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<document id="DEBAF61073D45A33EF689A559D895CE2" ID-CLB-Dataset="22221" ID-DOI="http://dx.doi.org/10.5852/ejt.2013.45" ID-GBIF-Dataset="a66592b8-63a8-4c2d-9471-e58ddb2c0559" ID-ISSN="2118-9773" ID-Zenodo-Dep="3822710" ID-ZooBank="41B10E4D-7DAB-40CA-A8FE-4ECA078E04A3" checkinTime="1589313835598" checkinUser="carolina" docAuthor="Zahniser, James N. &amp; Dietrich, Chris H." docDate="2013" docId="6903BC00A312FFB8AC6DE05C2AADFDEC" docLanguage="en" docName="ejt-2013-45.pdf.imf" docOrigin="European Journal of Taxonomy 45" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.12:EJT.2018-.journal_article" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article" docStyleVersion="12" docTitle="Arthropoda Dmitriev 2002" docType="treatment" docVersion="8" lastPageNumber="95" masterDocId="953AC478A34EFFE6AE00E1572B5EFFD9" masterDocTitle="A review of the tribes of Deltocephalinae (Hemiptera: Auchenorrhyncha: Cicadellidae)" masterLastPageNumber="211" masterPageNumber="1" pageNumber="93" updateTime="1685912909012" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="A75B8519CD5D5C2B9CE1A58596560882">A review of the tribes of Deltocephalinae (Hemiptera: Auchenorrhyncha: Cicadellidae)</mods:title>
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<mods:namePart id="B54E8DE6B60C14F8618196E8BEA66ED6">Zahniser, James N.</mods:namePart>
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<mods:affiliation id="86621F81579F02AEFC371E475C9DC821">Illinois Natural History Survey, Prairie Research Institute, University of Illinois at Urbana-Champaign, 1816 S. Oak St., Champaign, IL 61820, United States of America &amp; urn: lsid: zoobank. org: author: C 44 D 6 E 44 - FA 1 C- 4 B 29 - B 7 BB-FAF 5940 CD 225 &amp; corresponding author e-mail: zahniser @ illinois. edu</mods:affiliation>
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<mods:affiliation id="DE6B13E77075652C69C3DCD4777194A2">Illinois Natural History Survey, Prairie Research Institute, University of Illinois at Urbana-Champaign, 1816 S. Oak St., Champaign, IL 61820, United States of America &amp; urn: lsid: zoobank. org: author: 82 FCB 86 C- 54 B 4 - 456 A-AE 5 E-D 7847 D 271 CB 9</mods:affiliation>
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<subSubSection id="A9B05E9DA312FFBAAC6DE05C2899FEFC" box="[621,967,267,293]" pageId="92" pageNumber="93" type="nomenclature">
<paragraph id="E1150D16A312FFBAAC6DE05C2899FEFC" blockId="92.[621,967,267,329]" box="[621,967,267,293]" pageId="92" pageNumber="93">
<heading id="BA5DBA7AA312FFBAAC6DE05C2899FEFC" box="[621,967,267,293]" pageId="92" pageNumber="93">
<taxonomicName id="26AA7695A312FFBAAC6DE05C2899FEFC" ID-CoL="8JKD5" authority="Dmitriev, 2002" authorityName="Dmitriev" authorityYear="2002" box="[621,967,267,293]" kingdom="Animalia" pageId="92" pageNumber="93" phylum="Arthropoda" rank="subTribe">
Glossocratina
<bibRefCitation id="853B70E7A312FFBAAD15E05C2899FEFC" author="Dmitriev D. A." box="[789,967,267,293]" pageId="92" pageNumber="93" pagination="975 - 1002" refId="ref85974" refString="Dmitriev D. A. 2002. Larvae of the leafhopper subfamily Deltocephalinae (Homoptera, Cicadellidae) from European part of Russia and adjacent territories. I. A key to tribes and review of the tribes Drabescini, Scaphytopiini, Hecalini, Limotettigini, and Opsiini. Entomologicheskoe Obozrenie 81 (3): 627 - 661. [In Russian, translated into English in Entomological Review 82 (8): 975 - 1002]" type="journal article" year="2002">Dmitriev, 2002</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="A9B05E9DA312FFBAACF0E079281DFE90" box="[752,835,302,329]" pageId="92" pageNumber="93" type="description">
<paragraph id="E1150D16A312FFBAACF0E079281DFE90" blockId="92.[621,967,267,329]" box="[752,835,302,329]" pageId="92" pageNumber="93">
<figureCitation id="79911193A312FFBAACF0E079281DFE90" box="[752,835,302,329]" captionStart="Fig" captionStartId="93.[189,232,1930,1956]" captionTargetBox="[233,1358,267,1885]" captionTargetId="figure@93.[231,1362,265,1892]" captionTargetPageId="93" captionText="Fig. 27. Hecalini (Glossocratina). Glossocratus afzelii (Stål, 1854). AI. Standard views (see Material &amp; Methods). G. Connective and style shown with aedeagus. J. Dorsal view of ♂ abdominal sternites I-III (partial view of sternite III). s1ap = apodeme of sternite I, s2ap = apodeme of sternite II." figureDoi="http://doi.org/10.5281/zenodo.3822772" httpUri="https://zenodo.org/record/3822772/files/figure.png" pageId="92" pageNumber="93">Fig. 27</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="A9B05E9DA312FFBAAEBDE02729D1FE53" box="[189,655,368,394]" pageId="92" pageNumber="93" type="type_taxon">
<paragraph id="E1150D16A312FFBAAEBDE02729D1FE53" blockId="92.[189,655,368,394]" box="[189,655,368,394]" pageId="92" pageNumber="93">
<typeStatus id="3E11B3B4A312FFBAAEBDE0272BA9FE53" box="[189,247,368,394]" pageId="92" pageNumber="93">Type</typeStatus>
genus:
<taxonomicName id="26AA7695A312FFBAAF52E02729D5FE53" authority="Fieber, 1866" authorityName="Fieber" authorityYear="1866" box="[338,651,368,394]" class="Insecta" family="Cicadellidae" genus="Glossocratus" kingdom="Animalia" order="Hemiptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="D3DED104A312FFBAAF52E0272AB3FE53" box="[338,493,368,394]" italics="true" pageId="92" pageNumber="93">Glossocratus</emphasis>
Fieber, 1866
</taxonomicName>
.
</paragraph>
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<subSubSection id="A9B05E9DA312FFBAAEBDE0E62A57FD14" pageId="92" pageNumber="93" type="diagnosis">
<paragraph id="E1150D16A312FFBAAEBDE0E62A68FE12" blockId="92.[189,310,433,459]" box="[189,310,433,459]" pageId="92" pageNumber="93">
<heading id="BA5DBA7AA312FFBAAEBDE0E62A68FE12" bold="true" box="[189,310,433,459]" fontSize="11" level="3" pageId="92" pageNumber="93" reason="3">
<emphasis id="D3DED104A312FFBAAEBDE0E62A68FE12" bold="true" box="[189,310,433,459]" pageId="92" pageNumber="93">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="E1150D16A312FFBAAEBDE0B72A57FD14" blockId="92.[189,1399,480,717]" pageId="92" pageNumber="93">
Glossocratina are medium sized to large brownish leafhoppers. They are distinguished from
<taxonomicName id="26AA7695A312FFBAAAEBE0B72E0DFE23" box="[1259,1363,480,506]" family="Null" higherTaxonomySource="Manual Input" pageId="92" pageNumber="97" rank="subTribe" subTribe="Hecalina">Hecalina</taxonomicName>
by the laterofrontal sutures keeled above the antennal pits, gena sometimes without a fine erect seta beside the laterofrontal suture, apodemes of male sternite II not as developed as in
<taxonomicName id="26AA7695A312FFBAAA57E3712F9CFD99" box="[1111,1218,550,576]" family="Null" higherTaxonomySource="Manual Input" pageId="92" pageNumber="97" rank="subTribe" subTribe="Hecalina">Hecalina</taxonomicName>
, ovipositor not protruding far beyond the pygofer apex, and second valvula with teeth. Nymphs can be distinguished from
<taxonomicName id="26AA7695A312FFBAAEFDE33A2A3BFD5E" box="[253,357,621,647]" family="Null" higherTaxonomySource="Manual Input" pageId="92" pageNumber="97" rank="subTribe" subTribe="Hecalina">Hecalina</taxonomicName>
by the oblique lateral carina extending from antennal pit to margin of head, outer margin of the gena angulate, anal tube with a long apical process, and body covered with short setae (
<bibRefCitation id="853B70E7A312FFBAAB05E3C72BA3FD14" author="Dmitriev D. A." pageId="92" pageNumber="93" pagination="975 - 1002" refId="ref85974" refString="Dmitriev D. A. 2002. Larvae of the leafhopper subfamily Deltocephalinae (Homoptera, Cicadellidae) from European part of Russia and adjacent territories. I. A key to tribes and review of the tribes Drabescini, Scaphytopiini, Hecalini, Limotettigini, and Opsiini. Entomologicheskoe Obozrenie 81 (3): 627 - 661. [In Russian, translated into English in Entomological Review 82 (8): 975 - 1002]" type="journal article" year="2002">Dmitriev, 2002</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A9B05E9DA312FFBAAEBDE3A32F5AF852" pageId="92" pageNumber="93" type="description">
<paragraph id="E1150D16A312FFBAAEBDE3A32A10FCD7" blockId="92.[189,334,756,782]" box="[189,334,756,782]" pageId="92" pageNumber="93">
<heading id="BA5DBA7AA312FFBAAEBDE3A32A10FCD7" bold="true" box="[189,334,756,782]" fontSize="11" level="3" pageId="92" pageNumber="93" reason="3">
<emphasis id="D3DED104A312FFBAAEBDE3A32A10FCD7" bold="true" box="[189,334,756,782]" pageId="92" pageNumber="93">Description</emphasis>
</heading>
</paragraph>
<paragraph id="E1150D16A312FFBAAEBDE27328F1FBEA" blockId="92.[189,1399,803,1076]" pageId="92" pageNumber="93">HEAD. Head subequal to or wider than pronotum. Discal portion of crown radially or longitudinally striate or shagreen. Anterior margin of head foliaceous or with 2 or 3 parallel carinae. Frontoclypeus tumid or not; texture shagreen. Laterofrontal sutures keeled above antennal pits. Clypellus parallelsided, tapering apically, or widening apically; apex following or slightly surpassing normal curve of gena. Lorum subequal to or wider than clypellus near base. Antennal bases near middle or posteroventral (lower) corners of eyes. Antennae short, less than 1.5 x width of head. Gena strongly incised laterally (nearly forming right angle); with or without fine erect seta beside laterofrontal suture. Antennal ledges absent. Ocelli present; close to eyes; on anterior margin of head.</paragraph>
<paragraph id="E1150D16A312FFBAAEBDE50C2E36FBAC" blockId="92.[189,1384,1115,1141]" box="[189,1384,1115,1141]" pageId="92" pageNumber="93">THORAX. Pronotum lateral margin carinate; lateral margin as long as or longer than basal width of eye.</paragraph>
<paragraph id="E1150D16A312FFBAAEBDE5CA28E9FB24" blockId="92.[189,1399,1180,1277]" pageId="92" pageNumber="93">WINGS. Forewing brachypterous to macropterous; appendix restricted to anal margin or large, extending around wing apex; with 3 anteapical cells; veins not raised; without reflexed costal veins; A1-A2 crossvein absent or present; apical venation not highly reticulate.</paragraph>
<paragraph id="E1150D16A312FFBAAEBDE4722F0CFA71" blockId="92.[189,1399,1316,1448]" pageId="92" pageNumber="93">
LEGS. Profemur with
<collectionCode id="87BB95D3A312FFBAAFBFE4732AADFAE7" box="[447,499,1316,1342]" country="Australia" name="Australian Museum" pageId="92" pageNumber="93" type="Museum">AM</collectionCode>
1 seta only or with
<collectionCode id="87BB95D3A312FFBAACE1E473284BFAE7" box="[737,789,1316,1342]" country="Australia" name="Australian Museum" pageId="92" pageNumber="93" type="Museum">AM</collectionCode>
1 and with one or more additional proximal setae; intercalary row setae not in one row or scattered; row AV with relatively long macrosetae. Protibia dorsal surface rounded, convex. Metafemur apex macrosetae 2+2+1 or 2+2+1 and with additional proximal macrosetae. Metatarsomere I expanded apically; plantar setae simple, tapered.
</paragraph>
<paragraph id="E1150D16A312FFBAAEBDE4872E29F965" blockId="92.[189,1399,1487,1725]" pageId="92" pageNumber="93">
<collectionCode id="87BB95D3A312FFBAAEBDE4872BA1FA3E" box="[189,255,1488,1514]" pageId="92" pageNumber="93">MALE</collectionCode>
GENITALIA. Apodemes of male sternite II not as long as those of sternite I (sternite II apodemes less developed than in
<taxonomicName id="26AA7695A312FFBAAFCFE4A52963F9D4" box="[463,573,1522,1549]" family="Null" higherTaxonomySource="Manual Input" pageId="92" pageNumber="97" rank="subTribe" subTribe="Hecalina">Hecalina</taxonomicName>
). Valve articulated with pygofer; with short point of articulation with pygofer. Pygofer dorsoapical margin incised to near mid-length; basolateral membranous cleft present; macrosetae well differentiated into several rows. Subgenital plates free from each other; articulated with valve; macrosetae uniseriate laterally. Style broadly bilobed basally, median anterior lobe pronounced. Basal processes of the aedeagus/connective absent. Aedeagus without basal hinge; with a single shaft and gonopore. Connective anterior arms somewhat divergent,
<emphasis id="D3DED104A312FFBAAD89E7F428C0F964" bold="true" box="[905,926,1699,1725]" pageId="92" pageNumber="93">Y</emphasis>
- or
<emphasis id="D3DED104A312FFBAADCBE7F428BEF964" bold="true" box="[971,992,1699,1725]" pageId="92" pageNumber="93">U</emphasis>
-shaped; articulated with aedeagus.
</paragraph>
<paragraph id="E1150D16A312FFBAAEBDE7B32F5AF852" blockId="92.[189,1398,1764,1931]" pageId="92" pageNumber="93">FEMALE GENITALIA. Pygofer with numerous macrosetae. Ovipositor not protruding far beyond pygofer apex. First valvula not strongly convex; dorsal sculpturing pattern granulose or maculose; sculpturing reaching dorsal margin; without distinctly delimited ventroapical sculpturing. Second valvula very broad; gradually or abruptly broadened medially or subapically; with or without dorsal median tooth; teeth on apical 1/3 or more; teeth small, regularly or irregularly shaped.</paragraph>
</subSubSection>
<caption id="B5D55D9EA313FFBBAEBDE6DD2E5CF832" ID-DOI="http://doi.org/10.5281/zenodo.3822772" ID-Zenodo-Dep="3822772" httpUri="https://zenodo.org/record/3822772/files/figure.png" pageId="93" pageNumber="94" startId="93.[189,232,1930,1956]" targetBox="[233,1358,267,1885]" targetPageId="93">
<paragraph id="E1150D16A313FFBBAEBDE6DD2E5CF832" blockId="93.[189,1399,1930,2027]" pageId="93" pageNumber="94">
<emphasis id="D3DED104A313FFBBAEBDE6DD2A47F87D" bold="true" box="[189,281,1930,1956]" pageId="93" pageNumber="94">Fig. 27.</emphasis>
<taxonomicName id="26AA7695A313FFBBAF22E6DD2ADBF87D" box="[290,389,1930,1956]" class="Insecta" family="Cicadellidae" kingdom="Animalia" order="Hemiptera" pageId="93" pageNumber="40" phylum="Arthropoda" rank="tribe" tribe="Hecalini">Hecalini</taxonomicName>
(Glossocratina).
<taxonomicName id="26AA7695A313FFBBAC52E6DD288AF87C" authority="(Stal, 1854)" baseAuthorityName="Stal" baseAuthorityYear="1854" box="[594,980,1930,1957]" class="Insecta" family="Cicadellidae" genus="Glossocratus" kingdom="Animalia" order="Hemiptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="afzelii">
<emphasis id="D3DED104A313FFBBAC52E6DD2863F87D" box="[594,829,1930,1956]" italics="true" pageId="93" pageNumber="94">Glossocratus afzelii</emphasis>
(Stål, 1854)
</taxonomicName>
.
<emphasis id="D3DED104A313FFBBADE0E6DD2F51F87D" bold="true" box="[992,1039,1930,1956]" pageId="93" pageNumber="94">AI</emphasis>
. Standard views (see Material &amp; Methods).
<emphasis id="D3DED104A313FFBBAF5EE6FA2A2BF81E" bold="true" box="[350,373,1965,1991]" pageId="93" pageNumber="94">G</emphasis>
. Connective and style shown with aedeagus.
<emphasis id="D3DED104A313FFBBAD97E6F928F8F811" bold="true" box="[919,934,1966,1992]" pageId="93" pageNumber="94">J</emphasis>
. Dorsal view of ♂ abdominal sternites I-III (partial view of sternite III).
<emphasis id="D3DED104A313FFBBAC45E6862922F832" box="[581,636,2001,2027]" italics="true" pageId="93" pageNumber="94">s1ap</emphasis>
= apodeme of sternite I,
<emphasis id="D3DED104A313FFBBADA2E6862884F832" box="[930,986,2001,2027]" italics="true" pageId="93" pageNumber="94">s2ap</emphasis>
= apodeme of sternite II.
</paragraph>
</caption>
<subSubSection id="A9B05E9DA310FFB8AEBDE05B2FBBFE8C" pageId="94" pageNumber="95" type="distribution">
<paragraph id="E1150D16A310FFB8AEBDE05B2ABAFEFE" blockId="94.[189,484,268,295]" box="[189,484,268,295]" pageId="94" pageNumber="95">
<heading id="BA5DBA7AA310FFB8AEBDE05B2ABAFEFE" bold="true" box="[189,484,268,295]" fontSize="11" level="3" pageId="94" pageNumber="95" reason="3">
<emphasis id="D3DED104A310FFB8AEBDE05B2ABAFEFE" bold="true" box="[189,484,268,295]" pageId="94" pageNumber="95">Geography and ecology</emphasis>
</heading>
</paragraph>
<paragraph id="E1150D16A310FFB8AEBDE06C2FBBFE8C" blockId="94.[189,1253,315,342]" box="[189,1253,315,342]" pageId="94" pageNumber="95">
Distribution: Palearctic, Afrotropical and
<collectingRegion id="236EC3F4A310FFB8ACA5E06C285BFE8C" box="[677,773,315,341]" country="Republic of the Congo" name="Orientale" pageId="94" pageNumber="95">Oriental</collectingRegion>
regions. Glossocratina are grass feeders.
</paragraph>
</subSubSection>
<subSubSection id="A9B05E9DA310FFB8AEBDE02A2AADFDEC" pageId="94" pageNumber="95" type="discussion">
<paragraph id="E1150D16A310FFB8AEBDE02A2A71FE4E" blockId="94.[189,303,381,407]" box="[189,303,381,407]" pageId="94" pageNumber="95">
<heading id="BA5DBA7AA310FFB8AEBDE02A2A71FE4E" bold="true" box="[189,303,381,407]" fontSize="11" level="3" pageId="94" pageNumber="95" reason="3">
<emphasis id="D3DED104A310FFB8AEBDE02A2A71FE4E" bold="true" box="[189,303,381,407]" pageId="94" pageNumber="95">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="E1150D16A310FFB8AEBDE0FC286CFE1C" blockId="94.[189,818,427,454]" box="[189,818,427,454]" pageId="94" pageNumber="95">
Glossocratina contains the
<typeStatus id="3E11B3B4A310FFB8AFFAE0FB2973FE1F" box="[506,557,428,454]" pageId="94" pageNumber="95">type</typeStatus>
genus and 29 species.
</paragraph>
<paragraph id="E1150D16A310FFB8AEBDE0BA2AD6FDDE" blockId="94.[189,392,493,519]" box="[189,392,493,519]" pageId="94" pageNumber="95">
<heading id="BA5DBA7AA310FFB8AEBDE0BA2AD6FDDE" bold="true" box="[189,392,493,519]" fontSize="11" level="3" pageId="94" pageNumber="95" reason="3">
<emphasis id="D3DED104A310FFB8AEBDE0BA2AD6FDDE" bold="true" box="[189,392,493,519]" pageId="94" pageNumber="95">Included genera</emphasis>
</heading>
</paragraph>
<paragraph id="E1150D16A310FFB8AEBDE34C2AADFDEC" blockId="94.[189,499,539,565]" box="[189,499,539,565]" pageId="94" pageNumber="95">
<taxonomicName id="26AA7695A310FFB8AEBDE34C2AADFDEC" authority="Fieber, 1866" authorityName="Fieber" authorityYear="1866" box="[189,499,539,565]" class="Insecta" family="Cicadellidae" genus="Glossocratus" kingdom="Animalia" order="Hemiptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="genus">
<emphasis id="D3DED104A310FFB8AEBDE34C2A06FDEC" box="[189,344,539,565]" italics="true" pageId="94" pageNumber="95">Glossocratus</emphasis>
Fieber, 1866
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>