271 lines
35 KiB
XML
271 lines
35 KiB
XML
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<mods:title id="3831CF40B5E862F910225F3888F973DC">Grebennikovius, a new genus to accommodate Epactoides basilewskyi (Balthasar, 1960) (Coleoptera: Scarabaeidae: Deltochilini)</mods:title>
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<mods:namePart id="04239867DEB8A5EF9F33E97DDC7A4EC8">Mlambo, Sukoluhle</mods:namePart>
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<mods:namePart id="D3E888121004A643F6B120268F7B1999">Scholtz, Clarke H.</mods:namePart>
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<mods:namePart id="99C6365FF296D974C9A754A0F77D9F39">Deschodt, Christian M.</mods:namePart>
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<mods:title id="4B37FC2804729662215976AD95B84D0F">Zootaxa</mods:title>
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<mods:part id="A87EE5551E8F07393FEC26C88120C4E4">
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<mods:date id="969F0A556BDEB084C245EDCFFAA02B4C">2019</mods:date>
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<treatment id="7A59030FFF9C2F6406D18BF8FC61F828" ID-DOI="http://doi.org/10.5281/zenodo.5945014" ID-GBIF-Taxon="154530339" ID-Zenodo-Dep="5945014" LSID="urn:lsid:zoobank.org:act:DE0F195B-0553-45C7-B01C-61C0E0BE77C5" httpUri="http://treatment.plazi.org/id/7A59030FFF9C2F6406D18BF8FC61F828" lastPageId="3" lastPageNumber="116" pageId="0" pageNumber="113">
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<subSubSection id="BAEAE192FF9C2F6706D18BF8FC1BF955" pageId="0" pageNumber="113" type="nomenclature">
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<paragraph id="F24FB219FF9C2F6706D18BF8FBE3F936" blockId="0.[151,1435,1695,2030]" box="[151,1113,1695,1722]" pageId="0" pageNumber="113">
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<heading id="A9070575FF9C2F6706D18BF8FBE3F936" bold="true" box="[151,1113,1695,1722]" fontSize="11" level="1" pageId="0" pageNumber="113" reason="1">
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<emphasis id="C0846E0BFF9C2F6706D18BF8FBE3F936" bold="true" box="[151,1113,1695,1722]" pageId="0" pageNumber="113">
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<taxonomicName id="35F0C99AFF9C2F6706D18BF8FD74F936" ID-CoL="4QYW" authority="Mlambo, Scholtz & Deschodt" authorityName="Mlambo & Scholtz & Deschodt" authorityYear="2019" box="[151,718,1695,1722]" class="Insecta" family="Scarabaeidae" genus="Grebennikovius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="113" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="C0846E0BFF9C2F6706D18BF8FEEDF935" bold="true" box="[151,343,1695,1721]" italics="true" pageId="0" pageNumber="113">Grebennikovius</emphasis>
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Mlambo, Scholtz & Deschodt
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</taxonomicName>
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,
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<taxonomicNameLabel id="DBB7D370FF9C2F67049D8BC7FCE7F936" box="[731,861,1696,1722]" pageId="0" pageNumber="113" rank="genus">new genus</taxonomicNameLabel>
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(
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<figureCitation id="6ACBAE9CFF9C2F67052A8BC7FC7AF936" box="[876,960,1696,1722]" captionStart="FIGURE 1" captionStartId="1.[151,244,1267,1289]" captionTargetBox="[171,1437,228,1221]" captionTargetId="figure@1.[151,1437,213,1246]" captionTargetPageId="1" captionText="FIGURE 1. A, Habitus of Grebennikovius basilewskyi (Balthasar, 1960) new combination, paratype male; B, habitus of Epactoides humberti (Paulian, 1975), holotype; C, anterolateral view of elytra of G. basilewskyi showing the carina on the elytral edge. Scale bar = 1 mm for Figs. A, B; Fig. C not to scale." httpUri="https://zenodo.org/record/2618193/files/figure.png" pageId="0" pageNumber="113">Figs. 1</figureCitation>
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,
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<figureCitation id="6ACBAE9CFF9C2F6705888BC7FC66F936" box="[974,988,1696,1722]" captionStart="FIGURE 2" captionStartId="2.[151,244,1964,1986]" captionTargetBox="[307,1294,222,1911]" captionTargetId="figure@2.[257,1316,191,1936]" captionTargetPageId="2" captionText="FIGURE 2. Aedeagus in dorsal (A), right lateral (B) and left lateral (C, D) view. A–C, Grebennikovius basilewskyi (Balthasar, 1960) new combination; D, Epactoides humberti (Paulian, 1975). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/2618195/files/figure.png" pageId="0" pageNumber="113">2</figureCitation>
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A–C, 3, 4)
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="F24FB219FF9C2F6706D18BA5FC1BF955" blockId="0.[151,1435,1695,2030]" box="[151,929,1729,1753]" pageId="0" pageNumber="113">
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Type-species:
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<taxonomicName id="35F0C99AFF9C2F6707768BA6FD67F955" authority="Balthasar, 1960" authorityName="Balthasar" authorityYear="1960" box="[304,733,1729,1753]" class="Insecta" family="Scarabaeidae" genus="Phacosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="113" phylum="Arthropoda" rank="species" species="basilewskyi">
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<emphasis id="C0846E0BFF9C2F6707768BA6FD94F955" box="[304,558,1729,1753]" italics="true" pageId="0" pageNumber="113">Phacosoma basilewskyi</emphasis>
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<bibRefCitation id="9661CFE8FF9C2F6704728BA5FD67F955" author="Balthasar, V." box="[564,733,1730,1753]" pageId="0" pageNumber="113" pagination="55 - 66" refId="ref1725" refString="Balthasar, V. (1960) Mission zoologique de l'I. R. S. A. C. en Afrique orientale (P. Basilewsky et N. Leleup, 1957). XLI. Coleoptera Scarabaeidae: Scarabaeini, Coprini, Onitini et Oniticellini. Annales du Musee Royal de l'Afrique Centrale, Serie 8 °, Sciences Zoologiques, 88, 55 - 66." type="journal article" year="1960">Balthasar, 1960</bibRefCitation>
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</taxonomicName>
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,
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<emphasis id="C0846E0BFF9C2F6704AE8BA5FC23F955" bold="true" box="[744,921,1730,1753]" pageId="0" pageNumber="113">here designated</emphasis>
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.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="BAEAE192FF9C2F6606D18B81FDA4F8AC" lastPageId="1" lastPageNumber="114" pageId="0" pageNumber="113" type="description">
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<paragraph id="F24FB219FF9C2F6706D18B81FBB2F977" blockId="0.[151,1435,1695,2030]" box="[151,1032,1766,1787]" pageId="0" pageNumber="113">
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http://zoobank.org/
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<uri id="8661BE1BFF9C2F6707128B81FBB2F977" box="[340,1032,1766,1787]" pageId="0" pageNumber="113">
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urn:lsid:zoobank.org:act:
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<uuid id="865688CCFF9C2F6704068B81FBB2F977" box="[576,1032,1766,1787]" pageId="0" pageNumber="113">DE0F195B-0553-45C7-B01C-61C0E0BE77C5</uuid>
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</paragraph>
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<paragraph id="F24FB219FF9C2F6706818A60FBA7F892" blockId="0.[151,1435,1695,2030]" box="[199,1053,1798,1822]" pageId="0" pageNumber="113">
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Composition.
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<taxonomicName id="35F0C99AFF9C2F6707188A61FCF2F892" authority="(Balthasar, 1960)" authorityName="Mlambo & Scholtz & Deschodt" authorityYear="2019" baseAuthorityName="Balthasar" baseAuthorityYear="1960" box="[350,840,1798,1822]" class="Insecta" family="Scarabaeidae" genus="Grebennikovius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="113" phylum="Arthropoda" rank="species" species="basilewskyi" status="comb. nov.">
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<emphasis id="C0846E0BFF9C2F6707188A61FD3DF892" box="[350,647,1798,1822]" italics="true" pageId="0" pageNumber="113">Grebennikovius basilewskyi</emphasis>
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(
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<bibRefCitation id="9661CFE8FF9C2F6704D38A60FCFAF892" author="Balthasar, V." box="[661,832,1799,1822]" pageId="0" pageNumber="113" pagination="55 - 66" refId="ref1725" refString="Balthasar, V. (1960) Mission zoologique de l'I. R. S. A. C. en Afrique orientale (P. Basilewsky et N. Leleup, 1957). XLI. Coleoptera Scarabaeidae: Scarabaeini, Coprini, Onitini et Oniticellini. Annales du Musee Royal de l'Afrique Centrale, Serie 8 °, Sciences Zoologiques, 88, 55 - 66." type="journal article" year="1960">Balthasar, 1960</bibRefCitation>
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)
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</taxonomicName>
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,
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<emphasis id="C0846E0BFF9C2F6705158A60FBACF892" bold="true" box="[851,1046,1799,1822]" pageId="0" pageNumber="113">
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<taxonomicNameLabel id="DBB7D370FF9C2F6705158A60FBACF892" box="[851,1046,1799,1822]" pageId="0" pageNumber="113" rank="species">new combination</taxonomicNameLabel>
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</emphasis>
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.
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</paragraph>
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<paragraph id="F24FB219FF9C2F6706818A4DFCFCF80A" blockId="0.[151,1435,1695,2030]" pageId="0" pageNumber="113">
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<emphasis id="C0846E0BFF9C2F6706818A4DFEE8F8CD" bold="true" box="[199,338,1834,1857]" pageId="0" pageNumber="113">Description.</emphasis>
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Length. 3.0–
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<quantity id="35081FFCFF9C2F6707AB8A4DFDFDF8CD" box="[493,583,1834,1857]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="0" pageNumber="113" unit="mm" value="3.5">3.5 mm</quantity>
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. Body small in size, oval shaped in dorsal view; head narrowing strongly anteriorly; elytra narrowing posteriorly but ending blunt and with notch at sutural interval; head and pronotum the same dark brown colour with elytra, pygidium and legs lighter brown.
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</paragraph>
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<paragraph id="F24FB219FF9C2F6706D18AF5FE14F862" blockId="0.[151,1435,1695,2030]" pageId="0" pageNumber="113">Head. Clypeus bidentate, denticles triangulate and upturned; sides with very slight notch where clypeo genal suture ends; surface with small, dense rounded punctures; punctures on clypeus smaller than those on genae and frons; punctures with very fine associated setae.</paragraph>
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<caption id="A68FE291FF9D2F6606D18994FD95FACF" httpUri="https://zenodo.org/record/2618193/files/figure.png" pageId="1" pageNumber="114" startId="1.[151,244,1267,1289]" targetBox="[171,1437,228,1221]" targetPageId="1">
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<paragraph id="F24FB219FF9D2F6606D18994FD95FACF" blockId="1.[151,1436,1267,1347]" pageId="1" pageNumber="114">
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<emphasis id="C0846E0BFF9D2F6606D18994FEB7FA85" bold="true" box="[151,269,1267,1289]" pageId="1" pageNumber="114">FIGURE 1.</emphasis>
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A, Habitus of
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<taxonomicName id="35F0C99AFF9D2F6607E68994FCE1FA85" authority="(Balthasar, 1960)" authorityName="Mlambo & Scholtz & Deschodt" authorityYear="2019" baseAuthorityName="Balthasar" baseAuthorityYear="1960" box="[416,859,1267,1289]" class="Insecta" family="Scarabaeidae" genus="Grebennikovius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="114" phylum="Arthropoda" rank="species" species="basilewskyi" status="comb. nov.">
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<emphasis id="C0846E0BFF9D2F6607E68994FD17FA85" box="[416,685,1267,1289]" italics="true" pageId="1" pageNumber="114">Grebennikovius basilewskyi</emphasis>
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(Balthasar, 1960)
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</taxonomicName>
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<taxonomicNameLabel id="DBB7D370FF9D2F6605248993FBB1FA85" box="[866,1035,1268,1289]" pageId="1" pageNumber="114" rank="species">new combination</taxonomicNameLabel>
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, paratype male; B, habitus of
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<taxonomicName id="35F0C99AFF9D2F6603778994FE30FAAA" authority="(Paulian, 1975)" authorityName="Paulian" authorityYear="1975" baseAuthorityName="Paulian" baseAuthorityYear="1975" class="Insecta" family="Scarabaeidae" genus="Epactoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="114" phylum="Arthropoda" rank="species" species="humberti">
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<emphasis id="C0846E0BFF9D2F6603778994FF54FAAA" italics="true" pageId="1" pageNumber="114">Epactoides humberti</emphasis>
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(Paulian, 1975)
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</taxonomicName>
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, holotype; C, anterolateral view of elytra of
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<taxonomicName id="35F0C99AFF9D2F6605718877FC7AFAAA" authorityName="Mlambo & Scholtz & Deschodt" authorityYear="2019" baseAuthorityName="Balthasar" baseAuthorityYear="1960" box="[823,960,1296,1318]" class="Insecta" family="Scarabaeidae" genus="Grebennikovius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="114" phylum="Arthropoda" rank="species" species="basilewskyi">
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<emphasis id="C0846E0BFF9D2F6605718877FC7AFAAA" box="[823,960,1296,1318]" italics="true" pageId="1" pageNumber="114">G. basilewskyi</emphasis>
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</taxonomicName>
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showing the carina on the elytral edge. Scale bar = 1 mm for Figs. A, B; Fig. C not to scale.
