692 lines
83 KiB
XML
692 lines
83 KiB
XML
<document id="F60CC43D766DC056BD5CBB75B14A0401" ID-CLB-Dataset="39179" ID-DOI="10.11646/zootaxa.4107.1.3" ID-GBIF-Dataset="be2b4a90-7526-4c63-a5f4-9fe399ab2dac" ID-ISSN="1175-5326" ID-Zenodo-Dep="271350" ID-ZooBank="21FB8099-278A-4F44-B207-717683A0D5B0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461910688042" checkinUser="plazi" docAuthor="Freyhof, Jörg, Geiger, Matthias F., Golzarianpour, Kiavash & Patimar, Rahman" docDate="2016" docId="59377E3DB76BFFF6FF7CFADBFA6DFDFE" docLanguage="en" docName="zootaxa.4107.1.3.pdf.imf" docOrigin="Zootaxa 4107 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sasanidus Freyhof, Geiger, Golzarianpour & Patimar, 2016, new genus" docType="treatment" docVersion="9" lastPageNumber="76" masterDocId="A50E0645B76EFFFDFFEBFFA7FFFBFFE6" masterDocTitle="Sasanidus, a new generic name for Noemacheilus kermanshahensis Bănărescu & Nalbant, with discussion of Ilamnemacheilus and Schistura (Teleostei; Nemacheilidae)" masterLastPageNumber="80" masterPageNumber="65" pageNumber="70" updateTime="1698662070806" updateUser="ExternalLinkService">
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<mods:mods id="860D472BE74D4DD2A4A7CE9A3C37B907" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="2103E232A1A07BD6A1E128D69689B18D">
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<mods:title id="90C94F73484F71346CD8EF05DCD8407A">Sasanidus, a new generic name for Noemacheilus kermanshahensis Bănărescu & Nalbant, with discussion of Ilamnemacheilus and Schistura (Teleostei; Nemacheilidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="DB74375EBBCB083000A42ADFA4383139" type="personal">
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<mods:role id="3966AAAFC24FBA779FB9B2266746AAC4">
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<mods:roleTerm id="25CA45C46933C0E062D5839B0B0B378A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E41653B8F307A9316DF5B44B90F2E595">Freyhof, Jörg</mods:namePart>
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</mods:name>
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<mods:roleTerm id="D51D9837E2F751C2F91E6C5E4988CCE5">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0DB0546F8BC9181724EAF2F2B1C9E146">Geiger, Matthias F.</mods:namePart>
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</mods:name>
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<mods:roleTerm id="17596DC733EF4493EA52046F11747C7A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="CBF75399FB65F7345980E5BC0AC35F66">Golzarianpour, Kiavash</mods:namePart>
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</mods:name>
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<mods:name id="F066D193752486216BBE66A8585C3F5E" type="personal">
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<mods:role id="0E01DF3CECE4C028AA1046121C56D584">
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<mods:roleTerm id="57EF5A93840417CC809AA3B4FADBF583">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3413D51BFACB3F72E46FCF831A007629">Patimar, Rahman</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="537BA85AACABEAE7BDFA72B5E61B4C1F">text</mods:typeOfResource>
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<mods:titleInfo id="B30D20015CC2237DE8F93A25C6BE9DE6">
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<mods:title id="2AD99662CF034EC38CE24B0251F270AB">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="E229A476FEF888EAD2A629A24E4B6B8E">
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<mods:date id="0D9D8F4B8402F9C79E6340A039221EFC">2016</mods:date>
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<mods:detail id="40483322ED138B7B3E9B9A56E46A358C" type="volume">
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<mods:number id="9AB9BF088A272E947307A3932FF5CA86">4107</mods:number>
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<mods:detail id="C5C8460045962808E94FA70E60F39022" type="issue">
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<mods:number id="63545598FC5CE0BC12780D890094C63E">1</mods:number>
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<mods:start id="E6A50A6D5160FB448798B6F1A7EE5C63">65</mods:start>
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<mods:end id="8178492D6A65D10FB4F1A5D25FCFC32C">80</mods:end>
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<mods:classification id="87DAC85B7F3F12B005F2DC8528BDED8F">journal article</mods:classification>
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<mods:identifier id="19FE6A6427DBEC8F9062DFFB64DF398C" type="DOI">10.11646/zootaxa.4107.1.3</mods:identifier>
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<mods:identifier id="75483B31D7D2B276A7CD4102F82862AA" type="GBIF-Dataset">be2b4a90-7526-4c63-a5f4-9fe399ab2dac</mods:identifier>
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<mods:identifier id="66CE8A6A3B8A458CD5EE34DC39FCCAEF" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="077C1A400B343E7EAC0D8E9874249EC7" type="Zenodo-Dep">271350</mods:identifier>
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<treatment id="59377E3DB76BFFF6FF7CFADBFA6DFDFE" ID-DOI="http://doi.org/10.5281/zenodo.6057150" ID-GBIF-Taxon="119651895" ID-Zenodo-Dep="6057150" LSID="urn:lsid:plazi:treatment:59377E3DB76BFFF6FF7CFADBFA6DFDFE" httpUri="http://treatment.plazi.org/id/59377E3DB76BFFF6FF7CFADBFA6DFDFE" lastPageId="11" lastPageNumber="76" pageId="5" pageNumber="70">
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<subSubSection id="99849CA0B76BFFF8FF7CFADBFF0EFA5C" pageId="5" pageNumber="70" type="nomenclature">
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<paragraph id="D121CF2BB76BFFF8FF7CFADBFE6DFA70" blockId="5.[151,406,1404,1466]" box="[151,406,1404,1430]" pageId="5" pageNumber="70">
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<heading id="8A697847B76BFFF8FF7CFADBFE6DFA70" bold="true" box="[151,406,1404,1430]" fontSize="10" level="2" pageId="5" pageNumber="70" reason="2">
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||
<emphasis id="E3EA1339B76BFFF8FF7CFADBFE6DFA70" bold="true" box="[151,406,1404,1430]" pageId="5" pageNumber="70">
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||
<taxonomicName id="169EB4A8B76BFFF8FF7CFADBFEF6FA73" box="[151,269,1404,1429]" class="Actinopterygii" family="Nemacheilidae" genus="Sasanidus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="70" phylum="Chordata" rank="genus" status="gen. nov.">
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<emphasis id="E3EA1339B76BFFF8FF7CFADBFEF6FA73" bold="true" box="[151,269,1404,1429]" italics="true" pageId="5" pageNumber="70">Sasanidus</emphasis>
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||
</taxonomicName>
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||
,
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||
<taxonomicNameLabel id="F8D9AE42B76BFFF8FEF0FADAFE6DFA70" box="[283,406,1405,1430]" pageId="5" pageNumber="70" rank="genus">new genus</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="D121CF2BB76BFFF8FF7CFA06FF0EFA5C" blockId="5.[151,406,1404,1466]" box="[151,245,1441,1466]" pageId="5" pageNumber="70">
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||
<figureCitation id="49A5D3AEB76BFFF8FF7CFA06FF0EFA5C" box="[151,245,1441,1466]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="6.[151,244,670,691]" captionStartId-1="6.[151,244,1626,1647]" captionStartId-2="7.[151,244,1835,1856]" captionTargetBox-0="[150,1436,191,649]" captionTargetBox-1="[151,1436,776,1603]" captionTargetBox-2="[151,1435,191,1814]" captionTargetId-0="figure@6.[150,1436,191,649]" captionTargetId-1="figure@6.[151,1436,776,1605]" captionTargetId-2="figure@7.[151,1435,191,1814]" captionTargetPageId-0="6" captionTargetPageId-1="6" captionTargetPageId-2="7" captionText-0="FIGURE 2. Sasanidus kermanshahensis, about 45 mm SL, not preserved; Iran: Haram-Abad stream." captionText-1="FIGURE 3. Sasanidus kermanshahensis, Iran: a, FCKG 806, 70 mm SL; stream Ab-alashtar. Black spots are parasites; b, FCKG 800, 42 mm SL; spring Pashow; c, FCKG 804, 70 mm SL; Gamasiab River at Baba-Rostam." captionText-2="FIGURE 4. Sasanidus kermanshahensis, FSJF 3213, 63 mm SL; Iran: Gamasiab River." httpUri-0="https://zenodo.org/record/271352/files/figure.png" httpUri-1="https://zenodo.org/record/271353/files/figure.png" httpUri-2="https://zenodo.org/record/271354/files/figure.png" pageId="5" pageNumber="70">Fig. 2–4</figureCitation>
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||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="99849CA0B76BFFFBFF7CFA4EFEDFF8D8" lastPageId="6" lastPageNumber="71" pageId="5" pageNumber="70" type="materials_examined">
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||
<paragraph id="D121CF2BB76BFFF8FF7CFA4EFC3AFA19" blockId="5.[151,961,1513,1535]" box="[151,961,1513,1535]" pageId="5" pageNumber="70">
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||
<emphasis id="E3EA1339B76BFFF8FF7CFA4EFED9FA19" bold="true" box="[151,290,1513,1535]" pageId="5" pageNumber="70">
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||
<typeStatus id="0E257189B76BFFF8FF7CFA4EFF37FA19" box="[151,204,1513,1535]" pageId="5" pageNumber="70">Type</typeStatus>
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species.
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</emphasis>
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||
<taxonomicName id="169EB4A8B76BFFF8FEC2FA4EFC46FA19" authority="Banarescu & Nalbant, 1966:151" authorityName="Banarescu & Nalbant" authorityPageNumber="151" authorityYear="1966" box="[297,957,1513,1535]" family="Nemacheilidae" genus="Noemacheilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="70" phylum="Chordata" rank="species" species="kermanshahensis">
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<emphasis id="E3EA1339B76BFFF8FEC2FA4EFD96FA19" box="[297,621,1513,1535]" italics="true" pageId="5" pageNumber="70">Noemacheilus kermanshahensis</emphasis>
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||
<bibRefCitation id="B50FB2DAB76BFFF8FD98FA4DFC69FA19" author="Banarescu" box="[627,914,1513,1535]" pageId="5" pageNumber="70" refString="Banarescu, P. G. & Nalbant. T. T. (1966) The 3 rd Danish Expedition to Central Asia. Zoological results 34. Cobitidae (Pisces) from Afghanistan and Iran. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening i Kobenhavn, 129, 149 - 186." type="journal article" year="1966">Bănărescu & Nalbant, 1966</bibRefCitation>
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||
:151
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||
</taxonomicName>
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||
.
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||
</paragraph>
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||
<paragraph id="D121CF2BB76BFFF8FF7CF98BFEE4F812" blockId="5.[151,1437,1580,2036]" pageId="5" pageNumber="70">
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||
<emphasis id="E3EA1339B76BFFF8FF7CF98BFE7EF9A3" bold="true" box="[151,389,1580,1605]" pageId="5" pageNumber="70">Material examined.</emphasis>
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||
All from
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<collectingCountry id="A9898FBBB76BFFF8FDE9F98AFDC8F9A2" box="[514,563,1581,1604]" name="Iran" pageId="5" pageNumber="70">Iran</collectingCountry>
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||
:
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||
<collectionCode id="B78F57EEB76BFFF8FDA8F98BFD86F9A2" box="[579,637,1580,1604]" pageId="5" pageNumber="70">FSJF</collectionCode>
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||
3213, 5,
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||
<quantity id="166662CEB76BFFF8FD1FF98BFC89F9A2" box="[756,882,1580,1605]" metricMagnitude="-2" metricUnit="m" metricValue="5.45" metricValueMax="6.3" metricValueMin="4.6" pageId="5" pageNumber="70" unit="mm" value="54.5" valueMax="63.0" valueMin="46.0">46–63 mm</quantity>
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||
SL; Hamadan prov.: Gamasiab River,
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||
<geoCoordinate id="B4AAA9ECB76BFFF8FAABF98BFA67F9A2" box="[1344,1436,1580,1604]" direction="north" orientation="latitude" pageId="5" pageNumber="70" precision="925" value="34.216667">34°13'N</geoCoordinate>
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||
<geoCoordinate id="B4AAA9ECB76BFFF8FF7CF9F7FF08F98E" box="[151,243,1616,1640]" direction="east" orientation="longitude" pageId="5" pageNumber="70" precision="925" value="48.283333">48°17'E</geoCoordinate>
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||
. —
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||
<collectionCode id="B78F57EEB76BFFF8FECEF9F7FEA4F98E" box="[293,351,1616,1640]" pageId="5" pageNumber="70">FSJF</collectionCode>
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||
3234, 5,
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||
<quantity id="166662CEB76BFFF8FE26F9E8FDB2F981" box="[461,585,1615,1640]" metricMagnitude="-2" metricUnit="m" metricValue="3.75" metricValueMax="4.5" metricValueMin="3.0" pageId="5" pageNumber="70" unit="mm" value="37.5" valueMax="45.0" valueMin="30.0">30–45 mm</quantity>
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||
SL; Kermanshah prov.: Sepidbarg River west of Javanrud,
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||
<geoCoordinate id="B4AAA9ECB76BFFF8FB1EF9F7FA67F98E" box="[1269,1436,1616,1640]" direction="north" orientation="latitude" pageId="5" pageNumber="70" precision="1" value="34.805954">34°48'21.44"N</geoCoordinate>
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||
<geoCoordinate id="B4AAA9ECB76BFFF8FF7CF9D3FEC4F96B" box="[151,319,1652,1677]" direction="east" orientation="longitude" pageId="5" pageNumber="70" precision="1" value="46.4581">46°27'29.15"E</geoCoordinate>
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||
. —
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||
<collectionCode id="B78F57EEB76BFFF8FE91F9D3FE3EF96A" box="[378,453,1652,1676]" pageId="5" pageNumber="70">FCKG</collectionCode>
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||
802, 21,
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||
<quantity id="166662CEB76BFFF8FDAFF9D3FD3FF96A" box="[580,708,1652,1677]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="4.5" metricValueMin="3.5" pageId="5" pageNumber="70" unit="mm" value="40.0" valueMax="45.0" valueMin="35.0">35–45 mm</quantity>
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SL, Lorestan prov.: stream below Horro bridge at Dorood- Khoramabad highway,
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||
<geoCoordinate id="B4AAA9ECB76BFFF8FE74F93FFDC3F956" box="[415,568,1688,1712]" direction="north" orientation="latitude" pageId="5" pageNumber="70" precision="1" value="33.5205">33°31'13.8"N</geoCoordinate>
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||
<geoCoordinate id="B4AAA9ECB76BFFF8FDABF930FD21F956" box="[576,730,1687,1712]" direction="east" orientation="longitude" pageId="5" pageNumber="70" precision="1" value="48.771946">48°46'19.0"E</geoCoordinate>
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||
. —
|
||
<collectionCode id="B78F57EEB76BFFF8FCE0F93FFCADF956" box="[779,854,1688,1712]" pageId="5" pageNumber="70">FCKG</collectionCode>
|
||
806, 3,
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||
<quantity id="166662CEB76BFFF8FC5DF930FBCBF949" box="[950,1072,1687,1712]" metricMagnitude="-2" metricUnit="m" metricValue="5.75" metricValueMax="7.0" metricValueMin="4.5" pageId="5" pageNumber="70" unit="mm" value="57.5" valueMax="70.0" valueMin="45.0">45–70 mm</quantity>
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||
SL, Lorestan prov.: stream Ab- Alashtar on road from Alashtar to Khoramabad,
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||
<geoCoordinate id="B4AAA9ECB76BFFF8FD25F91BFC9CF933" box="[718,871,1724,1749]" direction="north" orientation="latitude" pageId="5" pageNumber="70" precision="1" value="33.786">33°47'09.6"N</geoCoordinate>
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||
<geoCoordinate id="B4AAA9ECB76BFFF8FC9AF91BFBF0F932" box="[881,1035,1724,1748]" direction="east" orientation="longitude" pageId="5" pageNumber="70" precision="1" value="48.20639">48°12'23.0"E</geoCoordinate>
|
||
. —
|
||
<collectionCode id="B78F57EEB76BFFF8FBD4F91BFB71F932" box="[1087,1162,1724,1748]" pageId="5" pageNumber="70">FCKG</collectionCode>
|
||
805, 4,
|
||
<quantity id="166662CEB76BFFF8FB05F91BFA90F932" box="[1262,1387,1724,1748]" metricMagnitude="-2" metricUnit="m" metricValue="4.25" metricValueMax="4.7" metricValueMin="3.8" pageId="5" pageNumber="70" unit="mm" value="42.5" valueMax="47.0" valueMin="38.0">38–47 mm</quantity>
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||
SL, Lorestan prov.: Sezar (Marboreh) River at Dorood,
|
||
<geoCoordinate id="B4AAA9ECB76BFFF8FD08F947FC87F91E" box="[739,892,1760,1784]" direction="north" orientation="latitude" pageId="5" pageNumber="70" precision="1" value="33.477474">33°28'38.9"N</geoCoordinate>
|
||
<geoCoordinate id="B4AAA9ECB76BFFF8FC6FF978FBF3F91E" box="[900,1032,1759,1784]" direction="east" orientation="longitude" pageId="5" pageNumber="70" precision="15" value="49.066387">49°03'59"E</geoCoordinate>
|
||
. —
|
||
<collectionCode id="B78F57EEB76BFFF8FBD2F947FB7FF91E" box="[1081,1156,1760,1784]" pageId="5" pageNumber="70">FCKG</collectionCode>
|
||
800, 40,
|
||
<quantity id="166662CEB76BFFF8FB1AF947FA97F911" box="[1265,1388,1760,1784]" metricMagnitude="-2" metricUnit="m" metricValue="5.45" metricValueMax="7.1" metricValueMin="3.8" pageId="5" pageNumber="70" unit="mm" value="54.5" valueMax="71.0" valueMin="38.0">38–71 mm</quantity>
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||
SL; Hamadan prov.: spring Pashow near
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||
<collectingCountry id="A9898FBBB76BFFF8FDDEF8A3FD95F8FB" box="[565,622,1796,1821]" name="Guyana" pageId="5" pageNumber="70">Gian</collectingCountry>
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||
, flowing to Gamasiab River,
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||
<geoCoordinate id="B4AAA9ECB76BFFF8FC53F8A3FBEFF8FA" box="[952,1044,1796,1820]" direction="north" orientation="latitude" pageId="5" pageNumber="70" precision="925" value="34.216667">34°13'N</geoCoordinate>
|
||
<geoCoordinate id="B4AAA9ECB76BFFF8FBF7F8A3FB83F8FA" box="[1052,1144,1796,1820]" direction="east" orientation="longitude" pageId="5" pageNumber="70" precision="925" value="48.283333">48°17'E</geoCoordinate>
|
||
. —
|
||
<collectionCode id="B78F57EEB76BFFF8FB4DF8A3FB0AF8FA" box="[1190,1265,1796,1820]" pageId="5" pageNumber="70">FCKG</collectionCode>
|
||
804, 1,
|
||
<quantity id="166662CEB76BFFF8FAA6F8A3FA67F8FA" box="[1357,1436,1796,1820]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="70" unit="mm" value="70.0">70 mm</quantity>
|
||
SL; Hamadan prov.: Gamasiab River at Baba-Rostam,
|
||
<geoCoordinate id="B4AAA9ECB76BFFF8FCE8F880FC66F8A6" box="[771,925,1831,1856]" direction="north" orientation="latitude" pageId="5" pageNumber="70" precision="1" value="34.132137">34°07'55.7"N</geoCoordinate>
|
||
<geoCoordinate id="B4AAA9ECB76BFFF8FC4FF880FBC6F8A6" box="[932,1085,1831,1856]" direction="east" orientation="longitude" pageId="5" pageNumber="70" precision="1" value="48.39864">48°23'55.1"E</geoCoordinate>
|
||
. —
|
||
<collectionCode id="B78F57EEB76BFFF8FB86F88FFB43F8A6" box="[1133,1208,1832,1856]" pageId="5" pageNumber="70">FCKG</collectionCode>
|
||
801, 21,
|
||
<quantity id="166662CEB76BFFF8FAC8F880FA66F8D9" box="[1315,1437,1831,1856]" metricMagnitude="-2" metricUnit="m" metricValue="3.85" metricValueMax="4.5" metricValueMin="3.2" pageId="5" pageNumber="70" unit="mm" value="38.5" valueMax="45.0" valueMin="32.0">32–45 mm</quantity>
|
||
SL; Hamadan prov.: Haram-Abad River at Ashmizan,
|
||
<geoCoordinate id="B4AAA9ECB76BFFF8FCF6F8EBFC81F882" box="[797,890,1868,1892]" direction="north" orientation="latitude" pageId="5" pageNumber="70" precision="925" value="34.116665">34°07'N</geoCoordinate>
|
||
<geoCoordinate id="B4AAA9ECB76BFFF8FC6DF8EBFC15F882" box="[902,1006,1868,1893]" direction="east" orientation="longitude" pageId="5" pageNumber="70" precision="925" value="49.416668">48°85' E</geoCoordinate>
|
||
. —
|
||
<collectionCode id="B78F57EEB76BFFF8FBCCF8EBFB89F882" box="[1063,1138,1868,1892]" pageId="5" pageNumber="70">FCKG</collectionCode>
|
||
807, 20,
|
||
<quantity id="166662CEB76BFFF8FB01F8EBFA93F882" box="[1258,1384,1868,1893]" metricMagnitude="-2" metricUnit="m" metricValue="3.85" metricValueMax="4.5" metricValueMin="3.2" pageId="5" pageNumber="70" unit="mm" value="38.5" valueMax="45.0" valueMin="32.0">32–45 mm</quantity>
|
||
SL; Hamadan prov.: stream seyyed Shahab, Bujin village, Asad-abad,
|
||
<geoCoordinate id="B4AAA9ECB76BFFF8FC71F8D7FBC8F86E" box="[922,1075,1904,1928]" direction="north" orientation="latitude" pageId="5" pageNumber="70" precision="1" value="34.80303">34°48'10.9"N</geoCoordinate>
|
||
<geoCoordinate id="B4AAA9ECB76BFFF8FBD6F8D7FB39F86E" box="[1085,1218,1904,1928]" direction="east" orientation="longitude" pageId="5" pageNumber="70" precision="15" value="48.045">48°02'42"E</geoCoordinate>
|
||
. —
|
||
<collectionCode id="B78F57EEB76BFFF8FB1DF8D7FABAF86E" box="[1270,1345,1904,1928]" pageId="5" pageNumber="70">FCKG</collectionCode>
|
||
803, 2,
|
||
<quantity id="166662CEB76BFFF8FF7CF833FEEAF84A" box="[151,273,1940,1965]" metricMagnitude="-2" metricUnit="m" metricValue="3.75" metricValueMax="4.0" metricValueMin="3.5" pageId="5" pageNumber="70" unit="mm" value="37.5" valueMax="40.0" valueMin="35.0">35–40 mm</quantity>
|
||
SL; Lorestan prov.: stream Kakareza at Kakareza village,
|
||
<geoCoordinate id="B4AAA9ECB76BFFF8FC5BF833FBA9F84A" box="[944,1106,1940,1965]" direction="north" orientation="latitude" pageId="5" pageNumber="70" precision="1" value="33.709972">33°42'35.9" N</geoCoordinate>
|
||
<geoCoordinate id="B4AAA9ECB76BFFF8FBB0F833FB0FF84B" box="[1115,1268,1940,1965]" direction="east" orientation="longitude" pageId="5" pageNumber="70" precision="1" value="48.277416">48°16'38.7"E</geoCoordinate>
|
||
. —
|
||
<collectionCode id="B78F57EEB76BFFF8FACEF833FA8BF84A" box="[1317,1392,1940,1964]" pageId="5" pageNumber="70">FCKG</collectionCode>
|
||
not catalogued, 6, 35-
|
||
<quantity id="166662CEB76BFFF8FE81F81FFE3BF836" box="[362,448,1976,2001]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="5" pageNumber="70" unit="mm" value="45.0">45 mm</quantity>
|
||
SL; Lorestan prov.: Houzian river, road Aligudarz toward Gahar lake,
|
||
<geoCoordinate id="B4AAA9ECB76BFFF8FAF3F81FFA67F836" box="[1304,1436,1976,2000]" direction="north" orientation="latitude" pageId="5" pageNumber="70" precision="15" value="33.35861">33°21'31"N</geoCoordinate>
|
||
<geoCoordinate id="B4AAA9ECB76BFFF8FF7CF87BFEE0F812" box="[151,283,2012,2036]" direction="east" orientation="longitude" pageId="5" pageNumber="70" precision="15" value="49.74">49°44'24"E</geoCoordinate>
|
||
.
|
||
</paragraph>
|
||
<caption id="85E19FA3B768FFFBFF7CFD39FB99FD55" box="[151,1122,670,691]" httpUri="https://zenodo.org/record/271352/files/figure.png" pageId="6" pageNumber="71" targetBox="[150,1436,191,649]" targetPageId="6">
|
||
<paragraph id="D121CF2BB768FFFBFF7CFD39FB99FD55" blockId="6.[151,1122,670,691]" box="[151,1122,670,691]" pageId="6" pageNumber="71">
|
||
<emphasis id="E3EA1339B768FFFBFF7CFD39FEF7FD55" bold="true" box="[151,268,670,691]" pageId="6" pageNumber="71">FIGURE 2.</emphasis>
|
||
<taxonomicName id="169EB4A8B768FFFBFEF9FD39FDE6FD55" box="[274,541,670,691]" class="Actinopterygii" family="Nemacheilidae" genus="Sasanidus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="71" phylum="Chordata" rank="species" species="kermanshahensis">
|
||
<emphasis id="E3EA1339B768FFFBFEF9FD39FDE6FD55" box="[274,541,670,691]" italics="true" pageId="6" pageNumber="71">Sasanidus kermanshahensis</emphasis>
|
||
</taxonomicName>
|
||
, about 45 mm SL, not preserved; Iran: Haram-Abad stream.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="85E19FA3B768FFFBFF7CF9FDFC19F96A" httpUri="https://zenodo.org/record/271353/files/figure.png" pageId="6" pageNumber="71" targetBox="[151,1436,776,1603]" targetPageId="6">
|
||
<paragraph id="D121CF2BB768FFFBFF7CF9FDFC19F96A" blockId="6.[151,1436,1626,1676]" pageId="6" pageNumber="71">
|
||
<emphasis id="E3EA1339B768FFFBFF7CF9FDFEF7F989" bold="true" box="[151,268,1626,1647]" pageId="6" pageNumber="71">FIGURE 3.</emphasis>
|
||
<taxonomicName id="169EB4A8B768FFFBFEF9F9FDFDE5F989" box="[274,542,1626,1647]" class="Actinopterygii" family="Nemacheilidae" genus="Sasanidus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="71" phylum="Chordata" rank="species" species="kermanshahensis">
|
||
<emphasis id="E3EA1339B768FFFBFEF9F9FDFDE5F989" box="[274,542,1626,1647]" italics="true" pageId="6" pageNumber="71">Sasanidus kermanshahensis</emphasis>
|
||
</taxonomicName>
|
||
, Iran: a, FCKG 806, 70 mm SL; stream Ab-alashtar. Black spots are parasites; b, FCKG 800, 42 mm SL; spring Pashow; c, FCKG 804, 70 mm SL; Gamasiab River at Baba-Rostam.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D121CF2BB768FFFBFF2CF97AFEDFF8D8" blockId="6.[151,1436,1757,2034]" pageId="6" pageNumber="71">
|
||
<emphasis id="E3EA1339B768FFFBFF2CF97AFD07F913" bold="true" box="[199,764,1757,1782]" pageId="6" pageNumber="71">Material used in the molecular genetic analysis.</emphasis>
|
||
<collectionCode id="B78F57EEB768FFFBFCE8F979FCC6F910" box="[771,829,1758,1782]" pageId="6" pageNumber="71">FSJF</collectionCode>
|
||
DNA-1997;
|
||
<collectingCountry id="A9898FBBB768FFFBFC24F979FBFBF913" box="[975,1024,1758,1781]" name="Iran" pageId="6" pageNumber="71">Iran</collectingCountry>
|
||
: Kermanshah prov.: Sepidbarg River west of Javanrud,
|
||
<geoCoordinate id="B4AAA9ECB768FFFBFE61F8A6FDCAF8FF" box="[394,561,1793,1817]" direction="north" orientation="latitude" pageId="6" pageNumber="71" precision="1" value="34.805954">34°48'21.44"N</geoCoordinate>
|
||
<geoCoordinate id="B4AAA9ECB768FFFBFDADF8A6FD12F8FC" box="[582,745,1793,1818]" direction="east" orientation="longitude" pageId="6" pageNumber="71" precision="1" value="46.4581">46°27'29.15"E</geoCoordinate>
|
||
(GenBank accession numbers:
|
||
<accessionNumber id="CECD52C8B768FFFBFB6AF8A6FAFAF8FC" box="[1153,1281,1793,1818]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KU928261" pageId="6" pageNumber="71">KU928261</accessionNumber>
|
||
,
|
||
<accessionNumber id="CECD52C8B768FFFBFAF2F8A6FA63F8FF" box="[1305,1432,1793,1817]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KU928288" pageId="6" pageNumber="71">KU928288</accessionNumber>
|
||
,
|
||
<accessionNumber id="CECD52C8B768FFFBFF7CF881FEE2F8D8" box="[151,281,1830,1854]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KU928289" pageId="6" pageNumber="71">KU928289</accessionNumber>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="99849CA0B768FFF5FF2CF8EEFB1AFDFE" lastPageId="8" lastPageNumber="73" pageId="6" pageNumber="71" type="diagnosis">
|
||
<paragraph id="D121CF2BB768FFFAFF2CF8EEFA67F817" blockId="6.[151,1436,1757,2034]" lastBlockId="7.[151,1436,1937,2034]" lastPageId="7" lastPageNumber="72" pageId="6" pageNumber="71">
|
||
<emphasis id="E3EA1339B768FFFBFF2CF8EEFEBAF884" bold="true" box="[199,321,1865,1890]" pageId="6" pageNumber="71">Diagnosis.</emphasis>
|
||
<taxonomicName id="169EB4A8B768FFFBFEA5F8EEFE3BF884" box="[334,448,1865,1890]" class="Actinopterygii" family="Nemacheilidae" genus="Sasanidus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="71" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B768FFFBFEA5F8EEFE3BF884" box="[334,448,1865,1890]" italics="true" pageId="6" pageNumber="71">Sasanidus</emphasis>
|
||
</taxonomicName>
|
||
is not diagnosed by any unique external character state, but by a combination of characters.
|
||
<taxonomicName id="169EB4A8B768FFFBFEFDF8CAFE7CF860" box="[278,391,1901,1926]" class="Actinopterygii" family="Nemacheilidae" genus="Sasanidus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="71" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B768FFFBFEFDF8CAFE7CF860" box="[278,391,1901,1926]" italics="true" pageId="6" pageNumber="71">Sasanidus</emphasis>
|
||
</taxonomicName>
|
||
was earlier placed in the genus
|
||
<taxonomicName id="169EB4A8B768FFFBFD07F8CAFC48F860" box="[748,947,1901,1926]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="71" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B768FFFBFD07F8CAFC48F860" box="[748,947,1901,1926]" italics="true" pageId="6" pageNumber="71">Oxynoemacheilus</emphasis>
|
||
</taxonomicName>
|
||
and is distinguished from all species in that genus and also from all species of
|
||
<taxonomicName id="169EB4A8B768FFFBFDC8F836FD1EF84C" box="[547,741,1937,1962]" class="Actinopterygii" family="Nemacheilidae" genus="Seminemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="71" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B768FFFBFDC8F836FD1EF84C" box="[547,741,1937,1962]" italics="true" pageId="6" pageNumber="71">Seminemacheilus</emphasis>
|
||
</taxonomicName>
|
||
by the absence of an external sexual dimorphism (vs. longer pectoral fin, nuptial tubercles on fins, head and back in male). The prominent, irregularly shaped midlateral stripe, being characteristic for
|
||
<taxonomicName id="169EB4A8B768FFFBFE4BF87DFD86F814" box="[416,637,2009,2034]" class="Actinopterygii" family="Nemacheilidae" genus="Sasanidus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="71" phylum="Chordata" rank="species" species="kermanshahensis">
|
||
<emphasis id="E3EA1339B768FFFBFE4BF87DFD86F814" box="[416,637,2009,2034]" italics="true" pageId="6" pageNumber="71">S. kermanshahensis</emphasis>
|
||
</taxonomicName>
|
||
, is rarely found in species of
|
||
<taxonomicName id="169EB4A8B768FFFBFC23F87EFB74F814" box="[968,1167,2009,2034]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="71" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B768FFFBFC23F87EFB74F814" box="[968,1167,2009,2034]" italics="true" pageId="6" pageNumber="71">Oxynoemacheilus</emphasis>
|
||
</taxonomicName>
|
||
, but present in a few species (
|
||
<taxonomicName id="169EB4A8B769FFFAFF13F835FE8CF84F" box="[248,375,1938,1961]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="72" phylum="Chordata" rank="species" species="angorae">
|
||
<emphasis id="E3EA1339B769FFFAFF13F835FE8CF84F" box="[248,375,1938,1961]" italics="true" pageId="7" pageNumber="72">
|
||
<collectingCountry id="A9898FBBB769FFFAFF13F835FEF4F84F" box="[248,271,1938,1961]" name="Iceland" pageId="7" pageNumber="72">O</collectingCountry>
|
||
. angorae
|
||
</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="169EB4A8B769FFFAFE6DF835FDCBF84C" box="[390,560,1937,1962]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="72" phylum="Chordata" rank="species" species="phoxinoides">
|
||
<emphasis id="E3EA1339B769FFFAFE6DF835FDCBF84C" box="[390,560,1937,1962]" italics="true" pageId="7" pageNumber="72">
|
||
<collectingCountry id="A9898FBBB769FFFAFE6DF835FE66F84F" box="[390,413,1938,1961]" name="Iceland" pageId="7" pageNumber="72">O</collectingCountry>
|
||
. phoxinoides
|
||
</emphasis>
|
||
</taxonomicName>
|
||
). Not all individuals of
|
||
<taxonomicName id="169EB4A8B769FFFAFCD5F835FBE0F84C" box="[830,1051,1937,1962]" class="Actinopterygii" family="Nemacheilidae" genus="Sasanidus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="72" phylum="Chordata" rank="species" species="kermanshahensis">
|
||
<emphasis id="E3EA1339B769FFFAFCD5F835FBE0F84C" box="[830,1051,1937,1962]" italics="true" pageId="7" pageNumber="72">S. kermanshahensis</emphasis>
|
||
</taxonomicName>
|
||
show the characteristic midlateral stripe, which is often restricted to the caudal peduncle, and some have a mottled or marbled colour pattern (
|
||
<figureCitation id="49A5D3AEB769FFFAFAA1F812FA74F828" box="[1354,1423,1973,1998]" captionStart="FIGURE 2" captionStartId="6.[151,244,670,691]" captionTargetBox="[150,1436,191,649]" captionTargetId="figure@6.[150,1436,191,649]" captionTargetPageId="6" captionText="FIGURE 2. Sasanidus kermanshahensis, about 45 mm SL, not preserved; Iran: Haram-Abad stream." httpUri="https://zenodo.org/record/271352/files/figure.png" pageId="7" pageNumber="72">Fig. 2</figureCitation>
|
||
).
|
||
<taxonomicName id="169EB4A8B769FFFAFF7CF87EFEF2F814" box="[151,265,2009,2034]" class="Actinopterygii" family="Nemacheilidae" genus="Sasanidus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="72" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B769FFFAFF7CF87EFEF2F814" box="[151,265,2009,2034]" italics="true" pageId="7" pageNumber="72">Sasanidus</emphasis>
|
||
</taxonomicName>
|
||
is also distinguished from
|
||
<taxonomicName id="169EB4A8B769FFFAFDA4F87EFCEAF814" box="[591,785,2009,2034]" class="Actinopterygii" family="Nemacheilidae" genus="Seminemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="72" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B769FFFAFDA4F87EFCEAF814" box="[591,785,2009,2034]" italics="true" pageId="7" pageNumber="72">Seminemacheilus</emphasis>
|
||
</taxonomicName>
|
||
by having one epural bone in the caudal skeleton (vs.
|
||
</paragraph>
|
||
<caption id="85E19FA3B769FFFAFF7CF88CFC18F8A6" box="[151,995,1835,1856]" httpUri="https://zenodo.org/record/271354/files/figure.png" pageId="7" pageNumber="72" targetBox="[151,1435,191,1814]" targetPageId="7">
|
||
<paragraph id="D121CF2BB769FFFAFF7CF88CFC18F8A6" blockId="7.[151,995,1835,1856]" box="[151,995,1835,1856]" pageId="7" pageNumber="72">
|
||
<emphasis id="E3EA1339B769FFFAFF7CF88CFEF7F8A6" bold="true" box="[151,268,1835,1856]" pageId="7" pageNumber="72">FIGURE 4.</emphasis>
|
||
<taxonomicName id="169EB4A8B769FFFAFEF9F88CFDE6F8A6" box="[274,541,1835,1856]" class="Actinopterygii" family="Nemacheilidae" genus="Sasanidus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="72" phylum="Chordata" rank="species" species="kermanshahensis">
|
||
<emphasis id="E3EA1339B769FFFAFEF9F88CFDE6F8A6" box="[274,541,1835,1856]" italics="true" pageId="7" pageNumber="72">Sasanidus kermanshahensis</emphasis>
|
||
</taxonomicName>
|
||
, FSJF 3213, 63 mm SL; Iran: Gamasiab River.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D121CF2BB766FFF5FF7CFF30FB1AFDFE" blockId="8.[151,1437,151,536]" pageId="8" pageNumber="73">
|
||
absent).
|
||
<taxonomicName id="169EB4A8B766FFF5FF13FF30FE92FF56" box="[248,361,151,176]" class="Actinopterygii" family="Nemacheilidae" genus="Sasanidus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="73" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B766FFF5FF13FF30FE92FF56" box="[248,361,151,176]" italics="true" pageId="8" pageNumber="73">Sasanidus</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName id="169EB4A8B766FFF5FD86FF30FD0BFF56" box="[621,752,151,176]" class="Actinopterygii" family="Nemacheilidae" genus="Paracobitis" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="73" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B766FFF5FD86FF30FD0BFF56" box="[621,752,151,176]" italics="true" pageId="8" pageNumber="73">Paracobitis</emphasis>
|
||
</taxonomicName>
|
||
by the absence of a dorsal adipose crest (vs. presence), the absence of a dark brown or black bar or blotch at the caudal-fin base (vs. presence) and the presence of a prominent whitish or yellowish triangular patch on the posteriormost upper and lower caudal peduncle (vs. absence).
|
||
<taxonomicName id="169EB4A8B766FFF5FF7CFEA3FEF2FEFB" box="[151,265,260,285]" class="Actinopterygii" family="Nemacheilidae" genus="Sasanidus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="73" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B766FFF5FF7CFEA3FEF2FEFB" box="[151,265,260,285]" italics="true" pageId="8" pageNumber="73">Sasanidus</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName id="169EB4A8B766FFF5FDEDFEA3FD21FEFB" box="[518,730,260,285]" class="Actinopterygii" family="Nemacheilidae" genus="Turcinoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="73" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B766FFF5FDEDFEA3FD21FEFB" box="[518,730,260,285]" italics="true" pageId="8" pageNumber="73">Turcinoemacheilus</emphasis>
|
||
</taxonomicName>
|
||
by having the pelvic-fin origin clearly behind a vertical of the dorsal-fin origin (vs. pelvic-fin origin in front of a vertical of the dorsal-fin origin) and the anus situated about one eye diameter in front of the anal-fin origin (vs. more than two eye diameters).
|
||
<taxonomicName id="169EB4A8B766FFF5FBC6FEEBFB65FE83" box="[1069,1182,332,357]" class="Actinopterygii" family="Nemacheilidae" genus="Sasanidus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="73" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B766FFF5FBC6FEEBFB65FE83" box="[1069,1182,332,357]" italics="true" pageId="8" pageNumber="73">Sasanidus</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName id="169EB4A8B766FFF5FF7CFEC8FECDFE6E" box="[151,310,367,392]" class="Actinopterygii" family="Nemacheilidae" genus="Paraschistura" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="73" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B766FFF5FF7CFEC8FECDFE6E" box="[151,310,367,392]" italics="true" pageId="8" pageNumber="73">Paraschistura</emphasis>
|
||
</taxonomicName>
|
||
by lacking the black jet spot at the dorsal-fin origin (vs. presence), having a marbled colour pattern with a prominent, irregularly shaped midlateral stripe (vs. lateral bars often dissociated into irregularly shaped blotches or faded on flank in front of the dorsal-fin origin) and a slightly emarginate caudal fin (vs. deeply emarginate or forked), no dark brown or black bar or blotch at caudal-fin base (vs. presence) and prominent whitish or yellowish triangular patch on the posteriormost upper and lower caudal peduncle (vs. absence).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="85E19FA3B766FFF5FF7CFACDFE9BFA7A" httpUri="https://zenodo.org/record/271355/files/figure.png" pageId="8" pageNumber="73" targetBox="[151,1435,587,1364]" targetPageId="8">
|
||
<paragraph id="D121CF2BB766FFF5FF7CFACDFE9BFA7A" blockId="8.[151,1423,1386,1436]" pageId="8" pageNumber="73">
|
||
<emphasis id="E3EA1339B766FFF5FF7CFACDFEF7FA99" bold="true" box="[151,268,1386,1407]" pageId="8" pageNumber="73">FIGURE 5.</emphasis>
|
||
<taxonomicName id="169EB4A8B766FFF5FEF9FACDFE96FA99" box="[274,365,1386,1407]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="73" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B766FFF5FEF9FACDFE96FA99" box="[274,365,1386,1407]" italics="true" pageId="8" pageNumber="73">Schistura</emphasis>
|
||
</taxonomicName>
|
||
sp.,
|
||
<emphasis id="E3EA1339B766FFF5FE71FACDFE5DFA99" bold="true" box="[410,422,1386,1407]" pageId="8" pageNumber="73">a</emphasis>
|
||
, about 70 mm SL, not preserved;
|
||
<emphasis id="E3EA1339B766FFF5FD07FACDFD02FA99" bold="true" box="[748,761,1386,1407]" pageId="8" pageNumber="73">b</emphasis>
|
||
, FSJF 3514, 65 mm SL; India: confluence of Aasan and Giri Rivers. Photos: Nikhil Sood.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="85E19FA3B766FFF5FF7CF8BAFC6DF8D4" box="[151,918,1821,1842]" httpUri="https://zenodo.org/record/271356/files/figure.png" pageId="8" pageNumber="73" targetBox="[151,1435,1484,1798]" targetPageId="8">
|
||
<paragraph id="D121CF2BB766FFF5FF7CF8BAFC6DF8D4" blockId="8.[151,918,1821,1842]" box="[151,918,1821,1842]" pageId="8" pageNumber="73">
|
||
<emphasis id="E3EA1339B766FFF5FF7CF8BAFEF7F8D4" bold="true" box="[151,268,1821,1842]" pageId="8" pageNumber="73">FIGURE 6.</emphasis>
|
||
<taxonomicName id="169EB4A8B766FFF5FEF9F8BAFE33F8D4" box="[274,456,1821,1842]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="73" phylum="Chordata" rank="species" species="rupecula">
|
||
<emphasis id="E3EA1339B766FFF5FEF9F8BAFE33F8D4" box="[274,456,1821,1842]" italics="true" pageId="8" pageNumber="73">Schistura rupecula</emphasis>
|
||
</taxonomicName>
|
||
, SMF 8993, syntype, 71 mm SL; India: Shimla.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="99849CA0B766FFF5FF2CF825FBDDF805" pageId="8" pageNumber="73" type="etymology">
|
||
<paragraph id="D121CF2BB766FFF5FF2CF825FBDDF805" blockId="8.[151,1436,1922,2019]" pageId="8" pageNumber="73">
|
||
<emphasis id="E3EA1339B766FFF5FF2CF825FEB5F87D" bold="true" box="[199,334,1922,1947]" pageId="8" pageNumber="73">Etymology.</emphasis>
|
||
The genus is named for the Sassanid Empire, which was recognized as one of the leading regional powers for a period of more than 400 years. During Late Antiquity, the Sasanian Empire is considered to have been one of Iran's most important and influential historical periods. Gender masculine.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="99849CA0B767FFF7FF2CFF30FDE0FF56" lastPageId="10" lastPageNumber="75" pageId="9" pageNumber="74" type="distribution">
|
||
<paragraph id="D121CF2BB767FFF4FF2CFF30FDA4FF1E" blockId="9.[151,1436,151,248]" pageId="9" pageNumber="74">
|
||
<emphasis id="E3EA1339B767FFF4FF2CFF30FE9BFF56" bold="true" box="[199,352,151,176]" pageId="9" pageNumber="74">Distribution.</emphasis>
|
||
<taxonomicName id="169EB4A8B767FFF4FE8CFF30FD5BFF56" box="[359,672,151,176]" class="Actinopterygii" family="Nemacheilidae" genus="Sasanidus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="74" phylum="Chordata" rank="species" species="kermanshahensis">
|
||
<emphasis id="E3EA1339B767FFF4FE8CFF30FD5BFF56" box="[359,672,151,176]" italics="true" pageId="9" pageNumber="74">Sasanidus kermanshahensis</emphasis>
|
||
</taxonomicName>
|
||
is known from headwaters of the Karkheh and Dez Rivers. The Dez is a tributary to the Karun and both the Karkheh and Karun flowi from the Iranian Zagros Mountains westwards into the lowermost Tigris (
|
||
<bibRefCitation id="B50FB2DAB767FFF4FE2BFF78FDAAFF1E" author="Abdoli" box="[448,593,223,248]" pageId="9" pageNumber="74" refString="Abdoli, A. (2000) The Inland Water Fishes of Iran. Iranian Museum of Nature and Wildlife, Tehran. 378 pp. [in Farsi]" type="book" year="2000">Abdoli 2000</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="85E19FA3B767FFF4FF7CF89BFE89F86A" httpUri="https://zenodo.org/record/271357/files/figure.png" pageId="9" pageNumber="74" targetBox="[151,1436,299,1830]" targetPageId="9">
|
||
<paragraph id="D121CF2BB767FFF4FF7CF89BFE89F86A" blockId="9.[151,1436,1852,1932]" pageId="9" pageNumber="74">
|
||
<emphasis id="E3EA1339B767FFF4FF7CF89BFEF7F8B7" bold="true" box="[151,268,1852,1873]" pageId="9" pageNumber="74">FIGURE 7.</emphasis>
|
||
<taxonomicName id="169EB4A8B767FFF4FEF9F89BFE96F8B7" box="[274,365,1852,1873]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="74" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B767FFF4FEF9F89BFE96F8B7" box="[274,365,1852,1873]" italics="true" pageId="9" pageNumber="74">Schistura</emphasis>
|
||
</taxonomicName>
|
||
spp.: India: Karnataka:
|
||
<emphasis id="E3EA1339B767FFF4FDB8F89BFDA4F8B7" bold="true" box="[595,607,1852,1873]" pageId="9" pageNumber="74">a</emphasis>
|
||
, FSJF 3515, 50 mm SL; stream between Mukkodlu to Surlabhi;
|
||
<emphasis id="E3EA1339B767FFF4FB27F89BFB22F8B7" bold="true" box="[1228,1241,1852,1873]" pageId="9" pageNumber="74">b</emphasis>
|
||
, FSJF 3516, 36 mm SL; stream in Bhagamandala fields;
|
||
<emphasis id="E3EA1339B767FFF4FE13F8FDFDF8F889" bold="true" box="[504,515,1882,1903]" pageId="9" pageNumber="74">c</emphasis>
|
||
, FSJF 3517, 30 mm SL; rafting place at Barapol River;
|
||
<emphasis id="E3EA1339B767FFF4FBCFF8FDFBCAF889" bold="true" box="[1060,1073,1882,1903]" pageId="9" pageNumber="74">d</emphasis>
|
||
, FSJF 3518, 52 mm SL; River above Kumadhara waterfalls.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D121CF2BB764FFF7FF2CFF30FDE0FF56" blockId="10.[151,1437,151,2013]" box="[199,539,151,176]" pageId="10" pageNumber="75">
|
||
<emphasis id="E3EA1339B764FFF7FF2CFF30FE6AFF56" bold="true" box="[199,401,151,176]" pageId="10" pageNumber="75">Included species.</emphasis>
|
||
Monotypic.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="99849CA0B764FFF6FF2CFF1BFA6DFDFE" lastPageId="11" lastPageNumber="76" pageId="10" pageNumber="75" type="discussion">
|
||
<paragraph id="D121CF2BB764FFF7FF2CFF1BFE1AFCDE" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="75">
|
||
<emphasis id="E3EA1339B764FFF7FF2CFF1BFEB7FF33" bold="true" box="[199,332,188,213]" pageId="10" pageNumber="75">Discussion.</emphasis>
|
||
The COI barcode sequences analysed here and the very limited species coverage of the speciesrich family
|
||
<taxonomicName id="169EB4A8B764FFF7FEF0FF78FE38FF1E" box="[283,451,223,248]" class="Actinopterygii" family="Nemacheilidae" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="75" phylum="Chordata" rank="family">Nemacheilidae</taxonomicName>
|
||
do not allow strong phylogenetic conclusions to be made from the dataset presented in this study. Therefore, we do not speculate on the relationships of the different genera of nemacheilid loaches included in this study. The
|
||
<typeStatus id="0E257189B764FFF7FE08FE80FDBEFEA6" box="[483,581,295,320]" pageId="10" pageNumber="75" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="169EB4A8B764FFF7FD9EFE80FC4EFEA6" box="[629,949,295,320]" class="Actinopterygii" family="Nemacheilidae" genus="Sasanidus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="75" phylum="Chordata" rank="species" species="kermanshahensis">
|
||
<emphasis id="E3EA1339B764FFF7FD9EFE80FC4EFEA6" box="[629,949,295,320]" italics="true" pageId="10" pageNumber="75">Sasanidus kermanshahensis</emphasis>
|
||
</taxonomicName>
|
||
was illustrated by
|
||
<bibRefCitation id="B50FB2DAB764FFF7FB74FE8FFEE8FE83" author="Banarescu" pageId="10" pageNumber="75" refString="Banarescu, P. G. & Nalbant. T. T. (1966) The 3 rd Danish Expedition to Central Asia. Zoological results 34. Cobitidae (Pisces) from Afghanistan and Iran. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening i Kobenhavn, 129, 149 - 186." type="journal article" year="1966">Bănărescu & Nalbant (1966:152)</bibRefCitation>
|
||
and the species can be immediately identified by its body shape and colour pattern. The osteology of
|
||
<taxonomicName id="169EB4A8B764FFF7FF7CFEC8FEF2FE6E" box="[151,265,367,392]" class="Actinopterygii" family="Nemacheilidae" genus="Sasanidus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="75" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B764FFF7FF7CFEC8FEF2FE6E" box="[151,265,367,392]" italics="true" pageId="10" pageNumber="75">Sasanidus</emphasis>
|
||
</taxonomicName>
|
||
has recently been described by
|
||
<bibRefCitation id="B50FB2DAB764FFF7FD6BFEC8FC75FE6E" box="[640,910,367,392]" pageId="10" pageNumber="75" refString="Mafakheri, P., Eagderi, S., Farahmand, H. & Mosavii-Sabet, H. (2015) Descriptive Osteology of Oxynoemacheilus kermanshahensis (Banarescu and Nalbant, 1966) (Cypriniformes, Nemacheilidae). Croatian Journal of Fisheries, 73, 115 - 123. http: // dx. doi. org / 10.14798 / 73.3.840" type="journal article">
|
||
Mafakheri
|
||
<emphasis id="E3EA1339B764FFF7FD15FED6FCC1FE6E" box="[766,826,367,392]" italics="true" pageId="10" pageNumber="75">et al.</emphasis>
|
||
(2015)
|
||
</bibRefCitation>
|
||
and osteological characters mentioned below were examined by us based on our own materials.
|
||
<taxonomicName id="169EB4A8B764FFF7FD2CFE33FCC2FE4B" box="[711,825,404,429]" class="Actinopterygii" family="Nemacheilidae" genus="Sasanidus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="75" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B764FFF7FD2CFE33FCC2FE4B" box="[711,825,404,429]" italics="true" pageId="10" pageNumber="75">Sasanidus</emphasis>
|
||
</taxonomicName>
|
||
belong to a group of nemacheilid genera distinguished by the absence of the preethmoid I in the ethmoid region of the skull. Other genera in this group are
|
||
<taxonomicName id="169EB4A8B764FFF7FB06FE10FA6EFE36" box="[1261,1429,439,464]" class="Actinopterygii" family="Nemacheilidae" genus="Acanthocobitis" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="75" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B764FFF7FB06FE10FA6EFE36" box="[1261,1429,439,464]" italics="true" pageId="10" pageNumber="75">Acanthocobitis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="169EB4A8B764FFF7FF7CFE7BFEAFFE13" box="[151,340,476,501]" class="Actinopterygii" family="Nemacheilidae" genus="Afronemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="75" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B764FFF7FF7CFE7BFEAFFE13" box="[151,340,476,501]" italics="true" pageId="10" pageNumber="75">Afronemacheilus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="169EB4A8B764FFF7FE87FE7BFE29FE13" box="[364,466,476,501]" class="Actinopterygii" family="Nemacheilidae" genus="Dzihunia" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="75" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B764FFF7FE87FE7BFE29FE13" box="[364,466,476,501]" italics="true" pageId="10" pageNumber="75">Dzihunia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="169EB4A8B764FFF7FE07FE7BFD72FE13" box="[492,649,476,501]" class="Actinopterygii" family="Nemacheilidae" genus="Indoreonectes" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="75" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B764FFF7FE07FE7BFD72FE13" box="[492,649,476,501]" italics="true" pageId="10" pageNumber="75">Indoreonectes</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="169EB4A8B764FFF7FD48FE7BFCE7FE13" box="[675,796,476,501]" class="Actinopterygii" family="Nemacheilidae" genus="Iskandaria" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="75" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B764FFF7FD48FE7BFCE7FE13" box="[675,796,476,501]" italics="true" pageId="10" pageNumber="75">Iskandaria</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="169EB4A8B764FFF7FCDEFE7BFBF0FE13" box="[821,1035,476,501]" class="Actinopterygii" family="Nemacheilidae" genus="Mesonoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="75" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B764FFF7FCDEFE7BFBF0FE13" box="[821,1035,476,501]" italics="true" pageId="10" pageNumber="75">Mesonoemacheilus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="169EB4A8B764FFF7FBCFFE7BFB4EFE13" box="[1060,1205,476,501]" class="Actinopterygii" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="75" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B764FFF7FBCFFE7BFB4EFE13" box="[1060,1205,476,501]" italics="true" pageId="10" pageNumber="75">Nemacheilus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="169EB4A8B764FFF7FB25FE7BFA6EFE13" box="[1230,1429,476,501]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="75" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B764FFF7FB25FE7BFA6EFE13" box="[1230,1429,476,501]" italics="true" pageId="10" pageNumber="75">Oxynoemacheilus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="169EB4A8B764FFF7FF7CFE58FEE1FDFE" box="[151,282,511,536]" class="Actinopterygii" family="Nemacheilidae" genus="Paracobitis" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="75" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B764FFF7FF7CFE58FEE1FDFE" box="[151,282,511,536]" italics="true" pageId="10" pageNumber="75">Paracobitis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="169EB4A8B764FFF7FEC3FE58FE3CFDFE" box="[296,455,511,536]" class="Actinopterygii" family="Nemacheilidae" genus="Paraschistura" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="75" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B764FFF7FEC3FE58FE3CFDFE" box="[296,455,511,536]" italics="true" pageId="10" pageNumber="75">Paraschistura</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="169EB4A8B764FFF7FE3EFE58FDC4FDFE" box="[469,575,511,536]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="75" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B764FFF7FE3EFE58FDC4FDFE" box="[469,575,511,536]" italics="true" pageId="10" pageNumber="75">Schistura</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="169EB4A8B764FFF7FDA6FE58FCF4FDFE" box="[589,783,511,536]" class="Actinopterygii" family="Nemacheilidae" genus="Seminemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="75" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B764FFF7FDA6FE58FCF4FDFE" box="[589,783,511,536]" italics="true" pageId="10" pageNumber="75">Seminemacheilus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="169EB4A8B764FFF7FCACFE58FB5CFDFE" authority="Prokofiev 2009" authorityName="Prokofiev" authorityYear="2009" box="[839,1191,511,536]" class="Actinopterygii" family="Nemacheilidae" genus="Troglocobitis" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="75" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B764FFF7FCACFE58FC20FDFE" box="[839,987,511,536]" italics="true" pageId="10" pageNumber="75">Troglocobitis</emphasis>
|
||
(
|
||
<bibRefCitation id="B50FB2DAB764FFF7FC01FE58FB64FDFE" author="Prokofiev" box="[1002,1183,511,536]" pageId="10" pageNumber="75" refString="Prokofiev, A. M. (2009) Problems of the classification and phylogeny of Nemacheiline loaches of the group lacking the preethmoid I (Cypriniformes: Balitoridae: Nemacheilinae). Journal of Ichthyology, 49, 874 - 898. http: // dx. doi. org / 10.1134 / S 0032945209100051" type="journal article" year="2009">Prokofiev 2009</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. It remains unclear, if this group of genera forms a monophyletic group of related species.
|
||
<taxonomicName id="169EB4A8B764FFF7FC3BFD83FB76FDDB" box="[976,1165,548,573]" class="Actinopterygii" family="Nemacheilidae" genus="Afronemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="75" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B764FFF7FC3BFD83FB76FDDB" box="[976,1165,548,573]" italics="true" pageId="10" pageNumber="75">Afronemacheilus</emphasis>
|
||
</taxonomicName>
|
||
occurs in East Africa,
|
||
<taxonomicName id="169EB4A8B764FFF7FF7CFDE0FED0FD86" box="[151,299,583,608]" class="Actinopterygii" family="Nemacheilidae" genus="Troglocobitis" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="75" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B764FFF7FF7CFDE0FED0FD86" box="[151,299,583,608]" italics="true" pageId="10" pageNumber="75">Troglocobitis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="169EB4A8B764FFF7FED2FDE0FE5BFD86" box="[313,416,583,608]" class="Actinopterygii" family="Nemacheilidae" genus="Dzihunia" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="75" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B764FFF7FED2FDE0FE5BFD86" box="[313,416,583,608]" italics="true" pageId="10" pageNumber="75">Dzihunia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="169EB4A8B764FFF7FE3CFDE0FDABFD86" box="[471,592,583,608]" class="Actinopterygii" family="Nemacheilidae" genus="Iskandaria" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="75" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B764FFF7FE3CFDE0FDABFD86" box="[471,592,583,608]" italics="true" pageId="10" pageNumber="75">Iskandaria</emphasis>
|
||
</taxonomicName>
|
||
are found in the Aral Sea basin in Central Asia.
|
||
<taxonomicName id="169EB4A8B764FFF7FB9FFDE0FB1DFD86" box="[1140,1254,583,608]" class="Actinopterygii" family="Nemacheilidae" genus="Sasanidus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="75" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B764FFF7FB9FFDE0FB1DFD86" box="[1140,1254,583,608]" italics="true" pageId="10" pageNumber="75">Sasanidus</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName id="169EB4A8B764FFF7FF3EFDCBFE69FD63" box="[213,402,620,645]" class="Actinopterygii" family="Nemacheilidae" genus="Afronemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="75" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B764FFF7FF3EFDCBFE69FD63" box="[213,402,620,645]" italics="true" pageId="10" pageNumber="75">Afronemacheilus</emphasis>
|
||
</taxonomicName>
|
||
by having a single epural bone in the caudal skeleton (vs. absent). It is distinguished from
|
||
<taxonomicName id="169EB4A8B764FFF7FF7CFD28FED7FD4E" box="[151,300,655,680]" class="Actinopterygii" family="Nemacheilidae" genus="Troglocobitis" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="75" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B764FFF7FF7CFD28FED7FD4E" box="[151,300,655,680]" italics="true" pageId="10" pageNumber="75">Troglocobitis</emphasis>
|
||
</taxonomicName>
|
||
by having a bony swim-bladder capsule (vs. absent) and from
|
||
<taxonomicName id="169EB4A8B764FFF7FBDBFD28FB6CFD4E" box="[1072,1175,655,680]" class="Actinopterygii" family="Nemacheilidae" genus="Dzihunia" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="75" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B764FFF7FBDBFD28FB6CFD4E" box="[1072,1175,655,680]" italics="true" pageId="10" pageNumber="75">Dzihunia</emphasis>
|
||
</taxonomicName>
|
||
by having simple lip structures (vs. lower lip with pillow-like elements interrupted by deep folds) and a smooth skin (vs. with small, spine-like epidermal tubercles).
|
||
<taxonomicName id="169EB4A8B764FFF7FDEEFD70FD8CFD16" box="[517,631,727,752]" class="Actinopterygii" family="Nemacheilidae" genus="Sasanidus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="75" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B764FFF7FDEEFD70FD8CFD16" box="[517,631,727,752]" italics="true" pageId="10" pageNumber="75">Sasanidus</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName id="169EB4A8B764FFF7FC92FD70FC09FD16" box="[889,1010,727,752]" class="Actinopterygii" family="Nemacheilidae" genus="Iskandaria" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="75" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B764FFF7FC92FD70FC09FD16" box="[889,1010,727,752]" italics="true" pageId="10" pageNumber="75">Iskandaria</emphasis>
|
||
</taxonomicName>
|
||
by having the body fully covered by scales (vs. no scales), a deep body and caudal peduncle (vs. very slender and shallow) and no muscular, swollen cheeks in males (vs. present).
|
||
</paragraph>
|
||
<paragraph id="D121CF2BB764FFF7FF2CFCE3FDB8FBD2" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="75">
|
||
There are few studies about the genera of nemacheilid loaches of the Middle East (
|
||
<bibRefCitation id="B50FB2DAB764FFF7FB88FCE3FA6CFCBB" author="Banarescu" box="[1123,1431,836,861]" pageId="10" pageNumber="75" refString="Banarescu, P. G. & Nalbant, T. T. (1995) A generical classification of Nemacheilinae with description of two new genera (Teleostei: Cypriniformes: Cobitidae). Travaux du Museum d'Histoire Naturelle " Gr. Antipa, 35, 429 - 495." type="journal article" year="1995">Bănărescu & Nalbant 1995</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="B50FB2DAB764FFF7FF7CFCC0FEC8FC66" author="Kottelat" box="[151,307,871,896]" pageId="10" pageNumber="75" refString="Kottelat, M. (1990) Indochinese nemacheilines. A revision of nemacheiline loaches (Pisces: Cypriniformes) of Thailand, Burma, Laos, Cambodia and southern Viet Nam. Verlag Dr. Friedrich Pfeil, Munchen, 262 pp." type="book" year="1990">Kottelat 1990</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="B50FB2DAB764FFF7FED4FCC0FE15FC66" author="Prokofiev" box="[319,494,871,896]" pageId="10" pageNumber="75" refString="Prokofiev, A. M. (2009) Problems of the classification and phylogeny of Nemacheiline loaches of the group lacking the preethmoid I (Cypriniformes: Balitoridae: Nemacheilinae). Journal of Ichthyology, 49, 874 - 898. http: // dx. doi. org / 10.1134 / S 0032945209100051" type="journal article" year="2009">Prokofiev 2009</bibRefCitation>
|
||
) and there are also few studies presenting molecular data on nemacheilid loaches of that area (Esmaeili
|
||
<emphasis id="E3EA1339B764FFF7FE9CFC2AFE49FC43" box="[375,434,908,933]" italics="true" pageId="10" pageNumber="75">et al.</emphasis>
|
||
2014; Freyhof
|
||
<emphasis id="E3EA1339B764FFF7FD8CFC2AFD61FC43" box="[615,666,908,933]" italics="true" pageId="10" pageNumber="75">et al</emphasis>
|
||
. 2014;
|
||
<bibRefCitation id="B50FB2DAB764FFF7FD1FFC2BFC23FC43" author="Freyhof" box="[756,984,908,933]" pageId="10" pageNumber="75" refString="Freyhof, J., Sayyadzadeh, G., Esmaeili, H. R. & Geiger, M. (2015) Review of the genus Paraschistura from Iran with description of six new species (Teleostei: Nemacheilidae). Ichthyological Exploration of Freshwaters, 26, 1 - 48." type="journal article" year="2015">
|
||
Freyhof
|
||
<emphasis id="E3EA1339B764FFF7FCBDFC2AFC71FC43" box="[854,906,908,933]" italics="true" pageId="10" pageNumber="75">et al</emphasis>
|
||
. 2015
|
||
</bibRefCitation>
|
||
). There is no comprehensive molecular study on the nemacheilid loaches of
|
||
<collectingCountry id="A9898FBBB764FFF7FDABFC08FD81FC2E" box="[576,634,943,968]" name="India" pageId="10" pageNumber="75">India</collectingCountry>
|
||
or South-East Asia. Here, we bring all known genera of nemacheilid loaches from the Middle East together in one molecular dataset for the first time. Furthermore, we were able to add COI sequence data from representatives of all genera of nemacheilid loaches recognised as valid from western
|
||
<collectingCountry id="A9898FBBB764FFF7FF7CFBBBFF2AFBD3" box="[151,209,1052,1077]" name="India" pageId="10" pageNumber="75">India</collectingCountry>
|
||
and
|
||
<collectingCountry id="A9898FBBB764FFF7FEECFBBBFE9DFBD3" box="[263,358,1052,1077]" name="Pakistan" pageId="10" pageNumber="75">Pakistan</collectingCountry>
|
||
by
|
||
<bibRefCitation id="B50FB2DAB764FFF7FE7BFBBBFDC4FBD2" author="Kottelat" box="[400,575,1052,1077]" pageId="10" pageNumber="75" refString="Kottelat, M. (2012) Conspectus cobitidum: an inventory of the loaches of the world (Teleostei: Cypriniformes: Cobitoidei). The Raffles Bulletin of Zoology, 26 (Supplement), 1 - 199." type="journal article" year="2012">Kottelat (2012)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="D121CF2BB764FFF7FF2CFBE7FD0CF9A9" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="75">
|
||
However, a phylogenetic study cannot be based solely on the molecular method applied here, because the resolution of this single mitochondrial marker is insufficient to reliably resolve the relationships between the different genera studied. DNA barcode data do not offer a phylogenetic signal sufficiently strong to resolve evolutionary relationships at higher taxonomic levels (
|
||
<bibRefCitation id="B50FB2DAB764FFF7FD12FB0BFC0CFB23" author="Hajibabaei" box="[761,1015,1196,1221]" pageId="10" pageNumber="75" refString="Hajibabaei, M., Singer, G. A. & Hickey, D. A. (2006) Benchmarking DNA barcodes: an assessment using available primate sequences. Genome, 49 (7), 851 - 854. http: // dx. doi. org / 10.1139 / G 06 - 025" type="journal article" year="2006">
|
||
Hajibabaei
|
||
<emphasis id="E3EA1339B764FFF7FC90FB0AFC4FFB23" box="[891,948,1196,1221]" italics="true" pageId="10" pageNumber="75">et al.</emphasis>
|
||
2006
|
||
</bibRefCitation>
|
||
), which is clearly reflected in the low bootstrap support values of the inner nodes (
|
||
<figureCitation id="49A5D3AEB764FFF7FD5EFB68FD07FB0E" box="[693,764,1231,1256]" captionStart="FIGURE 1" captionStartId="4.[151,244,1839,1860]" captionTargetBox="[408,1181,206,1802]" captionTargetId="figure@4.[392,1191,191,1811]" captionTargetPageId="4" captionText="FIGURE 1. Maximum Likelihood estimation of the relationships based on the mitochondrial COI barcode region (Tamura 3 - parameter model, discrete Gamma distribution for rate differences with 5 categories + G parameter = 0.4706 and invariable site proportion + I = 58.22 %). Nucleotide positions with less than 100 % site coverage were eliminated resulting in 562 analysed positions. The tree is drawn to scale, with branch lengths measured in number of substitutions per site. Numbers of nodes indicate bootstrap values from 1000 pseudoreplicates and low values have deliberately been included to demonstrate the lack of evolutionary signal to resolve the relationships between supra-specific taxa." httpUri="https://zenodo.org/record/271351/files/figure.png" pageId="10" pageNumber="75">Fig. 1</figureCitation>
|
||
). Although analysed by phylogenetic-tree reconstruction methods, barcode-based trees should therefore not be interpreted as phylogenetic trees (
|
||
<bibRefCitation id="B50FB2DAB764FFF7FB61FB53FA6BFAEA" author="Hajibabaei" box="[1162,1424,1268,1293]" pageId="10" pageNumber="75" refString="Hajibabaei, M., Singer, G. A., Hebert, P. D. & Hickey, D. A. (2007) DNA barcoding: how it complements taxonomy, molecular phylogenetics and population genetics. TRENDS in Genetics, 23 (4), 167 - 172. http: // dx. doi. org / 10.1016 / j. tig. 2007.02.001" type="journal article" year="2007">
|
||
Hajibabaei
|
||
<emphasis id="E3EA1339B764FFF7FAE4FB52FAB1FAEB" box="[1295,1354,1268,1293]" italics="true" pageId="10" pageNumber="75">et al.</emphasis>
|
||
2007
|
||
</bibRefCitation>
|
||
). The method is well-suited to assign specimens to species, supported by high bootstrap values of nodes towards the tips, and mark conspicuous populations or taxa. In our study, a ML tree helped to identify
|
||
<taxonomicName id="169EB4A8B764FFF7FB73FA9BFAF1FAB3" box="[1176,1290,1340,1365]" class="Actinopterygii" family="Nemacheilidae" genus="Sasanidus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="75" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B764FFF7FB73FA9BFAF1FAB3" box="[1176,1290,1340,1365]" italics="true" pageId="10" pageNumber="75">Sasanidus</emphasis>
|
||
</taxonomicName>
|
||
as candidate taxon for a more comprehensive investigation, and aided with taxonomic sampling, putting the mitochondrial lineage of these fishes into a broader phylogenetic and biogeographic context. Taking all concerns into consideration, our data does not even place
|
||
<taxonomicName id="169EB4A8B764FFF7FD6BFA00FD09FA26" box="[640,754,1447,1472]" class="Actinopterygii" family="Nemacheilidae" genus="Sasanidus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="75" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B764FFF7FD6BFA00FD09FA26" box="[640,754,1447,1472]" italics="true" pageId="10" pageNumber="75">Sasanidus</emphasis>
|
||
</taxonomicName>
|
||
close to one of the genera described from the Middle East or
|
||
<collectingCountry id="A9898FBBB764FFF7FF7CFA6BFF28FA03" box="[151,211,1484,1509]" name="India" pageId="10" pageNumber="75">India</collectingCountry>
|
||
, supporting the hypothesis that it represents a distinct genus of its own. However, we do not exclude that other species of
|
||
<taxonomicName id="169EB4A8B764FFF7FEA4FA48FE3AF9EE" box="[335,449,1519,1544]" class="Actinopterygii" family="Nemacheilidae" genus="Sasanidus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="75" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B764FFF7FEA4FA48FE3AF9EE" box="[335,449,1519,1544]" italics="true" pageId="10" pageNumber="75">Sasanidus</emphasis>
|
||
</taxonomicName>
|
||
might occur in
|
||
<collectingCountry id="A9898FBBB764FFF7FD98FA48FD29F9EE" box="[627,722,1519,1544]" name="Pakistan" pageId="10" pageNumber="75">Pakistan</collectingCountry>
|
||
or
|
||
<collectingCountry id="A9898FBBB764FFF7FD13FA48FCCFF9EE" box="[760,820,1519,1544]" name="India" pageId="10" pageNumber="75">India</collectingCountry>
|
||
, where they might currently be identified as species of
|
||
<taxonomicName id="169EB4A8B764FFF7FF7CF9B3FEFAF9CB" box="[151,257,1556,1581]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="75" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B764FFF7FF7CF9B3FEFAF9CB" box="[151,257,1556,1581]" italics="true" pageId="10" pageNumber="75">Schistura</emphasis>
|
||
</taxonomicName>
|
||
. Nevertheless, none of the species described by
|
||
<bibRefCitation id="B50FB2DAB764FFF7FCCDF9B3FC02F9CA" box="[806,1017,1556,1581]" pageId="10" pageNumber="75" refString="Mirza, M. R., Nalbant, T. T. & Banarescu, P. M. (1981) A review of the genus Schistura in Pakistan with description of new species and subspecies (Pisces, Cobitidae, Noemacheilinae). Bijdragen tot de Dierkunde, 51, 105 - 130." type="journal article">Mirza et al. (1981)</bibRefCitation>
|
||
from
|
||
<collectingCountry id="A9898FBBB764FFF7FBABF9B3FB64F9CB" box="[1088,1183,1556,1581]" name="Pakistan" pageId="10" pageNumber="75">Pakistan</collectingCountry>
|
||
and by
|
||
<bibRefCitation id="B50FB2DAB764FFF7FB11F9B2FA67F9CA" author="Menon" box="[1274,1436,1556,1580]" pageId="10" pageNumber="75" refString="Menon, A. G. K. (1987) The fauna of India and the adjacent countries. Pisces. Vol. IV. Teleostei - Cobitoidea. Part 1. Homalopteridae. Zoological Survey of India, 1987, i - x + 1 - 259, pls. 1 - 16." type="journal article" year="1987">Menon (1987)</bibRefCitation>
|
||
from
|
||
<collectingCountry id="A9898FBBB764FFF7FF3FF990FEF5F9B6" box="[212,270,1591,1616]" name="India" pageId="10" pageNumber="75">India</collectingCountry>
|
||
are even superficially similar to
|
||
<taxonomicName id="169EB4A8B764FFF7FD95F990FD14F9B6" box="[638,751,1591,1616]" class="Actinopterygii" family="Nemacheilidae" genus="Sasanidus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="75" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B764FFF7FD95F990FD14F9B6" box="[638,751,1591,1616]" italics="true" pageId="10" pageNumber="75">Sasanidus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="D121CF2BB764FFF6FF2CF9FAFBFAFF1E" blockId="10.[151,1437,151,2013]" lastBlockId="11.[151,1436,151,536]" lastPageId="11" lastPageNumber="76" pageId="10" pageNumber="75">
|
||
It is well known that
|
||
<taxonomicName id="169EB4A8B764FFF7FE50F9FBFDDEF993" box="[443,549,1628,1653]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="75" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B764FFF7FE50F9FBFDDEF993" box="[443,549,1628,1653]" italics="true" pageId="10" pageNumber="75">Schistura</emphasis>
|
||
</taxonomicName>
|
||
is a polyphyletic assemblage of species (
|
||
<bibRefCitation id="B50FB2DAB764FFF7FC10F9FBFACEF993" author="Banarescu" box="[1019,1333,1628,1653]" pageId="10" pageNumber="75" refString="Banarescu, P. G. & Nalbant, T. T. (1995) A generical classification of Nemacheilinae with description of two new genera (Teleostei: Cypriniformes: Cobitidae). Travaux du Museum d'Histoire Naturelle " Gr. Antipa, 35, 429 - 495." type="journal article" year="1995">Bănărescu & Nalbant 1995</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="B50FB2DAB764FFF7FAA9F9FBFF28F97E" author="Kottelat" pageId="10" pageNumber="75" refString="Kottelat, M. (1990) Indochinese nemacheilines. A revision of nemacheiline loaches (Pisces: Cypriniformes) of Thailand, Burma, Laos, Cambodia and southern Viet Nam. Verlag Dr. Friedrich Pfeil, Munchen, 262 pp." type="book" year="1990">Kottelat 1990</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="B50FB2DAB764FFF7FF0AF9D8FE6EF97E" author="Prokofiev" box="[225,405,1663,1688]" pageId="10" pageNumber="75" refString="Prokofiev, A. M. (2009) Problems of the classification and phylogeny of Nemacheiline loaches of the group lacking the preethmoid I (Cypriniformes: Balitoridae: Nemacheilinae). Journal of Ichthyology, 49, 874 - 898. http: // dx. doi. org / 10.1134 / S 0032945209100051" type="journal article" year="2009">Prokofiev 2009</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="B50FB2DAB764FFF7FE4EF9D8FDBCF97E" author="Kottelat" box="[421,583,1663,1688]" pageId="10" pageNumber="75" refString="Kottelat, M. (2012) Conspectus cobitidum: an inventory of the loaches of the world (Teleostei: Cypriniformes: Cobitoidei). The Raffles Bulletin of Zoology, 26 (Supplement), 1 - 199." type="journal article" year="2012">Kottelat 2012</bibRefCitation>
|
||
). To exclude the possibility that
|
||
<taxonomicName id="169EB4A8B764FFF7FC2CF9D8FBC2F97E" box="[967,1081,1663,1688]" class="Actinopterygii" family="Nemacheilidae" genus="Sasanidus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="75" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B764FFF7FC2CF9D8FBC2F97E" box="[967,1081,1663,1688]" italics="true" pageId="10" pageNumber="75">Sasanidus</emphasis>
|
||
</taxonomicName>
|
||
is a member of
|
||
<taxonomicName id="169EB4A8B764FFF7FAEBF9D8FA91F97E" box="[1280,1386,1663,1688]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="75" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B764FFF7FAEBF9D8FA91F97E" box="[1280,1386,1663,1688]" italics="true" pageId="10" pageNumber="75">Schistura</emphasis>
|
||
</taxonomicName>
|
||
, we examined two
|
||
<typeStatus id="0E257189B764FFF7FED4F902FE5AF95B" box="[319,417,1701,1725]" pageId="10" pageNumber="75" type="syntype">syntypes</typeStatus>
|
||
of
|
||
<taxonomicName id="169EB4A8B764FFF7FE21F902FDB7F95B" box="[458,588,1700,1725]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="75" phylum="Chordata" rank="species" species="rupecula">
|
||
<emphasis id="E3EA1339B764FFF7FE21F902FDB7F95B" box="[458,588,1700,1725]" italics="true" pageId="10" pageNumber="75">S. rupecula</emphasis>
|
||
</taxonomicName>
|
||
, the
|
||
<typeStatus id="0E257189B764FFF7FD6DF902FD4DF95B" box="[646,694,1701,1725]" pageId="10" pageNumber="75">type</typeStatus>
|
||
species of
|
||
<taxonomicName id="169EB4A8B764FFF7FCD2F903FC58F95B" box="[825,931,1700,1725]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="75" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B764FFF7FCD2F903FC58F95B" box="[825,931,1700,1725]" italics="true" pageId="10" pageNumber="75">Schistura</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="169EB4A8B764FFF7FC58F903FB72F95B" box="[947,1161,1700,1725]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="75" phylum="Chordata" rank="species" species="rupecula">
|
||
<emphasis id="E3EA1339B764FFF7FC58F903FB72F95B" box="[947,1161,1700,1725]" italics="true" pageId="10" pageNumber="75">Schistura rupecula</emphasis>
|
||
</taxonomicName>
|
||
was described from the city of Shimla in Himachal Pradesh,
|
||
<collectingCountry id="A9898FBBB764FFF7FDD0F960FD8FF906" box="[571,628,1735,1760]" name="India" pageId="10" pageNumber="75">India</collectingCountry>
|
||
(
|
||
<bibRefCitation id="B50FB2DAB764FFF7FD6FF960FCB5F906" author="McClelland" box="[644,846,1735,1760]" pageId="10" pageNumber="75" refString="McClelland, J. (1838) Observations on six new species of Cyprinidae, with an outline of a new classification of the family. Journal of the Asiatic Society of Bengal, 7, 941 - 948, pls. 55 - 56." type="journal article" year="1838">McClelland 1838</bibRefCitation>
|
||
). The stream in Shimla is highly polluted today and no loaches could be found (Nikhil Sood, Bangalore; Rajeev Raghavan,
|
||
<collectingCountry id="A9898FBBB764FFF7FC55F94BFBE9F8E3" box="[958,1042,1772,1797]" name="India" pageId="10" pageNumber="75">Cochin</collectingCountry>
|
||
, pers. comm.). The stream flowing through Shimla is a headwater stream of the Aasan River, which flows to the Giri River, itself being a tributary to the Ganges. We include here one DNA sequence of a loach from the lowermost Aasan River from our own material, just at the confluence with the Giri River. We compared the series (FSJF 3514;
|
||
<figureCitation id="49A5D3AEB764FFF7FB47F8F0FB09F896" box="[1196,1266,1879,1904]" captionStart="FIGURE 4" captionStartId="7.[151,244,1835,1856]" captionTargetBox="[151,1435,191,1814]" captionTargetId="figure@7.[151,1435,191,1814]" captionTargetPageId="7" captionText="FIGURE 4. Sasanidus kermanshahensis, FSJF 3213, 63 mm SL; Iran: Gamasiab River." httpUri="https://zenodo.org/record/271354/files/figure.png" pageId="10" pageNumber="75">Fig. 4</figureCitation>
|
||
) to which this individual belongs, with the description and figure by
|
||
<bibRefCitation id="B50FB2DAB764FFF7FD14F8DBFBF1F872" author="McClelland" box="[767,1034,1916,1941]" pageId="10" pageNumber="75" refString="McClelland, J. (1838) Observations on six new species of Cyprinidae, with an outline of a new classification of the family. Journal of the Asiatic Society of Bengal, 7, 941 - 948, pls. 55 - 56." type="journal article" year="1838">McClelland (1838:948)</bibRefCitation>
|
||
, two
|
||
<typeStatus id="0E257189B764FFF7FBA2F8DAFB50F873" box="[1097,1195,1917,1941]" pageId="10" pageNumber="75" type="syntype">syntypes</typeStatus>
|
||
of
|
||
<taxonomicName id="169EB4A8B764FFF7FB39F8DAFAA8F873" box="[1234,1363,1916,1941]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="75" phylum="Chordata" rank="species" species="rupecula">
|
||
<emphasis id="E3EA1339B764FFF7FB39F8DAFAA8F873" box="[1234,1363,1916,1941]" italics="true" pageId="10" pageNumber="75">S. rupecula</emphasis>
|
||
</taxonomicName>
|
||
(SMF 8993;
|
||
<figureCitation id="49A5D3AEB764FFF7FF0BF807FEDDF85F" box="[224,294,1952,1977]" captionStart="FIGURE 6" captionStartId="8.[151,244,1821,1842]" captionTargetBox="[151,1435,1484,1798]" captionTargetId="figure@8.[151,1436,1484,1799]" captionTargetPageId="8" captionText="FIGURE 6. Schistura rupecula, SMF 8993, syntype, 71 mm SL; India: Shimla." httpUri="https://zenodo.org/record/271356/files/figure.png" pageId="10" pageNumber="75">Fig. 6</figureCitation>
|
||
) and an illustration of a possible
|
||
<typeStatus id="0E257189B764FFF7FD44F806FCFDF85F" box="[687,774,1953,1977]" pageId="10" pageNumber="75" type="syntype">syntype</typeStatus>
|
||
of
|
||
<taxonomicName id="169EB4A8B764FFF7FCDAF806FC49F85E" box="[817,946,1953,1976]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="75" phylum="Chordata" rank="species" species="montana">
|
||
<emphasis id="E3EA1339B764FFF7FCDAF806FC49F85E" box="[817,946,1953,1976]" italics="true" pageId="10" pageNumber="75">S. montana</emphasis>
|
||
</taxonomicName>
|
||
(BMNH 1860.3.19.118), a second species described from Shimla by
|
||
<bibRefCitation id="B50FB2DAB764FFF7FE55F863FD33F83A" author="McClelland" box="[446,712,1988,2013]" pageId="10" pageNumber="75" refString="McClelland, J. (1838) Observations on six new species of Cyprinidae, with an outline of a new classification of the family. Journal of the Asiatic Society of Bengal, 7, 941 - 948, pls. 55 - 56." type="journal article" year="1838">McClelland (1838:948)</bibRefCitation>
|
||
. All these loaches have 5½ branched anal-fin rays, 8+8 branched caudal-fin rays, bars on the flank and a moderately deep emarginate caudal fin. Despite being geographically very close to the
|
||
<typeStatus id="0E257189B765FFF6FECDFF1AFEADFF33" box="[294,342,189,213]" pageId="11" pageNumber="76">type</typeStatus>
|
||
locality of
|
||
<taxonomicName id="169EB4A8B765FFF6FE0BFF1AFD98FF33" box="[480,611,188,213]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="76" phylum="Chordata" rank="species" species="rupecula">
|
||
<emphasis id="E3EA1339B765FFF6FE0BFF1AFD98FF33" box="[480,611,188,213]" italics="true" pageId="11" pageNumber="76">S. rupecula</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="169EB4A8B765FFF6FD75FF1AFCE4FF32" box="[670,799,189,212]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="76" phylum="Chordata" rank="species" species="montana">
|
||
<emphasis id="E3EA1339B765FFF6FD75FF1AFCE4FF32" box="[670,799,189,212]" italics="true" pageId="11" pageNumber="76">S. montana</emphasis>
|
||
</taxonomicName>
|
||
, the fishes from Aasan River could not definitively be identified as one of these two species and are referred to as
|
||
<taxonomicName id="169EB4A8B765FFF6FCD9FF78FC60FF1E" box="[818,923,223,248]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="76" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B765FFF6FCD9FF78FC60FF1E" box="[818,923,223,248]" italics="true" pageId="11" pageNumber="76">Schistura</emphasis>
|
||
</taxonomicName>
|
||
sp. here.
|
||
</paragraph>
|
||
<paragraph id="D121CF2BB765FFF6FF2CFEA2FA6DFDFE" blockId="11.[151,1436,151,536]" pageId="11" pageNumber="76">
|
||
In addition to one DNA sequence of
|
||
<taxonomicName id="169EB4A8B765FFF6FD8AFEA3FD30FEFB" box="[609,715,260,285]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="76" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B765FFF6FD8AFEA3FD30FEFB" box="[609,715,260,285]" italics="true" pageId="11" pageNumber="76">Schistura</emphasis>
|
||
</taxonomicName>
|
||
sp. from the Aasan River, we included the COI sequence data of three
|
||
<taxonomicName id="169EB4A8B765FFF6FF3EFE80FEC4FEA6" box="[213,319,295,320]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="76" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B765FFF6FF3EFE80FEC4FEA6" box="[213,319,295,320]" italics="true" pageId="11" pageNumber="76">Schistura</emphasis>
|
||
</taxonomicName>
|
||
spp. From the same area and 26 sequences of fishes identified as belonging to the genus ‘
|
||
<taxonomicName id="169EB4A8B765FFF6FAC2FE80FA68FEA6" box="[1321,1427,295,320]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="76" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B765FFF6FAC2FE80FA68FEA6" box="[1321,1427,295,320]" italics="true" pageId="11" pageNumber="76">Schistura</emphasis>
|
||
</taxonomicName>
|
||
’ from NCBI GenBank (
|
||
<tableCitation id="9C1CFA90B765FFF6FE70FEEBFE0BFE82" box="[411,496,332,357]" captionStart="TABLE 1" captionStartId="2.[151,231,261,282]" captionText="TABLE 1. List of COI sequences of nemacheilid loaches downloaded from NCBI GenBank." pageId="11" pageNumber="76">Table 1</tableCitation>
|
||
). We sequenced four additional ‘
|
||
<taxonomicName id="169EB4A8B765FFF6FC87FEEBFC2DFE83" box="[876,982,332,357]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="76" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B765FFF6FC87FEEBFC2DFE83" box="[876,982,332,357]" italics="true" pageId="11" pageNumber="76">Schistura</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<taxonomicName id="169EB4A8B765FFF6FC03FEEAFB96FE82" box="[1000,1133,333,356]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="76" phylum="Chordata" rank="species" species="semiarmata">
|
||
<emphasis id="E3EA1339B765FFF6FC03FEEAFB96FE82" box="[1000,1133,333,356]" italics="true" pageId="11" pageNumber="76">semiarmata</emphasis>
|
||
</taxonomicName>
|
||
and 11 individuals of four unidentified species of ‘
|
||
<taxonomicName id="169EB4A8B765FFF6FE4CFEC8FDEAFE6E" box="[423,529,367,392]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="76" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B765FFF6FE4CFEC8FDEAFE6E" box="[423,529,367,392]" italics="true" pageId="11" pageNumber="76">Schistura</emphasis>
|
||
</taxonomicName>
|
||
’ from the Western Ghats that form a separate evolutionary lineage (
|
||
<figureCitation id="49A5D3AEB765FFF6FAFEFEC8FAA1FE6E" box="[1301,1370,367,392]" captionStart="FIGURE 7" captionStartId="9.[151,244,1852,1873]" captionTargetBox="[151,1436,299,1830]" captionTargetId="figure@9.[151,1436,299,1831]" captionTargetPageId="9" captionText="FIGURE 7. Schistura spp.: India: Karnataka: a, FSJF 3515, 50 mm SL; stream between Mukkodlu to Surlabhi; b, FSJF 3516, 36 mm SL; stream in Bhagamandala fields; c, FSJF 3517, 30 mm SL; rafting place at Barapol River; d, FSJF 3518, 52 mm SL; River above Kumadhara waterfalls." httpUri="https://zenodo.org/record/271357/files/figure.png" pageId="11" pageNumber="76">Fig. 7</figureCitation>
|
||
). The various loaches tentatively placed in the genus
|
||
<taxonomicName id="169EB4A8B765FFF6FD4EFE33FCF4FE4B" box="[677,783,404,429]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="76" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B765FFF6FD4EFE33FCF4FE4B" box="[677,783,404,429]" italics="true" pageId="11" pageNumber="76">Schistura</emphasis>
|
||
</taxonomicName>
|
||
and analysed in this study are not closely related and their clustering implies that several undescribed genera might be involved. Importantly, none of the mitochondrial lineages designated as ‘
|
||
<taxonomicName id="169EB4A8B765FFF6FE4EFE7BFDF4FE13" box="[421,527,476,501]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="76" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B765FFF6FE4EFE7BFDF4FE13" box="[421,527,476,501]" italics="true" pageId="11" pageNumber="76">Schistura</emphasis>
|
||
</taxonomicName>
|
||
’ in
|
||
<figureCitation id="49A5D3AEB765FFF6FDABFE7BFD5BFE12" box="[576,672,476,501]" captionStart="FIGURE 7" captionStartId="9.[151,244,1852,1873]" captionTargetBox="[151,1436,299,1830]" captionTargetId="figure@9.[151,1436,299,1831]" captionTargetPageId="9" captionText="FIGURE 7. Schistura spp.: India: Karnataka: a, FSJF 3515, 50 mm SL; stream between Mukkodlu to Surlabhi; b, FSJF 3516, 36 mm SL; stream in Bhagamandala fields; c, FSJF 3517, 30 mm SL; rafting place at Barapol River; d, FSJF 3518, 52 mm SL; River above Kumadhara waterfalls." httpUri="https://zenodo.org/record/271357/files/figure.png" pageId="11" pageNumber="76">Figure 7</figureCitation>
|
||
are closely related to
|
||
<taxonomicName id="169EB4A8B765FFF6FC4BFE7BFBE9FE13" box="[928,1042,476,501]" class="Actinopterygii" family="Nemacheilidae" genus="Sasanidus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="76" phylum="Chordata" rank="genus">
|
||
<emphasis id="E3EA1339B765FFF6FC4BFE7BFBE9FE13" box="[928,1042,476,501]" italics="true" pageId="11" pageNumber="76">Sasanidus</emphasis>
|
||
</taxonomicName>
|
||
. Future studies on the diversity of loaches in
|
||
<collectingCountry id="A9898FBBB765FFF6FEE6FE58FE97FDFE" box="[269,364,511,536]" name="Pakistan" pageId="11" pageNumber="76">Pakistan</collectingCountry>
|
||
and
|
||
<collectingCountry id="A9898FBBB765FFF6FE49FE58FE27FDFE" box="[418,476,511,536]" name="India" pageId="11" pageNumber="76">India</collectingCountry>
|
||
are encouraged to resolve their generic structure and the biogeographic relationships.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |