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<document id="5C733C045BE79C26BECBBBCA153188CD" ID-DOI="10.25674/415" ID-ISSN="2509-9523" ID-Zenodo-Dep="13194032" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722631177589" checkinUser="felipe" docAuthor="Seifert, Bernhard" docDate="2024" docId="577887E6FF87FFDD6B325E59FEBE309D" docLanguage="en" docName="SoilOrg.96.2.113-144.pdf" docOrigin="Soil Organisms (London, England) 96 (2)" docSource="http://dx.doi.org/10.25674/415" docStyle="DocumentStyle:EA2ECAC87EC21C07AB805D5B9B4601F2.2:SoilOrg.2018-.journal_article" docStyleId="EA2ECAC87EC21C07AB805D5B9B4601F2" docStyleName="SoilOrg.2018-.journal_article" docStyleVersion="2" docTitle="Cardiocondyla yemeni Collingwood &amp; Agosti 1996" docType="treatment" docVersion="2" lastPageNumber="136" masterDocId="AB41FF9EFF91FFCA681B5A5FFFA8347E" masterDocTitle="A taxonomic revision of the ants of the Cardiocondyla wroughtonii group (Hymenoptera: Formicidae) with a checklist of the Cardiocondyla species of the world" masterLastPageNumber="144" masterPageNumber="113" pageNumber="135" updateTime="1722679537643" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="9A3E12F985292A20011FDE8FBDB43DC7">A taxonomic revision of the ants of the Cardiocondyla wroughtonii group (Hymenoptera: Formicidae) with a checklist of the Cardiocondyla species of the world</mods:title>
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<subSubSection id="97CB657BFF87FFDC6B325E59FBAD3041" pageId="22" pageNumber="135" type="nomenclature">
<paragraph id="DF6E36F0FF87FFDC6B325E59FACE3060" blockId="22.[809,1382,1030,1055]" box="[809,1382,1030,1055]" pageId="22" pageNumber="135">
<heading id="8426819CFF87FFDC6B325E59FACE3060" box="[809,1382,1030,1055]" centered="true" fontSize="10" level="2" pageId="22" pageNumber="135" reason="4">
<taxonomicName id="18D14D73FF87FFDC6B325E59FACE3060" authority="Collingwood &amp; Agosti 1996" authorityName="Collingwood &amp; Agosti" authorityYear="1996" box="[809,1382,1030,1055]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="135" phylum="Arthropoda" rank="species" species="yemeni">
<emphasis id="EDA5EAE2FF87FFDC6B325E59FACE3060" bold="true" box="[809,1382,1030,1055]" pageId="22" pageNumber="135">
<emphasis id="EDA5EAE2FF87FFDC6B325E59FB8B3061" bold="true" box="[809,1059,1030,1055]" italics="true" pageId="22" pageNumber="135">Cardiocondyla yemeni</emphasis>
Collingwood &amp; Agosti 1996
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="DF6E36F0FF87FFDC6B325E77FBAD3041" blockId="22.[809,1029,1063,1088]" box="[809,1029,1063,1088]" pageId="22" pageNumber="135">
<heading id="8426819CFF87FFDC6B325E77FBAD3041" bold="true" box="[809,1029,1063,1088]" fontSize="10" level="2" pageId="22" pageNumber="135" reason="3">
<emphasis id="EDA5EAE2FF87FFDC6B325E77FBAD3041" bold="true" box="[809,1029,1063,1088]" pageId="22" pageNumber="135">
[
<typeStatus id="006A8852FF87FFDC6B2B5E77FCCC303E" box="[816,868,1064,1088]" pageId="22" pageNumber="135">type</typeStatus>
investigation]
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="97CB657BFF87FFDC6B325E17FAF13195" pageId="22" pageNumber="135" type="description">
<paragraph id="DF6E36F0FF87FFDC6B325E17FBC13138" blockId="22.[786,1399,1096,1975]" pageId="22" pageNumber="135">
This taxon has been described from
<collectingCountry id="A7C67660FF87FFDC6C855E16FB45301E" box="[1182,1261,1097,1120]" name="Yemen" pageId="22" pageNumber="135">Yemen</collectingCountry>
. Investigated wasonetypeworkerlabelledinCollingwoodshandwriting
<collectingCountry id="A7C67660FF87FFDC6B0D5ED5FCD730DC" box="[790,895,1162,1186]" name="Yemen" pageId="22" pageNumber="135">YEMEN</collectingCountry>
:
<collectingRegion id="1D15F812FF87FFDC6B9C5ED5FC4D30DC" box="[903,997,1162,1186]" country="Yemen" name="San'a'" pageId="22" pageNumber="135">SANAA</collectingRegion>
(sandy path)
<date id="AB6F1030FF87FFDC6C695ED5FB4230DC" box="[1138,1258,1162,1186]" pageId="22" pageNumber="135" value="1993-03-05">05 03 1993</date>
, Typus and
<taxonomicName id="18D14D73FF87FFDC6B0D5EF4FC1030BD" authorityName="Emery" authorityYear="1869" box="[790,952,1195,1219]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="135" phylum="Arthropoda" rank="genus">Cardiocondyla</taxonomicName>
yemeni; one topotypical worker, without postpetiole and gaster, labelled
<emphasis id="EDA5EAE2FF87FFDC6C735E92FAC6309A" box="[1128,1390,1228,1252]" italics="true" pageId="22" pageNumber="135">
<collectingCountry id="A7C67660FF87FFDC6C735E92FB68309A" box="[1128,1216,1229,1252]" name="Yemen" pageId="22" pageNumber="135">YEMEN</collectingCountry>
<collectingRegion id="1D15F812FF87FFDC6CDD5E93FAA7309D" box="[1222,1295,1228,1251]" country="Yemen" name="San'a'" pageId="22" pageNumber="135">Sanaa</collectingRegion>
17 III 93
</emphasis>
(day not clearly legible, other possible reading
<emphasis id="EDA5EAE2FF87FFDC6D195EB2FAC4317B" box="[1282,1388,1261,1285]" italics="true" pageId="22" pageNumber="135">27 III 93</emphasis>
) and
<taxonomicName id="18D14D73FF87FFDC6B5F5F51FB953158" authorityName="Seifert" authorityYear="2024" box="[836,1085,1294,1318]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="135" phylum="Arthropoda" rank="species" species="yemeni" status="sp. nov.">Cardiocondyla yemeni</taxonomicName>
<taxonomicNameLabel id="F6965799FF87FFDC6C5E5F51FBDF315B" box="[1093,1143,1294,1317]" pageId="22" pageNumber="135" rank="species">n.sp.</taxonomicNameLabel>
Collingwood &amp; Agosti 1996; depository SMN Görlitz.
</paragraph>
<paragraph id="DF6E36F0FF87FFDC6B325F10FB9731F6" blockId="22.[786,1399,1096,1975]" pageId="22" pageNumber="135">
All material examined. Morphometrically investigated were only the
<typeStatus id="006A8852FF87FFDC6BB05F2EFC7331F7" box="[939,987,1393,1417]" pageId="22" pageNumber="135">type</typeStatus>
samples.
</paragraph>
<paragraph id="DF6E36F0FF87FFDC6B325FCEFAF13195" blockId="22.[786,1399,1096,1975]" pageId="22" pageNumber="135">
<emphasis id="EDA5EAE2FF87FFDC6B325FCEFBAA31D7" bold="true" box="[809,1026,1425,1449]" pageId="22" pageNumber="135">Geographic range</emphasis>
. Known so far only from the Arabian Peninsula:
<collectingCountry id="A7C67660FF87FFDC6C1D5FECFBF931B4" box="[1030,1105,1459,1482]" name="Yemen" pageId="22" pageNumber="135">Yemen</collectingCountry>
and
<collectingCountry id="A7C67660FF87FFDC6CB85FEDFB4D31B4" box="[1187,1253,1458,1482]" name="Oman" pageId="22" pageNumber="135">Oman</collectingCountry>
(by image evaluation of CASENT0922296 in www.antweb.org).
</paragraph>
</subSubSection>
<subSubSection id="97CB657BFF87FFDD6B325FABFDC730DC" lastPageId="23" lastPageNumber="136" pageId="22" pageNumber="135" type="diagnosis">
<paragraph id="DF6E36F0FF87FFDC6B325FABFADF33C9" blockId="22.[786,1399,1096,1975]" pageId="22" pageNumber="135">
<emphasis id="EDA5EAE2FF87FFDC6B325FABFC323272" bold="true" box="[809,922,1524,1548]" pageId="22" pageNumber="135">Diagnosis</emphasis>
: --Worker (
<tableCitation id="9253034BFF87FFDC6C3B5FABFBC33272" box="[1056,1131,1524,1548]" captionStart="Table 2" captionStartId="26.[150,203,1476,1497]" captionText="Table 2. Workers of the C. wroughtonii group with mean FL/FR&gt; 1.040 and originating from Africa and theArabian Peninsula. Sequence of data: arithmetic mean ± standard deviation [minimum, maximum]." pageId="22" pageNumber="135">Tab. 2</tableCitation>
,
<figureCitation id="47EA2A75FF87FFDC6C615FABFB5A3272" box="[1146,1266,1524,1548]" captionStart="Figure 1115" captionStartId="8.[151,215,1755,1777]" captionTargetBox="[151,1399,259,1706]" captionTargetPageId="8" captionText="Figure 1115. Cardiocondyla yemeni, type, Sanaa / Yemen, 5 March 1993. (11) head in dorsal view; (12) lateral view; (13) dorsal view; (14) head surface between inner eye margin and paramedian vertex. (15) anterior dorsum of 1st gaster tergite. Figure 16. Cardiocondyla nana, holotype, Ulu Temburong / Brunei 22 February 1982; from www.antweb.org CASENT0901757, images by Ryan Perry." figureDoi="http://doi.org/10.5281/zenodo.13194036" httpUri="https://zenodo.org/record/13194036/files/figure.png" pageId="22" pageNumber="135">Figs 1115</figureCitation>
, key). Very small, CS 377 µm. Head very long, CL/CW 1.260. Median third of anterior clypeal margin straight, median third of occipital margin slightly concave. Postocular distance large, PoOc/CL 0.453. Frons relatively narrow (FRS/CS 0.242); frontal carinae slightly diverging frontal of FRS level (FL/FR 1.063), caudal of FRS level parallel or slightly converging. Eye rather small, EYE/ CS 0.238. Scape very short, SL/CS 0.724. Metanotal depression nearly absent (MGr/CS 0.55 %). Propodeal spines short (SP/CS 0.103) and almost triangular in lateral view, their axis in profile deviating from longitudinal mesosomal axis by 25°, in dorsal view not diverging and their bases relatively narrow (SPBA/CS 0.266).
</paragraph>
<paragraph id="DF6E36F0FF86FFDD68A65AAEFDA4301E" blockId="23.[189,802,241,1251]" pageId="23" pageNumber="136">
Petiole rather narrow and moderately high (PeW/CS 0.276, PeH/CS 0.336); in profile with a concave anterior face and a semicircular dorsum; in dorsal view with an almost globular node, which is only slightly longer than wide. Postpetiole narrow and low (PpW/CS 0.430, PpH/CS 0.268); postpetiolar sternite anterolaterally with a rounded lobiform protrusion on each side which clearly elevates above the level of median surface of sternite; the surface of the sternite thus appearing deeply concave in frontal view. Postpetiole in dorsal view with a straight to slightly concave anterior margin and convex sides. Lateral parts of clypeus each with 23 rugulae, central part slightly carinulate-reticulate. Frontal laminae with weak microsculpture, consisting of an irregular mixture of microrugulate, foveolate and microreticulate elements. Whole vertex foveolate, mediad of the eyes with densely-arranged, flat-bottomed foveolae of 1416 µm diameter. The largest foveolae show a well-demarcated but flat central tubercule of 68 µm diameter around the hair bases (
<figureCitation id="47EA2A75FF86FFDD6AD9593DFCBD3704" box="[706,789,866,890]" captionStart="Figure 1115" captionStartId="8.[151,215,1755,1777]" captionTargetBox="[151,1399,259,1706]" captionTargetPageId="8" captionText="Figure 1115. Cardiocondyla yemeni, type, Sanaa / Yemen, 5 March 1993. (11) head in dorsal view; (12) lateral view; (13) dorsal view; (14) head surface between inner eye margin and paramedian vertex. (15) anterior dorsum of 1st gaster tergite. Figure 16. Cardiocondyla nana, holotype, Ulu Temburong / Brunei 22 February 1982; from www.antweb.org CASENT0901757, images by Ryan Perry." figureDoi="http://doi.org/10.5281/zenodo.13194036" httpUri="https://zenodo.org/record/13194036/files/figure.png" pageId="23" pageNumber="136">Fig. 14</figureCitation>
). Whole surface of mesosoma foveolate-microreticulate; petiole and postpetiolar tergite microreticulate. First gaster tergite with moderately long and dense pubescence (PLG/CS 6.78 %, sqPDG 3.76) and a weakly developed microreticulum, with the hair bases placed in microfoveolae of 46 µm diameter. All body parts yellowish or yellowish-brown.
</paragraph>
<paragraph id="DF6E36F0FF86FFDD68CF5E36FDC730DC" blockId="23.[189,802,241,1251]" pageId="23" pageNumber="136">
<emphasis id="EDA5EAE2FF86FFDD68CF5E36FE7230FF" bold="true" box="[212,474,1129,1153]" pageId="23" pageNumber="136">Taxonomic comments.</emphasis>
For separation from closely related
<taxonomicName id="18D14D73FF86FFDD69165ED5FDAA30DC" box="[269,514,1162,1186]" class="Insecta" family="Formicidae" genus="Cardiocondyla" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="136" phylum="Arthropoda" rank="species" species="yoruba">
<emphasis id="EDA5EAE2FF86FFDD69165ED5FDAA30DC" box="[269,514,1162,1186]" italics="true" pageId="23" pageNumber="136">Cardiocondyla yoruba</emphasis>
</taxonomicName>
see there.
</paragraph>
</subSubSection>
<subSubSection id="97CB657BFF86FFDD68CF5EF4FEBE309D" pageId="23" pageNumber="136" type="biology_ecology">
<paragraph id="DF6E36F0FF86FFDD68CF5EF4FEBE309D" blockId="23.[189,802,241,1251]" pageId="23" pageNumber="136">
<emphasis id="EDA5EAE2FF86FFDD68CF5EF4FE9830BD" bold="true" box="[212,304,1195,1219]" pageId="23" pageNumber="136">Biology.</emphasis>
As habitat was reported a sandy path in Saana /
<collectingCountry id="A7C67660FF86FFDD68DF5E93FEBB309D" box="[196,275,1228,1251]" name="Yemen" pageId="23" pageNumber="136">Yemen</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
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