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</paragraph>
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</caption>
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<paragraph id="F24FB219FF9D2F660681880EFA85FA4A" blockId="1.[151,1436,1385,1963]" pageId="1" pageNumber="114">Pronotum. Somewhat convex; densely punctate with small punctures, punctures with associated thin recurved seta; posterior and anterior margins curving anteriorly, almost parallel, anteriomedial area slightly curving anteriorly; lateral margins with obtuse angel about one third from anterior angles; margin at anteriolateral angles appear flattened.</paragraph>
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<paragraph id="F24FB219FF9D2F66068188B6FED0F9DC" blockId="1.[151,1436,1385,1963]" pageId="1" pageNumber="114">Elytra. Striae with round punctures without setae, punctures closer together than their diameter; interstriae with surface smooth with single or double row of very dense setae each. Small humeral umbone present. Eight striae; stria V with small apical tubercle; stria VIII carinate, sides below carina slanted inward; sutural interval in posterior half appear significantly raised.</paragraph>
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<paragraph id="F24FB219FF9D2F6606818B3BFB3CF91A" blockId="1.[151,1436,1385,1963]" pageId="1" pageNumber="114">Mesometasternum with small punctures, punctures one puncture diameter apart. Mesometasternal suture faint and slightly curved anteriorly, area directly posterior of mesometasternal suture without punctures.</paragraph>
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<paragraph id="F24FB219FF9D2F6606818BC6FAFDF934" blockId="1.[151,1436,1385,1963]" box="[199,1351,1697,1720]" pageId="1" pageNumber="114">Pygidium. Surface with small punctures with short associated setae; median tubercle present, otherwise flat.</paragraph>
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<paragraph id="F24FB219FF9D2F6606818BA3FAA1F957" blockId="1.[151,1436,1385,1963]" box="[199,1307,1732,1755]" pageId="1" pageNumber="114">Legs. Front tibia with three denticles on the outer edge. First article of hind tarsi longer than the second.</paragraph>
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<paragraph id="F24FB219FF9D2F6606D18B80FDA4F8AC" blockId="1.[151,1436,1385,1963]" pageId="1" pageNumber="114">
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Aedeagus. Fairly symmetrical, see
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<figureCitation id="6ACBAE9CFF9D2F6604588B80FDD0F972" box="[542,618,1767,1790]" captionStart="FIGURE 2" captionStartId="2.[151,244,1964,1986]" captionTargetBox="[307,1294,222,1911]" captionTargetId="figure@2.[257,1316,191,1936]" captionTargetPageId="2" captionText="FIGURE 2. Aedeagus in dorsal (A), right lateral (B) and left lateral (C, D) view. A–C, Grebennikovius basilewskyi (Balthasar, 1960) new combination; D, Epactoides humberti (Paulian, 1975). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/2618195/files/figure.png" pageId="1" pageNumber="114">Figs. 2</figureCitation>
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A–C. Parameres half the length of the phallobase, apex of the parameres sloping downward towards a flat tip.
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</paragraph>
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</subSubSection>
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<subSubSection id="BAEAE192FF9D2F6606818A4BFEEEF8EA" pageId="1" pageNumber="114" type="distribution">
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<paragraph id="F24FB219FF9D2F6606818A4BFEEEF8EA" blockId="1.[151,1436,1385,1963]" pageId="1" pageNumber="114">
|
||
<emphasis id="C0846E0BFF9D2F6606818A4BFEE3F8CF" bold="true" box="[199,345,1836,1859]" pageId="1" pageNumber="114">Distribution.</emphasis>
|
||
This monotypic genus is currently only known from the isolated forests in the Uluguru Mountains in
|
||
<collectingCountry id="8AE7F289FF9D2F6606D18A28FF4DF8EA" box="[151,247,1871,1894]" name="Tanzania" pageId="1" pageNumber="114">Tanzania</collectingCountry>
|
||
(
|
||
<figureCitation id="6ACBAE9CFF9D2F6607438A28FEFCF8EA" box="[261,326,1871,1894]" captionStart="FIGURE 3" captionStartId="3.[151,244,1498,1520]" captionTargetBox="[150,1436,191,1477]" captionTargetId="figure@3.[150,1436,191,1477]" captionTargetPageId="3" captionText="FIGURE 3. Map showing the location of the Uluguru Mountains (black) in Tanzania. Light grey indicates mountains and dark grey indicates the great lakes (modified from Mlambo et al. 2014)." httpUri="https://zenodo.org/record/2618197/files/figure.png" pageId="1" pageNumber="114">Fig. 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="BAEAE192FF9D2F6606818A16FBCFF827" pageId="1" pageNumber="114" type="etymology">
|
||
<paragraph id="F24FB219FF9D2F6606818A16FBCFF827" blockId="1.[151,1436,1385,1963]" pageId="1" pageNumber="114">
|
||
<emphasis id="C0846E0BFF9D2F6606818A16FEFDF804" bold="true" box="[199,327,1905,1928]" pageId="1" pageNumber="114">Etymology.</emphasis>
|
||
<taxonomicName id="35F0C99AFF9D2F6607098A17FE4DF804" authority="Mlambo & Scholtz & Deschodt, 2019" authorityName="Mlambo & Scholtz & Deschodt" authorityYear="2019" box="[335,503,1904,1928]" class="Insecta" family="Scarabaeidae" genus="Grebennikovius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="114" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="C0846E0BFF9D2F6607098A17FE4DF804" box="[335,503,1904,1928]" italics="true" pageId="1" pageNumber="114">Grebennikovius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="DBB7D370FF9D2F6607B88A16FDCBF804" box="[510,625,1905,1928]" pageId="1" pageNumber="114" rank="genus">new genus</taxonomicNameLabel>
|
||
is named in honour of Vasily Grebennikov, who collected fresh specimens of this new genus for the molecular study by
|
||
<bibRefCitation id="9661CFE8FF9D2F66043E8AF3FCDAF827" author="Mlambo, S. & Sole, C. L. & Scholtz, C. H." box="[632,864,1939,1963]" pageId="1" pageNumber="114" pagination="115 - 120" refId="ref1794" refString="Mlambo, S., Sole, C. L. & Scholtz, C. H. (2014) Affinities of the Canthonini dung beetles of the Eastern Arc Mountains. Organisms, Diversity and Evolution, 14, 115 - 120. https: // doi. org / 10.1007 / s 13127 - 013 - 0158 - y" type="journal article" year="2014">
|
||
Mlambo
|
||
<emphasis id="C0846E0BFF9D2F66049C8AF3FCB5F827" box="[730,783,1939,1963]" italics="true" pageId="1" pageNumber="114">et al.</emphasis>
|
||
(2014)
|
||
</bibRefCitation>
|
||
. The gender is masculine.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="A68FE291FF9E2F6506D18ACBFC2FF852" httpUri="https://zenodo.org/record/2618195/files/figure.png" pageId="2" pageNumber="115" startId="2.[151,244,1964,1986]" targetBox="[307,1294,222,1911]" targetPageId="2">
|
||
<paragraph id="F24FB219FF9E2F6506D18ACBFC2FF852" blockId="2.[151,1436,1964,2015]" pageId="2" pageNumber="115">
|
||
<emphasis id="C0846E0BFF9E2F6506D18ACBFEB6F84E" bold="true" box="[151,268,1964,1986]" pageId="2" pageNumber="115">FIGURE 2.</emphasis>
|
||
Aedeagus in dorsal (A), right lateral (B) and left lateral (C, D) view. A–C,
|
||
<taxonomicName id="35F0C99AFF9E2F6505A28ACBFA26F84D" authority="(Balthasar, 1960)" authorityName="Mlambo & Scholtz & Deschodt" authorityYear="2019" baseAuthorityName="Balthasar" baseAuthorityYear="1960" box="[996,1436,1964,1986]" class="Insecta" family="Scarabaeidae" genus="Grebennikovius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="115" phylum="Arthropoda" rank="species" species="basilewskyi" status="comb. nov.">
|
||
<emphasis id="C0846E0BFF9E2F6505A28ACBFB55F84E" box="[996,1263,1964,1986]" italics="true" pageId="2" pageNumber="115">Grebennikovius basilewskyi</emphasis>
|
||
(Balthasar, 1960)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="DBB7D370FF9E2F6506D18AAEFEFAF852" box="[151,320,1993,2014]" pageId="2" pageNumber="115" rank="species">new combination</taxonomicNameLabel>
|
||
; D,
|
||
<taxonomicName id="35F0C99AFF9E2F6507208AAEFD70F852" authority="(Paulian, 1975)" authorityName="Paulian" authorityYear="1975" baseAuthorityName="Paulian" baseAuthorityYear="1975" box="[358,714,1993,2015]" class="Insecta" family="Scarabaeidae" genus="Epactoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="115" phylum="Arthropoda" rank="species" species="humberti">
|
||
<emphasis id="C0846E0BFF9E2F6507208AAEFD97F853" box="[358,557,1993,2015]" italics="true" pageId="2" pageNumber="115">Epactoides humberti</emphasis>
|
||
(Paulian, 1975)
|
||
</taxonomicName>
|
||
. Scale bar = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="A68FE291FF9F2F6406D188BDFD5DF981" httpUri="https://zenodo.org/record/2618197/files/figure.png" pageId="3" pageNumber="116" startId="3.[151,244,1498,1520]" targetBox="[150,1436,191,1477]" targetPageId="3">
|
||
<paragraph id="F24FB219FF9F2F6406D188BDFD5DF981" blockId="3.[151,1436,1498,1549]" pageId="3" pageNumber="116">
|
||
<emphasis id="C0846E0BFF9F2F6406D188BDFEB7FA7C" bold="true" box="[151,269,1498,1520]" pageId="3" pageNumber="116">FIGURE 3.</emphasis>
|
||
Map showing the location of the Uluguru Mountains (black) in Tanzania. Light grey indicates mountains and dark grey indicates the great lakes (modified from Mlambo
|
||
<emphasis id="C0846E0BFF9F2F640435889FFD19F981" box="[627,675,1527,1549]" italics="true" pageId="3" pageNumber="116">et al.</emphasis>
|
||
2014).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="BAEAE192FF9F2F6406818B54FC61F828" pageId="3" pageNumber="116" type="diagnosis">
|
||
<paragraph id="F24FB219FF9F2F6406818B54FC61F828" blockId="3.[151,1436,1587,1956]" pageId="3" pageNumber="116">
|
||
<emphasis id="C0846E0BFF9F2F6406818B54FE78F9C6" bold="true" box="[199,450,1587,1610]" pageId="3" pageNumber="116">Differential diagnosis.</emphasis>
|
||
The combination of the hind tarsus with the basal segments approximately twice as long as the next segment and the stria VIII being carinate with the sides below the carina slanting inward will separate this new genus from all other Afrotropical small deltochiline genera. Although the body shape of
|
||
<taxonomicName id="35F0C99AFF9F2F64020B8B10FB4FF903" authority="Mlambo & Scholtz & Deschodt, 2019" authorityName="Mlambo & Scholtz & Deschodt" authorityYear="2019" box="[1101,1269,1655,1679]" class="Insecta" family="Scarabaeidae" genus="Grebennikovius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="116" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="C0846E0BFF9F2F64020B8B10FB4FF903" box="[1101,1269,1655,1679]" italics="true" pageId="3" pageNumber="116">Grebennikovius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="DBB7D370FF9F2F6402BA8B1FFAD4F903" box="[1276,1390,1656,1679]" pageId="3" pageNumber="116" rank="genus">new genus</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="35F0C99AFF9F2F6406D18BFDFEB7F93E" authorityName="Olsoufieff" authorityYear="1947" box="[151,269,1690,1714]" class="Insecta" family="Scarabaeidae" genus="Epactoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="116" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C0846E0BFF9F2F6406D18BFDFEB7F93E" box="[151,269,1690,1714]" italics="true" pageId="3" pageNumber="116">Epactoides</emphasis>
|
||
</taxonomicName>
|
||
are quite similar in appearance,
|
||
<taxonomicName id="35F0C99AFF9F2F6404C78BFDFC93F93E" authority="Mlambo & Scholtz & Deschodt, 2019" authorityName="Mlambo & Scholtz & Deschodt" authorityYear="2019" box="[641,809,1690,1714]" class="Insecta" family="Scarabaeidae" genus="Grebennikovius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="116" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="C0846E0BFF9F2F6404C78BFDFC93F93E" box="[641,809,1690,1714]" italics="true" pageId="3" pageNumber="116">Grebennikovius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="DBB7D370FF9F2F6405728BFCFC13F93E" box="[820,937,1691,1714]" pageId="3" pageNumber="116" rank="genus">new genus</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="35F0C99AFF9F2F6402058BFDFB03F93E" authorityName="Olsoufieff" authorityYear="1947" box="[1091,1209,1690,1714]" class="Insecta" family="Scarabaeidae" genus="Epactoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="116" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C0846E0BFF9F2F6402058BFDFB03F93E" box="[1091,1209,1690,1714]" italics="true" pageId="3" pageNumber="116">Epactoides</emphasis>
|
||
</taxonomicName>
|
||
by having a clearly bidentate clypeus while the clypeus of
|
||
<taxonomicName id="35F0C99AFF9F2F64047C8BDAFD0AF959" authorityName="Olsoufieff" authorityYear="1947" box="[570,688,1725,1749]" class="Insecta" family="Scarabaeidae" genus="Epactoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="116" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C0846E0BFF9F2F64047C8BDAFD0AF959" box="[570,688,1725,1749]" italics="true" pageId="3" pageNumber="116">Epactoides</emphasis>
|
||
</taxonomicName>
|
||
is quadridentate (
|
||
<figureCitation id="6ACBAE9CFF9F2F6405348BDAFC45F958" box="[882,1023,1725,1748]" captionStart="FIGURE 1" captionStartId="1.[151,244,1267,1289]" captionTargetBox="[171,1437,228,1221]" captionTargetId="figure@1.[151,1437,213,1246]" captionTargetPageId="1" captionText="FIGURE 1. A, Habitus of Grebennikovius basilewskyi (Balthasar, 1960) new combination, paratype male; B, habitus of Epactoides humberti (Paulian, 1975), holotype; C, anterolateral view of elytra of G. basilewskyi showing the carina on the elytral edge. Scale bar = 1 mm for Figs. A, B; Fig. C not to scale." httpUri="https://zenodo.org/record/2618193/files/figure.png" pageId="3" pageNumber="116">Figs. 1A, 1B</figureCitation>
|
||
). Elytra of
|
||
<taxonomicName id="35F0C99AFF9F2F64023D8BDAFA99F959" authority="Mlambo & Scholtz & Deschodt, 2019" authorityName="Mlambo & Scholtz & Deschodt" authorityYear="2019" box="[1147,1315,1725,1749]" class="Insecta" family="Scarabaeidae" genus="Grebennikovius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="116" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="C0846E0BFF9F2F64023D8BDAFA99F959" box="[1147,1315,1725,1749]" italics="true" pageId="3" pageNumber="116">Grebennikovius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="DBB7D370FF9F2F64036C8BDAFA26F958" box="[1322,1436,1725,1748]" pageId="3" pageNumber="116" rank="genus">new genus</taxonomicNameLabel>
|
||
are blunt and have a notch at the posterior end whereas that of
|
||
<taxonomicName id="35F0C99AFF9F2F6405178BB8FC7DF97B" authorityName="Olsoufieff" authorityYear="1947" box="[849,967,1759,1783]" class="Insecta" family="Scarabaeidae" genus="Epactoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="116" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C0846E0BFF9F2F6405178BB8FC7DF97B" box="[849,967,1759,1783]" italics="true" pageId="3" pageNumber="116">Epactoides</emphasis>
|
||
</taxonomicName>
|
||
are sharp.
|
||
<taxonomicName id="35F0C99AFF9F2F6402028BB8FB56F97B" authority="Mlambo & Scholtz & Deschodt, 2019" authorityName="Mlambo & Scholtz & Deschodt" authorityYear="2019" box="[1092,1260,1759,1783]" class="Insecta" family="Scarabaeidae" genus="Grebennikovius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="116" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="C0846E0BFF9F2F6402028BB8FB56F97B" box="[1092,1260,1759,1783]" italics="true" pageId="3" pageNumber="116">Grebennikovius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="DBB7D370FF9F2F6402B28B87FAD2F97B" box="[1268,1384,1760,1783]" pageId="3" pageNumber="116" rank="genus">new genus</taxonomicNameLabel>
|
||
also superficially resembles some species of the southern African genus
|
||
<taxonomicName id="35F0C99AFF9F2F6405C68A64FC61F896" authorityName="Dejean" authorityYear="1833" box="[896,987,1795,1818]" class="Insecta" family="Scarabaeidae" genus="Epirinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="116" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C0846E0BFF9F2F6405C68A64FC61F896" box="[896,987,1795,1818]" italics="true" pageId="3" pageNumber="116">Epirinus</emphasis>
|
||
</taxonomicName>
|
||
. However,
|
||
<taxonomicName id="35F0C99AFF9F2F64021A8A65FABEF896" authority="Mlambo & Scholtz & Deschodt, 2019" authorityName="Mlambo & Scholtz & Deschodt" authorityYear="2019" box="[1116,1284,1794,1818]" class="Insecta" family="Scarabaeidae" genus="Grebennikovius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="116" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="C0846E0BFF9F2F64021A8A65FABEF896" box="[1116,1284,1794,1818]" italics="true" pageId="3" pageNumber="116">Grebennikovius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="DBB7D370FF9F2F64034B8A64FA3BF896" box="[1293,1409,1795,1818]" pageId="3" pageNumber="116" rank="genus">new genus</taxonomicNameLabel>
|
||
is generally smaller and it is carinate at the stria VIII with the area below the carina slanted inwardly (
|
||
<figureCitation id="6ACBAE9CFF9F2F6403458A42FAE3F8B0" box="[1283,1369,1829,1852]" captionStart="FIGURE 1" captionStartId="1.[151,244,1267,1289]" captionTargetBox="[171,1437,228,1221]" captionTargetId="figure@1.[151,1437,213,1246]" captionTargetPageId="1" captionText="FIGURE 1. A, Habitus of Grebennikovius basilewskyi (Balthasar, 1960) new combination, paratype male; B, habitus of Epactoides humberti (Paulian, 1975), holotype; C, anterolateral view of elytra of G. basilewskyi showing the carina on the elytral edge. Scale bar = 1 mm for Figs. A, B; Fig. C not to scale." httpUri="https://zenodo.org/record/2618193/files/figure.png" pageId="3" pageNumber="116">Fig. 1C</figureCitation>
|
||
). The aedeagus of
|
||
<taxonomicName id="35F0C99AFF9F2F6407648A2FFEC7F8D3" authorityName="Dejean" authorityYear="1833" box="[290,381,1864,1887]" class="Insecta" family="Scarabaeidae" genus="Epirinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="116" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C0846E0BFF9F2F6407648A2FFEC7F8D3" box="[290,381,1864,1887]" italics="true" pageId="3" pageNumber="116">Epirinus</emphasis>
|
||
</taxonomicName>
|
||
is asymmetrical (see
|
||
<emphasis id="C0846E0BFF9F2F6404368A2EFD2CF8EC" box="[624,662,1865,1888]" italics="true" pageId="3" pageNumber="116">e.g.</emphasis>
|
||
, aedeagus of
|
||
<taxonomicName id="35F0C99AFF9F2F6405748A2EFC08F8D3" baseAuthorityName="Paulian" baseAuthorityYear="1975" box="[818,946,1863,1888]" class="Insecta" family="Scarabaeidae" genus="Epactoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="116" phylum="Arthropoda" rank="species" species="humberti">
|
||
<emphasis id="C0846E0BFF9F2F6405748A2EFC08F8D3" box="[818,946,1863,1888]" italics="true" pageId="3" pageNumber="116">E. humberti</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="6ACBAE9CFF9F2F6405858A2FFBA2F8D3" box="[963,1048,1864,1887]" captionStart="FIGURE 2" captionStartId="2.[151,244,1964,1986]" captionTargetBox="[307,1294,222,1911]" captionTargetId="figure@2.[257,1316,191,1936]" captionTargetPageId="2" captionText="FIGURE 2. Aedeagus in dorsal (A), right lateral (B) and left lateral (C, D) view. A–C, Grebennikovius basilewskyi (Balthasar, 1960) new combination; D, Epactoides humberti (Paulian, 1975). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/2618195/files/figure.png" pageId="3" pageNumber="116">Fig. 2D</figureCitation>
|
||
) whereas the aedeagus of this new genus is symmetrical. Like several other deltochiline genera (
|
||
<bibRefCitation id="9661CFE8FF9F2F6405738A0CFBF3F80E" author="Scholtz, C. H. & Howden, H. F." box="[821,1097,1899,1922]" pageId="3" pageNumber="116" pagination="75 - 119" refId="ref1989" refString="Scholtz, C. H. & Howden, H. F. (1987) A revision of the African Cathonina [sic] (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of the Entomological Society of Southern Africa, 50, 75 - 119." type="journal article" year="1987">Scholtz & Howden 1987</bibRefCitation>
|
||
),
|
||
<taxonomicName id="35F0C99AFF9F2F6402188A0DFABCF80E" authority="Mlambo & Scholtz & Deschodt, 2019" authorityName="Mlambo & Scholtz & Deschodt" authorityYear="2019" box="[1118,1286,1898,1922]" class="Insecta" family="Scarabaeidae" genus="Grebennikovius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="116" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="C0846E0BFF9F2F6402188A0DFABCF80E" box="[1118,1286,1898,1922]" italics="true" pageId="3" pageNumber="116">Grebennikovius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="DBB7D370FF9F2F6403488A0CFA3BF80E" box="[1294,1409,1899,1922]" pageId="3" pageNumber="116" rank="genus">new genus</taxonomicNameLabel>
|
||
is monotypic and restricted to a specific forest. It has been collected in leaf litter.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |