721 lines
107 KiB
XML
721 lines
107 KiB
XML
<document id="5E6B71B2B0CB1064DE7D7C2FD072C5E6" ID-CLB-Dataset="92411" ID-DOI="10.1080/00222933.2022.2049389" ID-GBIF-Dataset="4cebde67-ce4c-402d-b8d5-901a5a528fab" ID-ISSN="1464-5262" ID-Zenodo-Dep="6758291" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1656311063267" checkinUser="plazi" docAuthor="Leever, Ellen M., Daniels, Savel R., Soma, Julia B. & Cumberlidge, Neil" docDate="2022" docId="9529DB19FFE3FFAF3A6CFF34FBD18844" docLanguage="en" docName="JNATHIST.56.1-4.241-263.pdf" docOrigin="Journal of Natural History 56 (1 - 4)" docSource="http://dx.doi.org/10.1080/00222933.2022.2049389" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Vahatra ambohitra Leever & Daniels & Soma & Cumberlidge 2022, comb. nov." docType="treatment" docVersion="9" lastPageNumber="254" masterDocId="6910A361FFE9FFA23B75FFBAFFF98911" masterDocTitle="Two new genera and a new species of freshwater crabs from northern Madagascar: Vahatra gen. nov. for Foza ambohitra Cumberlidge and Meyer, 2009, and Toamasina gen. nov. for Toamasina clarki sp. nov. (Brachyura: Potamoidea: Deckeniidae)" masterLastPageNumber="263" masterPageNumber="241" pageNumber="251" updateTime="1699350272557" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="7E584D29B569A961E59DDB1ADEA23A6F">Two new genera and a new species of freshwater crabs from northern Madagascar: Vahatra gen. nov. for Foza ambohitra Cumberlidge and Meyer, 2009, and Toamasina gen. nov. for Toamasina clarki sp. nov. (Brachyura: Potamoidea: Deckeniidae)</mods:title>
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<mods:namePart id="5F793D0BC97BA428FEE286A1A58A0A36">Leever, Ellen M.</mods:namePart>
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<mods:namePart id="72AC803152E284139332EDCFE62DFB18">Daniels, Savel R.</mods:namePart>
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<mods:namePart id="5F010EE8D11D2112090E53D16866AFD3">Soma, Julia B.</mods:namePart>
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<mods:title id="75F904F29631D7822B185D53AE2678A5">Journal of Natural History</mods:title>
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<mods:date id="CF1A55C4D3059E6561E5B367DF3FB27B">2022</mods:date>
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<treatment id="9529DB19FFE3FFAF3A6CFF34FBD18844" ID-DOI="http://doi.org/10.5281/zenodo.6762505" ID-GBIF-Taxon="197158407" ID-Zenodo-Dep="6762505" LSID="urn:lsid:plazi:treatment:9529DB19FFE3FFAF3A6CFF34FBD18844" httpUri="http://treatment.plazi.org/id/9529DB19FFE3FFAF3A6CFF34FBD18844" lastPageId="13" lastPageNumber="254" pageId="10" pageNumber="251">
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<paragraph id="1D3F6A0FFFE3FFA83A6CFF34FBF289B9" blockId="10.[281,1035,142,168]" box="[281,1035,142,168]" pageId="10" pageNumber="251">
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<heading id="4677DD63FFE3FFA83A6CFF34FBF289B9" box="[281,1035,142,168]" centered="true" fontSize="11" level="1" pageId="10" pageNumber="251" reason="6">
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<taxonomicName id="DA80118CFFE3FFA83A6CFF34FC8689B9" authority="(Cumberlidge and Meyer 2009)" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" baseAuthorityName="Cumberlidge and Meyer" baseAuthorityYear="2009" box="[281,895,142,168]" class="Malacostraca" family="Potamonautidae" genus="Vahatra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="251" phylum="Arthropoda" rank="species" species="ambohitra" status="comb. nov.">
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<emphasis id="2FF4B61DFFE3FFA83A6CFF34FDF189B9" bold="true" box="[281,520,142,168]" pageId="10" pageNumber="251">Vahatra ambohitra</emphasis>
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(
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<bibRefCitation id="791117FEFFE3FFA83963FF34FC8E89B9" author="Cumberlidge N & Meyer KS" box="[534,887,142,168]" pageId="10" pageNumber="251" pagination="77 - 89" refId="ref11416" refString="Cumberlidge N, Meyer KS. 2009. A new species of Foza Reed & Cumberlidge, 2006 from northern Madagascar (Decapoda, Brachyura, Potamoidea, Potamonautidae), with a redescription of F. goudoti (H. Milne Edwards, 1853) comb. n., and comments on Skelosophusa prolixa Ng & Takeda, 1994. ZooKeys. 18: 77 - 89. doi: 10.3897 / zookeys. 18.102." type="journal article" year="2009">Cumberlidge and Meyer 2009</bibRefCitation>
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)
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</taxonomicName>
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<taxonomicNameLabel id="34C70B66FFE3FFA838F2FF34FBF289B9" box="[903,1035,142,168]" pageId="10" pageNumber="251" rank="species">comb. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="559A3984FFE3FFA83A3FFF79FC0389CD" box="[330,1018,195,220]" pageId="10" pageNumber="251" type="description">
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<paragraph id="1D3F6A0FFFE3FFA83A3FFF79FC0389CD" blockId="10.[330,1018,195,220]" box="[330,1018,195,220]" pageId="10" pageNumber="251">
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(
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<figureCitation id="85BB768AFFE3FFA83A24FF79FE4A89CD" box="[337,435,195,220]" captionStart="Figure 1" captionStartId="1.[160,225,1227,1250]" captionTargetBox="[163,1153,142,1196]" captionTargetId="figure-116@1.[163,1153,142,1196]" captionTargetPageId="1" captionText="Figure 1. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar, views of the cephalothorax: a, dorsal, c, frontal, e, ventral. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar, views of the cephalothorax: b, dorsal; d, frontal, and f, ventral. Scale bar: a, c, e = 8.4 mm; b, d, f = 8.8 mm." figureDoi="http://doi.org/10.5281/zenodo.6758293" httpUri="https://zenodo.org/record/6758293/files/figure.png" pageId="10" pageNumber="251">Figures 1</figureCitation>
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(a,c,e), 2(a,c,e), 3(a), 4(a,b,e,f), 5(a,b,c), 6;
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<tableCitation id="50025FB4FFE3FFA8380FFF79FC0889CD" box="[890,1009,195,220]" captionStart-0="Table 1" captionStart-1="Table 2" captionStart-2="Table 3" captionStart-3="Table 4" captionStartId-0="3.[224,281,142,165]" captionStartId-1="17.[160,216,142,165]" captionStartId-2="18.[160,216,142,165]" captionStartId-3="18.[160,216,534,557]" captionTargetBox-0="[235,821,235,1031]" captionTargetPageId-1="17" captionTargetPageId-2="18" captionTargetPageId-3="18" captionText-0="Table 1. Updated list of all known Malagasy freshwater crabs (Family Deckeniidae Ortmann, 1897, Subfamily Hydrothelphusinae Bott, 1955) from 1853 to July 2021. The" captionText-1="Table 2. Measurements (in mm) of the terminal article of the mandibular palp (MPTA) and the anterior lobe of the terminal article of the mandibular palp (MPAL), and size class for five species of Malagasy freshwater crabs that are either currently (or were formerly) assigned to Foza. The relative length of the MPAL and the MPTA is also shown, along with the size classes for the MPAL." captionText-2="Table 3. Total length of the five articles (ischium, merus, carpus, propodus, dactylus) of the cheliped (P1) and the four ambulatory legs (P2–5) in relation to the carapace width (CW) for five species of Malagasy freshwater crabs that either are currently or were formerly assigned to Foza. The sum of all four ambulatory legs divided by the carapace width is also shown (ΣP2–5/CW), along with the size classes for the ambulatory legs." captionText-3="Table 4. Gonopod 1 (G1) measurements (taken on the ventral side) showing lengths of the G1 terminal article (TA) and the G1 subterminal article (SA) in mm for five species of Malagasy freshwater crabs that either are currently or were formerly assigned to Foza. The relative length of the G1 TA and the G1 SA is also shown, along with the size classes for the G1 TA." httpUri-0="http://table.plazi.org/id/49FF3A87FFEAFFA13B95FF34FC1A89F1" httpUri-1="http://table.plazi.org/id/49FF3A87FFF8FFB33BD5FF34FC3A89EC" httpUri-2="http://table.plazi.org/id/49FF3A87FFFBFFB03BD5FF34FE20880B" pageId="10" pageNumber="251" tableUuid-0="49FF3A87FFEAFFA13B95FF34FC1A89F1" tableUuid-1="49FF3A87FFF8FFB33BD5FF34FC3A89EC" tableUuid-2="49FF3A87FFFBFFB03BD5FF34FE20880B">Tables 1–4</tableCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="559A3984FFE3FFA83BD5FEB2FE908855" pageId="10" pageNumber="251" type="reference_group">
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<paragraph id="1D3F6A0FFFE3FFA83BD5FEB2FE908855" blockId="10.[160,1155,264,324]" pageId="10" pageNumber="251">
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<treatmentCitationGroup id="3D904D21FFE3FFA83BD5FEB2FE908855" pageId="10" pageNumber="251">
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<treatmentCitation id="9C214C1EFFE3FFA83BD5FEB2FD068830" ID-DOI="http://doi.org/10.5281/zenodo.3791531" ID-Zenodo-Dep="3791531" LSID="urn:lsid:zoobank.org:act:1007E7DF-D87E-41A9-AB9D-B3628DB72C7D" author="Cumberlidge N & Meyer KS" box="[160,767,264,289]" httpUri="http://treatment.plazi.org/id/03EF536EFFC5466FFF173FCFFEBCF9EB" page="84" pageId="10" pageNumber="251" year="2009">
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<taxonomicName id="DA80118CFFE3FFA83BD5FEB2FD068830" authority="Cumberlidge and Meyer, 2009: 79 - 84" authorityName="Cumberlidge and Meyer" authorityPageNumber="79 - 84" authorityYear="2009" box="[160,767,264,289]" class="Malacostraca" family="Potamonautidae" genus="Foza" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="251" phylum="Arthropoda" rank="species" species="ambohitra">
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<emphasis id="2FF4B61DFFE3FFA83BD5FEB2FEB48830" box="[160,333,264,289]" italics="true" pageId="10" pageNumber="251">Foza ambohitra</emphasis>
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<bibRefCitation id="791117FEFFE3FFA83A20FEB2FD068830" author="Cumberlidge N & Meyer KS" box="[341,767,264,289]" pageId="10" pageNumber="251" pagination="77 - 89" refId="ref11416" refString="Cumberlidge N, Meyer KS. 2009. A new species of Foza Reed & Cumberlidge, 2006 from northern Madagascar (Decapoda, Brachyura, Potamoidea, Potamonautidae), with a redescription of F. goudoti (H. Milne Edwards, 1853) comb. n., and comments on Skelosophusa prolixa Ng & Takeda, 1994. ZooKeys. 18: 77 - 89. doi: 10.3897 / zookeys. 18.102." type="journal article" year="2009">Cumberlidge and Meyer, 2009: 79–84</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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, figs 1–3, tab. 1) (partim, not NMU PN 17–
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<date id="693E4CCFFFE3FFA83B99FE91FEA68855" box="[236,351,299,324]" pageId="10" pageNumber="251" value="2003-03-21">21.3.2003</date>
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).
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</treatmentCitationGroup>
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</paragraph>
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<subSubSection id="559A3984FFE3FFA83BD5FE3AFE9C8CCD" pageId="10" pageNumber="251" type="materials_examined">
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<emphasis id="2FF4B61DFFE3FFA83BD5FE3AFE86888B" bold="true" box="[160,383,384,410]" italics="true" pageId="10" pageNumber="251">Material examined</emphasis>
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</paragraph>
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<paragraph id="1D3F6A0FFFE3FFA83BD5FE18FEDA8B35" blockId="10.[160,1156,384,1311]" pageId="10" pageNumber="251">
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<emphasis id="2FF4B61DFFE3FFA83BD5FE18FEB288AD" bold="true" box="[160,331,418,444]" italics="true" pageId="10" pageNumber="251">Type material.</emphasis>
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<materialsCitation id="ADE86052FFE3FFA83A2EFE19FEDA8B35" ID-GBIF-Occurrence="3861175317" accessionNumber="B 30154, MT749728" collectingDate="2005-05" collectionCode="FW, ASL, MNHN" collectorName="R. Roy" country="Madagascar" county="Ambohitra" elevation="421" latitude="-12.45" location="Antsiranana Province" longLatPrecision="5" longitude="49.13333" municipality="Diana Region" pageId="10" pageNumber="251" specimenCode="CW 39.5, CL 32.4" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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:
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: adult
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(
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,
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, CH 16.3,
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),
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<location id="185F3CD4FFE3FFA83BD5FE7CFE6288CE" LSID="urn:lsid:plazi:treatment:9529DB19FFE3FFAF3A6CFF34FBD18844:185F3CD4FFE3FFA83BD5FE7CFE6288CE" box="[160,411,454,479]" country="Madagascar" county="Ambohitra" latitude="-12.45" longLatPrecision="5" longitude="49.13333" municipality="Diana Region" name="Antsiranana Province" pageId="10" pageNumber="251">Antsiranana Province</location>
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,
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<collectingMunicipality id="FD5BF075FFE3FFA83ADFFE7CFDB288CE" box="[426,587,454,479]" pageId="10" pageNumber="251">Diana Region</collectingMunicipality>
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, forest close to
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<collectingCounty id="F45E1283FFE3FFA83867FE7CFC7588CE" box="[786,908,454,479]" pageId="10" pageNumber="251">Ambohitra</collectingCounty>
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(formerly Joffreville) (
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,
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),
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</quantity>
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<collectionCode id="7B91F2CAFFE3FFA83961FE52FDB88B10" box="[532,577,488,513]" pageId="10" pageNumber="251">ASL</collectionCode>
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, coll.
