treatments-xml/data/67/90/B0/6790B0D3F01657DDBAEED4F3E892440A.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1147.93824" ID-Pensoft-Pub="1313-2970-1147-1" ID-Pensoft-UUID="FF2B6B317B505F9EA0E1000BDCD16CE7" ID-ZooBank="9BB9881B0076473D8E53155D37CA1F50" ModsDocID="1313-2970-1147-1" checkinTime="1676378625502" checkinUser="pensoft" docAuthor="Pall-Gergely, Barna, Hunyadi, Andras, Vermeulen, Jaap J., Grego, Jozef, Sutcharit, Chirasak, Reischuetz, Alexander, Dumrongrojwattana, Pongrat, Botta-Dukat, Zoltan, Oerstan, Aydin, Fekete, Judit &amp; Jochum, Adrienne" docDate="2023" docId="6790B0D3F01657DDBAEED4F3E892440A" docLanguage="en" docName="ZooKeys 1147: 1-177" docOrigin="ZooKeys 1147" docPubDate="2023-02-13" docSource="http://dx.doi.org/10.3897/zookeys.1147.93824" docTitle="Angustopila thersites Pall-Gergely &amp; Vermeulen 2023, sp. nov." docType="treatment" docUuid="C9601C1B-14C6-4AED-8DE9-36D7B0C3DAF4" docUuidSource="ZooBank" docVersion="1" id="FF2B6B317B505F9EA0E1000BDCD16CE7" lastPageNumber="1" masterDocId="FF2B6B317B505F9EA0E1000BDCD16CE7" masterDocTitle="Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae)" masterLastPageNumber="177" masterPageNumber="1" pageNumber="1" updateTime="1676378625502" updateUser="pensoft">
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<mods:titleInfo>
<mods:title>Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pall-Gergely, Barna</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6167-7221</mods:nameIdentifier>
<mods:affiliation>Centre for Agricultural Research, Plant Protection Institute, Eoetvoes Lorand Research Network, Herman Otto ut 15, H- 1022 Budapest, Hungary</mods:affiliation>
<mods:nameIdentifier type="email">pallgergely2@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hunyadi, Andras</mods:namePart>
<mods:affiliation>Adria setany 10 G 2 / 5., H- 1148 Budapest, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vermeulen, Jaap J.</mods:namePart>
<mods:affiliation>JK Art and Science, Lauwerbes 8, 2318 AT Leiden, Netherlands</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Grego, Jozef</mods:namePart>
<mods:affiliation>Horna Micina 219, SK- 97401 Banska Bystrica, Slovakia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sutcharit, Chirasak</mods:namePart>
<mods:affiliation>Animal Systematic Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Reischuetz, Alexander</mods:namePart>
<mods:affiliation>Puechhaimgasse 52, A- 3580 Horn, Austria</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Dumrongrojwattana, Pongrat</mods:namePart>
<mods:affiliation>Department of Biology, Faculty of Science, Burapha University, 169 Longhardbangsaen Road, Muang District, Chonburi, 20131, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Botta-Dukat, Zoltan</mods:namePart>
<mods:affiliation>Centre for Ecological Research, Institute of Ecology and Botany, Alkotmany 2 - 4, H- 2600, Vacratot, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Oerstan, Aydin</mods:namePart>
<mods:affiliation>12501 Milestone Manor Lane, Germantown, Maryland, 20876, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fekete, Judit</mods:namePart>
<mods:affiliation>University of Pannonia, Centre of Natural Science, Research Group of Limnology, Egyetem u. 10, H- 8200 Veszprem, Hungary &amp; Centre for Ecological Research, Institute of Aquatic Ecology, Department of Tisza Research, 18 / c Bem square, H- 4026 Debrecen, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jochum, Adrienne</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6624-6412</mods:nameIdentifier>
<mods:affiliation>Naturhistorisches Museum der Burgergemeinde Bern, CH- 3005 Bern, Switzerland &amp; Institute of Ecology and Evolution, University of Bern, CH- 3012 Bern, Switzerland &amp; Senckenberg Forschungsinstitut und Naturmuseum, 60325 Frankfurt am Main, Germany</mods:affiliation>
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<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-02-13</mods:number>
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<mods:number>1147</mods:number>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1147.93824</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1147.93824</mods:identifier>
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="https://zoobank.org/C9601C1B-14C6-4AED-8DE9-36D7B0C3DAF4" authority="Páll-Gergely &amp; Vermeulen" authorityName="Pall-Gergely &amp; Vermeulen" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Angustopila thersites" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="thersites" status="sp. nov.">
Angustopila thersites 
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; Vermeulen
</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Angustopila thersites Pall-Gergely &amp; Vermeulen, sp. nov. A paratype (RMNH 347761) B-H holotype (RMNH 347760). Apertural (A, B), lateral (C), ventral (D) and apical (E) sides of the shell; granular sculpture on the protoconch (F), ventral (G) and frontal (H) surface of the body whorl." figureDoi="10.3897/zookeys.1147.93824.figure20" httpUri="https://binary.pensoft.net/fig/808829" pageId="0" pageNumber="1">Figs 20</figureCitation>
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Angustopila thersites Pall-Gergely &amp; Vermeulen, sp. nov., sample WMVT. 0327 (RMNH 347764) A specimen 2 B-I specimen 1. Apertural (A, B), lateral (C), ventral (D) and apical (E) sides of the shell; granular sculpture on the protoconch (F), aperture (G), ventral (H) and frontal (I) surface of the body whorl." figureDoi="10.3897/zookeys.1147.93824.figure21" httpUri="https://binary.pensoft.net/fig/808830" pageId="0" pageNumber="1">, 21</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
: Vietnam • 1 empty shell (H: 0.85 mm, D: 0.86 mm); Quang Ninh Province, Halong Bay area, Phao Trong Island (locality code: WMVT.0333); approx. GPS code: 
<geoCoordinate degrees="20" direction="north" minutes="49.80" orientation="latitude" precision="9" value="20.83">20°49.80'N</geoCoordinate>
, 
<geoCoordinate degrees="107" direction="east" minutes="08.32" orientation="longitude" precision="9" value="107.138664">107°08.32'E</geoCoordinate>
; 11 Sep. 2003; W.J.M. Maassen leg.; RMNH 347760.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
: Vietnam • 9 adult shells; same data as for holotype; RMNH 347761 • 3 shells; Quang Ninh Province, Halong Bay area, Luoi Liem Island; 
<geoCoordinate degrees="20" direction="north" minutes="49.67" orientation="latitude" precision="9" value="20.827833">20°49.67'N</geoCoordinate>
, 
<geoCoordinate degrees="107" direction="east" minutes="09.90" orientation="longitude" precision="9" value="107.165">107°09.90'E</geoCoordinate>
(locality code: WMVT.0334); 12 Sep. 2003; W.J.M. Maassen leg.; RMNH 347762 • 1 shell; Haiphong Province, Cat Ba Island, Cave Qua Vang, around cave entrance, rocky limestone slope with low, rather mature forest; 
<geoCoordinate degrees="20" direction="north" minutes="48.64" orientation="latitude" precision="9" value="20.810667">20°48.64'N</geoCoordinate>
, 
<geoCoordinate degrees="107" direction="east" minutes="04.64" orientation="longitude" precision="9" value="107.07733">107°04.64'E</geoCoordinate>
; 100 m a.s.l.; 6 Jun. 2017; J.J. Vermeulen &amp; K. Anker leg.; JJV 16621 • 7 shells; Haiphong Province, Cat Ba Island, Cave Qua Vang, inside cave, large, little disturbed active cave with speleothems; 
<geoCoordinate degrees="20" direction="north" minutes="48.64" orientation="latitude" precision="9" value="20.810667">20°48.64'N</geoCoordinate>
, 
<geoCoordinate degrees="107" direction="east" minutes="04.64" orientation="longitude" precision="9" value="107.07733">107°04.64'E</geoCoordinate>
; 60 m a.s.l.; 6 Jun. 2017; J.J. Vermeulen &amp; K. Anker leg.; JJV 16620 • 8 shells; Haiphong Province, Cat Ba Island, Cave Xa Bac, around cave entrance; 
<geoCoordinate degrees="20" direction="north" minutes="50.07" orientation="latitude" precision="9" value="20.8345">20°50.07'N</geoCoordinate>
, 
<geoCoordinate degrees="106" direction="east" minutes="58.61" orientation="longitude" precision="9" value="106.97684">106°58.61'E</geoCoordinate>
; 30 m a.s.l.; 9 Jun. 2017; J.J. Vermeulen &amp; K. Anker leg.; JJV 16599 • 15 shells; Quang Ninh Province, Halong Bay area, Dao Bo Hon, Song Sot Cave, drift material washed together over sinkhole in cave; 
<geoCoordinate degrees="20" direction="north" minutes="50.83" orientation="latitude" precision="9" value="20.847166">20°50.83'N</geoCoordinate>
, 
<geoCoordinate degrees="107" direction="east" minutes="05.67" orientation="longitude" precision="9" value="107.0945">107°05.67'E</geoCoordinate>
; 2 Oct. 1998; J.J. Vermeulen &amp; A.J. Whitten leg.; JJV 17654 (ex JJV 6237) • 1 shell; Haiphong Province, Cat Ba Island, along trail from headquarters to Viet Hai, along Ao Eck, sampling starting at coordinates, until 0.5 km beyond Ao Eck; 
<geoCoordinate degrees="20" direction="north" minutes="48.07" orientation="latitude" precision="9" value="20.801167">20°48.07'N</geoCoordinate>
, 
<geoCoordinate degrees="107" direction="east" minutes="00.83" orientation="longitude" precision="9" value="107.01383">107°00.83'E</geoCoordinate>
; 100 m a.s.l.; 4 Jun. 2017; J.J. Vermeulen &amp; K. Anker leg.; JJV 16618 • 4 shells; Haiphong Province, Cat Ba Island, Cave Uy Ban, around cave entrance; 
<geoCoordinate degrees="20" direction="north" minutes="46.39" orientation="latitude" precision="9" value="20.773167">20°46.39'N</geoCoordinate>
, 
<geoCoordinate degrees="107" direction="east" minutes="00.88" orientation="longitude" precision="9" value="107.01466">107°00.88'E</geoCoordinate>
; 40 m a.s.l.; 9 Jun. 2017; J.J. Vermeulen &amp; K. Anker leg.; JJV 16623 • 7 shells; Haiphong Province, Halong Bay area, unnamed island off E Coast Cat Ba, south facing bay with beach and densely vegetated limestone scree slope; 
<geoCoordinate degrees="20" direction="north" minutes="45.19" orientation="latitude" precision="9" value="20.753166">20°45.19'N</geoCoordinate>
, 
<geoCoordinate degrees="107" direction="east" minutes="04.45" orientation="longitude" precision="9" value="107.074165">107°04.45'E</geoCoordinate>
(approximate GPS data); 1 Oct. 1998; J.J. Vermeulen &amp; K. Anker leg.; JJV 6236 • 2 shells; Haiphong Province, Cat Ba Island, large, somewhat disturbed, active cave with speleothems; 
<geoCoordinate degrees="20" direction="north" minutes="50.06" orientation="latitude" precision="9" value="20.834333">20°50.06'N</geoCoordinate>
, 
<geoCoordinate degrees="106" direction="east" minutes="55.91" orientation="longitude" precision="9" value="106.93183">106°55.91'E</geoCoordinate>
; 0-50 m a.s.l.; 7 Jun. 2017; J.J. Vermeulen &amp; K. Anker leg.; JJV 16622 • 4 shells; Quang Ninh Province, Halong Bay area, unnamed island 1.8 km W of S point Cong Tai Island, steep limestone slope bordering beach, dense vegetation; 
<geoCoordinate degrees="20" direction="north" minutes="52.23" orientation="latitude" precision="9" value="20.8705">20°52.23'N</geoCoordinate>
, 
<geoCoordinate degrees="107" direction="east" minutes="18.25" orientation="longitude" precision="9" value="107.30417">107°18.25'E</geoCoordinate>
; 3 Oct. 1998; J.J. Vermeulen &amp; A.J. Whitten leg.; JJV 6239 • 25 shells; Haiphong Province, Cat Ba Island, Cave Trung Trang, around cave entrance, steep limestone cliffs with vegetated ledges; 
<geoCoordinate degrees="20" direction="north" minutes="47.30" orientation="latitude" precision="9" value="20.788334">20°47.30'N</geoCoordinate>
, 
<geoCoordinate degrees="106" direction="east" minutes="59.84" orientation="longitude" precision="9" value="106.99733">106°59.84'E</geoCoordinate>
; 50 m a.s.l.; 6 Jun. 2017; J.J. Vermeulen &amp; K. Anker leg.; JJV 16619 • 5 shells; same data as for preceding; coll. HA.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Additional material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation collectingDate="2003-09-06" collectorName="W. J. M. Maassen" country="Vietnam" latitude="20.867834" location="Thay Cave on Congfo Island" longLatPrecision="12" longitude="107.201" municipality="Halong Bay area" specimenCount="1" stateProvince="Quang Ninh Province">
<collectingCountry name="Vietnam">Vietnam</collectingCountry>
• 2 j/b shells; same data as for holotype; RMNH 347763 • 7 shells; 
<collectingRegion country="Vietnam" name="Quang Ninh">Quang Ninh Province</collectingRegion>
, 
<collectingMunicipality>Halong Bay area</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:6790B0D3F01657DDBAEED4F3E892440A:755B93E71E55547E2A9F08261B3D7E87" country="Vietnam" latitude="20.867834" longLatPrecision="12" longitude="107.201" municipality="Halong Bay area" name="Thay Cave on Congfo Island" stateProvince="Quang Ninh Province">Thay Cave on Congfo Island</location>
, collected inside the cave; 
<geoCoordinate degrees="20" direction="north" minutes="52.07" orientation="latitude" precision="9" value="20.867834">20°52.07'N</geoCoordinate>
, 
<geoCoordinate degrees="107" direction="east" minutes="12.06" orientation="longitude" precision="9" value="107.201">107°12.06'E</geoCoordinate>
(locality code: WMVT.0327); 
<collectingDate value="2003-09-06">6 Sep. 2003</collectingDate>
; 
<collectorName>W.J.M. Maassen</collectorName>
leg.; RMNH 347764
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">A small to medium-sized, concave-conical species with irregularly growing whorls, strong spiral striation, very narrow umbilicus, and a strongly oblique, toothless aperture.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">Shell small to medium-sized for the genus, mostly wider than high or rarely higher than wide (see population WMVT.0327), concave-conical, tightly and asymmetrically coiled, seemingly globular due to narrowing last whorl; body whorl widest in apertural view (although slightly wider than penultimate whorl) or penultimate whorl widest; protoconch bulbous, consists of ca. 1.25 whorls, with weak spiral striation preceding the first teleoconch whorl; teleoconch with irregular radial growth lines and dense, strong, rather regular wavy spiral striation (ca. 22 spiral striae on body whorl in apertural view); whorls 3.5-3.75, rounded; aperture curved and strongly oblique to shell axis in lateral view; aperture ovate-oblong; aperture tucked tightly under inflated penultimate whorl in apertural view; peristome slightly expanded, not reflected; not attached to penultimate whorl; umbilicus very narrow, ca. 1/5 of shell width; aperture toothless.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Measurements (in mm).</paragraph>
<paragraph pageId="0" pageNumber="1">
H = 0.75-0.87, D = 0.81-0.93, H/D*100 = 87.2-101.2 (
<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
= 10), RUD = 23.5-24.1 (
<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
= 3) (populations except for WMVT.0327); H = 0.81-0.85, D = 0.87-0.93, H/D*100, = 102.4-114.8, RUD = 22.2-24.7 (
<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
= 2, WMVT.0327).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
The most similar species is 
<taxonomicName lsidName="A. megastoma" pageId="0" pageNumber="1" rank="species" species="megastoma">
<emphasis italics="true" pageId="0" pageNumber="1">A. megastoma</emphasis>
</taxonomicName>
sp. nov., which also occurs sympatrically with this species. Distinguishing these two species is sometime difficult because both have quite an irregular shell, and exhibit a rather large intraspecific variability. Nevertheless, there are several differences as follows: 
<taxonomicName lsidName="A. megastoma" pageId="0" pageNumber="1" rank="species" species="megastoma">
<emphasis italics="true" pageId="0" pageNumber="1">A. megastoma</emphasis>
</taxonomicName>
sp. nov. is larger, and has a generally lower spire; its aperture is less rounded (rather semilunar because the body whorl is more attached to the penultimate whorl); the spiral striation of 
<taxonomicName lsidName="A. megastoma" pageId="0" pageNumber="1" rank="species" species="megastoma">
<emphasis italics="true" pageId="0" pageNumber="1">A. megastoma</emphasis>
</taxonomicName>
sp. nov. is denser and much fainter, in fact it is nearly invisible under the stereo microscope (only visible with SEM); the last whorl of 
<taxonomicName lsidName="A. megastoma" pageId="0" pageNumber="1" rank="species" species="megastoma">
<emphasis italics="true" pageId="0" pageNumber="1">A. megastoma</emphasis>
</taxonomicName>
sp. nov. is wider compared to the penultimate whorl than that of 
<taxonomicName lsidName="A. thersites" pageId="0" pageNumber="1" rank="species" species="thersites">
<emphasis italics="true" pageId="0" pageNumber="1">A. thersites</emphasis>
</taxonomicName>
sp. nov. (in the latter species often the penultimate whorl is wider than the last one in apertural view). See also under 
<taxonomicName lsidName="A. tonkinospiroides" pageId="0" pageNumber="1" rank="species" species="tonkinospiroides">
<emphasis italics="true" pageId="0" pageNumber="1">A. tonkinospiroides</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
This new species is named after Thersites (
<emphasis italics="true" pageId="0" pageNumber="1">Θερσίτης</emphasis>
= the ugliest Greek soldier laying siege to Troy) referring to the irregularly growing whorls.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species is known from several sites in the Halong Bay Area, northern Vietnam (Fig. 
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Distribution of Angustopila species." figureDoi="10.3897/zookeys.1147.93824.figure22" httpUri="https://binary.pensoft.net/fig/808831" pageId="0" pageNumber="1">22</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1147.93824.figure20" httpUri="https://binary.pensoft.net/fig/808829" pageId="0" pageNumber="1" start="Figure 20" startId="F20">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 20.</emphasis>
<taxonomicName authorityName="Pall-Gergely &amp; Vermeulen" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila thersites" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="thersites">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila thersites</emphasis>
</taxonomicName>
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; Vermeulen, sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
paratype (RMNH 347761) 
<emphasis bold="true" pageId="0" pageNumber="1">B-H</emphasis>
holotype (RMNH 347760). Apertural (
<emphasis bold="true" pageId="0" pageNumber="1">A, B</emphasis>
), lateral (
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
), ventral (
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
) and apical (
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
) sides of the shell; granular sculpture on the protoconch (
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
), ventral (
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
) and frontal (
<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
) surface of the body whorl.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1147.93824.figure21" httpUri="https://binary.pensoft.net/fig/808830" pageId="0" pageNumber="1" start="Figure 21" startId="F21">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 21.</emphasis>
<taxonomicName authorityName="Pall-Gergely &amp; Vermeulen" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila thersites" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="thersites">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila thersites</emphasis>
</taxonomicName>
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; Vermeulen, sp. nov., sample WMVT.0327 (RMNH 347764) 
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
specimen 2 
<emphasis bold="true" pageId="0" pageNumber="1">B-I</emphasis>
specimen 1. Apertural (
<emphasis bold="true" pageId="0" pageNumber="1">A, B</emphasis>
), lateral (
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
), ventral (
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
) and apical (
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
) sides of the shell; granular sculpture on the protoconch (
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
), aperture (
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
), ventral (
<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
) and frontal (
<emphasis bold="true" pageId="0" pageNumber="1">I</emphasis>
) surface of the body whorl.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1147.93824.figure22" httpUri="https://binary.pensoft.net/fig/808831" pageId="0" pageNumber="1" start="Figure 22" startId="F22">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 22.</emphasis>
Distribution of 
<taxonomicName authorityName="Jochum, Slapnik &amp; Pall-Gergely" authorityYear="2014" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila</emphasis>
</taxonomicName>
species.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
Shells from sample WMVT.0327 are unusually high-spired and are less concave-conical (some of them are even conical) than typical 
<taxonomicName lsidName="A. thersites" pageId="0" pageNumber="1" rank="species" species="thersites">
<emphasis italics="true" pageId="0" pageNumber="1">A. thersites</emphasis>
</taxonomicName>
sp. nov. shells. Although the differences in shell shape between most shells of sample WMVT.0327 and other 
<taxonomicName lsidName="A. thersites" pageId="0" pageNumber="1" rank="species" species="thersites">
<emphasis italics="true" pageId="0" pageNumber="1">A. thersites</emphasis>
</taxonomicName>
sp. nov. shells are obvious, the shell with the lowest spire of sample WMVT.0327, and the shell with the highest spire of the other 
<taxonomicName lsidName="A. thersites" pageId="0" pageNumber="1" rank="species" species="thersites">
<emphasis italics="true" pageId="0" pageNumber="1">A. thersites</emphasis>
</taxonomicName>
sp. nov. samples are very similar. In addition, the strong spiral striation, the aperture shape, the narrow umbilicus, and the very oblique aperture suggests that they are conspecific with 
<taxonomicName lsidName="A. thersites" pageId="0" pageNumber="1" rank="species" species="thersites">
<emphasis italics="true" pageId="0" pageNumber="1">A. thersites</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</subSubSection>
</treatment>
</document>