243,270]" pageId="65" pageNumber="78" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B6DC9EABFFF0FFAAFF36743ED9E8FEAC" box="[189,400,281,303]" pageId="65" pageNumber="78" type="description">
<paragraph id="FE79CD20FFF0FFAAFF36743ED9E8FEAC" blockId="65.[189,442,243,303]" box="[189,400,281,303]" pageId="65" pageNumber="78">
(
<figureCitation id="66FDD1A5FFF0FFAAFF4A743ED940FEAC" box="[193,312,281,303]" captionStart="Figs 119–143" captionStartId="15.[151,191,1969,1991]" captionTargetBox="[151,1398,232,1928]" captionTargetId="figure-156@15.[151,1398,232,1940]" captionTargetPageId="15" captionText="Figs 119–143. Drusilla bifida (119–122), Rabdotodrusilla pectinata (123–125), Amaurodera arunica (126–127, 132), A. gilvios (128, 133), A. reticulata (129, 134), A. dentata (130, 135), A. parvoculata (131, 136), A. thailandensis (137–138), A. fasciata (139), A. meorum(140),A. disparicollis (141), A. spinans (142),andA. varicollis(143):forebody (119, 123, 126, 128–131); head and pronotum (120); antenna (121, 124, 132–143); abdomen (122, 125, 127). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6421190" httpUri="https://zenodo.org/record/6421190/files/figure.png" pageId="65" pageNumber="78">Figs 119–122</figureCitation>
,
<figureCitation id="66FDD1A5FFF0FFAAFEB4743ED9F4FEAC" box="[319,396,281,303]" captionStart-0="Figs 359–378" captionStart-1="Figs 379–395" captionStartId-0="27.[151,191,1971,1993]" captionStartId-1="28.[189,229,1943,1965]" captionTargetBox-0="[155,1396,235,1964]" captionTargetBox-1="[193,1428,232,1921]" captionTargetId-0="figure-155@27.[151,1398,232,1964]" captionTargetId-1="figure-160@28.[189,1436,232,1934]" captionTargetPageId-1="28" captionText-0="Figs 359–378. Orphnebius laticeps (359–361), O. biimpressus (362–364), O. ulcerosus (365–369), O. falagrioides (370–372), and Drusilla bifida (373–378): tergite VIII (359, 362, 368, 370, 374); female sternite VIII (360, 363, 371); spermatheca (361, 364, 372); median lobe of aedeagus in lateral and in ventral view (365–366, 376–377); paramere (367); male sternite VIII (369, 375); posteromedian portion or head (373); apex of ventral process of aedeagus (378). Scale bars: 359–372, 374–377: 0.2 mm; 373, 378: 0.1 mm." captionText-1="Figs 379–395. Drusilla bifida (379), Rabdotodrusilla pectinata (380–383), Amaurodera arunica (384–388), A. gilvios (389–390, 395), and A. reticulata (391–394): sutural portion of left elytron (379); median lobe of aedeagus in lateral and in ventral view (380– 381, 385–386, 389–392); male tergite VIII (382); male sternite VIII (383); male pronotum (384); spermatheca (387–388, 395); subapical portion of median lobe of aedeagus in dry preparation (393); subapical portion of median lobe of aedeagus in transparent light (394). Scale bars: 384: 0.5 mm; 379–383, 385–392, 395: 0.2 mm; 393–394: 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.6421218" figureDoi-1="http://doi.org/10.5281/zenodo.6421220" httpUri-0="https://zenodo.org/record/6421218/files/figure.png" httpUri-1="https://zenodo.org/record/6421220/files/figure.png" pageId="65" pageNumber="78">373–379</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B6DC9EABFFF0FFAAFF1C746ED987FE42" pageId="65" pageNumber="78" type="materials_examined">
<paragraph id="FE79CD20FFF0FFAAFF1C746ED987FE42" blockId="65.[151,756,329,449]" pageId="65" pageNumber="78">
<emphasis id="CCB21132FFF0FFAAFF1C746ED94AFEE2" bold="true" box="[151,306,329,353]" pageId="65" pageNumber="78">Type material</emphasis>
:
<materialsCitation id="4EAEC77DFFF0FFAAFEC8746ED987FE42" ID-GBIF-Occurrence="3455118427" collectingDate="2014-01-25" collectorName="Ob" country="Thailand" determinerName="V. Assing" elevation="800" latitude="20.033333" location="Doi Pha Hom Pok" longLatPrecision="1269" longitude="99.23333" municipality="Bhoo Muan" pageId="65" pageNumber="78" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
<typeStatus id="217D7382FFF0FFAAFEC8746ED9DEFEE2" box="[323,422,329,353]" pageId="65" pageNumber="78" type="holotype">Holotype</typeStatus>
<specimenCount id="E8C006A9FFF0FFAAFE3B746DD9BDFEE2" box="[432,453,330,353]" pageId="65" pageNumber="78" type="male">♂</specimenCount>
: “
<collectingCountry id="86D18DB0FFF0FFAAFE55746EDA38FEE2" box="[478,576,329,353]" name="Thailand" pageId="65" pageNumber="78">Thailand</collectingCountry>
[47] –
<location id="FB199BFBFFF0FFAAFD12746ED979FE02" LSID="urn:lsid:plazi:treatment:766F7C36FFF0FFAAFF3675D4DC89FC02:FB199BFBFFF0FFAAFD12746ED979FE02" country="Thailand" latitude="20.033333" longLatPrecision="1269" longitude="99.23333" municipality="Bhoo Muan" name="Doi Pha Hom Pok" pageId="65" pageNumber="78">Doi Pha Hom Pok</location>
,
<collectingMunicipality id="1E1D575AFFF0FFAAFE86744ED9F4FE02" box="[269,396,361,385]" pageId="65" pageNumber="78">Bhoo Muan</collectingMunicipality>
waterf.,
<geoCoordinate id="9BF2ABE7FFF0FFAAFE60744EDA39FE02" box="[491,577,361,385]" degrees="20" direction="north" minutes="02" orientation="latitude" pageId="65" pageNumber="78" precision="925" value="20.033333">20°02'N</geoCoordinate>
,
<geoCoordinate id="9BF2ABE7FFF0FFAAFDC5744EDAE7FE02" box="[590,671,361,385]" degrees="99" direction="east" minutes="14" orientation="longitude" pageId="65" pageNumber="78" precision="925" value="99.23333">99°14'E</geoCoordinate>
,
<quantity id="393E60C5FFF0FFAAFD20744EDA88FE02" box="[683,752,361,385]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="65" pageNumber="78" unit="m" value="800.0">
<elevation id="75EB2A13FFF0FFAAFD20744EDA88FE02" box="[683,752,361,385]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="65" pageNumber="78" unit="m" value="800.0">800 m</elevation>
</quantity>
, litter,
<date id="8A78EBE0FFF0FFAAFF5074AED947FE22" box="[219,319,393,417]" pageId="65" pageNumber="78" value="2014-01-25">
<collectingDate id="9A3C1208FFF0FFAAFF5074AED947FE22" box="[219,319,393,417]" pageId="65" pageNumber="78" value="2014-01-25">25.I.2014</collectingDate>
</date>
, leg.
<collectorName id="5333A8F6FFF0FFAAFEF574AED9D8FE22" box="[382,416,393,417]" pageId="65" pageNumber="78">Ob</collectorName>
/
<typeStatus id="217D7382FFF0FFAAFE3574AEDA55FE22" box="[446,557,393,417]" pageId="65" pageNumber="78" type="holotype">Holotypus</typeStatus>
<specimenCount id="E8C006A9FFF0FFAAFDBC74ADDA34FE22" box="[567,588,394,417]" pageId="65" pageNumber="78" type="male">♂</specimenCount>
<taxonomicName id="39C6B6A3FFF0FFAAFDDC74AEDA8CFE22" authority="Assing, 2016" authorityName="Assing" authorityYear="2016" box="[599,756,393,417]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="78" phylum="Arthropoda" rank="species" species="bifida" status="sp. nov.">
<emphasis id="CCB21132FFF0FFAAFDDC74AEDA8CFE22" box="[599,756,393,417]" italics="true" pageId="65" pageNumber="78">Drusilla bifida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D781AC49FFF0FFAAFF1C748DD8A8FE42" box="[151,208,426,449]" pageId="65" pageNumber="78" rank="species">sp. n.</taxonomicNameLabel>
, det.
<determinerName id="0BA521E4FFF0FFAAFE82748DD913FE42" box="[265,363,425,449]" pageId="65" pageNumber="78">V. Assing</determinerName>
2015” (cAss).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="B6DC9EABFFF0FFAAFF1C74CEDA1AFDA2" pageId="65" pageNumber="78" type="etymology">
<paragraph id="FE79CD20FFF0FFAAFF1C74CEDA1AFDA2" blockId="65.[151,756,489,545]" pageId="65" pageNumber="78">
<emphasis id="CCB21132FFF0FFAAFF1C74CED976FD82" bold="true" box="[151,270,489,513]" pageId="65" pageNumber="78">Etymology</emphasis>
: The specific epithet (Latin, adjective) alludes to the bifid ventral process of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="B6DC9EABFFF0FFAAFF1C776EDD4BFD62" pageId="65" pageNumber="78" type="description">
<paragraph id="FE79CD20FFF0FFAAFF1C776EDA1EFCA2" blockId="65.[149,757,585,2081]" pageId="65" pageNumber="78">
<emphasis id="CCB21132FFF0FFAAFF1C776ED961FDE2" bold="true" box="[151,281,585,609]" pageId="65" pageNumber="78">Description</emphasis>
: Body length
<quantity id="393E60C5FFF0FFAAFE4B776EDA6BFDE2" box="[448,531,585,609]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="65" pageNumber="78" unit="mm" value="4.7">4.7 mm</quantity>
; length of forebody
<quantity id="393E60C5FFF0FFAAFF1C774ED893FD02" box="[151,235,617,641]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="65" pageNumber="78" unit="mm" value="2.2">2.2 mm</quantity>
. Coloration: head blackish; pronotum and elytra dark-brown; scutellum paler brown; abdomen darkbrown, with segments III and VIII–X reddish-brown; legs dark-yellowish; antenna dark-brown with antennomeres I–II and the base of III reddish; maxillary palpi brown with the apical palpomere yellowish.
</paragraph>
<paragraph id="FE79CD20FFF0FFAAFF1C760ED9ABFAA2" blockId="65.[149,757,585,2081]" pageId="65" pageNumber="78">
Head (
<figureCitation id="66FDD1A5FFF0FFAAFF63760ED900FCC2" box="[232,376,809,833]" captionStart="Figs 119–143" captionStartId="15.[151,191,1969,1991]" captionTargetBox="[151,1398,232,1928]" captionTargetId="figure-156@15.[151,1398,232,1940]" captionTargetPageId="15" captionText="Figs 119–143. Drusilla bifida (119–122), Rabdotodrusilla pectinata (123–125), Amaurodera arunica (126–127, 132), A. gilvios (128, 133), A. reticulata (129, 134), A. dentata (130, 135), A. parvoculata (131, 136), A. thailandensis (137–138), A. fasciata (139), A. meorum(140),A. disparicollis (141), A. spinans (142),andA. varicollis(143):forebody (119, 123, 126, 128–131); head and pronotum (120); antenna (121, 124, 132–143); abdomen (122, 125, 127). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6421190" httpUri="https://zenodo.org/record/6421190/files/figure.png" pageId="65" pageNumber="78">Figs 119–120</figureCitation>
) transverse, 1.21 times as broad as long; posterior angles weakly marked, nearly obsolete; frons and sclerotized portion of clypeus each with a distinct smooth elevation; punctation fine, practically invisible in the pronounced microreticulation (
<figureCitation id="66FDD1A5FFF0FFAAFD04768EDA90FC42" box="[655,744,937,961]" captionStart="Figs 359–378" captionStartId="27.[151,191,1971,1993]" captionTargetBox="[155,1396,235,1964]" captionTargetId="figure-155@27.[151,1398,232,1964]" captionText="Figs 359–378. Orphnebius laticeps (359–361), O. biimpressus (362–364), O. ulcerosus (365–369), O. falagrioides (370–372), and Drusilla bifida (373–378): tergite VIII (359, 362, 368, 370, 374); female sternite VIII (360, 363, 371); spermatheca (361, 364, 372); median lobe of aedeagus in lateral and in ventral view (365–366, 376–377); paramere (367); male sternite VIII (369, 375); posteromedian portion or head (373); apex of ventral process of aedeagus (378). Scale bars: 359–372, 374–377: 0.2 mm; 373, 378: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6421218" httpUri="https://zenodo.org/record/6421218/files/figure.png" pageId="65" pageNumber="78">Fig. 373</figureCitation>
); dorsal surface nearly matt. Eyes large and strongly bulging, nearly twice as long as distance from posterior margin of eye to posterior constriction in dorsal view. Antenna (
<figureCitation id="66FDD1A5FFF0FFAAFF14710ED88EFBC2" box="[159,246,1065,1089]" captionStart="Figs 119–143" captionStartId="15.[151,191,1969,1991]" captionTargetBox="[151,1398,232,1928]" captionTargetId="figure-156@15.[151,1398,232,1940]" captionTargetPageId="15" captionText="Figs 119–143. Drusilla bifida (119–122), Rabdotodrusilla pectinata (123–125), Amaurodera arunica (126–127, 132), A. gilvios (128, 133), A. reticulata (129, 134), A. dentata (130, 135), A. parvoculata (131, 136), A. thailandensis (137–138), A. fasciata (139), A. meorum(140),A. disparicollis (141), A. spinans (142),andA. varicollis(143):forebody (119, 123, 126, 128–131); head and pronotum (120); antenna (121, 124, 132–143); abdomen (122, 125, 127). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6421190" httpUri="https://zenodo.org/record/6421190/files/figure.png" pageId="65" pageNumber="78">Fig. 121</figureCitation>
)
<quantity id="393E60C5FFF0FFAAFE87710ED925FBC2" box="[268,349,1065,1089]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="65" pageNumber="78" unit="mm" value="1.9">1.9 mm</quantity>
long and rather massive; antennomere III somewhat flattened, distinctly dilated apically, approximately twice as long as broad, and much longer than antennomere II; antennomeres IV distinctly and V–VI weakly oblong, VII aproximately as long as broad, VIII–X indistinctly transverse, and XI small, narrower than the preceding antennomeres and shorter than the combined length of IX and X.
</paragraph>
<paragraph id="FE79CD20FFF0FFAAFF1C700DDA39F9A2" blockId="65.[149,757,585,2081]" pageId="65" pageNumber="78">
Pronotum (
<figureCitation id="66FDD1A5FFF0FFAAFE9A700ED9D9FAC2" box="[273,417,1321,1345]" captionStart="Figs 119–143" captionStartId="15.[151,191,1969,1991]" captionTargetBox="[151,1398,232,1928]" captionTargetId="figure-156@15.[151,1398,232,1940]" captionTargetPageId="15" captionText="Figs 119–143. Drusilla bifida (119–122), Rabdotodrusilla pectinata (123–125), Amaurodera arunica (126–127, 132), A. gilvios (128, 133), A. reticulata (129, 134), A. dentata (130, 135), A. parvoculata (131, 136), A. thailandensis (137–138), A. fasciata (139), A. meorum(140),A. disparicollis (141), A. spinans (142),andA. varicollis(143):forebody (119, 123, 126, 128–131); head and pronotum (120); antenna (121, 124, 132–143); abdomen (122, 125, 127). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6421190" httpUri="https://zenodo.org/record/6421190/files/figure.png" pageId="65" pageNumber="78">Figs 119–120</figureCitation>
) 1.08 times as broad as long and as broad as head, very weakly convex in cross-section, broadest at anterior angles; disc coarsely and longitudinally rugosely sculptured, matt, only in antero-median portion less matt and with defined punctation; midline with very narrow sulcus reaching neither anterior nor posterior margins; disc with an indistinct elevation and a lateral oblong impression on either side.
</paragraph>
<paragraph id="FE79CD20FFF0FFAAFF1C730ED9ADF942" blockId="65.[149,757,585,2081]" pageId="65" pageNumber="78">
Elytra (
<figureCitation id="66FDD1A5FFF0FFAAFF63730ED947F9C2" box="[232,319,1577,1601]" captionStart="Figs 119–143" captionStartId="15.[151,191,1969,1991]" captionTargetBox="[151,1398,232,1928]" captionTargetId="figure-156@15.[151,1398,232,1940]" captionTargetPageId="15" captionText="Figs 119–143. Drusilla bifida (119–122), Rabdotodrusilla pectinata (123–125), Amaurodera arunica (126–127, 132), A. gilvios (128, 133), A. reticulata (129, 134), A. dentata (130, 135), A. parvoculata (131, 136), A. thailandensis (137–138), A. fasciata (139), A. meorum(140),A. disparicollis (141), A. spinans (142),andA. varicollis(143):forebody (119, 123, 126, 128–131); head and pronotum (120); antenna (121, 124, 132–143); abdomen (122, 125, 127). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6421190" httpUri="https://zenodo.org/record/6421190/files/figure.png" pageId="65" pageNumber="78">Fig. 119</figureCitation>
) 1.07 times as long as pronotum; punctation dense and moderately fine; interstices with pronounced microreticulation (
<figureCitation id="66FDD1A5FFF0FFAAFE71734EDA2CF902" box="[506,596,1641,1665]" captionStart="Figs 379–395" captionStartId="28.[189,229,1943,1965]" captionTargetBox="[193,1428,232,1921]" captionTargetId="figure-160@28.[189,1436,232,1934]" captionTargetPageId="28" captionText="Figs 379–395. Drusilla bifida (379), Rabdotodrusilla pectinata (380–383), Amaurodera arunica (384–388), A. gilvios (389–390, 395), and A. reticulata (391–394): sutural portion of left elytron (379); median lobe of aedeagus in lateral and in ventral view (380– 381, 385–386, 389–392); male tergite VIII (382); male sternite VIII (383); male pronotum (384); spermatheca (387–388, 395); subapical portion of median lobe of aedeagus in dry preparation (393); subapical portion of median lobe of aedeagus in transparent light (394). Scale bars: 384: 0.5 mm; 379–383, 385–392, 395: 0.2 mm; 393–394: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6421220" httpUri="https://zenodo.org/record/6421220/files/figure.png" pageId="65" pageNumber="78">Fig. 379</figureCitation>
). Hind wings present. Metatarsomere I slightly longer than the combined length of II and III.
</paragraph>
<paragraph id="FE79CD20FFF0FFAAFF1C73EED908F8E2" blockId="65.[149,757,585,2081]" pageId="65" pageNumber="78">
Abdomen (
<figureCitation id="66FDD1A5FFF0FFAAFE9873EED912F962" box="[275,362,1737,1761]" captionStart="Figs 119–143" captionStartId="15.[151,191,1969,1991]" captionTargetBox="[151,1398,232,1928]" captionTargetId="figure-156@15.[151,1398,232,1940]" captionTargetPageId="15" captionText="Figs 119–143. Drusilla bifida (119–122), Rabdotodrusilla pectinata (123–125), Amaurodera arunica (126–127, 132), A. gilvios (128, 133), A. reticulata (129, 134), A. dentata (130, 135), A. parvoculata (131, 136), A. thailandensis (137–138), A. fasciata (139), A. meorum(140),A. disparicollis (141), A. spinans (142),andA. varicollis(143):forebody (119, 123, 126, 128–131); head and pronotum (120); antenna (121, 124, 132–143); abdomen (122, 125, 127). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6421190" httpUri="https://zenodo.org/record/6421190/files/figure.png" pageId="65" pageNumber="78">Fig. 121</figureCitation>
) slightly narrower than elytra; sternites III–VI anteriorly with dense and posteriorly with sparser fine punctation; tergite VII with very fine and very sparse punctation; posterior margin of tergite VII with palisade fringe.
</paragraph>
<paragraph id="FE79CD20FFF0FFAAFF1C724DDBD0FE82" blockId="65.[149,757,585,2081]" lastBlockId="65.[794,936,233,257]" pageId="65" pageNumber="78">
<specimenCount id="E8C006A9FFF0FFAAFF1C724DD8D4F802" box="[151,172,1898,1921]" pageId="65" pageNumber="78" type="male">♂</specimenCount>
: tergite VIII (
<figureCitation id="66FDD1A5FFF0FFAAFEC7724ED9DCF802" box="[332,420,1897,1921]" captionStart="Figs 359–378" captionStartId="27.[151,191,1971,1993]" captionTargetBox="[155,1396,235,1964]" captionTargetId="figure-155@27.[151,1398,232,1964]" captionText="Figs 359–378. Orphnebius laticeps (359–361), O. biimpressus (362–364), O. ulcerosus (365–369), O. falagrioides (370–372), and Drusilla bifida (373–378): tergite VIII (359, 362, 368, 370, 374); female sternite VIII (360, 363, 371); spermatheca (361, 364, 372); median lobe of aedeagus in lateral and in ventral view (365–366, 376–377); paramere (367); male sternite VIII (369, 375); posteromedian portion or head (373); apex of ventral process of aedeagus (378). Scale bars: 359–372, 374–377: 0.2 mm; 373, 378: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6421218" httpUri="https://zenodo.org/record/6421218/files/figure.png" pageId="65" pageNumber="78">Fig. 374</figureCitation>
) transverse, posterior margin with distinct and broad concavity in the middle; sternite VIII (
<figureCitation id="66FDD1A5FFF0FFAAFE89728ED922F842" box="[258,346,1961,1985]" captionStart="Figs 359–378" captionStartId="27.[151,191,1971,1993]" captionTargetBox="[155,1396,235,1964]" captionTargetId="figure-155@27.[151,1398,232,1964]" captionText="Figs 359–378. Orphnebius laticeps (359–361), O. biimpressus (362–364), O. ulcerosus (365–369), O. falagrioides (370–372), and Drusilla bifida (373–378): tergite VIII (359, 362, 368, 370, 374); female sternite VIII (360, 363, 371); spermatheca (361, 364, 372); median lobe of aedeagus in lateral and in ventral view (365–366, 376–377); paramere (367); male sternite VIII (369, 375); posteromedian portion or head (373); apex of ventral process of aedeagus (378). Scale bars: 359–372, 374–377: 0.2 mm; 373, 378: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6421218" httpUri="https://zenodo.org/record/6421218/files/figure.png" pageId="65" pageNumber="78">Fig. 375</figureCitation>
) transverse and with convex posterior margin; median lobe of aedeagus (
<figureCitation id="66FDD1A5FFF0FFAAFE7072EEDAF2F862" box="[507,650,1993,2017]" captionStart="Figs 359–378" captionStartId="27.[151,191,1971,1993]" captionTargetBox="[155,1396,235,1964]" captionTargetId="figure-155@27.[151,1398,232,1964]" captionText="Figs 359–378. Orphnebius laticeps (359–361), O. biimpressus (362–364), O. ulcerosus (365–369), O. falagrioides (370–372), and Drusilla bifida (373–378): tergite VIII (359, 362, 368, 370, 374); female sternite VIII (360, 363, 371); spermatheca (361, 364, 372); median lobe of aedeagus in lateral and in ventral view (365–366, 376–377); paramere (367); male sternite VIII (369, 375); posteromedian portion or head (373); apex of ventral process of aedeagus (378). Scale bars: 359–372, 374–377: 0.2 mm; 373, 378: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6421218" httpUri="https://zenodo.org/record/6421218/files/figure.png" pageId="65" pageNumber="78">Figs 376–377</figureCitation>
)
<quantity id="393E60C5FFF0FFAAFD1D72EEDA8CF862" box="[662,756,1993,2017]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="65" pageNumber="78" unit="mm" value="0.53">0.53 mm</quantity>
long; ventral process apically bifid (
<figureCitation id="66FDD1A5FFF0FFAAFDA672CEDAF0F782" box="[557,648,2025,2049]" captionStart="Figs 359–378" captionStartId="27.[151,191,1971,1993]" captionTargetBox="[155,1396,235,1964]" captionTargetId="figure-155@27.[151,1398,232,1964]" captionText="Figs 359–378. Orphnebius laticeps (359–361), O. biimpressus (362–364), O. ulcerosus (365–369), O. falagrioides (370–372), and Drusilla bifida (373–378): tergite VIII (359, 362, 368, 370, 374); female sternite VIII (360, 363, 371); spermatheca (361, 364, 372); median lobe of aedeagus in lateral and in ventral view (365–366, 376–377); paramere (367); male sternite VIII (369, 375); posteromedian portion or head (373); apex of ventral process of aedeagus (378). Scale bars: 359–372, 374–377: 0.2 mm; 373, 378: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6421218" httpUri="https://zenodo.org/record/6421218/files/figure.png" pageId="65" pageNumber="78">Fig. 378</figureCitation>
); internal structures of distinctive shapes; paramere
<quantity id="393E60C5FFF0FFAAFDDE7D2EDACCF7A2" box="[597,692,2057,2081]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" pageId="65" pageNumber="78" unit="mm" value="0.47">0.47 mm</quantity>
long.
<specimenCount id="E8C006A9FFF0FFAAFC9175CDDB57FE82" box="[794,815,234,257]" pageId="65" pageNumber="78" type="female">♀</specimenCount>
: unknown.
</paragraph>
<paragraph id="FE79CD20FFF0FFAAFC91740EDD4BFD62" blockId="65.[791,1400,297,737]" pageId="65" pageNumber="78">
<emphasis id="CCB21132FFF0FFAAFC91740EDCD8FEC2" bold="true" box="[794,1184,297,321]" pageId="65" pageNumber="78">Comparative notes and comment</emphasis>
:
<taxonomicName id="39C6B6A3FFF0FFAAFB3E740EDD2FFEC2" authorityName="Assing" authorityYear="2016" box="[1205,1367,297,321]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="78" phylum="Arthropoda" rank="species" species="bifida">
<emphasis id="CCB21132FFF0FFAAFB3E740EDD2FFEC2" box="[1205,1367,297,321]" italics="true" pageId="65" pageNumber="78">Drusilla bifida</emphasis>
</taxonomicName>
is distinguished from other species recorded from
<collectingCountry id="86D18DB0FFF0FFAAFA9C746EDD0FFEE2" box="[1303,1399,329,353]" name="Thailand" pageId="65" pageNumber="78">Thailand</collectingCountry>
and neighbouring regions by the bifid ventral process of the aedeagus, the characteristic sculpture of the pronotum, and the smooth elevations on the frons and the clypeus, from most species also by the coloration, the massive antennae, and the pronounced microsculpture on the head and elytra. The median lobe of the aedeagus slightly resembles that of the much larger
<taxonomicName id="39C6B6A3FFF0FFAAFB10770DDB35FDE2" authority="PACE, 2005" authorityName="PACE" authorityYear="2005" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="78" phylum="Arthropoda" rank="species" species="umranicola">
<emphasis id="CCB21132FFF0FFAAFB10770DDD48FDC2" box="[1179,1328,553,577]" italics="true" pageId="65" pageNumber="78">D. umranicola</emphasis>
PACE, 2005
</taxonomicName>
from
<collectingCountry id="86D18DB0FFF0FFAAFC06776EDBBDFDE2" box="[909,965,585,609]" name="India" pageId="65" pageNumber="78">India</collectingCountry>
(
<collectingRegion id="3C0203C2FFF0FFAAFC5A776EDC30FDE2" box="[977,1096,585,609]" country="India" name="Meghalaya" pageId="65" pageNumber="78">Meghalaya</collectingRegion>
) and that of
<taxonomicName id="39C6B6A3FFF0FFAAFB47776DDC71FD02" authority="PACE, 2012" authorityName="PACE" authorityYear="2012" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="78" phylum="Arthropoda" rank="species" species="thaifuscicollis">
<emphasis id="CCB21132FFF0FFAAFB47776DDD0FFDE2" box="[1228,1399,585,609]" italics="true" pageId="65" pageNumber="78">D. thaifuscicollis</emphasis>
PACE, 2012 (
<collectingCountry id="86D18DB0FFF0FFAAFC16774EDC7AFD02" box="[925,1026,617,641]" name="Thailand" pageId="65" pageNumber="78">Thailand</collectingCountry>
)
</taxonomicName>
, from which
<taxonomicName id="39C6B6A3FFF0FFAAFB1C774DDC8AFD02" authorityName="Assing" authorityYear="2016" box="[1175,1266,617,641]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="78" phylum="Arthropoda" rank="species" species="bifida">
<emphasis id="CCB21132FFF0FFAAFB1C774DDC8AFD02" box="[1175,1266,617,641]" italics="true" pageId="65" pageNumber="78">D. bifida</emphasis>
</taxonomicName>
additionally differs by the nearly obsolete posterior angles of the head, much larger eyes, and darker and distinctly more massive antennae with a relatively shorter antennomere XI.
</paragraph>
</subSubSection>
<subSubSection id="B6DC9EABFFF0FFAAFC91762EDC89FC02" pageId="65" pageNumber="78" type="distribution">
<paragraph id="FE79CD20FFF0FFAAFC91762EDC89FC02" blockId="65.[794,1399,777,897]" pageId="65" pageNumber="78">
<emphasis id="CCB21132FFF0FFAAFC91762EDCF4FCA2" bold="true" box="[794,1164,777,801]" pageId="65" pageNumber="78">Distribution and natural history</emphasis>
: The type locality is situated in the extreme north of
<collectingCountry id="86D18DB0FFF0FFAAFB0C760EDC91FCC2" box="[1159,1257,809,833]" name="Thailand" pageId="65" pageNumber="78">Thailand</collectingCountry>
, close to the border with
<collectingCountry id="86D18DB0FFF0FFAAFC15766DDC70FCE1" box="[926,1032,842,866]" name="Myanmar" pageId="65" pageNumber="78">Myanmar</collectingCountry>
. The
<typeStatus id="217D7382FFF0FFAAFBC9766EDCD8FCE2" box="[1090,1184,841,865]" pageId="65" pageNumber="78" type="holotype">holotype</typeStatus>
was sifted from leaf litter near a waterfall at an altitude of
<specimenCount id="E8C006A9FFF0FFAAFB23764EDC89FC02" box="[1192,1265,873,897]" pageId="65" pageNumber="78" type="generic">
<quantity id="393E60C5FFF0FFAAFB23764EDC94FC02" box="[1192,1260,873,897]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="65" pageNumber="78" unit="m" value="800.0">800 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
</treatment>
</document>
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