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<collectorName id="B0750FD9FFE3FFA839F6FE52FD358B10" box="[643,716,488,513]" pageId="10" pageNumber="251">R. Roy</collectorName>
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<date id="693E4CCFFFE3FFA839ADFE52FCBE8B10" box="[728,839,488,513]" pageId="10" pageNumber="251" value="2005-05">
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<collectingDate id="797AB527FFE3FFA839ADFE52FCBE8B10" box="[728,839,488,513]" pageId="10" pageNumber="251" value="2005-05">May 2005</collectingDate>
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(
|
||
<collectionCode id="7B91F2CAFFE3FFA83823FE52FC5B8B10" box="[854,930,488,513]" country="France" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="10" pageNumber="251">MNHN</collectionCode>
|
||
B 30154) (GenBank
|
||
<accessionNumber id="02D3F7ECFFE3FFA83BD5FDB1FEE08B35" box="[160,281,523,548]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MT749728" pageId="10" pageNumber="251">MT749728</accessionNumber>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="1D3F6A0FFFE3FFA83BB5FD94FD7B8C0E" blockId="10.[160,1156,384,1311]" pageId="10" pageNumber="251">
|
||
<materialsCitation id="ADE86052FFE3FFA83BB5FD94FE368BC0" ID-GBIF-Occurrence="3861175318" collectionCode="FW, CW, ASL" country="Switzerland" elevation="200" latitude="-12.45" location="Ankavanana Forest" longLatPrecision="5" longitude="49.13333" municipality="Anivorano-Nord" pageId="10" pageNumber="251" specimenCode="CW 37.8, CL 29.6, CL 23.6" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
|
||
<typeStatus id="C23BD4ADFFE3FFA83BB5FD94FEC98B56" box="[192,304,558,583]" pageId="10" pageNumber="251" type="paratype">Paratypes</typeStatus>
|
||
: Madagascar: adult
|
||
<specimenCount id="0B86A186FFE3FFA83967FD94FDDF8B54" box="[530,550,558,581]" pageId="10" pageNumber="251" type="female">♀</specimenCount>
|
||
(
|
||
<specimenCode id="4D26C274FFE3FFA83940FD94FD6E8B56" box="[565,663,558,583]" collectionCode="CW" pageId="10" pageNumber="251">CW 37.8</specimenCode>
|
||
,
|
||
<specimenCode id="4D26C274FFE3FFA839D6FD94FD008B56" box="[675,761,558,583]" collectionCode="CL" pageId="10" pageNumber="251">CL 29.6</specimenCode>
|
||
, CH 15.5,
|
||
<collectionCode id="7B91F2CAFFE3FFA8381EFD94FC768B56" box="[875,911,558,583]" pageId="10" pageNumber="251">FW</collectionCode>
|
||
8.0 mm), subadult
|
||
<specimenCount id="0B86A186FFE3FFA83F1AFD94FB7A8B54" box="[1135,1155,558,581]" pageId="10" pageNumber="251" type="female">♀</specimenCount>
|
||
(
|
||
<collectionCode id="7B91F2CAFFE3FFA83BD0FDEAFF348B78" box="[165,205,592,617]" pageId="10" pageNumber="251">CW</collectionCode>
|
||
30.0,
|
||
<specimenCode id="4D26C274FFE3FFA83A78FDEAFE998B78" box="[269,352,592,617]" collectionCode="CL" pageId="10" pageNumber="251">CL 23.6</specimenCode>
|
||
,
|
||
<collectingCountry id="65972A9FFFE3FFA83A1CFDEAFE738B78" box="[361,394,592,617]" name="Switzerland" pageId="10" pageNumber="251">CH</collectingCountry>
|
||
12.7,
|
||
<collectionCode id="7B91F2CAFFE3FFA83ABCFDEAFE148B78" box="[457,493,592,617]" pageId="10" pageNumber="251">FW</collectionCode>
|
||
<quantity id="DA78C7EAFFE3FFA83A87FDEAFDB18B78" box="[498,584,592,617]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="10" pageNumber="251" unit="mm" value="6.6">6.6 mm</quantity>
|
||
), Antsiranana Province, Diana Region, Analamerana Special Reserve,
|
||
<location id="185F3CD4FFE3FFA83A2AFDC9FDC38B9D" LSID="urn:lsid:plazi:treatment:9529DB19FFE3FFAF3A6CFF34FBD18844:185F3CD4FFE3FFA83A2AFDC9FDC38B9D" box="[351,570,627,652]" country="Switzerland" latitude="-12.45" longLatPrecision="5" longitude="49.13333" municipality="Anivorano-Nord" name="Ankavanana Forest" pageId="10" pageNumber="251">Ankavanana Forest</location>
|
||
,
|
||
<quantity id="DA78C7EAFFE3FFA83933FDC9FD5B8B9D" box="[582,674,627,652]" metricMagnitude="4" metricUnit="m" metricValue="1.58" pageId="10" pageNumber="251" unit="km" value="15.8">15.8 km</quantity>
|
||
south-east of
|
||
<collectingMunicipality id="FD5BF075FFE3FFA8383EFDC9FC068B9D" box="[843,1023,627,652]" pageId="10" pageNumber="251">Anivorano-Nord</collectingMunicipality>
|
||
(
|
||
<geoCoordinate id="78B40CC8FFE3FFA83F7AFDC9FB798B9D" box="[1039,1152,627,652]" degrees="12.4500" direction="south" orientation="latitude" pageId="10" pageNumber="251" precision="5" value="-12.45">12.4500°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="78B40CC8FFE3FFA83BD5FD2CFED78BBE" box="[160,302,662,687]" degrees="49.133333" direction="east" orientation="longitude" pageId="10" pageNumber="251" precision="1" value="49.13333">49.133333°E</geoCoordinate>
|
||
), from partially disturbed mixed dry deciduous and humid forest, in early morning,
|
||
<quantity id="DA78C7EAFFE3FFA83A66FD02FEA78BC0" box="[275,350,696,721]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="251" unit="m" value="200.0">
|
||
<elevation id="96AD8D3CFFE3FFA83A66FD02FEA78BC0" box="[275,350,696,721]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="251" unit="m" value="200.0">200 m</elevation>
|
||
</quantity>
|
||
<collectionCode id="7B91F2CAFFE3FFA83A1FFD02FE6E8BC0" box="[362,407,696,721]" pageId="10" pageNumber="251">ASL</collectionCode>
|
||
, coll
|
||
</materialsCitation>
|
||
<materialsCitation id="ADE86052FFE3FFA83ABAFD02FE0F8BC0" ID-GBIF-Occurrence="3861175312" box="[463,502,696,721]" collectionCode="FW, CW, ASL" country="Switzerland" elevation="200" isTailEnumeration="true" latitude="-12.45" location="Ankavanana Forest" longLatPrecision="5" longitude="49.13333" municipality="Anivorano-Nord" pageId="10" pageNumber="251" specimenCount="1" typeStatus="paratype">. M</materialsCitation>
|
||
. Raheriarisena, S.M
|
||
<materialsCitation id="ADE86052FFE3FFA839ADFD02FE3E8A28" ID-GBIF-Occurrence="3861175316" collectingDate="2004-01-23" collectionCode="FW, CW, CL" collectorName="Goodman" country="Switzerland" elevation="200" isTailEnumeration="true" latitude="-12.45" location="Ankavanana Forest" longLatPrecision="5" longitude="49.13333" municipality="Anivorano-Nord" pageId="10" pageNumber="251" specimenCode="FMNH 11045, CW 38.4, CL 28.8" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
|
||
.
|
||
<collectorName id="B0750FD9FFE3FFA83992FD02FCA38BC0" box="[743,858,696,721]" pageId="10" pageNumber="251">Goodman</collectorName>
|
||
,
|
||
<date id="693E4CCFFFE3FFA8381CFD02FBD38BC0" box="[873,1066,696,721]" pageId="10" pageNumber="251" value="2004-01-23">
|
||
<collectingDate id="797AB527FFE3FFA8381CFD02FBD38BC0" box="[873,1066,696,721]" pageId="10" pageNumber="251" value="2004-01-23">23 January 2004</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCode id="4D26C274FFE3FFA83F4EFD02FF108BE5" collectionCode="FMNH" country="USA" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="10" pageNumber="251" type="Museum">FMNH 11045</specimenCode>
|
||
); adult
|
||
<specimenCount id="0B86A186FFE3FFA83A48FD61FEA88BE3" box="[317,337,731,754]" pageId="10" pageNumber="251" type="male">♂</specimenCount>
|
||
(
|
||
<specimenCode id="4D26C274FFE3FFA83A2AFD61FE388BE5" box="[351,449,731,756]" collectionCode="CW" pageId="10" pageNumber="251">CW 38.4</specimenCode>
|
||
,
|
||
<specimenCode id="4D26C274FFE3FFA83AB8FD61FDDA8BE5" box="[461,547,731,756]" collectionCode="CL" pageId="10" pageNumber="251">CL 28.8</specimenCode>
|
||
,
|
||
<collectingCountry id="65972A9FFFE3FFA8395AFD61FDA98BE5" box="[559,592,731,756]" name="Switzerland" pageId="10" pageNumber="251">CH</collectingCountry>
|
||
15.0,
|
||
<collectionCode id="7B91F2CAFFE3FFA839E0FD61FD408BE5" box="[661,697,731,756]" pageId="10" pageNumber="251">FW</collectionCode>
|
||
<quantity id="DA78C7EAFFE3FFA839B7FD61FCE38BE5" box="[706,794,731,756]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="10" pageNumber="251" unit="mm" value="8.3">8.3 mm</quantity>
|
||
), adult
|
||
<specimenCount id="0B86A186FFE3FFA83805FD61FC7D8BE3" box="[880,900,731,754]" pageId="10" pageNumber="251" type="female">♀</specimenCount>
|
||
(
|
||
<collectionCode id="7B91F2CAFFE3FFA838E6FD61FC428BE5" box="[915,955,731,756]" pageId="10" pageNumber="251">CW</collectionCode>
|
||
35.0,
|
||
<collectionCode id="7B91F2CAFFE3FFA83F74FD61FBE48BE5" box="[1025,1053,731,756]" pageId="10" pageNumber="251">CL</collectionCode>
|
||
28.0,
|
||
<collectingCountry id="65972A9FFFE3FFA83F17FD61FB7A8BE5" box="[1122,1155,731,756]" name="Switzerland" pageId="10" pageNumber="251">CH</collectingCountry>
|
||
14.3,
|
||
<collectionCode id="7B91F2CAFFE3FFA83BA8FD44FEF88A06" box="[221,257,766,791]" pageId="10" pageNumber="251">FW</collectionCode>
|
||
<quantity id="DA78C7EAFFE3FFA83A7CFD44FE9B8A06" box="[265,354,766,791]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="10" pageNumber="251" unit="mm" value="8.1">8.1 mm</quantity>
|
||
), same data as FMNH 11045, but found during night at edge of small stream at forest edge, coll
|
||
</materialsCitation>
|
||
<materialsCitation id="ADE86052FFE3FFA83AB3FC9AFE1A8A28" ID-GBIF-Occurrence="3861175327" box="[454,483,800,825]" collectingDate="2004-01-23" collectionCode="FW, CW, CL" collectorName="Goodman" country="Switzerland" elevation="200" isTailEnumeration="true" latitude="-12.45" location="Ankavanana Forest" longLatPrecision="5" longitude="49.13333" municipality="Anivorano-Nord" pageId="10" pageNumber="251" specimenCount="1" typeStatus="paratype">. H</materialsCitation>
|
||
<materialsCitation id="ADE86052FFE3FFA83A91FC9AFDF08A6E" ID-GBIF-Occurrence="3861175311" accessionNumber="MT749727" collectingDate="2004-01-23" collectionCode="FW" collectorName="Rakotondravony & GenBank" country="Switzerland" elevation="200" isTailEnumeration="true" latitude="-12.45" location="Ankavanana Forest" longLatPrecision="5" longitude="49.13333" municipality="Anivorano-Nord" pageId="10" pageNumber="251" specimenCode="FMNH 11046, CW 37.1, CL 29.2" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
|
||
.
|
||
<collectorName id="B0750FD9FFE3FFA83A9BFC9AFD508A28" box="[494,681,800,825]" pageId="10" pageNumber="251">Rakotondravony</collectorName>
|
||
,
|
||
<date id="693E4CCFFFE3FFA839C6FC9AFC938A28" box="[691,874,800,825]" pageId="10" pageNumber="251" value="2004-01-23">
|
||
<collectingDate id="797AB527FFE3FFA839C6FC9AFC938A28" box="[691,874,800,825]" pageId="10" pageNumber="251" value="2004-01-23">23 January 2004</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCode id="4D26C274FFE3FFA83803FC9AFBF28A28" box="[886,1035,800,825]" collectionCode="FMNH" country="USA" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="10" pageNumber="251" type="Museum">FMNH 11046</specimenCode>
|
||
) (
|
||
<collectorName id="B0750FD9FFE3FFA83F68FC9AFB7A8A28" box="[1053,1155,800,825]" pageId="10" pageNumber="251">GenBank</collectorName>
|
||
<accessionNumber id="02D3F7ECFFE3FFA83BD5FCF9FEE08A4D" box="[160,281,835,860]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MT749727" pageId="10" pageNumber="251">MT749727</accessionNumber>
|
||
); adult
|
||
<specimenCount id="0B86A186FFE3FFA83A1EFCF9FE868A4B" box="[363,383,835,858]" pageId="10" pageNumber="251" type="male">♂</specimenCount>
|
||
(
|
||
<specimenCode id="4D26C274FFE3FFA83AF9FCF9FE148A4D" box="[396,493,835,860]" collectionCode="CW" pageId="10" pageNumber="251">CW 37.1</specimenCode>
|
||
,
|
||
<specimenCode id="4D26C274FFE3FFA83A8DFCF9FDB48A4D" box="[504,589,835,860]" collectionCode="CL" pageId="10" pageNumber="251">CL 29.2</specimenCode>
|
||
,
|
||
<collectingCountry id="65972A9FFFE3FFA83922FCF9FD818A4D" box="[599,632,835,860]" name="Switzerland" pageId="10" pageNumber="251">CH</collectingCountry>
|
||
16.8,
|
||
<collectionCode id="7B91F2CAFFE3FFA839C9FCF9FD198A4D" box="[700,736,835,860]" pageId="10" pageNumber="251">FW</collectionCode>
|
||
<quantity id="DA78C7EAFFE3FFA83992FCF9FCC68A4D" box="[743,831,835,860]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="10" pageNumber="251" unit="mm" value="8.1">8.1 mm</quantity>
|
||
), same data as FMNH 11045, but found in late afternoon, coll
|
||
</materialsCitation>
|
||
<materialsCitation id="ADE86052FFE3FFA8397CFCDCFDDC8A6E" ID-GBIF-Occurrence="3861175305" accessionNumber="MT749727" box="[521,549,870,895]" collectingDate="2004-01-23" collectionCode="FW" collectorName="Rakotondravony & GenBank" country="Switzerland" elevation="200" isTailEnumeration="true" latitude="-12.45" location="Ankavanana Forest" longLatPrecision="5" longitude="49.13333" municipality="Anivorano-Nord" pageId="10" pageNumber="251" specimenCount="1" typeStatus="paratype">. S</materialsCitation>
|
||
.M
|
||
<materialsCitation id="ADE86052FFE3FFA8394BFCDCFDF88AD5" ID-GBIF-Occurrence="3861175302" accessionNumber="MT749727" collectingDate="2004-01-25" collectionCode="CW, FW" collectorName="Goodman" country="Switzerland" elevation="200" isTailEnumeration="true" latitude="-12.45" location="Ankavanana Forest" longLatPrecision="5" longitude="49.13333" municipality="Anivorano-Nord" pageId="10" pageNumber="251" specimenCode="FMNH 11050, CL 28.7, CW 33.2, CL 26.2" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
|
||
.
|
||
<collectorName id="B0750FD9FFE3FFA8393DFCDCFD438A6E" box="[584,698,870,895]" pageId="10" pageNumber="251">Goodman</collectorName>
|
||
,
|
||
<date id="693E4CCFFFE3FFA839B1FCDCFC828A6E" box="[708,891,870,895]" pageId="10" pageNumber="251" value="2004-01-25">
|
||
<collectingDate id="797AB527FFE3FFA839B1FCDCFC828A6E" box="[708,891,870,895]" pageId="10" pageNumber="251" value="2004-01-25">25 January 2004</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCode id="4D26C274FFE3FFA838F2FCDCFBE68A6E" box="[903,1055,870,895]" collectionCode="FMNH" country="USA" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="10" pageNumber="251" type="Museum">FMNH 11050</specimenCode>
|
||
); adult
|
||
<specimenCount id="0B86A186FFE3FFA83F1AFCDCFB7A8A6C" box="[1135,1155,870,893]" pageId="10" pageNumber="251" type="female">♀</specimenCount>
|
||
(
|
||
<collectionCode id="7B91F2CAFFE3FFA83BD0FC32FF348AB0" box="[165,205,904,929]" pageId="10" pageNumber="251">CW</collectionCode>
|
||
38.0,
|
||
<specimenCode id="4D26C274FFE3FFA83A65FC32FE9C8AB0" box="[272,357,904,929]" collectionCode="CL" pageId="10" pageNumber="251">CL 28.7</specimenCode>
|
||
,
|
||
<collectingCountry id="65972A9FFFE3FFA83A1AFC32FE698AB0" box="[367,400,904,929]" name="Switzerland" pageId="10" pageNumber="251">CH</collectingCountry>
|
||
15.7,
|
||
<collectionCode id="7B91F2CAFFE3FFA83AA6FC32FE0E8AB0" box="[467,503,904,929]" pageId="10" pageNumber="251">FW</collectionCode>
|
||
<quantity id="DA78C7EAFFE3FFA83A8BFC32FDAC8AB0" box="[510,597,904,929]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="10" pageNumber="251" unit="mm" value="8.5">8.5 mm</quantity>
|
||
), adult
|
||
<specimenCount id="0B86A186FFE3FFA839DDFC33FD458AB1" box="[680,700,905,928]" pageId="10" pageNumber="251" type="male">♂</specimenCount>
|
||
(
|
||
<specimenCode id="4D26C274FFE3FFA839BCFC32FCD08AB0" box="[713,809,904,929]" collectionCode="CW" pageId="10" pageNumber="251">CW 33.2</specimenCode>
|
||
,
|
||
<specimenCode id="4D26C274FFE3FFA83841FC32FC718AB0" box="[820,904,904,929]" collectionCode="CL" pageId="10" pageNumber="251">CL 26.2</specimenCode>
|
||
,
|
||
<collectingCountry id="65972A9FFFE3FFA838E6FC32FC4D8AB0" box="[915,948,904,929]" name="Switzerland" pageId="10" pageNumber="251">CH</collectingCountry>
|
||
14.5,
|
||
<collectionCode id="7B91F2CAFFE3FFA83883FC32FBE38AB0" box="[1014,1050,904,929]" pageId="10" pageNumber="251">FW</collectionCode>
|
||
<quantity id="DA78C7EAFFE3FFA83F54FC32FB808AB0" box="[1057,1145,904,929]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="10" pageNumber="251" unit="mm" value="7.5">7.5 mm</quantity>
|
||
), same data as FMNH 11050, coll
|
||
</materialsCitation>
|
||
<materialsCitation id="ADE86052FFE3FFA83975FC11FDDB8AD5" ID-GBIF-Occurrence="3861175322" accessionNumber="MT749727" box="[512,546,939,964]" collectingDate="2004-01-25" collectionCode="CW, FW" collectorName="Goodman" country="Switzerland" elevation="200" isTailEnumeration="true" latitude="-12.45" location="Ankavanana Forest" longLatPrecision="5" longitude="49.13333" municipality="Anivorano-Nord" pageId="10" pageNumber="251" specimenCount="1" typeStatus="paratype">. M</materialsCitation>
|
||
. Raheriarisena, S. M
|
||
<materialsCitation id="ADE86052FFE3FFA83875FC11FE518D18" ID-GBIF-Occurrence="3861175301" accessionNumber="MT749727" collectingDate="2004-01-25" collectionCode="FW" collectorName="Goodman" country="Switzerland" elevation="200" isTailEnumeration="true" latitude="-12.45" location="Ankavanana Forest" longLatPrecision="5" longitude="49.13333" municipality="Anivorano-Nord" pageId="10" pageNumber="251" specimenCode="FMNH 11051, CW 39.1, CL 30.7" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||
.
|
||
<collectorName id="B0750FD9FFE3FFA8387EFC11FC838AD5" box="[779,890,939,964]" pageId="10" pageNumber="251">Goodman</collectorName>
|
||
(
|
||
<specimenCode id="4D26C274FFE3FFA838F2FC11FBE78AD5" box="[903,1054,939,964]" collectionCode="FMNH" country="USA" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="10" pageNumber="251" type="Museum">FMNH 11051</specimenCode>
|
||
); adult
|
||
<specimenCount id="0B86A186FFE3FFA83F1AFC11FB7A8AD3" box="[1135,1155,939,962]" pageId="10" pageNumber="251" type="female">♀</specimenCount>
|
||
(
|
||
<specimenCode id="4D26C274FFE3FFA83BD0FC74FEFE8AF6" box="[165,263,974,999]" collectionCode="CW" pageId="10" pageNumber="251">CW 39.1</specimenCode>
|
||
,
|
||
<specimenCode id="4D26C274FFE3FFA83A67FC74FE9E8AF6" box="[274,359,974,999]" collectionCode="CL" pageId="10" pageNumber="251">CL 30.7</specimenCode>
|
||
,
|
||
<collectingCountry id="65972A9FFFE3FFA83A06FC74FE6D8AF6" box="[371,404,974,999]" name="Switzerland" pageId="10" pageNumber="251">CH</collectingCountry>
|
||
17.1,
|
||
<collectionCode id="7B91F2CAFFE3FFA83AADFC74FE058AF6" box="[472,508,974,999]" pageId="10" pageNumber="251">FW</collectionCode>
|
||
<quantity id="DA78C7EAFFE3FFA83971FC74FDA58AF6" box="[516,604,974,999]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="10" pageNumber="251" unit="mm" value="8.7">8.7 mm</quantity>
|
||
), same data as FMNH 11045, but found at forest edge during night, coll
|
||
</materialsCitation>
|
||
<materialsCitation id="ADE86052FFE3FFA83AD2FC4AFE468D18" ID-GBIF-Occurrence="3861175307" accessionNumber="MT749727" box="[423,447,1008,1033]" collectingDate="2004-01-25" collectionCode="FW" collectorName="Goodman" country="Switzerland" elevation="200" isTailEnumeration="true" latitude="-12.45" location="Ankavanana Forest" longLatPrecision="5" longitude="49.13333" municipality="Anivorano-Nord" pageId="10" pageNumber="251" specimenCount="1" typeStatus="paratype">. J</materialsCitation>
|
||
<materialsCitation id="ADE86052FFE3FFA83ACAFC4AFF338D5E" ID-GBIF-Occurrence="3861175308" accessionNumber="MT749727" collectingDate="2004-01-25" collectionCode="FW" collectorName="Ravino" country="Switzerland" elevation="200" isTailEnumeration="true" latitude="-12.45" location="Ankavanana Forest" longLatPrecision="5" longitude="49.13333" municipality="Anivorano-Nord" pageId="10" pageNumber="251" specimenCode="FMNH 11052, CW 41.4, CL 31.9" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||
.
|
||
<collectorName id="B0750FD9FFE3FFA83ABEFC4AFDE08D18" box="[459,537,1008,1033]" pageId="10" pageNumber="251">Ravino</collectorName>
|
||
,
|
||
<date id="693E4CCFFFE3FFA83951FC4AFD278D18" box="[548,734,1008,1033]" pageId="10" pageNumber="251" value="2004-01-25">
|
||
<collectingDate id="797AB527FFE3FFA83951FC4AFD278D18" box="[548,734,1008,1033]" pageId="10" pageNumber="251" value="2004-01-25">25 January 2004</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCode id="4D26C274FFE3FFA83999FC4AFC7C8D18" box="[748,901,1008,1033]" collectionCode="FMNH" country="USA" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="10" pageNumber="251" type="Museum">FMNH 11052</specimenCode>
|
||
); adult
|
||
<specimenCount id="0B86A186FFE3FFA838ADFC4BFC158D19" box="[984,1004,1009,1032]" pageId="10" pageNumber="251" type="female">♀</specimenCount>
|
||
(
|
||
<specimenCode id="4D26C274FFE3FFA8388FFC4AFBA58D18" box="[1018,1116,1008,1033]" collectionCode="CW" pageId="10" pageNumber="251">CW 41.4</specimenCode>
|
||
,
|
||
<specimenCode id="4D26C274FFE3FFA83F12FC4AFF288D3D" collectionCode="CL" pageId="10" pageNumber="251">CL 31.9</specimenCode>
|
||
,
|
||
<collectingCountry id="65972A9FFFE3FFA83BA9FBA9FF048D3D" box="[220,253,1043,1068]" name="Switzerland" pageId="10" pageNumber="251">CH</collectingCountry>
|
||
17.4,
|
||
<collectionCode id="7B91F2CAFFE3FFA83A4AFBA9FE9A8D3D" box="[319,355,1043,1068]" pageId="10" pageNumber="251">FW</collectionCode>
|
||
<quantity id="DA78C7EAFFE3FFA83A1FFBA9FE388D3D" box="[362,449,1043,1068]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="10" pageNumber="251" unit="mm" value="8.7">8.7 mm</quantity>
|
||
), same data as FMNH 11045, but found on ground during day, coll
|
||
</materialsCitation>
|
||
<materialsCitation id="ADE86052FFE3FFA83BBCFB8CFF108D5E" ID-GBIF-Occurrence="3861175319" accessionNumber="MT749727" box="[201,233,1078,1103]" collectingDate="2004-01-25" collectionCode="FW" collectorName="Ravino" country="Switzerland" elevation="200" isTailEnumeration="true" latitude="-12.45" location="Ankavanana Forest" longLatPrecision="5" longitude="49.13333" municipality="Anivorano-Nord" pageId="10" pageNumber="251" specimenCount="1" typeStatus="paratype">. M</materialsCitation>
|
||
. Raheriarisena, S.M
|
||
<materialsCitation id="ADE86052FFE3FFA83AC8FB8CFB9E8D60" ID-GBIF-Occurrence="3861175303" accessionNumber="MT749727" collectingDate="2004-01-26" collectionCode="FW" collectorName="Goodman" country="Switzerland" elevation="200" isTailEnumeration="true" latitude="-12.45" location="Ankavanana Forest" longLatPrecision="5" longitude="49.13333" municipality="Anivorano-Nord" pageId="10" pageNumber="251" specimenCode="FMNH 11054, CW 43.1, CL 34.2" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||
.
|
||
<collectorName id="B0750FD9FFE3FFA83AB3FB8CFDC18D5E" box="[454,568,1078,1103]" pageId="10" pageNumber="251">Goodman</collectorName>
|
||
,
|
||
<date id="693E4CCFFFE3FFA83935FB8CFD0D8D5E" box="[576,756,1078,1103]" pageId="10" pageNumber="251" value="2004-01-26">
|
||
<collectingDate id="797AB527FFE3FFA83935FB8CFD0D8D5E" box="[576,756,1078,1103]" pageId="10" pageNumber="251" value="2004-01-26">26 January 2004</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCode id="4D26C274FFE3FFA8398AFB8CFC6D8D5E" box="[767,916,1078,1103]" collectionCode="FMNH" country="USA" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="10" pageNumber="251" type="Museum">FMNH 11054</specimenCode>
|
||
); adult
|
||
<specimenCount id="0B86A186FFE3FFA83894FB8CFC0C8D5C" box="[993,1013,1078,1101]" pageId="10" pageNumber="251" type="female">♀</specimenCount>
|
||
(
|
||
<specimenCode id="4D26C274FFE3FFA83F75FB8CFBA68D5E" box="[1024,1119,1078,1103]" collectionCode="CW" pageId="10" pageNumber="251">CW 43.1</specimenCode>
|
||
,
|
||
<specimenCode id="4D26C274FFE3FFA83F12FB8CFF288D60" collectionCode="CL" pageId="10" pageNumber="251">CL 34.2</specimenCode>
|
||
,
|
||
<collectingCountry id="65972A9FFFE3FFA83BAFFBE2FF028D60" box="[218,251,1112,1137]" name="Switzerland" pageId="10" pageNumber="251">CH</collectingCountry>
|
||
18.3,
|
||
<collectionCode id="7B91F2CAFFE3FFA83A49FBE2FE998D60" box="[316,352,1112,1137]" pageId="10" pageNumber="251">FW</collectionCode>
|
||
<quantity id="DA78C7EAFFE3FFA83A13FBE2FE448D60" box="[358,445,1112,1137]" metricMagnitude="-3" metricUnit="m" metricValue="9.9" pageId="10" pageNumber="251" unit="mm" value="9.9">9.9 mm</quantity>
|
||
), same data as FMNH 11045, but found in late afternoon, coll
|
||
</materialsCitation>
|
||
<materialsCitation id="ADE86052FFE3FFA83F12FBE2FB798D60" ID-GBIF-Occurrence="3861175310" accessionNumber="MT749727" box="[1127,1152,1112,1137]" collectingDate="2004-01-26" collectionCode="FW" collectorName="Goodman" country="Switzerland" elevation="200" isTailEnumeration="true" latitude="-12.45" location="Ankavanana Forest" longLatPrecision="5" longitude="49.13333" municipality="Anivorano-Nord" pageId="10" pageNumber="251" specimenCount="1" typeStatus="paratype">. S</materialsCitation>
|
||
. M. Goodman, M
|
||
<materialsCitation id="ADE86052FFE3FFA83A23FBC1FC6C8D85" ID-GBIF-Occurrence="3861175324" box="[342,917,1147,1172]" collectingDate="2004-01-28" collectionCode="FMNH" collectorName="Raheriarisena" country="Switzerland" pageId="10" pageNumber="251" specimenCode="FMNH 11056" specimenCount="1" typeStatus="paratype">
|
||
.
|
||
<collectorName id="B0750FD9FFE3FFA83A14FBC1FE038D85" box="[353,506,1147,1172]" pageId="10" pageNumber="251">Raheriarisena</collectorName>
|
||
,
|
||
<date id="693E4CCFFFE3FFA83971FBC1FD458D85" box="[516,700,1147,1172]" pageId="10" pageNumber="251" value="2004-01-28">
|
||
<collectingDate id="797AB527FFE3FFA83971FBC1FD458D85" box="[516,700,1147,1172]" pageId="10" pageNumber="251" value="2004-01-28">28 January 2004</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCode id="4D26C274FFE3FFA839BCFBC1FC998D85" box="[713,864,1147,1172]" collectionCode="FMNH" country="USA" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="10" pageNumber="251" type="Museum">FMNH 11056</specimenCode>
|
||
); juv
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="ADE86052FFE3FFA838EAFBC1FBB48DED" ID-GBIF-Occurrence="3861175325" collectionCode="FW, ASL" country="Switzerland" county="Bobakindro River" elevation="40" latitude="-12.45" location="Antsiranana Province" longLatPrecision="5" longitude="49.13333" municipality="Foret d'Analabe" pageId="10" pageNumber="251" specimenCode="CW 22.8, CL 17.6" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
|
||
<specimenCount id="0B86A186FFE3FFA838EAFBC1FC4A8D83" box="[927,947,1147,1170]" pageId="10" pageNumber="251" type="male">♂</specimenCount>
|
||
(
|
||
<specimenCode id="4D26C274FFE3FFA838B5FBC1FBD88D85" box="[960,1057,1147,1172]" collectionCode="CW" pageId="10" pageNumber="251">CW 22.8</specimenCode>
|
||
,
|
||
<specimenCode id="4D26C274FFE3FFA83F5EFBC1FB798D85" box="[1067,1152,1147,1172]" collectionCode="CL" pageId="10" pageNumber="251">CL 17.6</specimenCode>
|
||
,
|
||
<collectingCountry id="65972A9FFFE3FFA83BD5FB24FF388DA6" box="[160,193,1182,1207]" name="Switzerland" pageId="10" pageNumber="251">CH</collectingCountry>
|
||
9.4,
|
||
<collectionCode id="7B91F2CAFFE3FFA83B89FB24FED98DA6" box="[252,288,1182,1207]" pageId="10" pageNumber="251">FW</collectionCode>
|
||
<quantity id="DA78C7EAFFE3FFA83A5FFB24FE7C8DA6" box="[298,389,1182,1207]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="10" pageNumber="251" unit="mm" value="4.7">4.7 mm</quantity>
|
||
),
|
||
<location id="185F3CD4FFE3FFA83AEFFB24FD748DA6" LSID="urn:lsid:plazi:treatment:9529DB19FFE3FFAF3A6CFF34FBD18844:185F3CD4FFE3FFA83AEFFB24FD748DA6" box="[410,653,1182,1207]" country="Switzerland" county="Bobakindro River" latitude="-12.45" longLatPrecision="5" longitude="49.13333" municipality="Foret d'Analabe" name="Antsiranana Province" pageId="10" pageNumber="251">Antsiranana Province</location>
|
||
, Analamerana Special Reserve, Ankavanana Forest,
|
||
<quantity id="DA78C7EAFFE3FFA83B84FB7AFEC58DC8" box="[241,316,1216,1241]" metricMagnitude="3" metricUnit="m" metricValue="8.6" pageId="10" pageNumber="251" unit="km" value="8.6">8.6 km</quantity>
|
||
south-east of
|
||
<location id="185F3CD4FFE3FFA83AAEFB7AFDB18DC8" LSID="urn:lsid:plazi:treatment:9529DB19FFE3FFAF3A6CFF34FBD18844:185F3CD4FFE3FFA83AAEFB7AFDB18DC8" box="[475,584,1216,1241]" country="Switzerland" county="Bobakindro River" latitude="-12.45" longLatPrecision="5" longitude="49.13333" municipality="Foret d'Analabe" name="Menagisy" pageId="10" pageNumber="251">Menagisy</location>
|
||
,
|
||
<collectingMunicipality id="FD5BF075FFE3FFA83927FB7AFCFF8DC8" box="[594,774,1216,1241]" pageId="10" pageNumber="251">Foret d’Analabe</collectingMunicipality>
|
||
, along
|
||
<collectingCounty id="F45E1283FFE3FFA83823FB7AFBED8DC8" box="[854,1044,1216,1241]" pageId="10" pageNumber="251">Bobakindro River</collectingCounty>
|
||
(
|
||
<geoCoordinate id="78B40CC8FFE3FFA83F54FB7AFF498DED" degrees="12.4500" direction="south" orientation="latitude" pageId="10" pageNumber="251" precision="5" value="-12.45">12.4500° S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="78B40CC8FFE3FFA83BCDFB59FEBF8DED" box="[184,326,1251,1276]" degrees="49.133333" direction="east" orientation="longitude" pageId="10" pageNumber="251" precision="1" value="49.13333">49.133333°E</geoCoordinate>
|
||
), from disturbed dry deciduous forest, in early morning,
|
||
<quantity id="DA78C7EAFFE3FFA838C4FB59FC118DED" box="[945,1000,1251,1276]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="10" pageNumber="251" unit="m" value="40.0">
|
||
<elevation id="96AD8D3CFFE3FFA838C4FB59FC118DED" box="[945,1000,1251,1276]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="10" pageNumber="251" unit="m" value="40.0">40 m</elevation>
|
||
</quantity>
|
||
<collectionCode id="7B91F2CAFFE3FFA8389BFB59FBE28DED" box="[1006,1051,1251,1276]" pageId="10" pageNumber="251">ASL</collectionCode>
|
||
, coll
|
||
</materialsCitation>
|
||
<materialsCitation id="ADE86052FFE3FFA83F38FB59FB918DED" ID-GBIF-Occurrence="3861175323" box="[1101,1128,1251,1276]" collectionCode="FW, ASL" country="Switzerland" county="Bobakindro River" elevation="40" isTailEnumeration="true" latitude="-12.45" location="Antsiranana Province" longLatPrecision="5" longitude="49.13333" municipality="Foret d'Analabe" pageId="10" pageNumber="251" specimenCount="1" typeStatus="paratype">. S</materialsCitation>
|
||
.M
|
||
<materialsCitation id="ADE86052FFE3FFA83FF5FB59FD878C0E" ID-GBIF-Occurrence="3861175306" collectingDate="2004-01-17" collectionCode="FMNH" collectorName="Goodman" country="Switzerland" pageId="10" pageNumber="251" specimenCode="FMNH 11060" specimenCount="1" typeStatus="paratype">
|
||
.
|
||
<collectorName id="B0750FD9FFE3FFA83BD5FABCFEEB8C0E" box="[160,274,1286,1311]" pageId="10" pageNumber="251">Goodman</collectorName>
|
||
,
|
||
<date id="693E4CCFFFE3FFA83A69FABCFE2D8C0E" box="[284,468,1286,1311]" pageId="10" pageNumber="251" value="2004-01-17">
|
||
<collectingDate id="797AB527FFE3FFA83A69FABCFE2D8C0E" box="[284,468,1286,1311]" pageId="10" pageNumber="251" value="2004-01-17">17 January 2004</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCode id="4D26C274FFE3FFA83A94FABCFD808C0E" box="[481,633,1286,1311]" collectionCode="FMNH" country="USA" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="10" pageNumber="251" type="Museum">FMNH 11060</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="1D3F6A0FFFE3FFA83BD5FAE0FE9C8CCD" blockId="10.[160,1157,1370,1500]" pageId="10" pageNumber="251">
|
||
<emphasis id="2FF4B61DFFE3FFA83BD5FAE0FEA58C65" bold="true" box="[160,348,1370,1396]" italics="true" pageId="10" pageNumber="251">Other material.</emphasis>
|
||
<materialsCitation id="ADE86052FFE3FFA83A19FAE1FF338CA8" ID-GBIF-Occurrence="3861175313" collectionCode="CL" country="Madagascar" latitude="-12.92" location="Ankarana Special Reserve" longLatPrecision="7" longitude="49.14" municipality="Diana region" pageId="10" pageNumber="251" specimenCode="CW 50, CW 43" specimenCount="3" specimenCount-adult="2" specimenCount-male="1" stateProvince="Antsiranana">
|
||
<collectingCountry id="65972A9FFFE3FFA83A19FAE1FE0E8C65" box="[364,503,1371,1396]" name="Madagascar" pageId="10" pageNumber="251">Madagascar</collectingCountry>
|
||
:
|
||
<specimenCount id="0B86A186FFE3FFA83970FAE1FDA08C65" box="[517,601,1371,1396]" pageId="10" pageNumber="251" type="adult">2 adult</specimenCount>
|
||
<specimenCount id="0B86A186FFE3FFA83910FAE1FD748C63" box="[613,653,1371,1394]" pageId="10" pageNumber="251" type="male">♂♂</specimenCount>
|
||
(
|
||
<specimenCode id="4D26C274FFE3FFA839EAFAE1FD088C65" box="[671,753,1371,1396]" collectionCode="CW" pageId="10" pageNumber="251">CW 50</specimenCode>
|
||
, Cl
|
||
<quantity id="DA78C7EAFFE3FFA83857FAE1FC758C65" box="[802,908,1371,1396]" metricMagnitude="-2" metricUnit="m" metricValue="3.16" pageId="10" pageNumber="251" unit="mm" value="31.6">31.6 mm</quantity>
|
||
;
|
||
<specimenCode id="4D26C274FFE3FFA838EEFAE1FC128C65" box="[923,1003,1371,1396]" collectionCode="CW" pageId="10" pageNumber="251">CW 43</specimenCode>
|
||
,
|
||
<collectionCode id="7B91F2CAFFE3FFA8388FFAE1FBEF8C65" box="[1018,1046,1371,1396]" pageId="10" pageNumber="251">CL</collectionCode>
|
||
<quantity id="DA78C7EAFFE3FFA83F57FAE1FB808C65" box="[1058,1145,1371,1396]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="251" unit="mm" value="30.0">30 mm</quantity>
|
||
),
|
||
<collectingRegion id="DF44A4EDFFE3FFA83BD5FAC4FE6E8C86" box="[160,407,1406,1431]" country="Madagascar" name="Antsiranana" pageId="10" pageNumber="251">Antsiranana Province</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="FD5BF075FFE3FFA83AD0FAC4FDC78C86" box="[421,574,1406,1431]" pageId="10" pageNumber="251">Diana region</collectingMunicipality>
|
||
,
|
||
<location id="185F3CD4FFE3FFA83939FAC4FC8C8C86" LSID="urn:lsid:plazi:treatment:9529DB19FFE3FFAF3A6CFF34FBD18844:185F3CD4FFE3FFA83939FAC4FC8C8C86" box="[588,885,1406,1431]" country="Madagascar" latitude="-12.92" longLatPrecision="7" longitude="49.14" municipality="Diana region" name="Ankarana Special Reserve" pageId="10" pageNumber="251" stateProvince="Antsiranana">Ankarana Special Reserve</location>
|
||
(
|
||
<geoCoordinate id="78B40CC8FFE3FFA838F2FAC4FC008C86" box="[903,1017,1406,1431]" degrees="12.9200" direction="south" orientation="latitude" pageId="10" pageNumber="251" precision="5" value="-12.92">12.9200°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="78B40CC8FFE3FFA83F7DFAC4FB828C86" box="[1032,1147,1406,1431]" degrees="49.1400" direction="east" orientation="longitude" pageId="10" pageNumber="251" precision="5" value="49.14">49.1400°E</geoCoordinate>
|
||
), coll
|
||
</materialsCitation>
|
||
<materialsCitation id="ADE86052FFE3FFA83BBFFA1AFF1C8CA8" ID-GBIF-Occurrence="3861175314" box="[202,229,1440,1465]" collectionCode="CL" country="Madagascar" isTailEnumeration="true" latitude="-12.92" location="Ankarana Special Reserve" longLatPrecision="7" longitude="49.14" municipality="Diana region" pageId="10" pageNumber="251" specimenCount="1" stateProvince="Antsiranana">. F</materialsCitation>
|
||
. Glaw, M. Franzen, J. Köhler and N
|
||
<materialsCitation id="ADE86052FFE3FFA8390FFA1AFE988CCD" ID-GBIF-Occurrence="3861175321" collectingDate="2008-02-13" collectionCode="ZSM" country="Madagascar" pageId="10" pageNumber="251" specimenCode="A20145001" specimenCount="1" stateProvince="Antsiranana">
|
||
. d’Cruze,
|
||
<date id="693E4CCFFFE3FFA8399EFA1AFC4D8CA8" box="[747,948,1440,1465]" pageId="10" pageNumber="251" value="2008-02-13">
|
||
<collectingDate id="797AB527FFE3FFA8399EFA1AFC4D8CA8" box="[747,948,1440,1465]" pageId="10" pageNumber="251" value="2008-02-13">13 February 2008</collectingDate>
|
||
</date>
|
||
(
|
||
<collectionCode id="7B91F2CAFFE3FFA838B6FA1AFC0F8CA8" box="[963,1014,1440,1465]" country="Germany" name="Bavarian State Collection of Zoology" pageId="10" pageNumber="251" type="Museum">ZSM</collectionCode>
|
||
<specimenCode id="4D26C274FFE3FFA83F75FA1AFB788CA8" box="[1024,1153,1440,1465]" collectionCode="ZSM" country="Germany" name="Bavarian State Collection of Zoology" pageId="10" pageNumber="251" type="Museum">A20145001</specimenCode>
|
||
, ZSM A20145002)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="559A3984FFE3FFA83BD5F9ADFECD8F45" box="[160,308,1559,1620]" pageId="10" pageNumber="251" type="diagnosis">
|
||
<paragraph id="1D3F6A0FFFE3FFA83BD5F9ADFEED8F20" blockId="10.[160,308,1559,1620]" box="[160,276,1559,1585]" pageId="10" pageNumber="251">
|
||
<emphasis id="2FF4B61DFFE3FFA83BD5F9ADFEED8F20" bold="true" box="[160,276,1559,1585]" italics="true" pageId="10" pageNumber="251">Diagnosis</emphasis>
|
||
</paragraph>
|
||
<paragraph id="1D3F6A0FFFE3FFA83BD5F981FECD8F45" blockId="10.[160,308,1559,1620]" box="[160,308,1595,1620]" pageId="10" pageNumber="251">As for genus.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="559A3984FFE2FFAE3BD5FF34FC8E89DA" lastPageId="12" lastPageNumber="253" pageId="11" pageNumber="252" type="description">
|
||
<paragraph id="1D3F6A0FFFE2FFA93BD5FF34FEB989B9" blockId="11.[160,1156,142,1520]" box="[160,320,142,168]" pageId="11" pageNumber="252">
|
||
<emphasis id="2FF4B61DFFE2FFA93BD5FF34FEB989B9" bold="true" box="[160,320,142,168]" italics="true" pageId="11" pageNumber="252">Redescription</emphasis>
|
||
</paragraph>
|
||
<paragraph id="1D3F6A0FFFE2FFA93BD5FF08FB948CE1" blockId="11.[160,1156,142,1520]" pageId="11" pageNumber="252">
|
||
Based on
|
||
<typeStatus id="C23BD4ADFFE2FFA93A64FF08FE8189DA" box="[273,376,178,203]" pageId="11" pageNumber="252" type="holotype">holotype</typeStatus>
|
||
, adult
|
||
<specimenCount id="0B86A186FFE2FFA93AB0FF08FE2089D8" box="[453,473,178,201]" pageId="11" pageNumber="252" type="male">♂</specimenCount>
|
||
. Carapace outline transversely oval, extremely high (
|
||
<collectingCountry id="65972A9FFFE2FFA93F44FF08FBAA89DA" box="[1073,1107,178,203]" name="Switzerland" pageId="11" pageNumber="252">CH</collectingCountry>
|
||
/CW 0.42); front narrow (FW/CW = 0.22), deflexed; entire carapace surface generally smooth. Epibranchial tooth small, acute, close to exorbital tooth, positioned in line with postorbital margin; lateral margin evenly curved outward, smooth, continuous with posterolateral margin; postfrontal crest faint, incomplete, not traversing entire carapace, epigastric crests faint, in line with postorbital margin, postorbital crests faint, ending before meeting exorbital tooth; cardiac, urogastric sulci faint, semicircular sulcus deep, cervical sulcus broad, faint, long, not meeting postorbital crest (
|
||
<figureCitation id="85BB768AFFE2FFA9399FFE1EFCBF88AC" box="[746,838,420,445]" captionStart="Figure 1" captionStartId="1.[160,225,1227,1250]" captionTargetBox="[163,1153,142,1196]" captionTargetId="figure-116@1.[163,1153,142,1196]" captionTargetPageId="1" captionText="Figure 1. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar, views of the cephalothorax: a, dorsal, c, frontal, e, ventral. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar, views of the cephalothorax: b, dorsal; d, frontal, and f, ventral. Scale bar: a, c, e = 8.4 mm; b, d, f = 8.8 mm." figureDoi="http://doi.org/10.5281/zenodo.6758293" httpUri="https://zenodo.org/record/6758293/files/figure.png" pageId="11" pageNumber="252">Figure 1</figureCitation>
|
||
(a)). Suborbital, subhepatic regions of branchiostegite smooth, pterygostomial region smooth except for small field of granules at junction of longitudinal, vertical sulci; vertical sulcus on branchiostegite curved, granular, running downward from base of epibranchial tooth to epimeral sulcus (
|
||
<figureCitation id="85BB768AFFE2FFA93BDDFD95FF048B59" box="[168,253,559,584]" captionStart="Figure 1" captionStartId="1.[160,225,1227,1250]" captionTargetBox="[163,1153,142,1196]" captionTargetId="figure-116@1.[163,1153,142,1196]" captionTargetPageId="1" captionText="Figure 1. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar, views of the cephalothorax: a, dorsal, c, frontal, e, ventral. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar, views of the cephalothorax: b, dorsal; d, frontal, and f, ventral. Scale bar: a, c, e = 8.4 mm; b, d, f = 8.8 mm." figureDoi="http://doi.org/10.5281/zenodo.6758293" httpUri="https://zenodo.org/record/6758293/files/figure.png" pageId="11" pageNumber="252">Figure 1</figureCitation>
|
||
(c)). Epistomial tooth triangular, deflexed, edges smooth (
|
||
<figureCitation id="85BB768AFFE2FFA9380EFD95FC288B59" box="[891,977,559,584]" captionStart="Figure 1" captionStartId="1.[160,225,1227,1250]" captionTargetBox="[163,1153,142,1196]" captionTargetId="figure-116@1.[163,1153,142,1196]" captionTargetPageId="1" captionText="Figure 1. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar, views of the cephalothorax: a, dorsal, c, frontal, e, ventral. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar, views of the cephalothorax: b, dorsal; d, frontal, and f, ventral. Scale bar: a, c, e = 8.4 mm; b, d, f = 8.8 mm." figureDoi="http://doi.org/10.5281/zenodo.6758293" httpUri="https://zenodo.org/record/6758293/files/figure.png" pageId="11" pageNumber="252">Figure 1</figureCitation>
|
||
(c)). Mandibular palp terminal article bilobed, anterior lobe on terminal article conspicuous, medium-sized (MPAL/MPTA = 0.4) (
|
||
<figureCitation id="85BB768AFFE2FFA93AFFFDCEFE1F8B9C" box="[394,486,628,653]" captionStart="Figure 2" captionStartId="6.[160,225,1616,1639]" captionTargetBox="[163,1153,142,1584]" captionTargetId="figure-3@6.[163,1153,142,1584]" captionTargetPageId="6" captionText="Figure 2. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154,from Diana Region,Madagascar:a, frontal view of mandible;c, superior view of mandible;e, outer view of left third maxilliped. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17– 21.3.2003,from Toamasina Province,Madagascar:b, dorsal view of mandible;d, superior view of mandible; f,outer view of left third maxilliped.Scale bar:a–d = 1.0 mm;e, f = 2.2 mm.The anterior lobeof the palphas been shaded yellow (A-D) to distinguish it from the long setae that line the margins of this lobe." figureDoi="http://doi.org/10.5281/zenodo.6758295" httpUri="https://zenodo.org/record/6758295/files/figure.png" pageId="11" pageNumber="252">Figure 2</figureCitation>
|
||
(a,c);
|
||
<tableCitation id="50025FB4FFE2FFA93954FDCEFD8A8B9C" box="[545,627,628,653]" captionStart="Table 2" captionStartId="17.[160,216,142,165]" captionTargetPageId="17" captionText="Table 2. Measurements (in mm) of the terminal article of the mandibular palp (MPTA) and the anterior lobe of the terminal article of the mandibular palp (MPAL), and size class for five species of Malagasy freshwater crabs that are either currently (or were formerly) assigned to Foza. The relative length of the MPAL and the MPTA is also shown, along with the size classes for the MPAL." httpUri="http://table.plazi.org/id/49FF3A87FFF8FFB33BD5FF34FC3A89EC" pageId="11" pageNumber="252" tableUuid="49FF3A87FFF8FFB33BD5FF34FC3A89EC">Table 2</tableCitation>
|
||
). Exopod of third maxilliped reaching to lower half of merus, exopod with short flagellum (less than half merus length), ischium with deep vertical sulcus, beginning distally close to medial margin, angled diagonally downward (
|
||
<figureCitation id="85BB768AFFE2FFA93B90FD66FEB98BE4" box="[229,320,732,757]" captionStart="Figure 2" captionStartId="6.[160,225,1616,1639]" captionTargetBox="[163,1153,142,1584]" captionTargetId="figure-3@6.[163,1153,142,1584]" captionTargetPageId="6" captionText="Figure 2. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154,from Diana Region,Madagascar:a, frontal view of mandible;c, superior view of mandible;e, outer view of left third maxilliped. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17– 21.3.2003,from Toamasina Province,Madagascar:b, dorsal view of mandible;d, superior view of mandible; f,outer view of left third maxilliped.Scale bar:a–d = 1.0 mm;e, f = 2.2 mm.The anterior lobeof the palphas been shaded yellow (A-D) to distinguish it from the long setae that line the margins of this lobe." figureDoi="http://doi.org/10.5281/zenodo.6758295" httpUri="https://zenodo.org/record/6758295/files/figure.png" pageId="11" pageNumber="252">Figure 2</figureCitation>
|
||
(e)). Sternal suture S1/2 short, faint; S2/3 completely traversing sternum; S3/ 4 broad, U-shaped, deepest at edges, not meeting anterior margin of sternopleonal cavity; anterior sternopleonal cavity lacking setae; S4/5 meeting pleon at telson/PL6 suture; S6/7 meeting midpoint of lateral margin of PL6; episternal sulci S4/E4, S5/E5, S6/E6, S7/E7 absent, smooth (
|
||
<figureCitation id="85BB768AFFE2FFA93A14FCDDFE438A91" box="[353,442,871,896]" captionStart="Figure 1" captionStartId="1.[160,225,1227,1250]" captionTargetBox="[163,1153,142,1196]" captionTargetId="figure-116@1.[163,1153,142,1196]" captionTargetPageId="1" captionText="Figure 1. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar, views of the cephalothorax: a, dorsal, c, frontal, e, ventral. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar, views of the cephalothorax: b, dorsal; d, frontal, and f, ventral. Scale bar: a, c, e = 8.4 mm; b, d, f = 8.8 mm." figureDoi="http://doi.org/10.5281/zenodo.6758293" httpUri="https://zenodo.org/record/6758293/files/figure.png" pageId="11" pageNumber="252">Figure 1</figureCitation>
|
||
(e)). Right chela largest, proximal half of cutting edge of fixed finger (pollex of propodus) with four large molars, small teeth distally; cutting edge of movable finger (dactylus) with two large teeth proximally, one large tooth midway (
|
||
<figureCitation id="85BB768AFFE2FFA93BDDFC75FEFF8AF9" box="[168,262,975,1000]" captionStart="Figure 4" captionStartId="8.[160,225,1234,1257]" captionTargetBox="[163,1153,142,1202]" captionTargetId="figure-58@8.[163,1153,142,1202]" captionTargetPageId="8" captionText="Figure 4. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar: a, frontal view of right chela; b, frontal view of left chela; e, inferior view of merus of right cheliped; f, dorsal view of carpus and merus of right cheliped. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar: c, frontal view of right chela; d, frontal view of left chela; g, right cheliped carpus and merus dorsal view; h, inferior view of merus of right cheliped. Scale bar: a–d = 5.0 mm; e, f = 4.0 mm; g, h = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6758301" httpUri="https://zenodo.org/record/6758301/files/figure.png" pageId="11" pageNumber="252">Figure 4</figureCitation>
|
||
(a)); left chela subequal to right chela, dentition same as right chela (
|
||
<figureCitation id="85BB768AFFE2FFA93F50FC75FB7A8AF9" box="[1061,1155,975,1000]" captionStart="Figure 4" captionStartId="8.[160,225,1234,1257]" captionTargetBox="[163,1153,142,1202]" captionTargetId="figure-58@8.[163,1153,142,1202]" captionTargetPageId="8" captionText="Figure 4. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar: a, frontal view of right chela; b, frontal view of left chela; e, inferior view of merus of right cheliped; f, dorsal view of carpus and merus of right cheliped. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar: c, frontal view of right chela; d, frontal view of left chela; g, right cheliped carpus and merus dorsal view; h, inferior view of merus of right cheliped. Scale bar: a–d = 5.0 mm; e, f = 4.0 mm; g, h = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6758301" httpUri="https://zenodo.org/record/6758301/files/figure.png" pageId="11" pageNumber="252">Figure 4</figureCitation>
|
||
(b)); lower margin of propodus of both chela slightly indented medially (
|
||
<figureCitation id="85BB768AFFE2FFA93892FC48FBBE8D1A" box="[999,1095,1010,1035]" captionStart="Figure 4" captionStartId="8.[160,225,1234,1257]" captionTargetBox="[163,1153,142,1202]" captionTargetId="figure-58@8.[163,1153,142,1202]" captionTargetPageId="8" captionText="Figure 4. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar: a, frontal view of right chela; b, frontal view of left chela; e, inferior view of merus of right cheliped; f, dorsal view of carpus and merus of right cheliped. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar: c, frontal view of right chela; d, frontal view of left chela; g, right cheliped carpus and merus dorsal view; h, inferior view of merus of right cheliped. Scale bar: a–d = 5.0 mm; e, f = 4.0 mm; g, h = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6758301" httpUri="https://zenodo.org/record/6758301/files/figure.png" pageId="11" pageNumber="252">Figure 4</figureCitation>
|
||
(a,b)). Distal tooth on inner margin of cheliped carpus large, pointed; proximal tooth significantly smaller, acute, followed by small granules (
|
||
<figureCitation id="85BB768AFFE2FFA939BCFB8DFCDC8D41" box="[713,805,1079,1104]" captionStart="Figure 4" captionStartId="8.[160,225,1234,1257]" captionTargetBox="[163,1153,142,1202]" captionTargetId="figure-58@8.[163,1153,142,1202]" captionTargetPageId="8" captionText="Figure 4. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar: a, frontal view of right chela; b, frontal view of left chela; e, inferior view of merus of right cheliped; f, dorsal view of carpus and merus of right cheliped. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar: c, frontal view of right chela; d, frontal view of left chela; g, right cheliped carpus and merus dorsal view; h, inferior view of merus of right cheliped. Scale bar: a–d = 5.0 mm; e, f = 4.0 mm; g, h = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6758301" httpUri="https://zenodo.org/record/6758301/files/figure.png" pageId="11" pageNumber="252">Figure 4</figureCitation>
|
||
(f)). Inferior margins of cheliped merus both lined with small, rounded teeth, distal tooth largest; superior side of cheliped merus roughened by granules, short striae; cheliped ischium margins smooth, rounded (
|
||
<figureCitation id="85BB768AFFE2FFA93BDDFB25FEFF8DA9" box="[168,262,1183,1208]" captionStart="Figure 4" captionStartId="8.[160,225,1234,1257]" captionTargetBox="[163,1153,142,1202]" captionTargetId="figure-58@8.[163,1153,142,1202]" captionTargetPageId="8" captionText="Figure 4. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar: a, frontal view of right chela; b, frontal view of left chela; e, inferior view of merus of right cheliped; f, dorsal view of carpus and merus of right cheliped. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar: c, frontal view of right chela; d, frontal view of left chela; g, right cheliped carpus and merus dorsal view; h, inferior view of merus of right cheliped. Scale bar: a–d = 5.0 mm; e, f = 4.0 mm; g, h = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6758301" httpUri="https://zenodo.org/record/6758301/files/figure.png" pageId="11" pageNumber="252">Figure 4</figureCitation>
|
||
(e)). Ambulatory legs P2–5 short (ΣP2–5/CW = 5.7) (
|
||
<figureCitation id="85BB768AFFE2FFA9382CFB25FC418DA9" box="[857,952,1183,1208]" captionStart="Figure 3" captionStartId="7.[160,225,797,820]" captionTargetBox="[163,1153,142,765]" captionTargetId="figure-326@7.[163,1153,142,765]" captionTargetPageId="7" captionText="Figure 3. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., paratype, adult ♀, CW 35.0 mm, FMNH 11046, from Analamerana Special Reserve, Madagascar, right ambulatory legs (a). Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar, right cheliped and ambulatory legs (b). Scale bar = 6.6 mm." figureDoi="http://doi.org/10.5281/zenodo.6758297" httpUri="https://zenodo.org/record/6758297/files/figure.png" pageId="11" pageNumber="252">Figure 3</figureCitation>
|
||
(a);
|
||
<tableCitation id="50025FB4FFE2FFA93897FB25FBCE8DA9" box="[994,1079,1183,1208]" captionStart="Table 3" captionStartId="18.[160,216,142,165]" captionTargetPageId="18" captionText="Table 3. Total length of the five articles (ischium, merus, carpus, propodus, dactylus) of the cheliped (P1) and the four ambulatory legs (P2–5) in relation to the carapace width (CW) for five species of Malagasy freshwater crabs that either are currently or were formerly assigned to Foza. The sum of all four ambulatory legs divided by the carapace width is also shown (ΣP2–5/CW), along with the size classes for the ambulatory legs." httpUri="http://table.plazi.org/id/49FF3A87FFFBFFB03BD5FF34FE20880B" pageId="11" pageNumber="252" tableUuid="49FF3A87FFFBFFB03BD5FF34FE20880B">Table 3</tableCitation>
|
||
). Male pleon plus telson triangular, tapered distally, widest at PL3, narrowest at telson; PL6 relatively broad, with convex lateral margins; telson outline with straight lateral margins, apex rounded (
|
||
<figureCitation id="85BB768AFFE2FFA93A3FFABDFE598C31" box="[330,416,1287,1312]" captionStart="Figure 1" captionStartId="1.[160,225,1227,1250]" captionTargetBox="[163,1153,142,1196]" captionTargetId="figure-116@1.[163,1153,142,1196]" captionTargetPageId="1" captionText="Figure 1. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar, views of the cephalothorax: a, dorsal, c, frontal, e, ventral. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar, views of the cephalothorax: b, dorsal; d, frontal, and f, ventral. Scale bar: a, c, e = 8.4 mm; b, d, f = 8.8 mm." figureDoi="http://doi.org/10.5281/zenodo.6758293" httpUri="https://zenodo.org/record/6758293/files/figure.png" pageId="11" pageNumber="252">Figure 1</figureCitation>
|
||
(e)). Sulcus between G1 TA, G1 SA faint on ventral side, distinct on dorsal side; G1 TA medium length (G1 TA/G1 SA = 0.3), dorsal side glabrous, slim, broadest proximally, distal half curving upward, tip broad; G1 SA with distinct raised rounded shoulder on external margin near G1 TA-SA junction (
|
||
<figureCitation id="85BB768AFFE2FFA938E5FAD5FC0B8C99" box="[912,1010,1391,1416]" captionStart="Figure 5" captionStartId="9.[160,225,1401,1424]" captionTargetBox="[344,971,142,1369]" captionTargetId="figure-3@9.[344,972,142,1370]" captionTargetPageId="9" captionText="Figure 5. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar. a, ventral view of left G1; b, dorsal view of left G1; and c, ventral view of left G2. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar: d, ventral view of left G1; e, dorsal view of left G1; and f, ventral view of left G2. Scale bar: a, b = 5.0 mm; c = 2.9 mm; d, e = 2.5 mm; f = 2.9 mm." figureDoi="http://doi.org/10.5281/zenodo.6758303" httpUri="https://zenodo.org/record/6758303/files/figure.png" pageId="11" pageNumber="252">Figure 5</figureCitation>
|
||
(a,b)). Broad, trapezoid membrane (DM) on dorsal side of G1 TA-SA junction; DM superior margin diagonal, inferior margin J-shaped, lateral margin broad, mesial margin narrow (
|
||
<figureCitation id="85BB768AFFE2FFA93F52FA0EFB7A8CDC" box="[1063,1155,1460,1485]" captionStart="Figure 5" captionStartId="9.[160,225,1401,1424]" captionTargetBox="[344,971,142,1369]" captionTargetId="figure-3@9.[344,972,142,1370]" captionTargetPageId="9" captionText="Figure 5. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar. a, ventral view of left G1; b, dorsal view of left G1; and c, ventral view of left G2. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar: d, ventral view of left G1; e, dorsal view of left G1; and f, ventral view of left G2. Scale bar: a, b = 5.0 mm; c = 2.9 mm; d, e = 2.5 mm; f = 2.9 mm." figureDoi="http://doi.org/10.5281/zenodo.6758303" httpUri="https://zenodo.org/record/6758303/files/figure.png" pageId="11" pageNumber="252">Figure 5</figureCitation>
|
||
(b)). G2 TA long (G2 TA/SA = 0.6), flagellum-like with distinct distal curve (
|
||
<figureCitation id="85BB768AFFE2FFA9389DFA6DFBBD8CE1" box="[1000,1092,1495,1520]" captionStart="Figure 5" captionStartId="9.[160,225,1401,1424]" captionTargetBox="[344,971,142,1369]" captionTargetId="figure-3@9.[344,972,142,1370]" captionTargetPageId="9" captionText="Figure 5. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar. a, ventral view of left G1; b, dorsal view of left G1; and c, ventral view of left G2. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar: d, ventral view of left G1; e, dorsal view of left G1; and f, ventral view of left G2. Scale bar: a, b = 5.0 mm; c = 2.9 mm; d, e = 2.5 mm; f = 2.9 mm." figureDoi="http://doi.org/10.5281/zenodo.6758303" httpUri="https://zenodo.org/record/6758303/files/figure.png" pageId="11" pageNumber="252">Figure 5</figureCitation>
|
||
(c)).
|
||
</paragraph>
|
||
<paragraph id="1D3F6A0FFFE2FFA93BD5F996FF378F57" blockId="11.[160,1156,1580,1675]" box="[160,206,1580,1606]" pageId="11" pageNumber="252">
|
||
<emphasis id="2FF4B61DFFE2FFA93BD5F996FF378F57" bold="true" box="[160,206,1580,1606]" italics="true" pageId="11" pageNumber="252">Size</emphasis>
|
||
</paragraph>
|
||
<paragraph id="1D3F6A0FFFE2FFA93BD5F9F5FEB28F9A" blockId="11.[160,1156,1580,1675]" pageId="11" pageNumber="252">
|
||
Medium-sized species, largest known specimen CW
|
||
<quantity id="DA78C7EAFFE2FFA93996F9F5FCCA8F79" box="[739,819,1615,1640]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="252" unit="mm" value="50.0">50 mm</quantity>
|
||
, pubertal moult between CWs 37 and
|
||
<quantity id="DA78C7EAFFE2FFA93B80F9C8FEBE8F9A" box="[245,327,1650,1675]" metricMagnitude="-2" metricUnit="m" metricValue="4.3" pageId="11" pageNumber="252" unit="mm" value="43.0">43 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="1D3F6A0FFFE5FFAE3BD5FF34FF1789B9" blockId="12.[160,887,142,203]" box="[160,238,142,168]" pageId="12" pageNumber="253">
|
||
<emphasis id="2FF4B61DFFE5FFAE3BD5FF34FF1789B9" bold="true" box="[160,238,142,168]" italics="true" pageId="12" pageNumber="253">Colour</emphasis>
|
||
</paragraph>
|
||
<paragraph id="1D3F6A0FFFE5FFAE3BD5FF08FC8E89DA" blockId="12.[160,887,142,203]" box="[160,887,178,203]" pageId="12" pageNumber="253">Unknown in life. Preserved specimens are uniformly pale brown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="559A3984FFE5FFAE3BD5FEBCFBF388E1" pageId="12" pageNumber="253" type="distribution">
|
||
<paragraph id="1D3F6A0FFFE5FFAE3BD5FEBCFED58831" blockId="12.[160,1156,262,496]" box="[160,300,262,288]" pageId="12" pageNumber="253">
|
||
<emphasis id="2FF4B61DFFE5FFAE3BD5FEBCFED58831" bold="true" box="[160,300,262,288]" italics="true" pageId="12" pageNumber="253">Distribution</emphasis>
|
||
</paragraph>
|
||
<paragraph id="1D3F6A0FFFE5FFAE3BD5FE90FBF388E1" blockId="12.[160,1156,262,496]" pageId="12" pageNumber="253">
|
||
<taxonomicName id="DA80118CFFE5FFAE3BD5FE90FCD18852" authority="(Cumberlidge and Meyer, 2009) comb." authorityName="comb." authorityYear="2022" baseAuthorityName="Cumberlidge and Meyer" baseAuthorityYear="2009" box="[160,808,298,323]" class="Malacostraca" family="Potamonautidae" genus="Vahatra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="241" phylum="Arthropoda" rank="species" species="ambohitra" status="comb.">
|
||
<emphasis id="2FF4B61DFFE5FFAE3BD5FE90FE898852" box="[160,368,298,323]" italics="true" pageId="12" pageNumber="253">Vahatra ambohitra</emphasis>
|
||
(
|
||
<bibRefCitation id="791117FEFFE5FFAE3A0BFE90FD2A8852" author="Cumberlidge N & Meyer KS" box="[382,723,298,323]" pageId="12" pageNumber="253" pagination="77 - 89" refId="ref11416" refString="Cumberlidge N, Meyer KS. 2009. A new species of Foza Reed & Cumberlidge, 2006 from northern Madagascar (Decapoda, Brachyura, Potamoidea, Potamonautidae), with a redescription of F. goudoti (H. Milne Edwards, 1853) comb. n., and comments on Skelosophusa prolixa Ng & Takeda, 1994. ZooKeys. 18: 77 - 89. doi: 10.3897 / zookeys. 18.102." type="journal article" year="2009">Cumberlidge and Meyer, 2009</bibRefCitation>
|
||
)
|
||
<taxonomicNameLabel id="34C70B66FFE5FFAE3994FE90FCD18852" box="[737,808,298,323]" pageId="12" pageNumber="253" rank="species">comb.</taxonomicNameLabel>
|
||
</taxonomicName>
|
||
nov. has a distributional range that includes Ambohitra, the Ankarana Special Reserve and the Analamerana Special Reserve, all in the Diana Region of
|
||
<collectingRegion id="DF44A4EDFFE5FFAE3929FED5FCAE8899" box="[604,855,367,392]" country="Madagascar" name="Antsiranana" pageId="12" pageNumber="253">Antsiranana Province</collectingRegion>
|
||
in northern
|
||
<collectingCountry id="65972A9FFFE5FFAE388FFED5FB7D8899" box="[1018,1156,367,392]" name="Madagascar" pageId="12" pageNumber="253">Madagascar</collectingCountry>
|
||
(
|
||
<figureCitation id="85BB768AFFE5FFAE3BDDFE28FEFF88BA" box="[168,262,402,427]" captionStart="Figure 6" captionStartId="16.[160,225,1033,1056]" captionTargetBox="[165,1150,144,998]" captionTargetId="figure-69@16.[163,1153,142,1001]" captionTargetPageId="16" captionText="Figure 6. Map of Madagascar showing the distributional range of Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov. (black circles, not shown to scale), Toamasina clarki sp. nov. (white triangle in black square, not shown to scale), Foza raimundi Reed and Cumberlidge, 2006 (white asterisk with black margin), and Foza manonae Cumberlidge, Klaus, Meyer, and Koppin, 2015 (white square with black margin). Scale bar = 184 km." figureDoi="http://doi.org/10.5281/zenodo.6758305" httpUri="https://zenodo.org/record/6758305/files/figure.png" pageId="12" pageNumber="253">Figure 6</figureCitation>
|
||
). This distributional range no longer includes Akirindro Mountain in northern
|
||
<collectingRegion id="DF44A4EDFFE5FFAE3BD5FE0EFE7B88DC" box="[160,386,436,461]" country="Madagascar" name="Toamasina" pageId="12" pageNumber="253">
|
||
<taxonomicName id="DA80118CFFE5FFAE3BD5FE0EFEE188DC" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" box="[160,280,436,461]" class="Malacostraca" family="Potamonautidae" genus="Toamasina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="259" phylum="Arthropoda" rank="genus">Toamasina</taxonomicName>
|
||
Province
|
||
</collectingRegion>
|
||
because the specimen from that locality (NMU PN 17–
|
||
<date id="693E4CCFFFE5FFAE3886FE0EFB9D88DC" box="[1011,1124,436,461]" pageId="12" pageNumber="253" value="2003-03-21">21.3.2003</date>
|
||
) is not now recognised as belonging to this species (see remarks section below).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="559A3984FFE5FFAE3BD5FD91FEFA8AA1" pageId="12" pageNumber="253" type="materials_examined">
|
||
<paragraph id="1D3F6A0FFFE5FFAE3BD5FD91FECF8B54" blockId="12.[160,1155,555,651]" box="[160,310,555,581]" pageId="12" pageNumber="253">
|
||
<emphasis id="2FF4B61DFFE5FFAE3BD5FD91FECF8B54" bold="true" box="[160,310,555,581]" italics="true" pageId="12" pageNumber="253">Type locality</emphasis>
|
||
</paragraph>
|
||
<paragraph id="1D3F6A0FFFE5FFAE3BD5FDF5FD2D8B9A" blockId="12.[160,1155,555,651]" pageId="12" pageNumber="253">
|
||
<materialsCitation id="ADE86052FFE5FFAE3BD5FDF5FD298B9A" ID-GBIF-Occurrence="3861175315" collectionCode="ASL" country="Madagascar" elevation="421" latitude="-12.45" location="Ambohitra" longLatPrecision="1" longitude="49.13333" municipality="Diana Region" pageId="12" pageNumber="253" specimenCount="1" stateProvince="Antsiranana">
|
||
<collectingCountry id="65972A9FFFE5FFAE3BD5FDF5FED08B79" box="[160,297,591,616]" name="Madagascar" pageId="12" pageNumber="253">Madagascar</collectingCountry>
|
||
;
|
||
<collectingRegion id="DF44A4EDFFE5FFAE3A42FDF5FDD48B79" box="[311,557,591,616]" country="Madagascar" name="Antsiranana" pageId="12" pageNumber="253">Antsiranana Province</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="FD5BF075FFE5FFAE394EFDF5FD218B79" box="[571,728,591,616]" pageId="12" pageNumber="253">Diana Region</collectingMunicipality>
|
||
, forest close to
|
||
<location id="185F3CD4FFE5FFAE38ECFDF5FBE88B79" LSID="urn:lsid:plazi:treatment:9529DB19FFE3FFAF3A6CFF34FBD18844:185F3CD4FFE5FFAE38ECFDF5FBE88B79" box="[921,1041,591,616]" country="Madagascar" latitude="-12.45" longLatPrecision="1" longitude="49.13333" municipality="Diana Region" name="Ambohitra" pageId="12" pageNumber="253" stateProvince="Antsiranana">Ambohitra</location>
|
||
(formerly Joffreville) (
|
||
<geoCoordinate id="78B40CC8FFE5FFAE3A55FDC8FE548B9A" box="[288,429,626,651]" degrees="12.450000" direction="south" orientation="latitude" pageId="12" pageNumber="253" precision="1" value="-12.45">12.450000°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="78B40CC8FFE5FFAE3AC2FDC8FDBC8B9A" box="[439,581,626,651]" degrees="49.133333" direction="east" orientation="longitude" pageId="12" pageNumber="253" precision="1" value="49.13333">49.133333°E</geoCoordinate>
|
||
),
|
||
<quantity id="DA78C7EAFFE5FFAE3921FDC8FD628B9A" box="[596,667,626,651]" metricMagnitude="2" metricUnit="m" metricValue="4.21" pageId="12" pageNumber="253" unit="m" value="421.0">
|
||
<elevation id="96AD8D3CFFE5FFAE3921FDC8FD628B9A" box="[596,667,626,651]" metricMagnitude="2" metricUnit="m" metricValue="4.21" pageId="12" pageNumber="253" unit="m" value="421.0">421 m</elevation>
|
||
</quantity>
|
||
<collectionCode id="7B91F2CAFFE5FFAE39D6FDC8FD298B9A" box="[675,720,626,651]" pageId="12" pageNumber="253">ASL</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="1D3F6A0FFFE5FFAE3BD5FD7CFF008BF1" blockId="12.[160,1156,710,944]" box="[160,249,710,736]" pageId="12" pageNumber="253">
|
||
<emphasis id="2FF4B61DFFE5FFAE3BD5FD7CFF008BF1" bold="true" box="[160,249,710,736]" italics="true" pageId="12" pageNumber="253">Habitat</emphasis>
|
||
</paragraph>
|
||
<paragraph id="1D3F6A0FFFE5FFAE3BD5FD50FEFA8AA1" blockId="12.[160,1156,710,944]" pageId="12" pageNumber="253">
|
||
<taxonomicName id="DA80118CFFE5FFAE3BD5FD50FCDC8A12" authority="(Cumberlidge and Meyer, 2009) comb." authorityName="comb." authorityYear="2022" baseAuthorityName="Cumberlidge and Meyer" baseAuthorityYear="2009" box="[160,805,746,771]" class="Malacostraca" family="Potamonautidae" genus="Vahatra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="241" phylum="Arthropoda" rank="species" species="ambohitra" status="comb.">
|
||
<emphasis id="2FF4B61DFFE5FFAE3BD5FD50FE898A12" box="[160,368,746,771]" italics="true" pageId="12" pageNumber="253">Vahatra ambohitra</emphasis>
|
||
(
|
||
<bibRefCitation id="791117FEFFE5FFAE3A08FD50FD288A12" author="Cumberlidge N & Meyer KS" box="[381,721,746,771]" pageId="12" pageNumber="253" pagination="77 - 89" refId="ref11416" refString="Cumberlidge N, Meyer KS. 2009. A new species of Foza Reed & Cumberlidge, 2006 from northern Madagascar (Decapoda, Brachyura, Potamoidea, Potamonautidae), with a redescription of F. goudoti (H. Milne Edwards, 1853) comb. n., and comments on Skelosophusa prolixa Ng & Takeda, 1994. ZooKeys. 18: 77 - 89. doi: 10.3897 / zookeys. 18.102." type="journal article" year="2009">Cumberlidge and Meyer, 2009</bibRefCitation>
|
||
)
|
||
<taxonomicNameLabel id="34C70B66FFE5FFAE39AAFD50FCDC8A12" box="[735,805,746,771]" pageId="12" pageNumber="253" rank="species">comb.</taxonomicNameLabel>
|
||
</taxonomicName>
|
||
nov. dwells on land, near small streams in partially disturbed mixed dry deciduous and humid forests of northern
|
||
<collectingCountry id="65972A9FFFE5FFAE3BD5FC95FED38A59" box="[160,298,815,840]" name="Madagascar" pageId="12" pageNumber="253">Madagascar</collectingCountry>
|
||
near the village of Ambohitra. The Ankarana Special Reserve and the Analamerana Special Reserve are both part of the Ankarana Highlands with dry deciduous and riverine forest that includes limestone formations (tsingy) with numerous caves and canyons.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="559A3984FFE5FFAF3BD5FC56FBD18844" lastPageId="13" lastPageNumber="254" pageId="12" pageNumber="253" type="discussion">
|
||
<paragraph id="1D3F6A0FFFE5FFAE3BD5FC56FEFF8D17" blockId="12.[160,1156,1004,1619]" box="[160,262,1004,1030]" pageId="12" pageNumber="253">
|
||
<emphasis id="2FF4B61DFFE5FFAE3BD5FC56FEFF8D17" bold="true" box="[160,262,1004,1030]" italics="true" pageId="12" pageNumber="253">Remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph id="1D3F6A0FFFE5FFAE3BD5FBB5FBF48DE9" blockId="12.[160,1156,1004,1619]" pageId="12" pageNumber="253">
|
||
The species was formerly assigned to
|
||
<taxonomicName id="DA80118CFFE5FFAE3915FBB5FD6A8D39" authorityName="Reed and Cumberlidge" authorityYear="2006" box="[608,659,1039,1064]" class="Malacostraca" family="Potamonautidae" genus="Foza" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="253" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2FF4B61DFFE5FFAE3915FBB5FD6A8D39" box="[608,659,1039,1064]" italics="true" pageId="12" pageNumber="253">Foza</emphasis>
|
||
</taxonomicName>
|
||
based on morphological characters but is removed from that genus based on molecular evidence and further morphological analysis.
|
||
<materialsCitation id="ADE86052FFE5FFAE3A7CFBEEFD108DC4" ID-GBIF-Occurrence="3861175326" accessionNumber="MT749727, MT749728" collectionCode="FMNH" collectorName="The & GenBank" location="Analamerana Special Reserve" pageId="12" pageNumber="253" specimenCode="FMNH 11045, 11046, 11050, 11051, 11052, 11054, 11056, FMNH 11046" specimenCount="2" typeStatus="paratype">
|
||
<collectorName id="B0750FD9FFE5FFAE3A7CFBEEFECA8D7C" box="[265,307,1108,1133]" pageId="12" pageNumber="253">The</collectorName>
|
||
<typeStatus id="C23BD4ADFFE5FFAE3A4BFBEEFE548D7C" box="[318,429,1108,1133]" pageId="12" pageNumber="253" type="paratype">paratypes</typeStatus>
|
||
from the
|
||
<location id="185F3CD4FFE5FFAE395EFBEEFC878D7C" LSID="urn:lsid:plazi:treatment:9529DB19FFE3FFAF3A6CFF34FBD18844:185F3CD4FFE5FFAE395EFBEEFC878D7C" box="[555,894,1108,1133]" name="Analamerana Special Reserve" pageId="12" pageNumber="253">Analamerana Special Reserve</location>
|
||
(
|
||
<specimenCode id="4D26C274FFE5FFAE38FAFBEEFBD28D7C" box="[911,1067,1108,1133]" collectionCode="FMNH" country="USA" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="12" pageNumber="253" type="Museum">FMNH 11045</specimenCode>
|
||
,
|
||
<specimenCode id="4D26C274FFE5FFAE3F4CFBEEFB788D7C" box="[1081,1153,1108,1133]" collectionCode="FMNH" country="USA" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="12" pageNumber="253" type="Museum">11046</specimenCode>
|
||
,
|
||
<specimenCode id="4D26C274FFE5FFAE3BD5FBCDFF1E8D81" box="[160,231,1143,1168]" collectionCode="FMNH" country="USA" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="12" pageNumber="253" type="Museum">11050</specimenCode>
|
||
,
|
||
<specimenCode id="4D26C274FFE5FFAE3B87FBCDFEC08D81" box="[242,313,1143,1168]" collectionCode="FMNH" country="USA" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="12" pageNumber="253" type="Museum">11051</specimenCode>
|
||
,
|
||
<specimenCode id="4D26C274FFE5FFAE3A31FBCDFE728D81" box="[324,395,1143,1168]" collectionCode="FMNH" country="USA" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="12" pageNumber="253" type="Museum">11052</specimenCode>
|
||
,
|
||
<specimenCode id="4D26C274FFE5FFAE3AE3FBCDFE248D81" box="[406,477,1143,1168]" collectionCode="FMNH" country="USA" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="12" pageNumber="253" type="Museum">11054</specimenCode>
|
||
,
|
||
<specimenCode id="4D26C274FFE5FFAE3A9CFBCDFDD48D81" box="[489,557,1143,1168]" collectionCode="FMNH" country="USA" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="12" pageNumber="253" type="Museum">11056</specimenCode>
|
||
and 11060) are retained in this species because one of these (
|
||
<specimenCode id="4D26C274FFE5FFAE3A71FB20FE6E8DA2" box="[260,407,1178,1203]" collectionCode="FMNH" country="USA" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="12" pageNumber="253" type="Museum">FMNH 11046</specimenCode>
|
||
) was sequenced (
|
||
<collectorName id="B0750FD9FFE5FFAE392CFB20FD468DA2" box="[601,703,1178,1203]" pageId="12" pageNumber="253">GenBank</collectorName>
|
||
<accessionNumber id="02D3F7ECFFE5FFAE39B1FB20FCC28DA2" box="[708,827,1178,1203]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MT749727" pageId="12" pageNumber="253">MT749727</accessionNumber>
|
||
) and is positioned next to the sequence (
|
||
<collectorName id="B0750FD9FFE5FFAE3A63FB06FE858DC4" box="[278,380,1212,1237]" pageId="12" pageNumber="253">GenBank</collectorName>
|
||
<accessionNumber id="02D3F7ECFFE5FFAE3AF7FB06FE008DC4" box="[386,505,1212,1237]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MT749728" pageId="12" pageNumber="253">MT749728</accessionNumber>
|
||
) for the
|
||
<typeStatus id="C23BD4ADFFE5FFAE3927FB06FD4F8DC4" box="[594,694,1212,1237]" pageId="12" pageNumber="253" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="DA80118CFFE5FFAE39ADFB06FD108DC4" authority="(Cumberlidge and Meyer, 2009)" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" baseAuthorityName="Cumberlidge and Meyer" baseAuthorityYear="2009" box="[728,745,1212,1237]" class="Malacostraca" family="Potamonautidae" genus="Vahatra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="253" phylum="Arthropoda" rank="species" species="ambohitra" status="comb. nov.">
|
||
<emphasis id="2FF4B61DFFE5FFAE39ADFB06FD108DC4" box="[728,745,1212,1237]" italics="true" pageId="12" pageNumber="253">V</emphasis>
|
||
</taxonomicName>
|
||
</materialsCitation>
|
||
<emphasis id="2FF4B61DFFE5FFAE399CFB06FC9D8DC4" box="[745,868,1212,1237]" italics="true" pageId="12" pageNumber="253">. ambohitra</emphasis>
|
||
(
|
||
<bibRefCitation id="791117FEFFE5FFAE381AFB06FF2E8DE9" author="Cumberlidge N & Meyer KS" pageId="12" pageNumber="253" pagination="77 - 89" refId="ref11416" refString="Cumberlidge N, Meyer KS. 2009. A new species of Foza Reed & Cumberlidge, 2006 from northern Madagascar (Decapoda, Brachyura, Potamoidea, Potamonautidae), with a redescription of F. goudoti (H. Milne Edwards, 1853) comb. n., and comments on Skelosophusa prolixa Ng & Takeda, 1994. ZooKeys. 18: 77 - 89. doi: 10.3897 / zookeys. 18.102." type="journal article" year="2009">Cumberlidge and Meyer, 2009</bibRefCitation>
|
||
)
|
||
<taxonomicNameLabel id="34C70B66FFE5FFAE3B93FB65FE9A8DE9" box="[230,355,1247,1272]" pageId="12" pageNumber="253" rank="species">comb. nov.</taxonomicNameLabel>
|
||
on the phylogenetic tree in
|
||
<bibRefCitation id="791117FEFFE5FFAE39D3FB65FC428DE9" author="Cumberlidge N & Soma JB & Leever EM & Daniels SR" box="[678,955,1247,1272]" pageId="12" pageNumber="253" pagination="584 - 599" refId="ref11613" refString="Cumberlidge N, Soma JB, Leever EM, Daniels SR. 2020. New lineages within the Malagasya freshwater crab fauna: Agora n. gen. for Thelphusa goudoti H. Milne Edwards, 1853, and a phytotelmic new species of Malagasya Cumberlidge & Sternberg, 2002 (Brachyura: Potamonautidae: Deckeniidae). Journal of Crustacean Biology. 40 (5): 584 - 599. doi: 10.1093 / jcbiol / ruaa 050." type="journal article" year="2020">Cumberlidge et al. (2020</bibRefCitation>
|
||
, fig. 1).
|
||
</paragraph>
|
||
<paragraph id="1D3F6A0FFFE5FFAE3BB5FAB8FE2D8F42" blockId="12.[160,1156,1004,1619]" pageId="12" pageNumber="253">
|
||
<taxonomicName id="DA80118CFFE5FFAE3BB5FAB8FE768C0A" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" baseAuthorityName="Cumberlidge and Meyer" baseAuthorityYear="2009" box="[192,399,1282,1307]" class="Malacostraca" family="Potamonautidae" genus="Vahatra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="253" phylum="Arthropoda" rank="species" species="ambohitra">
|
||
<emphasis id="2FF4B61DFFE5FFAE3BB5FAB8FE768C0A" box="[192,399,1282,1307]" italics="true" pageId="12" pageNumber="253">Vahatra ambohitra</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from
|
||
<taxonomicName id="DA80118CFFE5FFAE39B5FAB8FCC38C0A" authorityName="Reed and Cumberlidge" authorityYear="2006" box="[704,826,1282,1307]" class="Malacostraca" family="Potamonautidae" genus="Foza" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="253" phylum="Arthropoda" rank="species" species="raimundi">
|
||
<emphasis id="2FF4B61DFFE5FFAE39B5FAB8FCC38C0A" box="[704,826,1282,1307]" italics="true" pageId="12" pageNumber="253">F. raimundi</emphasis>
|
||
</taxonomicName>
|
||
mainly by the G1 TA which is glabrous on the dorsal side in
|
||
<taxonomicName id="DA80118CFFE5FFAE3A81FA9EFD788C2C" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" baseAuthorityName="Cumberlidge and Meyer" baseAuthorityYear="2009" box="[500,641,1316,1341]" class="Malacostraca" family="Potamonautidae" genus="Vahatra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="253" phylum="Arthropoda" rank="species" species="ambohitra">
|
||
<emphasis id="2FF4B61DFFE5FFAE3A81FA9EFD788C2C" box="[500,641,1316,1341]" italics="true" pageId="12" pageNumber="253">V. ambohitra</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="85BB768AFFE5FFAE39E5FA9EFD148C2C" box="[656,749,1316,1341]" captionStart="Figure 5" captionStartId="9.[160,225,1401,1424]" captionTargetBox="[344,971,142,1369]" captionTargetId="figure-3@9.[344,972,142,1370]" captionTargetPageId="9" captionText="Figure 5. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar. a, ventral view of left G1; b, dorsal view of left G1; and c, ventral view of left G2. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar: d, ventral view of left G1; e, dorsal view of left G1; and f, ventral view of left G2. Scale bar: a, b = 5.0 mm; c = 2.9 mm; d, e = 2.5 mm; f = 2.9 mm." figureDoi="http://doi.org/10.5281/zenodo.6758303" httpUri="https://zenodo.org/record/6758303/files/figure.png" pageId="12" pageNumber="253">Figure 5</figureCitation>
|
||
(b)), vs a G1 TA that is heavily setose on the dorsal side in
|
||
<taxonomicName id="DA80118CFFE5FFAE3AF2FAFDFDF88C71" authorityName="Reed and Cumberlidge" authorityYear="2006" box="[391,513,1351,1376]" class="Malacostraca" family="Potamonautidae" genus="Foza" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="253" phylum="Arthropoda" rank="species" species="raimundi">
|
||
<emphasis id="2FF4B61DFFE5FFAE3AF2FAFDFDF88C71" box="[391,513,1351,1376]" italics="true" pageId="12" pageNumber="253">F. raimundi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="85BB768AFFE5FFAE397AFAFDFD928C71" box="[527,619,1351,1376]" captionStart="Figure 7" captionStartId="19.[160,225,1121,1144]" captionTargetBox="[163,1153,142,1089]" captionTargetId="figure-84@19.[163,1153,142,1090]" captionTargetPageId="19" captionText="Figure 7. Foza raimundi Reed and Cumberlidge, 2006, adult ♂, CW 18.7 mm, FMNH 15336, from Marojejy National Park, Madagascar: a, dorsal view of cephalothorax; b, frontal view of cephalothorax; c, ventral view of cephalothorax; g, ventral view of left G1; h, dorsal view of left G1; i, ventral view of left G2. Foza manonae Cumberlidge, Klaus, Meyer, and Koppin, 2015, holotype, adult ♂, CW 43.07 mm, ZSM A20145003, from Ankarana Special Reserve, Madagascar: d, dorsal view of cephalothorax; e, frontal view of cephalothorax; f, ventral view of cephalothorax; j, ventral view of left G1; k, dorsal view of left G1; l, ventral view of left G2. Scale bar:a–c = 6.6 mm; d–f = 10.0 mm; g, h = 2.0 mm; i = 2.5 mm; j, k = 3.3 mm; l = 4.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6758309" httpUri="https://zenodo.org/record/6758309/files/figure.png" pageId="12" pageNumber="253">Figure 7</figureCitation>
|
||
(h)); by a lack of setae on the superior part of the pterygostomial region and on the anterior subpleonal cavity in
|
||
<taxonomicName id="DA80118CFFE5FFAE381FFAD0FC018C92" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" baseAuthorityName="Cumberlidge and Meyer" baseAuthorityYear="2009" box="[874,1016,1386,1411]" class="Malacostraca" family="Potamonautidae" genus="Vahatra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="253" phylum="Arthropoda" rank="species" species="ambohitra">
|
||
<emphasis id="2FF4B61DFFE5FFAE381FFAD0FC018C92" box="[874,1016,1386,1411]" italics="true" pageId="12" pageNumber="253">V. ambohitra</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="85BB768AFFE5FFAE3F72FAD0FBA78C92" box="[1031,1118,1386,1411]" captionStart="Figure 1" captionStartId="1.[160,225,1227,1250]" captionTargetBox="[163,1153,142,1196]" captionTargetId="figure-116@1.[163,1153,142,1196]" captionTargetPageId="1" captionText="Figure 1. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar, views of the cephalothorax: a, dorsal, c, frontal, e, ventral. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar, views of the cephalothorax: b, dorsal; d, frontal, and f, ventral. Scale bar: a, c, e = 8.4 mm; b, d, f = 8.8 mm." figureDoi="http://doi.org/10.5281/zenodo.6758293" httpUri="https://zenodo.org/record/6758293/files/figure.png" pageId="12" pageNumber="253">Figure 1</figureCitation>
|
||
(c);
|
||
<bibRefCitation id="791117FEFFE5FFAE3BD5FA36FDF18CB4" author="Cumberlidge N & Meyer KS" box="[160,520,1420,1445]" pageId="12" pageNumber="253" pagination="77 - 89" refId="ref11416" refString="Cumberlidge N, Meyer KS. 2009. A new species of Foza Reed & Cumberlidge, 2006 from northern Madagascar (Decapoda, Brachyura, Potamoidea, Potamonautidae), with a redescription of F. goudoti (H. Milne Edwards, 1853) comb. n., and comments on Skelosophusa prolixa Ng & Takeda, 1994. ZooKeys. 18: 77 - 89. doi: 10.3897 / zookeys. 18.102." type="journal article" year="2009">Cumberlidge and Meyer (2009</bibRefCitation>
|
||
, fig. 2B,C), vs both these regions distinctly setose in
|
||
<taxonomicName id="DA80118CFFE5FFAE3BD5FA15FEE48CD9" authorityName="Reed and Cumberlidge" authorityYear="2006" box="[160,285,1455,1480]" class="Malacostraca" family="Potamonautidae" genus="Foza" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="253" phylum="Arthropoda" rank="species" species="raimundi">
|
||
<emphasis id="2FF4B61DFFE5FFAE3BD5FA15FEE48CD9" box="[160,285,1455,1480]" italics="true" pageId="12" pageNumber="253">F. raimundi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="85BB768AFFE5FFAE3A5BFA15FE748CD9" box="[302,397,1455,1480]" captionStart="Figure 7" captionStartId="19.[160,225,1121,1144]" captionTargetBox="[163,1153,142,1089]" captionTargetId="figure-84@19.[163,1153,142,1090]" captionTargetPageId="19" captionText="Figure 7. Foza raimundi Reed and Cumberlidge, 2006, adult ♂, CW 18.7 mm, FMNH 15336, from Marojejy National Park, Madagascar: a, dorsal view of cephalothorax; b, frontal view of cephalothorax; c, ventral view of cephalothorax; g, ventral view of left G1; h, dorsal view of left G1; i, ventral view of left G2. Foza manonae Cumberlidge, Klaus, Meyer, and Koppin, 2015, holotype, adult ♂, CW 43.07 mm, ZSM A20145003, from Ankarana Special Reserve, Madagascar: d, dorsal view of cephalothorax; e, frontal view of cephalothorax; f, ventral view of cephalothorax; j, ventral view of left G1; k, dorsal view of left G1; l, ventral view of left G2. Scale bar:a–c = 6.6 mm; d–f = 10.0 mm; g, h = 2.0 mm; i = 2.5 mm; j, k = 3.3 mm; l = 4.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6758309" httpUri="https://zenodo.org/record/6758309/files/figure.png" pageId="12" pageNumber="253">Figure 7</figureCitation>
|
||
(b,c);
|
||
<bibRefCitation id="791117FEFFE5FFAE3ABFFA15FCE18CD9" author="Reed SK & Cumberlidge N" box="[458,792,1455,1480]" pageId="12" pageNumber="253" pagination="58 - 66" refId="ref12024" refString="Reed SK, Cumberlidge N. 2006. Foza raimundi, a new genus and species of potamonautid freshwater crab (Crustacea: Decapoda: Potamoidea) from northern Madagascar. Proc Biol Soc Wash. 119 (1): 58 - 66. doi: 10.2988 / 0006 - 324 X (2006) 119 [58: FRANGA] 2.0. CO; 2." type="journal article" year="2006">Reed and Cumberlidge (2006</bibRefCitation>
|
||
, fig. 1A, I); and by the movable and fixed fingers of the major chela in
|
||
<taxonomicName id="DA80118CFFE5FFAE3939FA68FD218CFA" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" baseAuthorityName="Cumberlidge and Meyer" baseAuthorityYear="2009" box="[588,728,1490,1515]" class="Malacostraca" family="Potamonautidae" genus="Vahatra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="253" phylum="Arthropoda" rank="species" species="ambohitra">
|
||
<emphasis id="2FF4B61DFFE5FFAE3939FA68FD218CFA" box="[588,728,1490,1515]" italics="true" pageId="12" pageNumber="253">V. ambohitra</emphasis>
|
||
</taxonomicName>
|
||
that have three and four large molars, respectively (
|
||
<figureCitation id="85BB768AFFE5FFAE3A46FA4EFE778F1C" box="[307,398,1524,1549]" captionStart="Figure 4" captionStartId="8.[160,225,1234,1257]" captionTargetBox="[163,1153,142,1202]" captionTargetId="figure-58@8.[163,1153,142,1202]" captionTargetPageId="8" captionText="Figure 4. Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov., holotype, adult ♂, CW 39.5 mm, MNHN B 30154, from Diana Region, Madagascar: a, frontal view of right chela; b, frontal view of left chela; e, inferior view of merus of right cheliped; f, dorsal view of carpus and merus of right cheliped. Toamasina clarki sp. nov., holotype, adult ♂, CW 43.0 mm, NMU PN 17–21.3.2003, from Toamasina Province, Madagascar: c, frontal view of right chela; d, frontal view of left chela; g, right cheliped carpus and merus dorsal view; h, inferior view of merus of right cheliped. Scale bar: a–d = 5.0 mm; e, f = 4.0 mm; g, h = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6758301" httpUri="https://zenodo.org/record/6758301/files/figure.png" pageId="12" pageNumber="253">Figure 4</figureCitation>
|
||
(a)), vs a movable finger with no large molar teeth, and a fixed finger with only one large molar on the pollex of the major chela in
|
||
<taxonomicName id="DA80118CFFE5FFAE38F1F9ADFBFA8F21" authorityName="Reed and Cumberlidge" authorityYear="2006" box="[900,1027,1559,1584]" class="Malacostraca" family="Potamonautidae" genus="Foza" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="253" phylum="Arthropoda" rank="species" species="raimundi">
|
||
<emphasis id="2FF4B61DFFE5FFAE38F1F9ADFBFA8F21" box="[900,1027,1559,1584]" italics="true" pageId="12" pageNumber="253">F. raimundi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="791117FEFFE5FFAE3F61F9ADFE8F8F42" author="Reed SK & Cumberlidge N" pageId="12" pageNumber="253" pagination="58 - 66" refId="ref12024" refString="Reed SK, Cumberlidge N. 2006. Foza raimundi, a new genus and species of potamonautid freshwater crab (Crustacea: Decapoda: Potamoidea) from northern Madagascar. Proc Biol Soc Wash. 119 (1): 58 - 66. doi: 10.2988 / 0006 - 324 X (2006) 119 [58: FRANGA] 2.0. CO; 2." type="journal article" year="2006">Reed and Cumberlidge, 2006</bibRefCitation>
|
||
, fig. 1e).
|
||
</paragraph>
|
||
<paragraph id="1D3F6A0FFFE4FFAF3BD5FF34FE7289B9" blockId="13.[160,1155,142,341]" box="[160,395,142,168]" pageId="13" pageNumber="254">
|
||
<emphasis id="2FF4B61DFFE4FFAF3BD5FF34FE7289B9" bold="true" box="[160,395,142,168]" italics="true" pageId="13" pageNumber="254">Conservation status</emphasis>
|
||
</paragraph>
|
||
<paragraph id="1D3F6A0FFFE4FFAF3BD5FF08FBD18844" blockId="13.[160,1155,142,341]" pageId="13" pageNumber="254">
|
||
The conservation status of
|
||
<taxonomicName id="DA80118CFFE4FFAF3AAFFF08FC1989DA" authority="(Cumberlidge and Meyer, 2009)" authorityName="Leever & Daniels & Soma & Cumberlidge" authorityYear="2022" baseAuthorityName="Cumberlidge and Meyer" baseAuthorityYear="2009" box="[474,992,178,203]" class="Malacostraca" family="Potamonautidae" genus="Vahatra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="254" phylum="Arthropoda" rank="species" species="ambohitra" status="comb. nov.">
|
||
<emphasis id="2FF4B61DFFE4FFAF3AAFFF08FD9389DA" box="[474,618,178,203]" italics="true" pageId="13" pageNumber="254">V. ambohitra</emphasis>
|
||
(
|
||
<bibRefCitation id="791117FEFFE4FFAF390FFF08FC2189DA" author="Cumberlidge N & Meyer KS" box="[634,984,178,203]" pageId="13" pageNumber="254" pagination="77 - 89" refId="ref11416" refString="Cumberlidge N, Meyer KS. 2009. A new species of Foza Reed & Cumberlidge, 2006 from northern Madagascar (Decapoda, Brachyura, Potamoidea, Potamonautidae), with a redescription of F. goudoti (H. Milne Edwards, 1853) comb. n., and comments on Skelosophusa prolixa Ng & Takeda, 1994. ZooKeys. 18: 77 - 89. doi: 10.3897 / zookeys. 18.102." type="journal article" year="2009">Cumberlidge and Meyer, 2009</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="34C70B66FFE4FFAF389CFF08FB9089DA" box="[1001,1129,178,203]" pageId="13" pageNumber="254" rank="species">comb. nov.</taxonomicNameLabel>
|
||
is assessed as Least Concern (LC) on the International Union for Conservation of Nature Red List (
|
||
<bibRefCitation id="791117FEFFE4FFAF3BA0FF4DFE518801" author="Cumberlidge N" box="[213,424,247,272]" pageId="13" pageNumber="254" refId="ref11129" refString="Cumberlidge N 2016. Foza ambohitra. The IUCN Red List of Threatened Species 2016: e. T 70870429 A 70915711. 10.2305 / IUCN. UK. 2016 - 3. RLTS. T 70870429 A 70915711. en. Downloaded on Downloaded on Jun 28." type="book" year="2016">Cumberlidge 2016</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="791117FEFFE4FFAF3AC2FF4DFD308801" author="Cumberlidge N & Rasamy Razanabolana J & Ranaivoson CH & Randrianasolo HH & Sayer C & Maiz-Tome L & Van Damme D & Darwall WRT" box="[439,713,247,272]" pageId="13" pageNumber="254" pagination="32 - 41" refId="ref11494" refString="Cumberlidge N, Rasamy Razanabolana J, Ranaivoson CH, Randrianasolo HH, Sayer C, Maiz-Tome L, Van Damme D, Darwall WRT. 2017. Updated extinction risk assessments of Madagascar's freshwater decapod crustaceans reveal fewer threatened species but more Data Deficient species. Malagasy Nature. 12: 32 - 41." type="journal article" year="2017">Cumberlidge et al. 2017</bibRefCitation>
|
||
), but this now needs to be reassessed because the present taxonomic revision significantly reduces the extent of occurrence of this species from
|
||
<quantity id="DA78C7EAFFE4FFAF3A10FE86FE3E8844" box="[357,455,316,341]" metricMagnitude="6" metricUnit="m" metricValue="5.874" pageId="13" pageNumber="254" unit="km" value="5874.0">5874 km</quantity>
|
||
2 (five locations) to
|
||
<quantity id="DA78C7EAFFE4FFAF39D7FE86FD0C8844" box="[674,757,316,341]" metricMagnitude="5" metricUnit="m" metricValue="9.35" pageId="13" pageNumber="254" unit="km" value="935.0">935 km</quantity>
|
||
2 (four locations) (
|
||
<figureCitation id="85BB768AFFE4FFAF38CAFE86FBE58844" box="[959,1052,316,341]" captionStart="Figure 6" captionStartId="16.[160,225,1033,1056]" captionTargetBox="[165,1150,144,998]" captionTargetId="figure-69@16.[163,1153,142,1001]" captionTargetPageId="16" captionText="Figure 6. Map of Madagascar showing the distributional range of Vahatra ambohitra (Cumberlidge and Meyer, 2009) comb. nov. (black circles, not shown to scale), Toamasina clarki sp. nov. (white triangle in black square, not shown to scale), Foza raimundi Reed and Cumberlidge, 2006 (white asterisk with black margin), and Foza manonae Cumberlidge, Klaus, Meyer, and Koppin, 2015 (white square with black margin). Scale bar = 184 km." figureDoi="http://doi.org/10.5281/zenodo.6758305" httpUri="https://zenodo.org/record/6758305/files/figure.png" pageId="13" pageNumber="254">Figure 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |