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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1178.105703" ID-Pensoft-Pub="1313-2970-1178-17" ID-Pensoft-UUID="26951753967254868073E2B13F9FDC60" ID-ZooBank="01B76CF7FF7A4CDFAAF5B4793EA932ED" ModsDocID="1313-2970-1178-17" checkinTime="1693642902469" checkinUser="pensoft" docAuthor="Penaherrera-R., Pedro" docDate="2023" docId="18C560DFF8E157DCA32C32EADCE98A6C" docLanguage="en" docName="ZooKeys 1178: 17-38" docOrigin="ZooKeys 1178" docPubDate="2023-09-01" docSource="http://dx.doi.org/10.3897/zookeys.1178.105703" docTitle="Cymbiapophysa homeroi Peñaherrera-R. 2023, sp. nov." docType="treatment" docUuid="B36F5C8C-9E34-4741-B70D-178D248FF378" docUuidSource="ZooBank" docVersion="1" id="26951753967254868073E2B13F9FDC60" lastPageNumber="17" masterDocId="26951753967254868073E2B13F9FDC60" masterDocTitle="Increasing knowledge of Cymbiapophysa Gabriel &amp; Sherwood, 2020 (Araneae, Theraphosidae): general distribution, key to species, and three new species from Ecuador" masterLastPageNumber="38" masterPageNumber="17" pageNumber="17" updateTime="1693642902469" updateUser="pensoft">
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<mods:title id="88A764B89D332D1D571582574DFE84BC">Increasing knowledge of Cymbiapophysa Gabriel &amp; Sherwood, 2020 (Araneae, Theraphosidae): general distribution, key to species, and three new species from Ecuador</mods:title>
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<mods:name id="EDF7EFC628E598FBE2735D60475A33E4" type="personal">
<mods:role id="1F7C376E2709D90E32F4C6D2C952B661">
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</mods:role>
<mods:namePart id="506F9B1AC90F34C0B1AAAE9ABA0E54FB">Penaherrera-R., Pedro</mods:namePart>
<mods:nameIdentifier id="34383901AB242C427DFC490BF25F9087" type="ORCID">https://orcid.org/0000-0001-9285-3403</mods:nameIdentifier>
<mods:affiliation id="517ECD72EAE580FB81129DCB79AB9B18">Universidad San Francisco de Quito USFQ, Colegio de Ciencias Biologicas y Ambientales, Instituto de Biodiversidad Tropical IBIOTROP, Laboratorio de Zoologia Terrestre, Museo de Zoologia, Quito 170901, Ecuador</mods:affiliation>
<mods:nameIdentifier id="B4C7A66529396D1B6C585EC5BCD1551D" type="email">pedropjpr5380@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="9FAFF2064751C564929AD18D3BDBB93D">text</mods:typeOfResource>
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<mods:title id="2886A917F4C17F26E0E126EB8785AE5D">ZooKeys</mods:title>
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<mods:part id="212771357A87FF771E19070ADC2AA5B8">
<mods:date id="6719E60C55C2A7C60AFFC407B75AD743">2023</mods:date>
<mods:detail id="C7607ABD7A906BBA1C75A90E53577DBE" type="pubDate">
<mods:number id="AA1DF159D1575566D953285445F93016">2023-09-01</mods:number>
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<mods:detail id="EAE78EA66554444656853BE2BC3009FF" type="volume">
<mods:number id="5EBDEFF62EE74B2AAD6CE25E5C9E0AF9">1178</mods:number>
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<mods:start id="87335EC59F687616593A4A4C7E71F05B">17</mods:start>
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<mods:classification id="0941EC3348638345A77FD470A80E21AA">journal article</mods:classification>
<mods:identifier id="FCCF66572C413BE663D7006BA8E2ECF4" type="DOI">http://dx.doi.org/10.3897/zookeys.1178.105703</mods:identifier>
<mods:identifier id="A9312D6D196045FD2BC4F45601826989" type="Pensoft-Pub">1313-2970-1178-17</mods:identifier>
<mods:identifier id="2FFD6EA42D71E8F9C7C95B7353E4384C" type="ZooBank">01B76CF7FF7A4CDFAAF5B4793EA932ED</mods:identifier>
<mods:identifier id="31F8F176D6E974292EF1D206541C56D8" type="Pensoft-UUID">26951753967254868073E2B13F9FDC60</mods:identifier>
</mods:mods>
<treatment id="18C560DFF8E157DCA32C32EADCE98A6C" LSID="urn:lsid:zoobank.org:act:B36F5C8C-9E34-4741-B70D-178D248FF378" httpUri="http://treatment.plazi.org/id/18C560DFF8E157DCA32C32EADCE98A6C" lastPageNumber="17" pageId="0" pageNumber="17">
<subSubSection id="04D3929464547A964C98ACBE8B3AAFF3" pageId="0" pageNumber="17" type="nomenclature">
<paragraph id="48FA2DA244214FF0B818E85B046DA487" pageId="0" pageNumber="17">
<taxonomicName id="5836C2EEA8564B995437763591EF27C7" LSID="https://zoobank.org/B36F5C8C-9E34-4741-B70D-178D248FF378" authority="Peñaherrera-R., 2023" authorityName="Peñaherrera-R." authorityYear="2023" class="Arachnida" family="Theraphosidae" genus="Cymbiapophysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymbiapophysa homeroi" order="Araneae" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="homeroi" status="sp. nov.">Cymbiapophysa homeroi</taxonomicName>
<taxonomicNameLabel id="1A0E27300A2607E104D397EAE849D40E" pageId="0" pageNumber="17">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="FB3C166D39DE3FB005EF0F8CD2FFE53C" pageId="0" pageNumber="17" type="description">
<paragraph id="8FF2855556A181A3D5961DBBFBDF66C5" pageId="0" pageNumber="17">
<figureCitation id="729FD10959E5C5A1CD603816F81CCFD2" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Leg I, metatarsus morphology, retrolateral view A Cymbiapophysa falconi sp. nov. male holotype (AE- 0005) B Cymbiapophysa homeroi sp. nov. male holotype (ZSFQ-i 11577) C Cymbiapophysa carmencita sp. nov. male holotype (ZSFQ-i 11578). Scale bars: 0.5 mm (A); 1 mm (B, C)." figureDoi="10.3897/zookeys.1178.105703.figure1" httpUri="https://binary.pensoft.net/fig/899940" pageId="0" pageNumber="17">Figs 1</figureCitation>
<figureCitation id="1CAFF80DBFE9A1092466D15BAA721462" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cymbiapophysa homeroi sp. nov. male holotype (ZSFQ-i 11577), palpal bulb (left hand side): A prolateral view B retrolateral view C dorsal view D ventral view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1178.105703.figure4" httpUri="https://binary.pensoft.net/fig/899943" pageId="0" pageNumber="17">, 4</figureCitation>
<figureCitation id="8E511132B7BE6304C46E25FB53B17B65" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Distribution of the genus Cymbiapophysa Gabriel &amp; Sherwood, 2020 including biogeographical regions of Ecuador and Colombia sensu Morrone 2014 and Cisneros-Heredia 2019. Blue circle = Valley of Yunguilla, type locality of C. falconi sp. nov.; Black circle = Reserve Rio Guajalito, type locality of C. homeroi sp. nov.; White circle = Padrewasi, type locality of C. carmencita sp. nov.; Yellow circle = Paramba, type locality of C. velox; Pink circle = Pambelar, historic record of C. velox; Dark green circle = Carondelet, type locality of C. seldeni; Light green circle = Reserva Natural Biotopo Selva Humeda, type locality of C. marimbai; Dark brown circle = Itsmina, possible occurrence of C. magna; Light brown circle = Andagoya, possible occurrence of C. magna." figureDoi="10.3897/zookeys.1178.105703.figure10" httpUri="https://binary.pensoft.net/fig/899949" pageId="0" pageNumber="17">, 10</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="6B56E33015C397E8F72B64B493F9A1E6" pageId="0" pageNumber="17" type="materials_examined">
<paragraph id="148A0F52DF655FF72E7064C658CAFB97" pageId="0" pageNumber="17">Examined material.</paragraph>
<paragraph id="6B01BEF19210E53010F843108C1C3C42" pageId="0" pageNumber="17">
<materialsCitation id="23F9E2968760F77B4FBA24B7FD702BB4" collectingDate="2015-11-26" collectorName="M. Costales" country="Republic of Ecuador" county="Province" elevation="2260" latitude="-0.2338" location="Parish of San Jose de Alluriquin" longLatPrecision="7" longitude="-78.7939" municipality="Canton Santo Domingo" specimenCount="1" specimenCount-male="1" stateProvince="Santo Domingo de los Tsachilas" typeStatus="Holotype">
<emphasis id="7F64BB94ABD47E5D26A789E58BAB98EE" bold="true" italics="true" pageId="0" pageNumber="17">
<typeStatus id="3E7C05C03063CF55217693FB0E87BDB9">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="BC3F9DF8735E973FE34AACC72F27D034" name="Ecuador">Republic of Ecuador</collectingCountry>
•
<specimenCount id="2692EBAFA25ABECCF65ED57127E63A5A" type="male">1 ♂</specimenCount>
;
<collectingCounty id="9E79451351B186FFAF46125FE140DE46">Province</collectingCounty>
of
<collectingRegion id="CBFF194F7C4E1F8A697DB42D97848FFE" country="Ecuador" name="Santo Domingo de los Tsachilas">Santo Domingo de los Tsachilas</collectingRegion>
,
<collectingMunicipality id="12CEE782AE2ED2F42ECE031FCA616EA6">Canton Santo Domingo</collectingMunicipality>
,
<location id="AF8C457322232DD354BC89A4128CF785" LSID="urn:lsid:plazi:treatment:18C560DFF8E157DCA32C32EADCE98A6C:AF8C457322232DD354BC89A4128CF785" country="Republic of Ecuador" county="Province" latitude="-0.2338" longLatPrecision="7" longitude="-78.7939" municipality="Canton Santo Domingo" name="Parish of San Jose de Alluriquin" stateProvince="Santo Domingo de los Tsachilas">
Parish of San
<normalizedToken id="77D0616DEE32B48AF23B7F6874A2A839" originalValue="José">Jose</normalizedToken>
de
<normalizedToken id="753789EFA2CDE2BB62893724D4899277" originalValue="Alluriquín">Alluriquin</normalizedToken>
</location>
,
<location id="A9AF074DCF5681A1393F40BB3C91A50D" LSID="urn:lsid:plazi:treatment:18C560DFF8E157DCA32C32EADCE98A6C:A9AF074DCF5681A1393F40BB3C91A50D" country="Republic of Ecuador" county="Province" latitude="-0.2338" longLatPrecision="7" longitude="-78.7939" municipality="Canton Santo Domingo" name="Reserve Rio Guajalito" stateProvince="Santo Domingo de los Tsachilas">
Reserve
<location id="2B2647E40FDC32397EC08F5F006E3306" LSID="urn:lsid:plazi:treatment:18C560DFF8E157DCA32C32EADCE98A6C:2B2647E40FDC32397EC08F5F006E3306" country="Republic of Ecuador" county="Province" latitude="-0.2338" longLatPrecision="7" longitude="-78.7939" municipality="Canton Santo Domingo" name="Rio Guajalito" stateProvince="Santo Domingo de los Tsachilas">Rio Guajalito</location>
</location>
, trail
<location id="CA5E7B439EA84A4F949328422972EA6A" LSID="urn:lsid:plazi:treatment:18C560DFF8E157DCA32C32EADCE98A6C:CA5E7B439EA84A4F949328422972EA6A" country="Republic of Ecuador" county="Province" latitude="-0.2338" longLatPrecision="7" longitude="-78.7939" municipality="Canton Santo Domingo" name="Los Espanoles" stateProvince="Santo Domingo de los Tsachilas">
Los
<normalizedToken id="CC26CAA40B645F248950B9B09B1617E5" originalValue="Españoles">Espanoles</normalizedToken>
</location>
;
<geoCoordinate id="6F140332E9B3A97F43527C90B0236907" degrees="0.2338" direction="south" orientation="latitude" precision="5" value="-0.2338">-0.2338</geoCoordinate>
,
<geoCoordinate id="691E3F9935B203DE4697C1358C100250" degrees="78.7939" direction="west" orientation="longitude" precision="5" value="-78.7939">-78.7939</geoCoordinate>
,
<elevation id="0E49815E3E781CC16A642F1F8BB53B46" metricMagnitude="3" metricUnit="m" metricValue="2.26" unit="m" value="2260.0">
<quantity id="68723D676ECFAB4D078DD2397921B44A" metricMagnitude="3" metricUnit="m" metricValue="2.26" unit="m" value="2260.0">2260 m</quantity>
a.s.l.
</elevation>
;
<collectingDate id="4DCDFE542BEE2BBB9E0E22007479F9FB" value="2015-11-26">26 November 2015</collectingDate>
;
<collectorName id="87941CFFFFE04891DBED3374D89E7D10">M. Costales</collectorName>
leg.; ZSFQ-i11577.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="926A21441EA832A3718D8FBC7B4E1C35" pageId="0" pageNumber="17" type="diagnosis">
<paragraph id="7515DC480D657186D4C9FF34AB12D393" pageId="0" pageNumber="17">Diagnosis.</paragraph>
<paragraph id="BBE2709BE6DB89E47F8AB4BA07BD42E7" pageId="0" pageNumber="17">
<taxonomicName id="9AA0D767A1E5259B3EFE9B1258ECE08B" class="Arachnida" family="Theraphosidae" genus="Cymbiapophysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymbiapophysa homeroi" order="Araneae" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="homeroi">
<emphasis id="E37BA54837055D2C65401AB01121228C" italics="true" pageId="0" pageNumber="17">Cymbiapophysa homeroi</emphasis>
</taxonomicName>
sp. nov. can be distinguished from other species by the morphology of the male palpal bulb from
<taxonomicName id="683E74075F80513B603B4E3A30BE1C26" lsidName="C. falconi" pageId="0" pageNumber="17" rank="species" species="falconi">
<emphasis id="8C387646A3D1CD4FC190C2E4C01406C5" italics="true" pageId="0" pageNumber="17">C. falconi</emphasis>
</taxonomicName>
sp. nov. by the presence of a continuous and slightly serrated PACK keel, PS keel as long as PI keel, and D developed (disjunct and distally slightly serrated PACK keel, PS keel longer than PI keel, and D weakly developed in
<taxonomicName id="0E9D19EFB92906B8FFCB98C60C749FBC" lsidName="C. falconi" pageId="0" pageNumber="17" rank="species" species="falconi">
<emphasis id="FF12C08D8FA79A9B2E9A063F82C5FF20" italics="true" pageId="0" pageNumber="17">C. falconi</emphasis>
</taxonomicName>
sp. nov.); from
<taxonomicName id="16AE82D5C1E95574263FD2A313B31A1D" lsidName="C. carmencita" pageId="0" pageNumber="17" rank="species" species="carmencita">
<emphasis id="A5F928B77083A4F7D29C7CCA7B15C310" italics="true" pageId="0" pageNumber="17">C. carmencita</emphasis>
</taxonomicName>
sp. nov. by the presence of a slightly serrated PACK keel and smooth PI keel, developed RS keel, weakly developed RI keel, weakly developed A keel, D developed, and absence of PAIK keel and Type III urticating setae (slightly serrated PACK and PI keels, absence of RS and RI keels, disjunct and developed A keel, developed PAIK keel, D weakly developed, and presence of Type III urticating setae in
<taxonomicName id="3AD6D735CE92C42619B2FF0C791C016E" lsidName="C. carmencita" pageId="0" pageNumber="17" rank="species" species="carmencita">
<emphasis id="9F6449966F5354FB440545083C68693B" italics="true" pageId="0" pageNumber="17">C. carmencita</emphasis>
</taxonomicName>
sp. nov.); from
<taxonomicName id="17C5D71C832742F90241707AB45B4572" lsidName="C. velox" pageId="0" pageNumber="17" rank="species" species="velox">
<emphasis id="D5E41CBE1A2733D440AF23A7366F5B77" italics="true" pageId="0" pageNumber="17">C. velox</emphasis>
</taxonomicName>
and
<taxonomicName id="5254D3FADD43D6AF53844D99892C4AC7" lsidName="C. yimana" pageId="0" pageNumber="17" rank="species" species="yimana">
<emphasis id="ECC6142073541C258A1B642F4CF373D0" italics="true" pageId="0" pageNumber="17">C. yimana</emphasis>
</taxonomicName>
by palpal bulb morphology with the presence of a continuous and slightly serrated PACK keel, D developed, and absence of Type III urticating setae (PACK keel(s) absent, D weakly developed, and presence of Type III urticating setae in
<taxonomicName id="B259FB79BB887CC17B0977355B66AF2A" lsidName="C. velox" pageId="0" pageNumber="17" rank="species" species="velox">
<emphasis id="81A76493DDA283E866A8FABA1C9FC287" italics="true" pageId="0" pageNumber="17">C. velox</emphasis>
</taxonomicName>
and
<taxonomicName id="B289734B6207C88653CE4F34500856BA" lsidName="C. yimana" pageId="0" pageNumber="17" rank="species" species="yimana">
<emphasis id="4311016F65775F5247CE66A16CC77D15" italics="true" pageId="0" pageNumber="17">C. yimana</emphasis>
</taxonomicName>
(
<bibRefCitation id="5C12C918E6FC1CB5155AD6314E399630" DOI="https://doi.org/10.13156/arac.2020.18.4.301" author="Gabriel, R" journalOrPublisher="Arachnology" pageId="0" pageNumber="17" pagination="301 - 316" refId="B17" refString="Gabriel, R, Sherwood, D, 2020. Revised taxonomic placement of Pseudhapalopus Strand, 1907, with notes on some related taxa (Araneae: Theraphosidae). Arachnology 18 (4): 301 - 316, DOI: https://doi.org/10.13156/arac.2020.18.4.301" title="Revised taxonomic placement of Pseudhapalopus Strand, 1907, with notes on some related taxa (Araneae: Theraphosidae)." url="https://doi.org/10.13156/arac.2020.18.4.301" volume="18" year="2020">Gabriel and Sherwood 2020</bibRefCitation>
;
<bibRefCitation id="0548C5FC7BA7B94DDEBBE080BB404238" DOI="https://doi.org/10.13156/arac.2021.18.8.838" author="Sherwood, D" journalOrPublisher="Arachnology" pageId="0" pageNumber="17" pagination="838 - 843" refId="B43" refString="Sherwood, D, Gabriel, R, Brescovit, AD, Lucas, SM, 2021a. A new species of Cymbiapophysa Gabriel &amp; Sherwood, 2020 from Colombia and a transfer from Proshapalopus Mello-Leitao, 1923 (Araneae: Theraphosidae). Arachnology 18 (8): 838 - 843, DOI: https://doi.org/10.13156/arac.2021.18.8.838" title="A new species of Cymbiapophysa Gabriel &amp; Sherwood, 2020 from Colombia and a transfer from Proshapalopus Mello-Leitao, 1923 (Araneae: Theraphosidae)." url="https://doi.org/10.13156/arac.2021.18.8.838" volume="18" year="2021 a">Sherwood et al. 2021a</bibRefCitation>
)); from
<taxonomicName id="052121263E483801606E92CA73CDE7A3" lsidName="C. marimbai" pageId="0" pageNumber="17" rank="species" species="marimbai">
<emphasis id="C46525D44D8D60B2DD1C8DA8716C7461" italics="true" pageId="0" pageNumber="17">C. marimbai</emphasis>
</taxonomicName>
,
<taxonomicName id="70209A002CC7E0E444DF1F834F13D4D8" lsidName="C. magna" pageId="0" pageNumber="17" rank="species" species="magna">
<emphasis id="19B39090F4F916EBEF5E4B64EFEE4042" italics="true" pageId="0" pageNumber="17">C. magna</emphasis>
</taxonomicName>
, and
<taxonomicName id="B57C2FDF38570C9C4259FF3DB043946F" lsidName="C. seldeni" pageId="0" pageNumber="17" rank="species" species="seldeni">
<emphasis id="71AB5097BFA65B35BB3A82329DB14355" italics="true" pageId="0" pageNumber="17">C. seldeni</emphasis>
</taxonomicName>
by the presence of a single slightly PACK keel and a RI keel, having the PS keel as long as PI keel, absence of a tibial apophysis, D developed, and absence of Type III urticating setae (two PACK keels present, RI keel absent, PS keel longer than PI, tibial apophysis present, D well-developed, and presence of Type III urticating setae in
<taxonomicName id="80F97BB532ACCCAC5E67D27F71F4D399" lsidName="C. marimbai" pageId="0" pageNumber="17" rank="species" species="marimbai">
<emphasis id="84BE106E9B2EDAC6F4396B3CB7C4B69A" italics="true" pageId="0" pageNumber="17">C. marimbai</emphasis>
</taxonomicName>
; PACK and RI keels absent, PS keel longer than PI, tibial apophysis present, D developed, and presence of Type III urticating setae in
<taxonomicName id="EBC5F7606D8C2B8A80836EF6EDF51FB7" lsidName="C. magna" pageId="0" pageNumber="17" rank="species" species="magna">
<emphasis id="AFFB549A879027CA389C3B9C74142FB5" italics="true" pageId="0" pageNumber="17">C. magna</emphasis>
</taxonomicName>
; PACK keel (s) absent, PS keel as long as PI keel, tibial apophysis present, and D weakly-developed in
<taxonomicName id="74D297FB1BBFAC45560F8CEC42E17E15" lsidName="C. seldeni" pageId="0" pageNumber="17" rank="species" species="seldeni">
<emphasis id="E421A5B1ACD7AEC798149D9387BF4A6F" italics="true" pageId="0" pageNumber="17">C. seldeni</emphasis>
</taxonomicName>
(
<bibRefCitation id="29C22B6D8DC704F8FF82AABC9A573B3A" DOI="https://doi.org/10.1080/03946975.2018.1493181" author="Perafan, C" journalOrPublisher="Tropical Zoology" pageId="0" pageNumber="17" pagination="200 - 213" refId="B32" refString="Perafan, C, Valencia-Cuellar, D, 2018. Proshapalopus marimbai, a new tarantula species (Mygalomorphae, Theraphosidae) and first genus record from Colombia. Tropical Zoology 31 (4): 200 - 213, DOI: https://doi.org/10.1080/03946975.2018.1493181" title="Proshapalopus marimbai, a new tarantula species (Mygalomorphae, Theraphosidae) and first genus record from Colombia." url="https://doi.org/10.1080/03946975.2018.1493181" volume="31" year="2018">
<normalizedToken id="538EEF745A22960FFE77D050B5A01FCE" originalValue="Perafán">Perafan</normalizedToken>
and
<normalizedToken id="11EA30A236E557A5460235CFAA8AD737" originalValue="Valencia-Cuéllar">Valencia-Cuellar</normalizedToken>
2018
</bibRefCitation>
;
<bibRefCitation id="2E2CF9306EF74325BB5115B4269F0CB0" DOI="https://doi.org/10.13156/arac.2021.18.8.838" author="Sherwood, D" journalOrPublisher="Arachnology" pageId="0" pageNumber="17" pagination="838 - 843" refId="B43" refString="Sherwood, D, Gabriel, R, Brescovit, AD, Lucas, SM, 2021a. A new species of Cymbiapophysa Gabriel &amp; Sherwood, 2020 from Colombia and a transfer from Proshapalopus Mello-Leitao, 1923 (Araneae: Theraphosidae). Arachnology 18 (8): 838 - 843, DOI: https://doi.org/10.13156/arac.2021.18.8.838" title="A new species of Cymbiapophysa Gabriel &amp; Sherwood, 2020 from Colombia and a transfer from Proshapalopus Mello-Leitao, 1923 (Araneae: Theraphosidae)." url="https://doi.org/10.13156/arac.2021.18.8.838" volume="18" year="2021 a">Sherwood et al. 2021a</bibRefCitation>
;
<bibRefCitation id="E2C0036DFB5B2D38FD3004F01DCCC9F4" DOI="https://doi.org/10.13156/arac.2023.19.5.777" author="Sherwood, D" journalOrPublisher="Arachnology" pageId="0" pageNumber="17" pagination="777 - 779" refId="B42" refString="Sherwood, D, Gabriel, R, 2023. Cymbiapophysa seldeni, a new theraphosine species from Ecuador (Araneae: Theraphosidae). Arachnology 19 (5): 777 - 779, DOI: https://doi.org/10.13156/arac.2023.19.5.777" title="Cymbiapophysa seldeni, a new theraphosine species from Ecuador (Araneae: Theraphosidae)." url="https://doi.org/10.13156/arac.2023.19.5.777" volume="19" year="2023">Sherwood and Gabriel 2023</bibRefCitation>
)). Additionally,
<taxonomicName id="3A5838BBD7830FB609ABE737262ECAAE" class="Arachnida" family="Theraphosidae" genus="Cymbiapophysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymbiapophysa homeroi" order="Araneae" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="homeroi">
<emphasis id="99C3EA24F23FE47C85A58DAB78B604DB" italics="true" pageId="0" pageNumber="17">Cymbiapophysa homeroi</emphasis>
</taxonomicName>
sp. nov. can further be distinguished from all other species, by the exception of
<taxonomicName id="AE6148D34E3A84D6B4F5BDE445A33A12" lsidName="C. carmencita" pageId="0" pageNumber="17" rank="species" species="carmencita">
<emphasis id="ED34B4FE6DE8E316CD4B43BA9E79A066" italics="true" pageId="0" pageNumber="17">C. carmencita</emphasis>
</taxonomicName>
sp. nov., by the presence of a well-developed PI keel (developed PI keel in
<taxonomicName id="4075ED4656D1440B7E21FD8FED6590D9" lsidName="C. magna" pageId="0" pageNumber="17" rank="species" species="magna">
<emphasis id="7F1736848F4C36E54AFB9998CB52CED7" italics="true" pageId="0" pageNumber="17">C. magna</emphasis>
</taxonomicName>
,
<taxonomicName id="A5662D0D9D8C094C6003D72513C7D324" lsidName="C. marimbai" pageId="0" pageNumber="17" rank="species" species="marimbai">
<emphasis id="C5B566F13F929466E8F6A5767B73F37A" italics="true" pageId="0" pageNumber="17">C. marimbai</emphasis>
</taxonomicName>
,
<taxonomicName id="7957D4A459B7F831383DDCE24500A147" lsidName="C. seldeni" pageId="0" pageNumber="17" rank="species" species="seldeni">
<emphasis id="4E6D325E184501896EA4DFF2EEAE9514" italics="true" pageId="0" pageNumber="17">C. seldeni</emphasis>
</taxonomicName>
, and
<taxonomicName id="C1D43711AE53805810FC8289247CC049" lsidName="C. yimana" pageId="0" pageNumber="17" rank="species" species="yimana">
<emphasis id="BE89B19740C357D4750ED80A327EA024" italics="true" pageId="0" pageNumber="17">C. yimana</emphasis>
</taxonomicName>
; weakly developed RI keel in
<taxonomicName id="84F1E000A7BC1AC6CFAD9E905B220B66" lsidName="C. falconi" pageId="0" pageNumber="17" rank="species" species="falconi">
<emphasis id="491933F048A57F83205AC80EF90DF228" italics="true" pageId="0" pageNumber="17">C. falconi</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName id="F55EB6CE56566048C8B96C5872A04E93" lsidName="C. velox" pageId="0" pageNumber="17" rank="species" species="velox">
<emphasis id="19CF9A107F26E3AA1E438B90BD43ADDC" italics="true" pageId="0" pageNumber="17">C. velox</emphasis>
</taxonomicName>
(
<bibRefCitation id="34D681CB47929F5969957658B3CED87B" DOI="https://doi.org/10.1080/03946975.2018.1493181" author="Perafan, C" journalOrPublisher="Tropical Zoology" pageId="0" pageNumber="17" pagination="200 - 213" refId="B32" refString="Perafan, C, Valencia-Cuellar, D, 2018. Proshapalopus marimbai, a new tarantula species (Mygalomorphae, Theraphosidae) and first genus record from Colombia. Tropical Zoology 31 (4): 200 - 213, DOI: https://doi.org/10.1080/03946975.2018.1493181" title="Proshapalopus marimbai, a new tarantula species (Mygalomorphae, Theraphosidae) and first genus record from Colombia." url="https://doi.org/10.1080/03946975.2018.1493181" volume="31" year="2018">
<normalizedToken id="73A786ACC52F8F68A9A20FE0EABEEC0F" originalValue="Perafán">Perafan</normalizedToken>
and
<normalizedToken id="FDACB91863F72809F85EC5918B2DBE2B" originalValue="Valencia-Cuéllar">Valencia-Cuellar</normalizedToken>
2018
</bibRefCitation>
;
<bibRefCitation id="B386129E8639F8D7B8418F581DB589FC" DOI="https://doi.org/10.13156/arac.2020.18.4.301" author="Gabriel, R" journalOrPublisher="Arachnology" pageId="0" pageNumber="17" pagination="301 - 316" refId="B17" refString="Gabriel, R, Sherwood, D, 2020. Revised taxonomic placement of Pseudhapalopus Strand, 1907, with notes on some related taxa (Araneae: Theraphosidae). Arachnology 18 (4): 301 - 316, DOI: https://doi.org/10.13156/arac.2020.18.4.301" title="Revised taxonomic placement of Pseudhapalopus Strand, 1907, with notes on some related taxa (Araneae: Theraphosidae)." url="https://doi.org/10.13156/arac.2020.18.4.301" volume="18" year="2020">Gabriel and Sherwood 2020</bibRefCitation>
;
<bibRefCitation id="8043D94C3D2DB03D96D3603A0B55DD31" DOI="https://doi.org/10.13156/arac.2021.18.8.838" author="Sherwood, D" journalOrPublisher="Arachnology" pageId="0" pageNumber="17" pagination="838 - 843" refId="B43" refString="Sherwood, D, Gabriel, R, Brescovit, AD, Lucas, SM, 2021a. A new species of Cymbiapophysa Gabriel &amp; Sherwood, 2020 from Colombia and a transfer from Proshapalopus Mello-Leitao, 1923 (Araneae: Theraphosidae). Arachnology 18 (8): 838 - 843, DOI: https://doi.org/10.13156/arac.2021.18.8.838" title="A new species of Cymbiapophysa Gabriel &amp; Sherwood, 2020 from Colombia and a transfer from Proshapalopus Mello-Leitao, 1923 (Araneae: Theraphosidae)." url="https://doi.org/10.13156/arac.2021.18.8.838" volume="18" year="2021 a">Sherwood et al. 2021a</bibRefCitation>
;
<bibRefCitation id="0B348A50A9EC5315A429EB6D935FBA7A" DOI="https://doi.org/10.13156/arac.2023.19.5.777" author="Sherwood, D" journalOrPublisher="Arachnology" pageId="0" pageNumber="17" pagination="777 - 779" refId="B42" refString="Sherwood, D, Gabriel, R, 2023. Cymbiapophysa seldeni, a new theraphosine species from Ecuador (Araneae: Theraphosidae). Arachnology 19 (5): 777 - 779, DOI: https://doi.org/10.13156/arac.2023.19.5.777" title="Cymbiapophysa seldeni, a new theraphosine species from Ecuador (Araneae: Theraphosidae)." url="https://doi.org/10.13156/arac.2023.19.5.777" volume="19" year="2023">Sherwood and Gabriel 2023</bibRefCitation>
)).
</paragraph>
</subSubSection>
<subSubSection id="4BB0C2EDEAAEE3BB52A58C54092AC006" pageId="0" pageNumber="17" type="description">
<paragraph id="A3A63F6A93D4E3FB9FD697556D7A1E8A" pageId="0" pageNumber="17">Description.</paragraph>
<paragraph id="0F9FC61DD7051C2603480762D384D67B" pageId="0" pageNumber="17">
<emphasis id="7BB0C1AE7CD3FE435869466FDFA74D8F" bold="true" italics="true" pageId="0" pageNumber="17">Male holotype</emphasis>
(ZSFQ-i11577): Total length including chelicerae: 24.75. Carapace: length 12.95, width 11.10. Caput: slightly raised. Ocular tubercle: raised, length 1.14, width 2.05. Eyes: AME&gt; ALE, AME&gt; PLE, PLE&gt; PME, anterior eye row straight, posterior row slightly recurved. Clypeus: narrow; clypeal fringe short. Fovea: straight. Chelicera: length 2.14, width 2.23. Abdomen: length 9.66, width 5.76. Maxilla with 112-146 cuspules covering approximately 60% of the proximal edge. Labium: length 1.80, width 2.00, with 52 cuspules most separated by 0.5-1.0
<normalizedToken id="CB48CD8E37411E2CD75A9B494AAAD0EB" originalValue="×">x</normalizedToken>
the width of a cuspule. Labio-sternal mounds joined along the entire base of the labium. Sternum: length 4.87, width 4.89, with three pairs of sigilla. Scopulation, leg measurements and spination unknown due to the fragile state of the specimen. Palpal cymbium with conical well-developed retrolateral apophysis (Fig.
<figureCitation id="D8EAF418DD9D9A280933859273AC6363" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Leg I, metatarsus morphology, retrolateral view A Cymbiapophysa falconi sp. nov. male holotype (AE- 0005) B Cymbiapophysa homeroi sp. nov. male holotype (ZSFQ-i 11577) C Cymbiapophysa carmencita sp. nov. male holotype (ZSFQ-i 11578). Scale bars: 0.5 mm (A); 1 mm (B, C)." figureDoi="10.3897/zookeys.1178.105703.figure1" httpUri="https://binary.pensoft.net/fig/899940" pageId="0" pageNumber="17">1B</figureCitation>
). Leg I lacking tibial apophyses. Femur III and palpal tibia laterally incrassate. Metatarsus I straight. Palpal bulb (Fig.
<figureCitation id="EB16F3A2E12BB42A61367186099AF0F4" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cymbiapophysa homeroi sp. nov. male holotype (ZSFQ-i 11577), palpal bulb (left hand side): A prolateral view B retrolateral view C dorsal view D ventral view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1178.105703.figure4" httpUri="https://binary.pensoft.net/fig/899943" pageId="0" pageNumber="17">4</figureCitation>
) with developed and triangular TH. PS, PACK, and PI keels well-developed; RS keel developed; RI and A keels weakly developed. PS as long as PI keel. Small and slightly serrated PACK keel extending less than a half the length of the embolus. PI visible in retrolateral and prolateral views. ER disjunct from PC forming a PR, PAR absent. PC narrow and slightly constricted in posterior half. D developed. Ventral face with rugulose area. Type Ia urticating setae present, Type III urticating setae absent. Colour: after thirteen years in preservative, generally with a brown colouration and setae with a pale grey colour.
</paragraph>
<caption id="459DF66FB2DA2DA200DDB118E8994E37" doi="10.3897/zookeys.1178.105703.figure4" httpUri="https://binary.pensoft.net/fig/899943" pageId="0" pageNumber="17" start="Figure 4" startId="F4">
<paragraph id="D734D4C23DD90153D2565847A57CA7C7" pageId="0" pageNumber="17">
<emphasis id="1584398652BA87252DF6D302D1F6246D" bold="true" pageId="0" pageNumber="17">Figure 4.</emphasis>
<taxonomicName id="7D0F3A61320A2E6A50F8FB628D4F1AA9" class="Arachnida" family="Theraphosidae" genus="Cymbiapophysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymbiapophysa homeroi" order="Araneae" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="homeroi">
<emphasis id="B788AB41D99008A157B0274A9D59B9F3" italics="true" pageId="0" pageNumber="17">Cymbiapophysa homeroi</emphasis>
</taxonomicName>
sp. nov. male holotype (ZSFQ-i11577), palpal bulb (left hand side):
<emphasis id="51801B1C70F23E57D85E992965D5B45A" bold="true" pageId="0" pageNumber="17">A</emphasis>
prolateral view
<emphasis id="6B1E01D6B382016FC430D506DACC371C" bold="true" pageId="0" pageNumber="17">B</emphasis>
retrolateral view
<emphasis id="0191F0C7E3F735089777095044361C71" bold="true" pageId="0" pageNumber="17">C</emphasis>
dorsal view
<emphasis id="2C3CB57971EDC28B30FF71D9C3E9E86E" bold="true" pageId="0" pageNumber="17">D</emphasis>
ventral view. Scale bars: 0.5 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="6095EF5F7EC3252E4E77BBE429FEF755" pageId="0" pageNumber="17" type="etymology">
<paragraph id="22658472F89128F020F64E093CA6C26D" pageId="0" pageNumber="17">Etymology.</paragraph>
<paragraph id="E63782AF387807B1436A51DFCD2E1D3F" pageId="0" pageNumber="17">
The specific epithet is a patronym for my father, Homero Giovanni
<normalizedToken id="F82E14BFDEA6B704F3D858F5101C9E79" originalValue="Peñaherrera">Penaherrera</normalizedToken>
Zavala, who has always been a main pillar of my life, and always supported my crazy ideas like keeping aquariums and unconventional animals inside the house, which helped me grow my curiosity about nature.
</paragraph>
</subSubSection>
<subSubSection id="9CCCE54967B2D488DD8351576A2CFB80" pageId="0" pageNumber="17" type="distribution">
<paragraph id="9FC2FC98321647BDD2BBC5F9B62D5621" pageId="0" pageNumber="17">Distribution.</paragraph>
<paragraph id="E0AE4825B7F028045A92CC7E49635450" pageId="0" pageNumber="17">
<taxonomicName id="54C77D9308800843745E3F95275006F9" class="Arachnida" family="Theraphosidae" genus="Cymbiapophysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymbiapophysa homeroi" order="Araneae" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="homeroi">
<emphasis id="4D516A554458BFDD44184B668C27F363" italics="true" pageId="0" pageNumber="17">Cymbiapophysa homeroi</emphasis>
</taxonomicName>
sp. nov. is only known from its type locality, on the Reserve Rio Guajalito, alongside the trail Los
<normalizedToken id="D05472FC373EC9E5741C636AFBE74BE5" originalValue="Españoles">Espanoles</normalizedToken>
, Province of Santo Domingo de los Tsachilas, north of the western mountain range of the Andean Cordillera of Ecuador, at 2261 m.
</paragraph>
</subSubSection>
<subSubSection id="5187505B82400939D1F2BD5E81670D68" pageId="0" pageNumber="17" type="ecology">
<paragraph id="62081E2F69DFE0D6418A0BE138FE631A" pageId="0" pageNumber="17">Ecology.</paragraph>
<paragraph id="A05C52D7CC293CCF274C0A724F380DE5" pageId="0" pageNumber="17">
The holotype of
<taxonomicName id="D5A861ED71A1DA9E72C382F8695198AD" class="Arachnida" family="Theraphosidae" genus="Cymbiapophysa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymbiapophysa homeroi" order="Araneae" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="homeroi">
<emphasis id="C5C32FF5E6525A32834F856EEC6C599F" italics="true" pageId="0" pageNumber="17">Cymbiapophysa homeroi</emphasis>
</taxonomicName>
sp. nov. was collected in the low montane evergreen forest of the Cordillera Occidental of the Andes of Ecuador, in the Northern Andes biogeographic province (Fig.
<figureCitation id="C454E4142025C9131995138B0FF3E484" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Distribution of the genus Cymbiapophysa Gabriel &amp; Sherwood, 2020 including biogeographical regions of Ecuador and Colombia sensu Morrone 2014 and Cisneros-Heredia 2019. Blue circle = Valley of Yunguilla, type locality of C. falconi sp. nov.; Black circle = Reserve Rio Guajalito, type locality of C. homeroi sp. nov.; White circle = Padrewasi, type locality of C. carmencita sp. nov.; Yellow circle = Paramba, type locality of C. velox; Pink circle = Pambelar, historic record of C. velox; Dark green circle = Carondelet, type locality of C. seldeni; Light green circle = Reserva Natural Biotopo Selva Humeda, type locality of C. marimbai; Dark brown circle = Itsmina, possible occurrence of C. magna; Light brown circle = Andagoya, possible occurrence of C. magna." figureDoi="10.3897/zookeys.1178.105703.figure10" httpUri="https://binary.pensoft.net/fig/899949" pageId="0" pageNumber="17">10</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="767F26964D205D33EF2CAC0F2F7578A5" pageId="0" pageNumber="17" type="remarks">
<paragraph id="D213FCD28E6A0F601DC002622A42F7D0" pageId="0" pageNumber="17">Remarks.</paragraph>
<paragraph id="F74360B8D5F5A1BB1CDB7AF1653EC463" pageId="0" pageNumber="17">
The specimen is in a fragile state; thus, I did not measure scopulation, leg lengths, leg spination, and posterior lateral spinnerets segments to prevent fragmentation and loss of leg and pedipalp segments. Nevertheless, the most important characteristics to diagnose this new species are presented. The type specimen was found in the didactic collection of the ZSFQ and the original specimen label indicates &quot;Guajalito. M.
<normalizedToken id="8E065DF45746694365621E9B28B92A51" originalValue="Costales’’">Costales''</normalizedToken>
. This specimen was collected by M. Costales during a student excursion to the Guajalito Reserve, managed by Vlastimil Zak, professor of Botany, in 2016 (Diego F. Cisneros-Heredia, in litt. 2023).
</paragraph>
</subSubSection>
</treatment>
</document>
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