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<mods:title id="A3986B83FEAF541C475BFE382BEEFADE">Rhantus fengi sp. n. from Xizang, China, and notes on Laccoporus nigritulus (Gschwendtner) (Coleoptera, Dytiscidae)</mods:title>
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<mods:namePart id="61D9E48B0C88EA8B2AF4F936B7BC3B05">Zhao, Shuang</mods:namePart>
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<mods:namePart id="2FAE63E593F7C40E17BDAFA3A801A558">Jia, Fenglong</mods:namePart>
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<mods:namePart id="4C2ECB35155F103F61CE68B8FD5FC7FC">Balke, Michael</mods:namePart>
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<treatment id="58DFEF222735FD2714C1FCFF8E4D5FFE" ID-GBIF-Taxon="159365013" LSID="urn:lsid:plazi:treatment:58DFEF222735FD2714C1FCFF8E4D5FFE" httpUri="http://treatment.plazi.org/id/58DFEF222735FD2714C1FCFF8E4D5FFE" lastPageId="7" lastPageNumber="67" pageId="4" pageNumber="65" scope_family="Dytiscidae" scope_order="Coleoptera">
<subSubSection id="B10BB3530417B8B7581D0AA0C04D7FEE" pageId="4" pageNumber="65" type="nomenclature">
<paragraph id="2B44716F85545C4DFA3B2C26D2B06011" pageId="4" pageNumber="65">
<taxonomicName id="6B6EEAB184C6C5A2DB66E8F245035082" ID-CoL="3RMD5" LSID="http://www.species-id.net/wiki/Laccoporus_nigritulus" authority="Gschwendtner, 1936" authorityName="Gschwendtner" authorityYear="1936" class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus nigritulus" order="Coleoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="nigritulus">Laccoporus nigritulus (Gschwendtner, 1936)</taxonomicName>
Figs 10-15
</paragraph>
</subSubSection>
<subSubSection id="749FA532F8DDB2E21A8C191F572E7356" pageId="4" pageNumber="65" type="reference_group">
<paragraph id="3B72DC2A81464907FC1F8F28F3ADA778" pageId="4" pageNumber="65">
<taxonomicName id="F0048FBFEF2155C12D9329A53A8118F4" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus apicicornis" order="Coleoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="apicicornis">Laccophilus apicicornis</taxonomicName>
var. nigritulus
<bibRefCitation id="CD8A708D89E3F2E581839E5B53ECCE49" author="Gschwendtner, L" journalOrPublisher="Records of the Indian Museum" pageId="7" pageNumber="68" pagination="365 - 374" title="Interessante und neue Schwimmkaefer des Indischen Museums in Calcutta" volume="37" year="1936">Gschwendtner 1936</bibRefCitation>
: 367.
</paragraph>
<paragraph id="5179063E35A4BAB41B314ABF04580BE4" pageId="4" pageNumber="65">
<taxonomicName id="21AA89F35451C955D600A04C8A76DC24" class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus nigritulus" order="Coleoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="nigritulus">Laccoporus nigritulus</taxonomicName>
(Gschwendtner):
<bibRefCitation id="067277C9860752A3CB6896728F1CE235" author="Vazirani, TG" journalOrPublisher="Current Science" pageId="7" pageNumber="68" pagination="563 - 564" title="On Laccoporus nigritulus (Gschwendtner) comb. n. (Coleoptera: Dytiscidae)." volume="39" year="1970">Vazirani 1970</bibRefCitation>
: 563-564 (new combination, new status);
<bibRefCitation id="DDB2C46D7DA1894F7C5FAF12FBCFD4B9" author="Nilsson, AN" journalOrPublisher="Apollo Books, Stenstrup" pageId="7" pageNumber="68" title="Dytiscidae. - World Catalogue of Insects. Vol. 3" year="2001">Nilsson 2001</bibRefCitation>
: 253.
</paragraph>
<paragraph id="ABD7E3C75E87D45B63305A96CD013D4D" pageId="4" pageNumber="65">
<taxonomicName id="2FCE0DB958A28FE0F2D2C897350859DB" class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus viator" order="Coleoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="viator">Laccoporus viator</taxonomicName>
<bibRefCitation id="9EF06DEC778BD62F48A37216A4C73CC5" author="Balfour-Browne, J" journalOrPublisher="The Annals and Magazine of Natural History (11)" pageId="7" pageNumber="68" pagination="97 - 114" title="A contribution to the study of the Dytiscidae. - I. (Coleoptera, Adephaga)." volume="3" year="1939">Balfour-Browne 1939</bibRefCitation>
: 104;
<bibRefCitation id="6DDD2BBBD9F10E4315A6EA508F0888DD" author="Nilsson, AN" journalOrPublisher="Apollo Books, Stenstrup" pageId="7" pageNumber="68" title="Dytiscidae. - World Catalogue of Insects. Vol. 3" year="2001">Nilsson 2001</bibRefCitation>
: 253; new synonymy.
</paragraph>
<paragraph id="4B4F772AC6293C8A8429C321FB8D663E" pageId="4" pageNumber="65">
<taxonomicName id="B1DE07F6510052FD1EE891FB3ED78CE7" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus" order="Coleoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="genus">Laccophilus</taxonomicName>
sp.:
<bibRefCitation id="B5BABBB48BB2FD90FE9E626F6B98823E" author="Nilsson, AN" editor="Jaech, MA" journalOrPublisher="Zoologisch-Botanische Gesellschaft in Oesterreich and Wiener Coleopterologenverein, Wien" pageId="7" pageNumber="68" pagination="35 - 96" title="Noteridae and Dytiscidae: Annotated checklist of the Noteridae and Dytiscidae of China (Coleoptera)" volumeTitle="Water beetles of China, Vol. 1" year="1995">Nilsson 1995</bibRefCitation>
: 71.
</paragraph>
</subSubSection>
<subSubSection id="F35840086E22A7F5C30F235352E66654" pageId="4" pageNumber="65" type="type localities">
<paragraph id="C414F5F0F23DB9B31C7FDB712819EE0C" pageId="4" pageNumber="65">Type localities:</paragraph>
<paragraph id="1DC57B2015944591EAA8B2D01D5C670B" pageId="4" pageNumber="65">
<taxonomicName id="C2478677B62EA95236151D918DFDE284" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus apicicornis" order="Coleoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="apicicornis">Laccophilus apicicornis</taxonomicName>
var. nigritulus: China: Tibet, Te-ring Gompa;
<taxonomicName id="A3FB047F7F86DACC8901B55D8B5FE3D5" class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus viator" order="Coleoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="viator">Laccoporus viator</taxonomicName>
: China: Tibet, Gyangtse.
</paragraph>
</subSubSection>
<subSubSection id="A432A10D5CC45F4C6C9B8E6BE1B42FF7" lastPageId="5" lastPageNumber="66" pageId="4" pageNumber="65" type="type material">
<paragraph id="10982AEB25262A530FCE1515E95B9AC0" pageId="4" pageNumber="65">Type material:</paragraph>
<paragraph id="848B7E9B779FE0A25C6BC105B9FCEF02" pageId="4" pageNumber="65">
<taxonomicName id="136AE52CFAAB3C9EFFD82AF894514E70" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus apicicornis" order="Coleoptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="apicicornis">Laccophilus apicicornis</taxonomicName>
var. nigritulus: 10 syntypes, Te-ring Gompa. 14,000 ft. Tibet (F.H. Stewart) in ZSIC [not studied].
<bibRefCitation id="E0D21427754F910A2C27520C97A9A667" author="Nilsson, AN" journalOrPublisher="Apollo Books, Stenstrup" pageId="7" pageNumber="68" title="Dytiscidae. - World Catalogue of Insects. Vol. 3" year="2001">Nilsson (2001)</bibRefCitation>
mentioned a lectotype designation by
<bibRefCitation id="8BD93B87765CA70B16A61DAB181C167D" author="Vazirani, TG" journalOrPublisher="Current Science" pageId="7" pageNumber="68" pagination="563 - 564" title="On Laccoporus nigritulus (Gschwendtner) comb. n. (Coleoptera: Dytiscidae)." volume="39" year="1970">Vazirani (1970)</bibRefCitation>
, but Vazirani mentioned that he studied the holotype and one paratype, the type status is therefore rather unclear and we could not access types from ZSIC to clarify the situation.
</paragraph>
<paragraph id="1234CED17ED7748E78E369092A7265EC" pageId="5" pageNumber="66">
<taxonomicName id="1054F635E49E67F5992425303AC0E4CE" class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus viator" order="Coleoptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="viator">
<pageBreakToken id="8379D4CCE23E46239F505EA3C2B9562F" pageId="5" pageNumber="66" start="start">Laccoporus</pageBreakToken>
viator
</taxonomicName>
: Holotype,
<normalizedToken id="72B7C3D2FADC3B62BFFFD9A16EBD3822" originalValue="„allotype”">&quot;allotype&quot;</normalizedToken>
and 10 other paratypes in BMNH. We studied 2 ♂♂ paratypes labelled:
<normalizedToken id="C546F7E52FBECE133452875CAB823230" originalValue="„Co-type”">&quot;Co-type&quot;</normalizedToken>
, &quot;Gyangtse. / 13,000 ft. / June 1904. / Tibet Expedit. / H.J. Walton. / 1903
<normalizedToken id="A0F46FA38F5F80A8ADE16321A817A99A" originalValue="–172.”">-172.&quot;</normalizedToken>
, &quot;Laccoporus / viator. B-B. / Co-type.&quot;.
</paragraph>
</subSubSection>
<subSubSection id="72B610E288FCD7D33B51339C74B0BFF0" pageId="5" pageNumber="66" type="other material examined">
<paragraph id="23EB4B598A15E346E1E407D170543D82" pageId="5" pageNumber="66">Other material examined:</paragraph>
<paragraph id="C1B373200C2F2F4B24E682D756193272" pageId="5" pageNumber="66">
20 exs (SYSU): 2 ♂♂CHINA: Xizang, Dingri (Tering), 4300 m, 2.vi.1974, leg. Xuezhong Zhang, Academia Sinica; 1 ♀ the same data as male; 1 ♀ Xizang, suburb of Rikaze, 3826 m, 20-23.vii.1986, leg. Geqiu Liang; 5 ♂♂, 3 ♀♀ Xizang, Rikaze, 3862 m, 4.viii.1986, leg. Geqiu Liang; 1 ♂ the same data as the former, with a label &quot;Laccophilus indicus?&quot;; 4 ♂♂, 3 ♀♀ Xizang, Mozhugongka County, 7.viii.1986, leg. Geqiu Liang; 8 exs (MTD, NMPC, CHF) &quot;Tibet, Yamtso-ufer bei / Nagartze, N28°58'31,9 / E90°24'6,0; 4450mNN; / 29.VII. 1998; leg. O.
<normalizedToken id="ABC8A2AF240831BC75CE51D12C7B1F42" originalValue="Jäger”">Jaeger&quot;</normalizedToken>
, &quot;Laccoporus nigritulus / (Gschwendtner) / Fery det. 1999&quot;; 11 exs (MTD, CHF) &quot;Tibet, Tingriebene / N28°34'39,7/E86°36'52,7; 4400 m / 3.-5.VIII. 1998, lg.
<normalizedToken id="CA5038E5BB6A6915D52E7CA66665B267" originalValue="Jäger“">Jaeger&quot;</normalizedToken>
, &quot;Laccoporus nigritulus / (Gschwendtner) / Fery det. 1999&quot;.
</paragraph>
</subSubSection>
<subSubSection id="974421BC02018C91EC0726184E5B79C8" pageId="5" pageNumber="66" type="redescription">
<paragraph id="47456DE7B72C225493028754592E6F3F" pageId="5" pageNumber="66">Redescription.</paragraph>
<paragraph id="A295E13E88487B5B323A7CC47BEDEF4B" pageId="5" pageNumber="66">
Measurements. Length 4.5-5.0 mm, width 2.5-2.6 mm (paratypes of
<taxonomicName id="040E7E0DFB2A104B96D4C7C7A16FAAF7" class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus viator" order="Coleoptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="viator">Laccoporus viator</taxonomicName>
: length 4.5-4.8 mm, width 2.3-2.6 mm).
</paragraph>
</subSubSection>
<subSubSection id="56300221AC574C4EAF420B2541FE6C97" lastPageId="6" lastPageNumber="67" pageId="5" pageNumber="66" type="colouration">
<paragraph id="27CAD3BFC6AF490C714219E627F52A4B" pageId="5" pageNumber="66">Colouration</paragraph>
<paragraph id="7492459CFA4EBF007E52EBC2E61F0357" lastPageId="6" lastPageNumber="67" pageId="5" pageNumber="66">
(Figs 10, 11, 15). Dorsum uniformly yellow brown. Head yellowish brown, with one short dark brown line near antennae. Antenna with antennomeres 1-4 and base of antennomeres 5-6 yellow brown, antennomeres 7-11 andapical parts of antennomeres 5-6 dark brown. Maxillary palpomeres yellow brown, apical palpomere apically dark. Labial palpomeres yellow brown, apical palpomere dark. Pronotum sometimes paler with darker median, anterior and posterior transverse bands. Elytra clearly darker than pronotum, but with the same color at base as pronotum. Ventral surface of head and thorax black. Abdomen black, ventrites 3-6 yellow brown along posterior margins (e.g. Fig. 15). Legs yellow brown with coxae black, posterior half of metacoxal process yellowstructures(see http://www.species-id.net/wiki/Laccoporus_nigritulus for high resolution images of surfaces). Head with surface sculpture consisting of small irregular polygonal meshes, with some sparse coarse and shallow punctures in posterior half. Meshes somewhat stronger on posterior portion. Clypeus with transverse series of large punctures present anteriorly. Labrum with very deep excavation, surface with short transverse meshes, anterior notch with dense and short white setae. Antenna filiform, not dilated. Pronotum more or less depressed laterally, without lateral bead, with small irregular polygonal meshes, sometimes very tiny punctures detectable. Posterior corner almost rectangular. Elytra with base as wide as posterior portion of pronotum, surface with irregular polygonal meshes in size and shape, sculpture more or less stronger than that on pronotum, tiny punctures very sparse and clearly detectable. Prosternal process thin and long,bisinuate in lateral view, almost reaching posterior margin of mesocoxae. Metaventrite medially with some coarse and strong punctures behind mesocoxae. Metacoxae and metasternum strongly and irregularly scratched, with dense longitudinal sculptures and sparse tiny punctures. Pro- and mesofemora with tuft of long setae at basal portion posteriorly. Metafemora with fine and dense sculpture and tiny punctures. Metatibia with inner spur longer than outer one. Pro- and mesotarsi smooth. Metatarsi smooth, with fine sculpture dorsally. First tarsomere of metatarsi almost as long as following two tarsomeres combined. Claws
<pageBreakToken id="06B41EFC279AE7110B96C41CD4BC8554" pageId="6" pageNumber="67" start="start">normally</pageBreakToken>
curved, protarsal claws longer than metatarsal claws. Metatarsomere 1 almost as long as metatarsomeres 2-3 combined. Claws regularly curved, proclaws longer than metaclaws.
</paragraph>
<caption id="965094D4338C4A92D236589AAA48D62A" pageId="6" pageNumber="67">
<paragraph id="0BF3210FBA2BBE870D29044A04EB9FEE" pageId="6" pageNumber="67">
Figures 10-15.
<taxonomicName id="8AFDE4D894360747C97D545C79A4DB82" class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus nigritulus" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="nigritulus">Laccoporus nigritulus</taxonomicName>
male 10 habitus, dorsal view 11 habitus, ventral view 12 left paramere 13 right paramere 14 median lobe of aedeagus, lateral view 15 ventral view of 6th ventrite.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="29028309650CE16C0E097B79FE22D715" pageId="6" pageNumber="67" type="male">
<paragraph id="77179F9AA5C06D1C9F9F789FC2581578" pageId="6" pageNumber="67">Male.</paragraph>
<paragraph id="421A713AE26138A466A6FE9CCEBFE33B" pageId="6" pageNumber="67">Protarsus not dilated. Median lobe and parameres as in Figs 12-14.</paragraph>
</subSubSection>
<subSubSection id="2EED11E81E7B6765BB2F691CFC4EBEAD" pageId="6" pageNumber="67" type="distribution">
<paragraph id="179F7BAD95CAB2191C7C4C451B2AEFA4" pageId="6" pageNumber="67">Distribution.</paragraph>
<paragraph id="4E8F5E664E1D3D6818FC6BEC5D02EC20" pageId="6" pageNumber="67">China: Xizang.</paragraph>
</subSubSection>
<subSubSection id="2C853BA12F7383D26BBA2760582366BF" lastPageId="7" lastPageNumber="68" pageId="6" pageNumber="67" type="remarks">
<paragraph id="8EB4758C974ACFF19441A6BF38462407" pageId="6" pageNumber="67">Remarks.</paragraph>
<paragraph id="29BD3E009C1D040B6DF156EDF205F381" pageId="6" pageNumber="67">
Genus
<taxonomicName id="D6A292997620ABF48BCD81148E1169AB" class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="genus">Laccoporus</taxonomicName>
was erected by
<bibRefCitation id="3EA2B2E0C22596525DB80ABCE11537BE" author="Balfour-Browne, J" journalOrPublisher="The Annals and Magazine of Natural History (11)" pageId="7" pageNumber="68" pagination="97 - 114" title="A contribution to the study of the Dytiscidae. - I. (Coleoptera, Adephaga)." volume="3" year="1939">Balfour-Browne (1939)</bibRefCitation>
with
<taxonomicName id="8B1DBD9393BE7221642CAB78454A3E43" class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus viator" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="viator">Laccoporus viator</taxonomicName>
Balfour-Browne, 1939 as the type species. This author had no knowledge about
<taxonomicName id="FE5B61F68B902DA1F75D25242AC9B785" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus apicornis" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="apicornis">Laccophilus apicornis</taxonomicName>
var. nigritulus Gschwendtner, 1936. The latter taxon was transferred to
<taxonomicName id="9A50D23F66D3138166D64BBB938B16CC" class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="genus">Laccoporus</taxonomicName>
and raised to species status by
<bibRefCitation id="3A4E6413BD15E3767689E9BED0177978" author="Vazirani, TG" journalOrPublisher="Current Science" pageId="7" pageNumber="68" pagination="563 - 564" title="On Laccoporus nigritulus (Gschwendtner) comb. n. (Coleoptera: Dytiscidae)." volume="39" year="1970">Vazirani (1970)</bibRefCitation>
who studied two syntypes housed in ZSIC.
<bibRefCitation id="C4B28C19E122630E7E50CE2F11F2C6E3" author="Vazirani, TG" journalOrPublisher="Current Science" pageId="7" pageNumber="68" pagination="563 - 564" title="On Laccoporus nigritulus (Gschwendtner) comb. n. (Coleoptera: Dytiscidae)." volume="39" year="1970">Vazirani (1970)</bibRefCitation>
had doubts that
<taxonomicName id="5982219F1A2923EEB0EB9121A8B7DC25" class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus viator" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="viator">Laccoporus viator</taxonomicName>
and
<taxonomicName id="A2F81753AA117C504F60DBDDFEE8609F" class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus nigritulus" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="nigritulus">Laccoporus nigritulus</taxonomicName>
were different species, but he could not study
<normalizedToken id="9483515D32EFDFFDB6D0A7FE489ED3FB" originalValue="Balfour-Browne’s">Balfour-Browne's</normalizedToken>
types. We have no access to specimens in ZSIC, but
<bibRefCitation id="AD0F16125605EEE1CDE54198AB732064" author="Vazirani, TG" journalOrPublisher="Current Science" pageId="7" pageNumber="68" pagination="563 - 564" title="On Laccoporus nigritulus (Gschwendtner) comb. n. (Coleoptera: Dytiscidae)." volume="39" year="1970">Vazirani (1970)</bibRefCitation>
provided an illustration of the ventral side of the median lobe of
<taxonomicName id="E8F7A6C866307EF22F2EF5C12CEC096E" class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus nigritulus" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="nigritulus">Laccoporus nigritulus</taxonomicName>
, which agrees with our specimens.
</paragraph>
<paragraph id="4185D86B51B2ADF81B35ECE0237228AD" pageId="6" pageNumber="67">
The types of
<taxonomicName id="65938EFD8BC31F62767535081E5B43E9" class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus viator" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="viator">Laccoporus viator</taxonomicName>
are from Gyangtse (Jiangzi). The types of
<taxonomicName id="AD57E8B127131FC97E9C92E1527DED74" class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus nigritulus" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="nigritulus">Laccoporus nigritulus</taxonomicName>
were collected from
<normalizedToken id="02C306FAEF7C15FD2EB2489D4F4FF4E0" originalValue="“Te-ring”">&quot;Te-ring&quot;</normalizedToken>
(Dingri) and
<normalizedToken id="83E8F623831719D3E37983E4593631AD" originalValue="“Gompa”">&quot;Gompa&quot;</normalizedToken>
(Gangba). The distance between these sites is only about 100-120 km (Fig. 16). The two latter sites are not far (about 90-120 km respectively) from Rikaze where some of our newly studied specimens originate from. The altiude of these three sites is 3900-4300 m.
</paragraph>
<caption id="3FA9661644508C67B1AAB042CE3D2EB5" pageId="6" pageNumber="67">
<paragraph id="5C44C26580F0D7F2048C6964004A2683" pageId="6" pageNumber="67">
Figure 16. Xizang Autonomous Region, collecting localities of
<taxonomicName id="EFF5A8D3DB2229BC2E2505A3A60D3858" class="Insecta" family="Dytiscidae" genus="Rhantus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhantus fengi" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="fengi">Rhantus fengi</taxonomicName>
and
<taxonomicName id="D7C3B402BFFED4320BCE68D576CBA953" class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus nigritulus" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="nigritulus">Laccoporus nigritulus</taxonomicName>
and notable geographic localities for orientation. Dingri = Te-ring (type locality of
<taxonomicName id="7803648738DB9FAA644C8A392083C814" class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus nigritulus" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="nigritulus">Laccoporus nigritulus</taxonomicName>
); Gangba = Gompa; Jiangzi = Gyangtse (type locality of
<taxonomicName id="8C6AFB766CEAA6848E31BDF46B73FC66" class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus viator" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="viator">Laccoporus viator</taxonomicName>
).
</paragraph>
</caption>
<paragraph id="560F6615EBF6BFA2BEBF9E27995E5BC3" lastPageId="7" lastPageNumber="68" pageId="6" pageNumber="67">
<bibRefCitation id="8C5CA7977763B4350630A096557F9FA4" author="Zeng, H" journalOrPublisher="Department of Biology, Sun Yat-sen University, Guangzhou" pageId="7" pageNumber="68" title="Taxonomy of Chinese Dytiscidae in the museums of China" year="1989">Zeng (1989)</bibRefCitation>
described &quot;
<taxonomicName id="B5F96DEB3E58BDD83E7FA5E2CA309AAE" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus thibetanus" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="thibetanus">Laccophilus thibetanus</taxonomicName>
sp. n.&quot; from Rikaze, Xizang in her PhD thesis (unpublished, in Chinese) (in the corrected thesis version, the name was
<pageBreakToken id="CEADBEA4C2250A05719C0F5CB14789AF" pageId="7" pageNumber="68" start="start">changed</pageBreakToken>
to
<taxonomicName id="5F2AE8F5D7D63FDB7C1928667426DD6B" class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus xizangensis" order="Coleoptera" pageId="7" pageNumber="68" phylum="Arthropoda" rank="species" species="xizangensis">Laccoporus xizangensis</taxonomicName>
sp. n. by herself, possibly acknowledging that Xizang is a more precise reference to the type locality than
<normalizedToken id="48594C74D16F59371A88A4E61E7DF95E" originalValue="“Tibet”">&quot;Tibet&quot;</normalizedToken>
).
<bibRefCitation id="6BD529BB7C004539F149F06CB0805A44" author="Nilsson, AN" editor="Jaech, MA" journalOrPublisher="Zoologisch-Botanische Gesellschaft in Oesterreich and Wiener Coleopterologenverein, Wien" pageId="7" pageNumber="68" pagination="35 - 96" title="Noteridae and Dytiscidae: Annotated checklist of the Noteridae and Dytiscidae of China (Coleoptera)" volumeTitle="Water beetles of China, Vol. 1" year="1995">Nilsson (1995: 71)</bibRefCitation>
recorded a &quot;
<taxonomicName id="DF7EAEA16DBFC03085D29272B6D1AAB2" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus" order="Coleoptera" pageId="7" pageNumber="68" phylum="Arthropoda" rank="genus">Laccophilus</taxonomicName>
sp.&quot; and stated that it is based on &quot;
<taxonomicName id="570FE554C5E611BE85B5172ED78BC18F" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus tibetanus" order="Coleoptera" pageId="7" pageNumber="68" phylum="Arthropoda" rank="species" species="tibetanus">Laccophilus tibetanus</taxonomicName>
Zeng, 1989: 4 (nom. nud., Tibet)&quot;. We found the specimens that have the same locality data as recorded by
<bibRefCitation id="732D43CFFF68C6ED1ACC1EC393F4A93A" author="Zeng, H" journalOrPublisher="Department of Biology, Sun Yat-sen University, Guangzhou" pageId="7" pageNumber="68" title="Taxonomy of Chinese Dytiscidae in the museums of China" year="1989">Zeng (1989)</bibRefCitation>
in SYSU. Although there is no specific name and no type designation attached to these specimens, we are confident that
<bibRefCitation id="14CDF6B925F1BD3968B4AF0307BC9C89" author="Zeng, H" journalOrPublisher="Department of Biology, Sun Yat-sen University, Guangzhou" pageId="7" pageNumber="68" title="Taxonomy of Chinese Dytiscidae in the museums of China" year="1989">Zeng (1989)</bibRefCitation>
based her unpublished
<taxonomicName id="29BB22D41FB57751C336634E20488695" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus xizangensis" order="Coleoptera" pageId="7" pageNumber="68" phylum="Arthropoda" rank="species" species="xizangensis">Laccophilus xizangensis</taxonomicName>
on these specimens. All the specimens are conspecific with
<taxonomicName id="EE1CA68547715F2984198B0176A84651" class="Insecta" family="Dytiscidae" genus="Laccoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccoporus nigritulus" order="Coleoptera" pageId="7" pageNumber="68" phylum="Arthropoda" rank="species" species="nigritulus">Laccoporus nigritulus</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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