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<document id="9381A08C81A2ABD8DE084DDCFDE91B39" ID-DOI="10.5281/zenodo.12807677" ID-ISSN="1313-9916" ID-Zenodo-Dep="12807677" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721835052394" checkinUser="felipe" docAuthor="Mitov, Plamen" docDate="2021" docId="1D602969B56FA531FF2A0B2CFBE9FD1C" docLanguage="en" docName="ZooNotes.2021.178.1-4.pdf" docOrigin="ZooNotes 2021 (178)" docStyle="DocumentStyle:F573CCA44A18459B99963FDCC5E47C08.7:ZooNotes.2009-2023.journal_article" docStyleId="F573CCA44A18459B99963FDCC5E47C08" docStyleName="ZooNotes.2009-2023.journal_article" docStyleVersion="7" docTitle="Graecophalangium marenzelleri Mitov 2021, comb. n." docType="treatment" docVersion="2" lastPageNumber="3" masterDocId="E1595111B56FA533FF970D02FFCBFFF7" masterDocTitle="A new member of the genus Graecophalangium Roewer, 1923 and the first generic record for the Turkish harvestman fauna (Arachnida: Opiliones)" masterLastPageNumber="4" masterPageNumber="1" pageNumber="1" updateTime="1721837030448" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="71F593120188D767C0B6E562CEC6F7AD">A new member of the genus Graecophalangium Roewer, 1923 and the first generic record for the Turkish harvestman fauna (Arachnida: Opiliones)</mods:title>
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<mods:namePart id="D89886175A252718B401E9B8690F1EDA">Mitov, Plamen</mods:namePart>
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<mods:title id="63F00FAA16A486FEDA2E4F7F1A2EBB2B">ZooNotes</mods:title>
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<mods:date id="5185A34C568900750FC8DD0AF9D9E8FC">2021</mods:date>
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<mods:number id="EF8981675875E1CE365712CD278400C6">2021-04-26</mods:number>
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<treatment id="1D602969B56FA531FF2A0B2CFBE9FD1C" LSID="urn:lsid:plazi:treatment:1D602969B56FA531FF2A0B2CFBE9FD1C" httpUri="http://treatment.plazi.org/id/1D602969B56FA531FF2A0B2CFBE9FD1C" lastPageId="2" lastPageNumber="3" pageId="0" pageNumber="1">
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<heading id="CE3E2F13B56FA533FF2A0B2CFC07F9B0" bold="true" box="[189,972,1582,1607]" fontSize="10" level="2" pageId="0" pageNumber="1" reason="2">
<emphasis id="A7BD446DB56FA533FF2A0B2CFC07F9B0" bold="true" box="[189,972,1582,1607]" pageId="0" pageNumber="1">
<taxonomicName id="52C9E3FCB56FA533FF2A0B2CFC87F9B0" ID-CoL="9X9NP" authority="(Nosek, 1905)" authorityName="Mitov" authorityYear="2021" baseAuthorityName="Nosek" baseAuthorityYear="1905" box="[189,844,1582,1607]" class="Arachnida" family="Phalangiidae" genus="Graecophalangium" kingdom="Animalia" order="Opiliones" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="marenzelleri" status="comb. nov.">
<emphasis id="A7BD446DB56FA533FF2A0B2CFD4FF9B0" bold="true" box="[189,644,1582,1607]" italics="true" pageId="0" pageNumber="1">Graecophalangium marenzelleri</emphasis>
(
<bibRefCitation id="F158E58EB56FA533FD010B2CFC89F9B0" author="Nosek, A." box="[662,834,1582,1607]" pageId="0" pageNumber="1" pagination="114 - 154" refId="ref2690" refString="Nosek, A. (1905) Araneiden, Opilionen und Chernetiden. In: Penther, A., Zederbauer, E. (Eds.), Ergebnisse einer naturwissenschaftliche Reise zum Erdschais-Dagh (Kleinasien). Annalen des Naturhistorischen Museums in Wien, 20: 114 - 154." type="journal article" year="1905">Nosek, 1905</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="BC8EF916B56FA533FCC20B2CFC07F9B0" box="[853,972,1582,1607]" pageId="0" pageNumber="1" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
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<paragraph id="9576987FB56FA533FF2A0B51FD01F99F" blockId="0.[189,1399,1582,1768]" box="[189,714,1618,1640]" pageId="0" pageNumber="1">
<treatmentCitationGroup id="B5D9BF51B56FA533FF2A0B51FD01F99F" box="[189,714,1618,1640]" pageId="0" pageNumber="1">
<treatmentCitation id="1468BE6EB56FA533FF2A0B51FD0FF99F" author="Nosek, A." box="[189,708,1618,1640]" page="153" pageId="0" pageNumber="1" year="1905">
<taxonomicName id="52C9E3FCB56FA533FF2A0B51FD0FF99F" ID-CoL="7VVF6" authority="Nosek, 1905: 152 - 153" authorityName="Nosek" authorityPageNumber="152 - 153" authorityYear="1905" box="[189,708,1618,1640]" class="Arachnida" family="Phalangiidae" genus="Egaenus" kingdom="Animalia" order="Opiliones" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="marenzelleri">
<emphasis id="A7BD446DB56FA533FF2A0B51FE7DF990" box="[189,438,1618,1640]" italics="true" pageId="0" pageNumber="1">Egaenus Marenzelleri</emphasis>
<bibRefCitation id="F158E58EB56FA533FE290B50FD0FF99F" author="Nosek, A." box="[446,708,1618,1640]" pageId="0" pageNumber="1" pagination="114 - 154" refId="ref2690" refString="Nosek, A. (1905) Araneiden, Opilionen und Chernetiden. In: Penther, A., Zederbauer, E. (Eds.), Ergebnisse einer naturwissenschaftliche Reise zum Erdschais-Dagh (Kleinasien). Annalen des Naturhistorischen Museums in Wien, 20: 114 - 154." type="journal article" year="1905">Nosek, 1905: 152-153</bibRefCitation>
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;
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</paragraph>
<paragraph id="9576987FB56FA533FF2A0B71FD58F970" blockId="0.[189,1399,1582,1768]" box="[189,659,1650,1672]" pageId="0" pageNumber="1">
<treatmentCitationGroup id="B5D9BF51B56FA533FF2A0B71FD58F970" box="[189,659,1650,1672]" pageId="0" pageNumber="1">
<taxonomicName id="52C9E3FCB56FA533FF2A0B71FE7DF970" ID-CoL="7VVF6" authorityName="Nosek" authorityYear="1905" box="[189,438,1650,1672]" class="Arachnida" family="Phalangiidae" genus="Egaenus" kingdom="Animalia" order="Opiliones" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="marenzelleri">
<emphasis id="A7BD446DB56FA533FF2A0B71FE7DF970" box="[189,438,1650,1672]" italics="true" pageId="0" pageNumber="1">Egaenus Marenzelleri</emphasis>
</taxonomicName>
:
<treatmentCitation id="1468BE6EB56FA533FE520B70FD46F970" author="Roewer, C. - F." box="[453,653,1650,1671]" page="11" pageId="0" pageNumber="1" year="1911">
<bibRefCitation id="F158E58EB56FA533FE520B70FD46F970" author="Roewer, C. - F." box="[453,653,1650,1671]" pageId="0" pageNumber="1" pagination="1 - 106" refId="ref2742" refString="Roewer, C. - F. (1911) Ubersicht der Genera der Subfamilie der Phalangiini der Opiliones Palpatores nebst Beschreibung einiger neuer Gattungen und Arten. Archiv fur Naturgeschichte (Berlin) I, 77 (Supplementheft 2): 1 - 106." type="journal article" year="1911">Roewer 1911: 11</bibRefCitation>
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;
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</paragraph>
<paragraph id="9576987FB56FA533FF2A0B91FDD0F93F" blockId="0.[189,1399,1582,1768]" pageId="0" pageNumber="1">
<treatmentCitationGroup id="B5D9BF51B56FA533FF2A0B91FDD0F93F" pageId="0" pageNumber="1">
<taxonomicName id="52C9E3FCB56FA533FF2A0B91FE77F950" ID-CoL="7VVF6" authorityName="Nosek" authorityYear="1905" box="[189,444,1682,1704]" class="Arachnida" family="Phalangiidae" genus="Egaenus" kingdom="Animalia" order="Opiliones" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="marenzelleri">
<emphasis id="A7BD446DB56FA533FF2A0B91FE77F950" box="[189,444,1682,1704]" italics="true" pageId="0" pageNumber="1">Egaenus marenzelleri</emphasis>
</taxonomicName>
:
<treatmentCitation id="1468BE6EB56FA533FE470B90FD7AF950" author="Roewer, C. - F." box="[464,689,1682,1703]" page="202" pageId="0" pageNumber="1" year="1912">
<bibRefCitation id="F158E58EB56FA533FE470B90FD7AF950" author="Roewer, C. - F." box="[464,689,1682,1703]" pageId="0" pageNumber="1" pagination="1 - 295" refId="ref2788" refString="Roewer, C. - F. (1912) Revision der Opiliones Palpatores (= Opiliones Plagiostethi). II. Teil: Familie der Phalangiidae. (Subfamilien: Sclerosomini, Oligolophini, Phalangiini). Abhandlungen aus dem Gebiete der Naturwissenschaften, herausgegeben vom Naturwissenschaftlichen Verein in Hamburg, 20 (1): 1 - 295." type="journal article" year="1912">Roewer 1912: 202</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="1468BE6EB56FA533FD550B90FC68F95F" author="Roewer, C. - F." box="[706,931,1682,1704]" page="869" pageId="0" pageNumber="1" year="1923">
<bibRefCitation id="F158E58EB56FA533FD550B90FC68F95F" author="Roewer, C. - F." box="[706,931,1682,1704]" pageId="0" pageNumber="1" refId="ref2849" refString="Roewer, C. - F. (1923) Die Weberknechte der Erde. Systematische Bearbeitung der bisher bekannten Opiliones. Gustav Fischer, Jena, 1116 pp." type="book" year="1923">Roewer 1923: 869</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="1468BE6EB56FA533FC230B90FB2DF95F" author="Bayram, A. &amp; Corak, I. &amp; Danisman, T. &amp; Sancak, Z. &amp; Yigit, N." box="[948,1254,1682,1704]" page="572" pageId="0" pageNumber="1" year="2010">
<bibRefCitation id="F158E58EB56FA533FC230B90FB2DF95F" author="Bayram, A. &amp; Corak, I. &amp; Danisman, T. &amp; Sancak, Z. &amp; Yigit, N." box="[948,1254,1682,1704]" pageId="0" pageNumber="1" pagination="563 - 585" refId="ref2302" refString="Bayram, A., Corak, I., Danisman, T., Sancak, Z. &amp; Yigit, N. (2010) Checklist of the harvestmen of Turkey (Arachnida: Opiliones). Munis Entomology &amp; Zoology, 5 (2): 563 - 585." type="journal article" year="2010">
Bayram
<emphasis id="A7BD446DB56FA533FB8B0B91FB91F950" box="[1052,1114,1682,1704]" italics="true" pageId="0" pageNumber="1">et al.</emphasis>
2010: 572
</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="1468BE6EB56FA533FB600B90FEE2F93F" author="Kurt, K. &amp; Eman, O. K. &amp; Demir, H. &amp; Seyyar, O." page="36" pageId="0" pageNumber="1" year="2010">
<bibRefCitation id="F158E58EB56FA533FB600B90FEE2F93F" author="Kurt, K. &amp; Eman, O. K. &amp; Demir, H. &amp; Seyyar, O." pageId="0" pageNumber="1" pagination="33 - 44" refId="ref2428" refString="Kurt, K., Eman, O. K., Demir, H. &amp; Seyyar, O. (2010) The Turkish Harvestmen (Opiliones) with zoogeographical remarks. Serket, 12 (2): 33 - 44." type="journal article" year="2010">
Kurt
<emphasis id="A7BD446DB56FA533FAAE0B91FABCF950" box="[1337,1399,1682,1704]" italics="true" pageId="0" pageNumber="1">et al.</emphasis>
2010: 36
</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="1468BE6EB56FA533FEA20BB0FDDFF93F" author="Muranyi, D." box="[309,532,1714,1736]" page="4" pageId="0" pageNumber="1" year="2015">
<bibRefCitation id="F158E58EB56FA533FEA20BB0FDDFF93F" author="Muranyi, D." box="[309,532,1714,1736]" pageId="0" pageNumber="1" pagination="4 - 13" refId="ref2641" refString="Muranyi, D. (2015) First record of the genus Graecophalangium Roewer, 1923 (Opiliones: Phalangiidae) from Albania, with redescription of G. militare (C. L. Koch, 1839). Ecologica Montenegrina, 4: 4 - 13." type="journal article" year="2015">Murányi 2015: 4-6</bibRefCitation>
</treatmentCitation>
;
</treatmentCitationGroup>
</paragraph>
<paragraph id="9576987FB56FA533FF2A0BD0FB06F910" blockId="0.[189,1399,1582,1768]" box="[189,1229,1746,1768]" pageId="0" pageNumber="1">
<treatmentCitationGroup id="B5D9BF51B56FA533FF2A0BD0FB06F910" box="[189,1229,1746,1768]" pageId="0" pageNumber="1">
<emphasis id="A7BD446DB56FA533FF2A0BD0FDF9F91F" box="[189,562,1746,1768]" italics="true" pageId="0" pageNumber="1">
<taxonomicName id="52C9E3FCB56FA533FF2A0BD0FDE0F910" ID-CoL="9X9NP" authorityName="Mitov" authorityYear="2021" baseAuthorityName="Nosek" baseAuthorityYear="1905" box="[189,555,1746,1767]" class="Arachnida" family="Phalangiidae" genus="Graecophalangium" kingdom="Animalia" order="Opiliones" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="marenzelleri">Graecophalangium marenzelleri</taxonomicName>
:
</emphasis>
<treatmentCitation id="1468BE6EB56FA533FDAE0BD0FD36F91F" author="Kurt, K." box="[569,765,1746,1768]" page="1622" pageId="0" pageNumber="1" year="2014">
<bibRefCitation id="F158E58EB56FA533FDAE0BD0FD36F91F" author="Kurt, K." box="[569,765,1746,1768]" pageId="0" pageNumber="1" pagination="1617 - 1631" refId="ref2395" refString="Kurt, K. (2014) Updated checklist of harvestmen (Arachnida: Opiliones) in Turkey. Archives of Biological Sciences, 66 (4): 1617 - 1631." type="journal article" year="2014">Kurt 2014: 1622</bibRefCitation>
</treatmentCitation>
(invalid proposal for new combination).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DDD3CBF4B56FA532FF2A0A1BFC37FE37" lastPageId="1" lastPageNumber="2" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="9576987FB56FA533FF2A0A1BFD72F7E6" blockId="0.[189,1399,1817,2066]" pageId="0" pageNumber="1">
<emphasis id="A7BD446DB56FA533FF2A0A1BFE80F8C5" bold="true" box="[189,331,1817,1842]" pageId="0" pageNumber="1">Diagnosis.</emphasis>
A relatively small (body length ca. 4.0 mm) and „unarmed‟ member of genus
<emphasis id="A7BD446DB56FA533FF2A0A3DFE7DF8A0" box="[189,438,1855,1879]" italics="true" pageId="0" pageNumber="1">
<taxonomicName id="52C9E3FCB56FA533FF2A0A3DFE7AF8A0" authorityName="Roewer" authorityYear="1923" box="[189,433,1855,1879]" class="Arachnida" family="Phalangiidae" genus="Graecophalangium" kingdom="Animalia" order="Opiliones" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Graecophalangium</taxonomicName>
,
</emphasis>
that differs by the outline of the shaft and glans of the penis (Figs 16- 18). Chelicerae frontal with only one small, bluntly conical apophysis on the fixed finger base (Figs 10, 12). Ocularium low and smooth, with only a few denticles and short setae (Fig. 4); pedipalps unarmed, with a very weak mediodistal apophysis on the patella; tarsal segments slightly curved (Figs 13-15). Legs smooth, only with setae; femur and tibia of the first leg weakly spindle-shaped (Fig. 7).
</paragraph>
<paragraph id="9576987FB56EA532FF2A0DCAFC37FE37" blockId="1.[189,1399,200,2092]" pageId="1" pageNumber="2">
<emphasis id="A7BD446DB56EA532FF2A0DCAFEF6FF16" bold="true" box="[189,317,200,225]" pageId="1" pageNumber="2">Affinities</emphasis>
According to the penis type (more slender and less sclerotized) and its structure,
<taxonomicName id="52C9E3FCB56EA532FF2A0DECFE4CFEF1" authorityName="Mitov" authorityYear="2021" baseAuthorityName="Nosek" baseAuthorityYear="1905" box="[189,391,238,262]" class="Arachnida" family="Phalangiidae" genus="Graecophalangium" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="marenzelleri">
<emphasis id="A7BD446DB56EA532FF2A0DECFE4CFEF1" box="[189,391,238,262]" italics="true" pageId="1" pageNumber="2">G. marenzelleri</emphasis>
</taxonomicName>
belongs most probably to the group of species related to
<taxonomicName id="52C9E3FCB56EA532FB430DECFEB5FEDC" authority="Martens, 1966" authorityName="Martens" authorityYear="1966" class="Arachnida" family="Phalangiidae" genus="Graecophalangium" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="cretaeum">
<emphasis id="A7BD446DB56EA532FB430DECFABCFEF1" box="[1236,1399,238,262]" italics="true" pageId="1" pageNumber="2">G. cretaeum</emphasis>
Martens, 1966
</taxonomicName>
(from Crete) and
<taxonomicName id="52C9E3FCB56EA532FDE70C16FC02FEDC" authority="Starega, 1973" authorityName="Starega" authorityYear="1973" box="[624,969,275,299]" class="Arachnida" family="Phalangiidae" genus="Graecophalangium" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="punicum">
<emphasis id="A7BD446DB56EA532FDE70C16FCCFFEDC" box="[624,772,276,299]" italics="true" pageId="1" pageNumber="2">G. punicum</emphasis>
Staręga, 1973
</taxonomicName>
(from
<collectingCountry id="EDDED8EFB56EA532FBB20C11FB52FEDC" box="[1061,1177,275,299]" name="Lebanon" pageId="1" pageNumber="2">Lebanon</collectingCountry>
). In lateral view, the thickness of the penis trunk increases evenly (without narrowing) from the distal to the basal part, as observed in both
<typeStatus id="4A7226DDB56EA532FD150C5CFD35FE81" box="[642,766,350,374]" pageId="1" pageNumber="2" type="holotype">holotypes</typeStatus>
of
<taxonomicName id="52C9E3FCB56EA532FCA30C5CFC1EFE81" authorityName="Martens" authorityYear="1966" box="[820,981,350,374]" class="Arachnida" family="Phalangiidae" genus="Graecophalangium" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="cretaeum">
<emphasis id="A7BD446DB56EA532FCA30C5CFC1EFE81" box="[820,981,350,374]" italics="true" pageId="1" pageNumber="2">G. cretaeum</emphasis>
</taxonomicName>
(NHMW inv. № 4355) and
<taxonomicName id="52C9E3FCB56EA532FACC0C5CFEE1FE6C" authorityName="Starega" authorityYear="1973" class="Arachnida" family="Phalangiidae" genus="Graecophalangium" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="punicum">
<emphasis id="A7BD446DB56EA532FACC0C5CFEE1FE6C" italics="true" pageId="1" pageNumber="2">G. punicum</emphasis>
</taxonomicName>
(inv. № 46/51, Museum and Institute of Zoology in Warszawa,
<collectingCountry id="EDDED8EFB56EA532FB100C81FB2FFE6C" box="[1159,1252,387,411]" name="Poland" pageId="1" pageNumber="2">Poland</collectingCountry>
, MIZW). In the terms of its habitus
<taxonomicName id="52C9E3FCB56EA532FE600CABFD72FE36" authorityName="Mitov" authorityYear="2021" baseAuthorityName="Nosek" baseAuthorityYear="1905" box="[503,697,425,449]" class="Arachnida" family="Phalangiidae" genus="Graecophalangium" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="marenzelleri">
<emphasis id="A7BD446DB56EA532FE600CABFD72FE36" box="[503,697,425,449]" italics="true" pageId="1" pageNumber="2">G. marenzelleri</emphasis>
</taxonomicName>
is closest to
<emphasis id="A7BD446DB56EA532FCF40CABFC37FE37" box="[867,1020,425,448]" italics="true" pageId="1" pageNumber="2">
<taxonomicName id="52C9E3FCB56EA532FCF40CABFC33FE37" authorityName="Starega" authorityYear="1973" box="[867,1016,425,448]" class="Arachnida" family="Phalangiidae" genus="Graecophalangium" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="punicum">G. punicum</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="DDD3CBF4B56EA531FE8A0CCFFD53FEF1" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2" type="description">
<paragraph id="9576987FB56EA532FE8A0CCFFDB1FE11" blockId="1.[189,1399,200,2092]" box="[285,634,461,486]" pageId="1" pageNumber="2">
<emphasis id="A7BD446DB56EA532FE8A0CCFFDB1FE11" bold="true" box="[285,634,461,486]" pageId="1" pageNumber="2">Redescription (Figs 1-21)</emphasis>
</paragraph>
<paragraph id="9576987FB56EA532FF2A0CF1FC3CFD56" blockId="1.[189,1399,200,2092]" pageId="1" pageNumber="2">
The following redescription is based on a single male specimen, labelled as the
<typeStatus id="4A7226DDB56EA532FB710CF1FA9FFDFC" box="[1254,1364,499,523]" pageId="1" pageNumber="2" type="holotype">holotype</typeStatus>
of
<taxonomicName id="52C9E3FCB56EA532FF2A0F18FD5AFDC6" authority="Nosek, 1905" authorityName="Nosek" authorityYear="1905" box="[189,657,537,561]" class="Arachnida" family="Phalangiidae" genus="Egaenus" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="marenzelleri">
<emphasis id="A7BD446DB56EA532FF2A0F18FE12FDC6" box="[189,473,537,561]" italics="true" pageId="1" pageNumber="2">Egaenus marenzelleri</emphasis>
<bibRefCitation id="F158E58EB56EA532FE7F0F1BFD5AFDC6" author="Nosek, A." box="[488,657,537,561]" pageId="1" pageNumber="2" pagination="114 - 154" refId="ref2690" refString="Nosek, A. (1905) Araneiden, Opilionen und Chernetiden. In: Penther, A., Zederbauer, E. (Eds.), Ergebnisse einer naturwissenschaftliche Reise zum Erdschais-Dagh (Kleinasien). Annalen des Naturhistorischen Museums in Wien, 20: 114 - 154." type="journal article" year="1905">Nosek, 1905</bibRefCitation>
</taxonomicName>
(NHMW inv. № 4355); Type locality:
<collectingCountry id="EDDED8EFB56EA532FB340F1BFAC9FDC6" box="[1187,1282,537,561]" name="Turkey" pageId="1" pageNumber="2">Turkey</collectingCountry>
, Central
<collectingCountry id="EDDED8EFB56EA532FF2A0F3CFEE6FDA1" box="[189,301,574,598]" name="Turkey" pageId="1" pageNumber="2">Anatolia</collectingCountry>
: “Erciyas Dag (Nordseite)” (=Mount Erciyes,
<geoCoordinate id="F0FDFEB8B56EA532FCEA0F3FFBE0FDA1" box="[893,1067,573,598]" degrees="38" direction="north" minutes="31" orientation="latitude" pageId="1" pageNumber="2" precision="1" seconds="51.6" value="38.531">38°3151.6″N</geoCoordinate>
,
<geoCoordinate id="F0FDFEB8B56EA532FBAB0F3FFB2DFDA1" box="[1084,1254,573,598]" degrees="35" direction="east" minutes="26" orientation="longitude" pageId="1" pageNumber="2" precision="1" seconds="49.2" value="35.447002">35°2649.2″E</geoCoordinate>
), “Ende
<date id="E177BEBFB56EA532FACC0F3CFECAFD8B" pageId="1" pageNumber="2" value="1902-05">V. 1902</date>
”, leg. A. Penther, det.
<bibRefCitation id="F158E58EB56EA532FDBA0F61FD13FD8C" author="Nosek, A." box="[557,728,611,635]" pageId="1" pageNumber="2" pagination="114 - 154" refId="ref2690" refString="Nosek, A. (1905) Araneiden, Opilionen und Chernetiden. In: Penther, A., Zederbauer, E. (Eds.), Ergebnisse einer naturwissenschaftliche Reise zum Erdschais-Dagh (Kleinasien). Annalen des Naturhistorischen Museums in Wien, 20: 114 - 154." type="journal article" year="1905">Nosek (1905)</bibRefCitation>
. All measurements are in mm. In square brackets are the figures from Nosek‟s (1905) original species description.
</paragraph>
<paragraph id="9576987FB56EA531FF2A0FAFFD53FEF1" blockId="1.[189,1399,200,2092]" lastBlockId="2.[189,1399,201,747]" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
<emphasis id="A7BD446DB56EA532FF2A0FAFFEC9FD31" bold="true" box="[189,258,685,710]" pageId="1" pageNumber="2">Body</emphasis>
(Figs 1-3) length (L): 3.97 (with anterior carapace edges = 4.15) [4.3]; prosoma L: 1.6 [1.6] and width (W): 2.76 [2.8]; abdomen L: 2.4 [2.7] and W: 2.75 [3.0]; carapace armed with a few acute black-tipped tubercles (bearing strong setae) at the frontal corners, laterally and in front of ocularium; similar tubercles arranged in lines are found on meso- and metapeltidium; on abdominal dorsum they are only a few; body venter with black-brown, sparse setae. Color (after more than 100 years in alcohol): body light brown/yellow-brown, brown saddle margins are clearly visible (Fig 1); its natural color is described by
<bibRefCitation id="F158E58EB56EA532FAB00E8CFE8CFC3B" author="Nosek, A." pageId="1" pageNumber="2" pagination="114 - 154" refId="ref2690" refString="Nosek, A. (1905) Araneiden, Opilionen und Chernetiden. In: Penther, A., Zederbauer, E. (Eds.), Ergebnisse einer naturwissenschaftliche Reise zum Erdschais-Dagh (Kleinasien). Annalen des Naturhistorischen Museums in Wien, 20: 114 - 154." type="journal article" year="1905">Nosek (1905: 39)</bibRefCitation>
. Supracheliceral lamellae are small, bearing one small tubercle.
<emphasis id="A7BD446DB56EA532FB370EB1FABCFC3B" bold="true" box="[1184,1399,947,972]" pageId="1" pageNumber="2">Ocular tubercle</emphasis>
(Figs 3-4) is smooth with a few denticles (5/6) and short setae (3/3) only; low (height=0.26) [0.3], length-width=0.52 [0.6]; with shallow longitudinal furrow; inner distance between the eyes 0.32 [0.3]; distance from the front of cephalothorax = 0.57. Epistome (Figs 5-6) L: 0.15- 0.2;
<emphasis id="A7BD446DB56EA532FF6F094AFE41FB96" bold="true" box="[248,394,1096,1121]" pageId="1" pageNumber="2">Chelicerae</emphasis>
(Figs 8-12) light yellow-brown, frontally with brown spots and with only one small bluntly conical apophysis on the fixed finger base (Figs 5, 9-10, 12); basal segment L: 1.47 [0.5] and maximum W (in dorsal view, Fig. 11): 0.58, dorsally and ventrally with several spine-tipped tubercles; distal segment L: 1.70 [1.8] and W (in frontal view): 0.54, frontally covered with stout spine-like setae; movable finger L: 0.71 [0.5].
<emphasis id="A7BD446DB56EA532FB8709DFFB5FFB01" bold="true" box="[1040,1172,1245,1270]" pageId="1" pageNumber="2">Pedipalps</emphasis>
unarmed, whiteyellowish (Figs 13-15); segment lengths: trochanter (Tr): 0.60, femur (Fe): 1.28 [1.3], patella (Pa): 0.71 [0.7], tibia (Ti): 0.83 [0.7], tarsus (Ta): 1.50 [1.6], total: 5.0; patella (0.37) and tibia (0.33) maximum width, in dorsal view (Fig. 15); claw L: 0.142. All segments bear setae. Tr ventrally with spine-tipped tubercles, Fe ventrally with stout spine-like setae; Pa with a very weak mediodistal apophysis (Fig. 15); Ti ventrally with several denticles; Ta slightly curved, ventrally with a field of micro-denticles.
<emphasis id="A7BD446DB56EA532FD4108BFFCDFFA21" bold="true" box="[726,788,1469,1494]" pageId="1" pageNumber="2">Legs</emphasis>
yellow with brown spots on patella and tibia; cylindrical (first leg femur and tibia weakly spindle-shaped, Figs 1, 7), smooth, with setae only; tarsi ending with a single claw. Ti I. distally on ventral side with small denticles and micro-denticles. Fe I. width (in lateral view): 0.6; segment lengths: Leg I. Tr: 0.53, Fe: 2.73 [2.5+0.2], Pa: 1.06 [1.2], Ti: 2.16 [2.1], metatarsus (Mt): 2.33 [2.4], Ta: 3.9, total: 12.71; Leg II. Tr: 0.50, Fe: 3.7 [3.5+0.3], Pa: 1.33 [1.3], Ti: 2.96 [2.8], Mt: 2.90 [3.0], Ta: 7.1, total: 18.49; Leg III. Tr: 0.43, Fe: 2.56 [2.2+0.25], Pa: 1.06 [1.1], Ti: 1.91 [1.75], Mt: 2.66 [2.9], Ta: 4.3, total: 12.92; Leg IV. Tr: 0.63, Fe: 3.83 [3.4+0.3], Pa: 1.26 [1.3], Ti: 2.6 [2.5], Mt: 4.23 [4.2], Ta: 6.0, total: 18.55.
<emphasis id="A7BD446DB56EA532FDAA0BEAFD43F8F6" bold="true" box="[573,648,1768,1793]" pageId="1" pageNumber="2">Penis</emphasis>
(Figs 16-21) light yellow to yellow-brown, L: 2.76 (= prosoma width); shaft L: 2.71; shaft (in dorsal view) with a distinct broad basis (W: 0.36; dorsally without a pronounced ridge and furrows, Fig. 21); distal part of the shaft forms long (0.61) „
<emphasis id="A7BD446DB56EA532FE8B0A58FE74F886" box="[284,447,1881,1905]" italics="true" pageId="1" pageNumber="2">wing paddle</emphasis>
‟ (! not a „
<emphasis id="A7BD446DB56EA532FDD90A58FD69F886" box="[590,674,1882,1905]" italics="true" pageId="1" pageNumber="2">spoon,</emphasis>
‟ such as in
<taxonomicName id="52C9E3FCB56EA532FCC00A5BFC73F886" authorityName="Silhavy" authorityYear="1965" box="[855,952,1881,1905]" class="Arachnida" family="Phalangiidae" genus="Rilaena" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="A7BD446DB56EA532FCC00A5BFC73F886" box="[855,952,1881,1905]" italics="true" pageId="1" pageNumber="2">Rilaena</emphasis>
</taxonomicName>
) with distal W: 0.186 (= stylus length), thereafter shaft narrows slightly towards the middle (0.13), then again widens slightly (0.17) to its extended basal part. In profile, the thickness of the penis-trunk increases evenly from its distal (0.07) towards the basal part (0.24). Glans elongated (L: 0.486), slightly concave dorsally (W: 0.114, in lateral view), with a relatively wide/thick upper (0.021) and lower (the keel) (0.027) glans wall (Fig. 18); glans with two pairs of setae (here hidden under artifacts) (Figs 18-20), distance between the base of setae ca. 0.04; stylus L: 0.186. Female is unknown.
</paragraph>
</subSubSection>
<subSubSection id="DDD3CBF4B56DA531FE8A0C10FBE9FD1C" pageId="2" pageNumber="3" type="discussion">
<paragraph id="9576987FB56DA531FE8A0C10FBE9FD1C" blockId="2.[189,1399,201,747]" pageId="2" pageNumber="3">
<emphasis id="A7BD446DB56DA531FE8A0C10FE6AFEDC" bold="true" box="[285,417,274,299]" pageId="2" pageNumber="3">Remarks:</emphasis>
<taxonomicName id="52C9E3FCB56DA531FE390C11FC81FEDC" authorityName="Mitov" authorityYear="2021" baseAuthorityName="Nosek" baseAuthorityYear="1905" box="[430,842,275,299]" class="Arachnida" family="Phalangiidae" genus="Graecophalangium" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="marenzelleri">
<emphasis id="A7BD446DB56DA531FE390C11FC81FEDC" box="[430,842,275,299]" italics="true" pageId="2" pageNumber="3">Graecophalangium marenzelleri</emphasis>
</taxonomicName>
is known at present only from the
<typeStatus id="4A7226DDB56DA531FAA80C16FABDFEDB" box="[1343,1398,276,300]" pageId="2" pageNumber="3">type</typeStatus>
locality and is the seventh member of the genus. I reported in a conference presentation (
<bibRefCitation id="F158E58EB56DA531FF520C5CFE91FE81" author="Mitov, P. G." box="[197,346,350,374]" pageId="2" pageNumber="3" refId="ref2473" refString="Mitov, P. G. (2008) Graecophalangium marenzelleri comb. nov. a new systematic position for Egaenus marenzelleri Nosek, 1905 (Opiliones, Phalangiidae). In: Nentwig, W., Schmidt, M. &amp; Kropf, C. (Eds.), 24 th European Congress of Arachnology 25. - 29. August 2008 in Bern, Switzerland, Book of abstracts. Natural History Museum Bern, University of Bern, Bern, p. 134." type="journal volume" year="2008">Mitov 2008</bibRefCitation>
), that
<taxonomicName id="52C9E3FCB56DA531FE260C5DFD0DFE81" authorityName="Mitov" authorityYear="2021" box="[433,710,350,374]" genus="Eganeus" pageId="2" pageNumber="3" rank="species" species="marenzelleri">
<emphasis id="A7BD446DB56DA531FE260C5DFD0DFE81" box="[433,710,350,374]" italics="true" pageId="2" pageNumber="3">Eganeus marenzelleri</emphasis>
</taxonomicName>
has to be regarded as species of
<taxonomicName id="52C9E3FCB56DA531FBEA0C5CFAA6FE81" authorityName="Roewer" authorityYear="1923" box="[1149,1389,350,374]" class="Arachnida" family="Phalangiidae" genus="Graecophalangium" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="A7BD446DB56DA531FBEA0C5CFAA6FE81" box="[1149,1389,350,374]" italics="true" pageId="2" pageNumber="3">Graecophalangium</emphasis>
</taxonomicName>
; however, the latter communication had not meet the criteria for publication of the International Code of Zoological Nomenclature (4
<superScript id="62BC3537B56DA531FCE40CABFC4EFE40" attach="left" box="[883,901,425,439]" fontSize="6" pageId="2" pageNumber="3">th</superScript>
edition).
<bibRefCitation id="F158E58EB56DA531FB800CABFB73FE36" author="Kurt, K." box="[1047,1208,425,449]" pageId="2" pageNumber="3" pagination="1617 - 1631" refId="ref2395" refString="Kurt, K. (2014) Updated checklist of harvestmen (Arachnida: Opiliones) in Turkey. Archives of Biological Sciences, 66 (4): 1617 - 1631." type="journal article" year="2014">Kurt (2014)</bibRefCitation>
inadvertently included this combination (without justification) in a published checklist, which is regarded here as an invalid taxonomic act since the transfer has not been proposed in a taxonomically valid publication. Thus, the present article is the first formal transfer of this species to the genus
<taxonomicName id="52C9E3FCB56DA531FE720F3CFD1DFDA1" authorityName="Roewer" authorityYear="1923" box="[485,726,574,598]" class="Arachnida" family="Phalangiidae" genus="Graecophalangium" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="A7BD446DB56DA531FE720F3CFD1DFDA1" box="[485,726,574,598]" italics="true" pageId="2" pageNumber="3">Graecophalangium</emphasis>
</taxonomicName>
as
<taxonomicName id="52C9E3FCB56DA531FC870F3CFC1CFDA1" authorityName="Mitov" authorityYear="2021" box="[784,983,574,598]" genus="Eganeus" pageId="2" pageNumber="3" rank="species" species="marenzelleri" status="comb. n.">
<emphasis id="A7BD446DB56DA531FC870F3CFC1CFDA1" box="[784,983,574,598]" italics="true" pageId="2" pageNumber="3">G. marenzelleri</emphasis>
</taxonomicName>
<emphasis id="A7BD446DB56DA531FC720F3FFBA8FDA1" bold="true" box="[997,1123,573,598]" pageId="2" pageNumber="3">
<taxonomicNameLabel id="BC8EF916B56DA531FC720F3FFBA8FDA1" box="[997,1123,573,598]" pageId="2" pageNumber="3" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
<bibRefCitation id="F158E58EB56DA531FBE50F3CFAD9FDA1" author="Kurt, K." box="[1138,1298,574,598]" pageId="2" pageNumber="3" pagination="1617 - 1631" refId="ref2395" refString="Kurt, K. (2014) Updated checklist of harvestmen (Arachnida: Opiliones) in Turkey. Archives of Biological Sciences, 66 (4): 1617 - 1631." type="journal article" year="2014">Kurt (2014)</bibRefCitation>
, in the same checklist for the genus
<taxonomicName id="52C9E3FCB56DA531FDA80F61FCFBFD8C" authorityName="Roewer" authorityYear="1923" box="[575,816,611,635]" class="Arachnida" family="Phalangiidae" genus="Graecophalangium" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="A7BD446DB56DA531FDA80F61FCFBFD8C" box="[575,816,611,635]" italics="true" pageId="2" pageNumber="3">Graecophalangium</emphasis>
</taxonomicName>
, also inadvertently included a species name for Turkish material mentioned in a conference presentation by
<bibRefCitation id="F158E58EB56DA531FBD60F8BFB25FD56" author="Mitov, P. G." box="[1089,1262,649,673]" pageId="2" pageNumber="3" pagination="16 - 21" refId="ref2562" refString="Mitov, P. G. (2009) On the genus Graecophalangium Roewer, 1923 (Opiliones, Phalangiidae). In: Chatzaki, M., Spiridopoulou, K. &amp; Stathi, I. (Eds.), 25 th European Congress of Arachnology, 16 - 21 August 2009, Alexandroupoli, Greece, Book of abstracts. Democritus University of Thrace, Natural History Museum of Crete, p. 81." type="journal article" year="2009">Mitov (2009)</bibRefCitation>
, which is nomen nudum as it has not yet been formally described. Therefore
<taxonomicName id="52C9E3FCB56DA531FB100FACFA9BFD31" authorityName="Mitov" authorityYear="2021" box="[1159,1360,686,710]" genus="Eganeus" pageId="2" pageNumber="3" rank="species" species="marenzelleri">
<emphasis id="A7BD446DB56DA531FB100FACFA9BFD31" box="[1159,1360,686,710]" italics="true" pageId="2" pageNumber="3">G. marenzelleri</emphasis>
</taxonomicName>
is formally the first generic record for the Turkish harvestman fauna.
</paragraph>
</subSubSection>
</treatment>
</document>