629 lines
85 KiB
XML
629 lines
85 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zse.97.67788" ID-GBIF-Dataset="4853559a-b2ce-4159-aaf1-a2625d333a25" ID-GBIF-Taxon="183822032" ID-Pensoft-Pub="1860-0743-2-393" ID-Pensoft-UUID="564A467951EF5C8A883587B138B7809E" ID-ZooBank="A32F2CCD3CB8457087D13B38CA555E3C" ModsDocID="1860-0743-97-2-393" checkinTime="1627415565744" checkinUser="pensoft" docAuthor="Framenau, Volker W., Vink, Cor J., Scharff, Nikolaj, Baptista, Renner L. C. & Castanheira, Pedro de S." docDate="2021" docId="A9F43A95863055BA9986778DACF539B1" docLanguage="en" docName="ZoosystEvol 97(2): 393-405" docOrigin="Zoosystematics and Evolution 97 (2)" docPubDate="2021-07-26" docSource="http://dx.doi.org/10.3897/zse.97.67788" docTitle="Novakiella Court & Forster, 1993, in Platnick 1993" docType="treatment" docVersion="3" id="564A467951EF5C8A883587B138B7809E" lastPageNumber="393" masterDocId="564A467951EF5C8A883587B138B7809E" masterDocTitle="Review of the Australian and New Zealand orb-weaving spider genus Novakiella (Araneae, Araneidae)" masterLastPageNumber="405" masterPageNumber="393" pageId="0" pageNumber="393" updateTime="1643429406696" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>Review of the Australian and New Zealand orb-weaving spider genus Novakiella (Araneae, Araneidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Framenau, Volker W.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7724-3831</mods:nameIdentifier>
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<mods:affiliation>Harry Butler Institute, Murdoch University, 90 South St, Murdoch, Western Australia 6150, Australia & Department of Terrestrial Zoology, Western Australian Museum, Locked Bag 49, Welshpool DC, Western Australia 6986, Australia & Centrum fuer Naturkunde (CeNak), Universitaet Hamburg, Martin-Luther-King-Platz 3, 20146, Hamburg, Germany</mods:affiliation>
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<mods:nameIdentifier type="email">volker.framenau@murdoch.edu.au</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Vink, Cor J.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4237-0117</mods:nameIdentifier>
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<mods:affiliation>Department of Pest-management and Conservation, PO Box 85084, Lincoln University, Lincoln 7647, Christchurch, New Zealand & Centrum fuer Naturkunde (CeNak), Universitaet Hamburg, Martin-Luther-King-Platz 3, 20146, Hamburg, Germany</mods:affiliation>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Scharff, Nikolaj</mods:namePart>
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<mods:affiliation>Zoology Section, Research and Collections, Natural History Museum of Denmark, University of Copenhagen, Copenhagen, Denmark</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Baptista, Renner L. C.</mods:namePart>
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<mods:affiliation>Laboratorio de Diversidade de Aracnideos, Universidade do Brasil / Universidade Federal do Rio de Janeiro. Av. Carlos Chagas Filho 373, 21941 - 902, Ilha do Fundao, Rio de Janeiro, Brazil</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Castanheira, Pedro de S.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-0623-1622</mods:nameIdentifier>
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<mods:affiliation>Harry Butler Institute, Murdoch University, 90 South St, Murdoch, Western Australia 6150, Australia & Laboratorio de Diversidade de Aracnideos, Universidade do Brasil / Universidade Federal do Rio de Janeiro. Av. Carlos Chagas Filho 373, 21941 - 902, Ilha do Fundao, Rio de Janeiro, Brazil</mods:affiliation>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Zoosystematics and Evolution</mods:title>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-07-26</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>97</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>2</mods:number>
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<mods:extent unit="page">
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<mods:start>393</mods:start>
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<mods:end>405</mods:end>
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</mods:extent>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zse.97.67788</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zse.97.67788</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1860-0743-2-393</mods:identifier>
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<mods:identifier type="ZooBank">A32F2CCD3CB8457087D13B38CA555E3C</mods:identifier>
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<mods:identifier type="Pensoft-UUID">564A467951EF5C8A883587B138B7809E</mods:identifier>
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<treatment ID-GBIF-Taxon="183822032" LSID="urn:lsid:plazi:treatment:A9F43A95863055BA9986778DACF539B1" httpUri="http://treatment.plazi.org/id/A9F43A95863055BA9986778DACF539B1" lastPageNumber="393" pageId="0" pageNumber="393">
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<subSubSection pageId="0" pageNumber="393" type="nomenclature">
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<paragraph pageId="0" pageNumber="393">
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Genus
|
||
<taxonomicName LSID="A9F43A95-8630-55BA-9986-778DACF539B1" authority="Court & Forster, 1993, in Platnick (1993)" authorityName="Court & Forster, 1993, in Platnick" authorityYear="1993" class="Arachnida" family="Araneidae" genus="Novakiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Novakiella" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="genus">Novakiella Court & Forster, 1993, in Platnick (1993)</taxonomicName>
|
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</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="393" type="reference_group">
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||
<paragraph pageId="0" pageNumber="393">
|
||
<taxonomicName class="Insecta" family="Araneidae" genus="Novakia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Novakia" order="Araneae" pageId="0" pageNumber="393" phylum="Mollusca" rank="genus">Novakia</taxonomicName>
|
||
Court & Forster, 1988: junior homonym of
|
||
<taxonomicName class="Insecta" family="Araneidae" genus="Novakia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Novakia" order="Araneae" pageId="0" pageNumber="393" phylum="Mollusca" rank="genus">Novakia</taxonomicName>
|
||
Strobl, 1893 (
|
||
<taxonomicName class="Insecta" family="Araneidae" genus="Novakia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Diptera" pageId="0" pageNumber="393" phylum="Mollusca" rank="order">Diptera</taxonomicName>
|
||
) and
|
||
<taxonomicName class="Insecta" family="Araneidae" genus="Novakia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Novakia" order="Araneae" pageId="0" pageNumber="393" phylum="Mollusca" rank="genus">Novakia</taxonomicName>
|
||
Tolmachoff, 1926 (
|
||
<taxonomicName family="Araneidae" genus="Novakia" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Araneae" pageId="0" pageNumber="393" phylum="Mollusca" rank="phylum">Mollusca</taxonomicName>
|
||
) is another junior homonym.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="393" type="type-species">
|
||
<paragraph pageId="0" pageNumber="393">Type-species.</paragraph>
|
||
<paragraph pageId="0" pageNumber="393">
|
||
<taxonomicName authority="Walckenaer, 1805" authorityName="Walckenaer" authorityYear="1805" class="Arachnida" family="Araneidae" genus="Epeira" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Epeira tri-tuberculata" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="species" species="tri-tuberculata">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Epeira tri-tuberculata</emphasis>
|
||
</taxonomicName>
|
||
Urquhart, 1887.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="393" type="diagnosis">
|
||
<paragraph pageId="0" pageNumber="393">Diagnosis.</paragraph>
|
||
<paragraph pageId="0" pageNumber="393">
|
||
The informal clade of the backobourkiines is well supported by the molecular phylogeny of
|
||
<bibRefCitation DOI="https://doi.org/10.1111/cla.12382" author="Scharff, N" journalOrPublisher="Cladistics" pageId="0" pageNumber="393" pagination="1 - 21" refId="B16" refString="Scharff, N, Coddington, JA, Blackledge, TA, Agnarsson, I, Framenau, VW, Szűts, T, Hayashi, CY, Dimitrov, D, 2020. Phylogeny of the orb-weaving spider family Araneidae (Araneae: Araneoidea). Cladistics 36 (1): 1 - 21, DOI: https://doi.org/10.1111/cla.12382" title="Phylogeny of the orb-weaving spider family Araneidae (Araneae: Araneoidea)." url="https://doi.org/10.1111/cla.12382" volume="36" year="2020">Scharff et al. (2020)</bibRefCitation>
|
||
, but the taxonomy and systematics of the species and genera within this clade are poorly resolved. Only three genera within the clade have been revised using modern taxonomic methods:
|
||
<taxonomicName authorityName="Joseph & Framenau" authorityYear="2012" class="Arachnida" family="Araneidae" genus="Plebs" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Plebs" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Plebs</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName class="Arachnida" family="Araneidae" genus="Backobourkia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Backobourkia" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Backobourkia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Framenau" authorityYear="2011" class="Arachnida" family="Araneidae" genus="Lariniophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lariniophora" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Lariniophora</emphasis>
|
||
</taxonomicName>
|
||
Framenau, 2011. The genera
|
||
<taxonomicName authorityName="L.Koch" authorityYear="1872" class="Arachnida" family="Araneidae" genus="Carepalxis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Carepalxis" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Carepalxis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Karsch" authorityYear="1878" class="Arachnida" family="Araneidae" genus="Acroaspis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Acroaspis" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Acroaspis</emphasis>
|
||
</taxonomicName>
|
||
have not been revised and their putative synapomorphies remain unknown. It is therefore difficult to diagnose
|
||
<taxonomicName authorityName="Court & Forster, 1993, in Platnick" authorityYear="1993" class="Arachnida" family="Araneidae" genus="Novakiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Novakiella" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Novakiella</emphasis>
|
||
</taxonomicName>
|
||
against these genera. Other Australian backobourkiines included in
|
||
<bibRefCitation DOI="https://doi.org/10.1111/cla.12382" author="Scharff, N" journalOrPublisher="Cladistics" pageId="0" pageNumber="393" pagination="1 - 21" refId="B16" refString="Scharff, N, Coddington, JA, Blackledge, TA, Agnarsson, I, Framenau, VW, Szűts, T, Hayashi, CY, Dimitrov, D, 2020. Phylogeny of the orb-weaving spider family Araneidae (Araneae: Araneoidea). Cladistics 36 (1): 1 - 21, DOI: https://doi.org/10.1111/cla.12382" title="Phylogeny of the orb-weaving spider family Araneidae (Araneae: Araneoidea)." url="https://doi.org/10.1111/cla.12382" volume="36" year="2020">Scharff et al. (2020)</bibRefCitation>
|
||
represent species that have clearly been misplaced in genera they do not belong to (i.e.,
|
||
<taxonomicName class="Arachnida" family="Araneidae" genus="Eriophora" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eriophora" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Eriophora</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName class="Arachnida" family="Araneidae" genus="Araneus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Araneus" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Araneus</emphasis>
|
||
</taxonomicName>
|
||
) and these represent undescribed genera (in that study listed as
|
||
<normalizedToken originalValue="“NGEN01”">"NGEN01"</normalizedToken>
|
||
for
|
||
<taxonomicName baseAuthorityName="Keyserling" baseAuthorityYear="1865" class="Arachnida" family="Araneidae" genus="Eriophora" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eriophora transmarina" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="species" species="transmarina">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Eriophora transmarina</emphasis>
|
||
</taxonomicName>
|
||
(Keyserling, 1865),
|
||
<normalizedToken originalValue="“NGEN02”">"NGEN02"</normalizedToken>
|
||
for
|
||
<taxonomicName baseAuthorityName="Main" baseAuthorityYear="1954" class="Arachnida" family="Araneidae" genus="Araneus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Araneus recherchensis" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="species" species="recherchensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Araneus recherchensis</emphasis>
|
||
</taxonomicName>
|
||
Main, 1954 and
|
||
<normalizedToken originalValue="“NGEN05”">"NGEN05"</normalizedToken>
|
||
for
|
||
<taxonomicName authorityName="Simon" authorityYear="1908" class="Arachnida" family="Araneidae" genus="Araneus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Araneus senicaudatus" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="species" species="senicaudatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Araneus senicaudatus</emphasis>
|
||
</taxonomicName>
|
||
Simon, 1908). Until these species have been revised and placed in new or existing genera,
|
||
<taxonomicName authorityName="Court & Forster, 1993, in Platnick" authorityYear="1993" class="Arachnida" family="Araneidae" genus="Novakiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Novakiella" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Novakiella</emphasis>
|
||
</taxonomicName>
|
||
cannot be diagnosed against them.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="393">
|
||
<taxonomicName authorityName="Court & Forster, 1993, in Platnick" authorityYear="1993" class="Arachnida" family="Araneidae" genus="Novakiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Novakiella" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Novakiella</emphasis>
|
||
</taxonomicName>
|
||
distinctly differs from the revised backobourkiine genera by overall somatic morphology. The abdomen is subtriangular with strong humeral humps (Figs
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Novakiella trituberculosa, male (WAM T 73571). A. Dorsal habitus; B. ventral habitus; C. Left pedipalp, mesal view; D. Left pedipalp, ventral view. Abbreviations: C, conductor; CL, conductor lobe; E, embolus; MA, median apophysis; P, paracymbium; Ra, radix; TA, terminal apophysis. Scale bars: A, B, 1 mm; C, D, 0.5 mm; E, 1.2 mm." figureDoi="10.3897/zse.97.67788.figure1" httpUri="https://binary.pensoft.net/fig/570631" pageId="0" pageNumber="393">1A</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Novakiella trituberculosa, female (WAM T 73556). A. Dorsal habitus; B. Ventral habitus; C. Epigyne, ventral view; D. Cleared epigyne, ventral view; E. Cleared epigyne, dorsal view. Abbreviations: CO, copulatory openings; S, scape; Sp, spermatheca. Scale bars: A, B, 2 mm; C-E, 0.1 mm." figureDoi="10.3897/zse.97.67788.figure3" httpUri="https://binary.pensoft.net/fig/570633" pageId="0" pageNumber="393">3A</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Novakiella boletus sp. nov., male holotype (MV K 9867). A. Dorsal habitus; B. Ventral habitus; C. Left tibia, ventral view; D. Left pedipalp, mesal view; E. Left pedipalp, ventral view, F. left pedipalp, apical view. Abbreviations: C, conductor; CL, conductor lobe; E, embolus; MA, median apophysis; P, paracymbium; Ra, radix; TA, terminal apophysis. Scale bars: A, B, 2 mm; C, 1 mm; D-F, 0.5 mm." figureDoi="10.3897/zse.97.67788.figure4" httpUri="https://binary.pensoft.net/fig/570634" pageId="0" pageNumber="393">4A</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Novakiella boletus sp. nov., female (SAM). A. Dorsal habitus; B. Ventral habitus; C. Epigyne, ventral view. Abbreviations: CO, copulatory openings; L, lips; S, scape. Scale bars: A, B, 2 mm, C, 0.2 mm." figureDoi="10.3897/zse.97.67788.figure6" httpUri="https://binary.pensoft.net/fig/570636" pageId="0" pageNumber="393">6A</figureCitation>
|
||
), while it is rounded with small humerals in
|
||
<taxonomicName class="Arachnida" family="Araneidae" genus="Backobourkia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Backobourkia" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Backobourkia</emphasis>
|
||
</taxonomicName>
|
||
(i.e.,
|
||
<bibRefCitation author="Framenau, VW" journalOrPublisher="Arthropod Systematics & Phylogeny" pageId="0" pageNumber="393" pagination="79 - 111" refId="B5" refString="Framenau, VW, Duperre, N, Blackledge, TA, Vink, CJ, 2010. Systematics of the new Australasian orb-weaving spider genus Backobourkia (Araneae: Araneidae: Araneinae). Arthropod Systematics & Phylogeny 68: 79 - 111" title="Systematics of the new Australasian orb-weaving spider genus Backobourkia (Araneae: Araneidae: Araneinae)." volume="68" year="2010">Framenau et al. 2010</bibRefCitation>
|
||
, fig. 5), slightly elongated in
|
||
<taxonomicName authorityName="Joseph & Framenau" authorityYear="2012" class="Arachnida" family="Araneidae" genus="Plebs" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Plebs" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Plebs</emphasis>
|
||
</taxonomicName>
|
||
(e.g.
|
||
<bibRefCitation DOI="https://doi.org/10.1111/j.1096-3642.2012.00845.x" author="Joseph, MM" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="393" pagination="279 - 341" refId="B7" refString="Joseph, MM, Framenau, VW, 2012. Systematic review of a new orb-weaving spider genus (Araneae: Araneidae), with special reference to the Australasian-Pacific and South-East Asian fauna. Zoological Journal of the Linnean Society 166: 279 - 341, DOI: https://doi.org/10.1111/j.1096-3642.2012.00845.x" title="Systematic review of a new orb-weaving spider genus (Araneae: Araneidae), with special reference to the Australasian-Pacific and South-East Asian fauna." url="https://doi.org/10.1111/j.1096-3642.2012.00845.x" volume="166" year="2012">Joseph and Framenau 2012</bibRefCitation>
|
||
, figs 6, 7, 10), and strongly elongated in
|
||
<taxonomicName authorityName="Framenau" authorityYear="2011" class="Arachnida" family="Araneidae" genus="Lariniophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lariniophora" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Lariniophora</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation DOI="https://doi.org/10.18195/issn.0312-3162.26(2).2011.191-201" author="Framenau, VW" journalOrPublisher="Records of the Western Australian Museum" pageId="0" pageNumber="393" pagination="191 - 201" refId="B4" refString="Framenau, VW, 2011. Lariniophora, a new monotypic orb-weaving spider genus from Australia (Araneae: Araneidae: Araneinae). Records of the Western Australian Museum 26 (2): 191 - 201, DOI: https://doi.org/10.18195/issn.0312-3162.26(2).2011.191-201" title="Lariniophora, a new monotypic orb-weaving spider genus from Australia (Araneae: Araneidae: Araneinae)." url="https://doi.org/10.18195/issn.0312-3162.26(2).2011.191-201" volume="26" year="2011">Framenau 2011</bibRefCitation>
|
||
, figs 2, 3). Males can be differentiated by the presence of a tibial apico-prolateral spur carrying a thick spine (or macroseta) on leg II (Figs
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Novakiella trituberculosa, male (WAM T 73571). A. Dorsal habitus; B. ventral habitus; C. Left pedipalp, mesal view; D. Left pedipalp, ventral view. Abbreviations: C, conductor; CL, conductor lobe; E, embolus; MA, median apophysis; P, paracymbium; Ra, radix; TA, terminal apophysis. Scale bars: A, B, 1 mm; C, D, 0.5 mm; E, 1.2 mm." figureDoi="10.3897/zse.97.67788.figure1" httpUri="https://binary.pensoft.net/fig/570631" pageId="0" pageNumber="393">1E</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Novakiella boletus sp. nov., male holotype (MV K 9867). A. Dorsal habitus; B. Ventral habitus; C. Left tibia, ventral view; D. Left pedipalp, mesal view; E. Left pedipalp, ventral view, F. left pedipalp, apical view. Abbreviations: C, conductor; CL, conductor lobe; E, embolus; MA, median apophysis; P, paracymbium; Ra, radix; TA, terminal apophysis. Scale bars: A, B, 2 mm; C, 1 mm; D-F, 0.5 mm." figureDoi="10.3897/zse.97.67788.figure4" httpUri="https://binary.pensoft.net/fig/570634" pageId="0" pageNumber="393">4C</figureCitation>
|
||
).
|
||
<taxonomicName class="Arachnida" family="Araneidae" genus="Verrucosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Verrucosa" order="Araneida" pageId="0" pageNumber="393" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Verrucosa</emphasis>
|
||
</taxonomicName>
|
||
McCook, 1888 and
|
||
<taxonomicName authorityName="L.Koch" authorityYear="1872" class="Arachnida" family="Araneidae" genus="Carepalxis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Carepalxis" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Carepalxis</emphasis>
|
||
</taxonomicName>
|
||
also have a spur on leg II, but in both genera it carries two spines (
|
||
<bibRefCitation DOI="https://doi.org/10.1636/0161-8202(2002)030[0527:KTTGOA]2.0.CO;2" author="Levi, HW" journalOrPublisher="Bulletin Museum of Comparative Zoology" pageId="0" pageNumber="393" refId="B10" refString="Levi, HW, 2002. Keys to the genera of araneid orbweavers (Araneae, Araneidae) of the Americas. Journal of Arachnology 30: 527-562. https://doi.org/10.1636/0161-8202(2002)030[0527:KTTGOA]2.0.CO;2" title="Keys to the genera of araneid orbweavers (Araneae, Araneidae) of the Americas. Journal of Arachnology 30: 527 - 562." url="https://doi.org/10.1636/0161-8202(2002)030[0527:KTTGOA]2.0.CO;2" year="2002">Levi 2002</bibRefCitation>
|
||
: p. 546;
|
||
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.3921.1.1" author="Lise, AA" journalOrPublisher="Zootaxa" pageId="0" pageNumber="393" pagination="1 - 105" refId="B11" refString="Lise, AA, Kesster, CC, Silva, ELC da, 2015. Revision of the orb-weaving spider genus Verrucosa McCook, 1888 (Araneae, Araneidae). Zootaxa 3921: 1 - 105, DOI: https://doi.org/10.11646/zootaxa.3921.1.1" title="Revision of the orb-weaving spider genus Verrucosa McCook, 1888 (Araneae, Araneidae)." url="https://doi.org/10.11646/zootaxa.3921.1.1" volume="3921" year="2015">Lise et al. 2015</bibRefCitation>
|
||
: p. 5; VWF pers. obs.). In addition,
|
||
<taxonomicName class="Arachnida" family="Araneidae" genus="Verrucosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Verrucosa" order="Araneida" pageId="0" pageNumber="393" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Verrucosa</emphasis>
|
||
</taxonomicName>
|
||
is limited to the Neotropics and not part of the backobourkiines (
|
||
<bibRefCitation DOI="https://doi.org/10.1111/cla.12382" author="Scharff, N" journalOrPublisher="Cladistics" pageId="0" pageNumber="393" pagination="1 - 21" refId="B16" refString="Scharff, N, Coddington, JA, Blackledge, TA, Agnarsson, I, Framenau, VW, Szűts, T, Hayashi, CY, Dimitrov, D, 2020. Phylogeny of the orb-weaving spider family Araneidae (Araneae: Araneoidea). Cladistics 36 (1): 1 - 21, DOI: https://doi.org/10.1111/cla.12382" title="Phylogeny of the orb-weaving spider family Araneidae (Araneae: Araneoidea)." url="https://doi.org/10.1111/cla.12382" volume="36" year="2020">Scharff et al. 2020</bibRefCitation>
|
||
). There are distinct differences in the male pedipalp morphology between
|
||
<taxonomicName authorityName="Court & Forster, 1993, in Platnick" authorityYear="1993" class="Arachnida" family="Araneidae" genus="Novakiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Novakiella" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Novakiella</emphasis>
|
||
</taxonomicName>
|
||
and other backobourkiines.
|
||
<taxonomicName authorityName="Court & Forster, 1993, in Platnick" authorityYear="1993" class="Arachnida" family="Araneidae" genus="Novakiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Novakiella" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Novakiella</emphasis>
|
||
</taxonomicName>
|
||
males have a stout median apophysis that is drawn out into a basally pointing acute projection (Figs
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Novakiella trituberculosa, male (WAM T 73571). A. Dorsal habitus; B. ventral habitus; C. Left pedipalp, mesal view; D. Left pedipalp, ventral view. Abbreviations: C, conductor; CL, conductor lobe; E, embolus; MA, median apophysis; P, paracymbium; Ra, radix; TA, terminal apophysis. Scale bars: A, B, 1 mm; C, D, 0.5 mm; E, 1.2 mm." figureDoi="10.3897/zse.97.67788.figure1" httpUri="https://binary.pensoft.net/fig/570631" pageId="0" pageNumber="393">1C</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Novakiella trituberculosa, male (WAM T 73571), expanded left pedipalp. A. Ventral view; B. Retrolateral view. Abbreviations: C, conductor; CL, conductor lobe; E, embolus; MA, median apophysis; P, paracymbium; Ra, radix; TA, terminal apophysis. Scale bars: 0.5 mm" figureDoi="10.3897/zse.97.67788.figure2" httpUri="https://binary.pensoft.net/fig/570632" pageId="0" pageNumber="393">2B</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Novakiella boletus sp. nov., male holotype (MV K 9867). A. Dorsal habitus; B. Ventral habitus; C. Left tibia, ventral view; D. Left pedipalp, mesal view; E. Left pedipalp, ventral view, F. left pedipalp, apical view. Abbreviations: C, conductor; CL, conductor lobe; E, embolus; MA, median apophysis; P, paracymbium; Ra, radix; TA, terminal apophysis. Scale bars: A, B, 2 mm; C, 1 mm; D-F, 0.5 mm." figureDoi="10.3897/zse.97.67788.figure4" httpUri="https://binary.pensoft.net/fig/570634" pageId="0" pageNumber="393">4D, E</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Novakiella boletus sp. nov., male holotype (MV K 9867), expanded left pedipalp. mesal view. Abbreviations: C, conductor; CL, conductor lobe; E, embolus; MA, median apophysis; Ra, radix; TA, terminal apophysis. Scale bar: 0.5 mm." figureDoi="10.3897/zse.97.67788.figure5" httpUri="https://binary.pensoft.net/fig/570635" pageId="0" pageNumber="393">5</figureCitation>
|
||
); in contrast, the median apophysis has a basal flange in
|
||
<taxonomicName class="Arachnida" family="Araneidae" genus="Backobourkia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Backobourkia" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Backobourkia</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation author="Framenau, VW" journalOrPublisher="Arthropod Systematics & Phylogeny" pageId="0" pageNumber="393" pagination="79 - 111" refId="B5" refString="Framenau, VW, Duperre, N, Blackledge, TA, Vink, CJ, 2010. Systematics of the new Australasian orb-weaving spider genus Backobourkia (Araneae: Araneidae: Araneinae). Arthropod Systematics & Phylogeny 68: 79 - 111" title="Systematics of the new Australasian orb-weaving spider genus Backobourkia (Araneae: Araneidae: Araneinae)." volume="68" year="2010">Framenau et al. 2010</bibRefCitation>
|
||
) (absent in
|
||
<taxonomicName authorityName="Court & Forster, 1993, in Platnick" authorityYear="1993" class="Arachnida" family="Araneidae" genus="Novakiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Novakiella" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Novakiella</emphasis>
|
||
</taxonomicName>
|
||
), is elongate transverse with two apical tips in
|
||
<taxonomicName authorityName="Joseph & Framenau" authorityYear="2012" class="Arachnida" family="Araneidae" genus="Plebs" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Plebs" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Plebs</emphasis>
|
||
</taxonomicName>
|
||
(Joseph & Framenau, 2012, e.g. figs 4B, 8A), and has a two-humped lobe in
|
||
<taxonomicName authorityName="Framenau" authorityYear="2011" class="Arachnida" family="Araneidae" genus="Lariniophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lariniophora" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Lariniophora</emphasis>
|
||
</taxonomicName>
|
||
(Framenau, 2011, fig. 4). All backobourkiines appear to have a basal extension of the conductor (discussed in
|
||
<bibRefCitation author="Framenau, VW" journalOrPublisher="Arthropod Systematics & Phylogeny" pageId="0" pageNumber="393" pagination="79 - 111" refId="B5" refString="Framenau, VW, Duperre, N, Blackledge, TA, Vink, CJ, 2010. Systematics of the new Australasian orb-weaving spider genus Backobourkia (Araneae: Araneidae: Araneinae). Arthropod Systematics & Phylogeny 68: 79 - 111" title="Systematics of the new Australasian orb-weaving spider genus Backobourkia (Araneae: Araneidae: Araneinae)." volume="68" year="2010">Framenau et al. 2010</bibRefCitation>
|
||
and there termed paramedian apophysis), but in
|
||
<taxonomicName authorityName="Court & Forster, 1993, in Platnick" authorityYear="1993" class="Arachnida" family="Araneidae" genus="Novakiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Novakiella" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Novakiella</emphasis>
|
||
</taxonomicName>
|
||
it is very different to all other described backobourkiines and much more conspicuous; we here propose a new term, conductor lobe (CL), which extends apically well past the radix (Figs
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Novakiella trituberculosa, male (WAM T 73571). A. Dorsal habitus; B. ventral habitus; C. Left pedipalp, mesal view; D. Left pedipalp, ventral view. Abbreviations: C, conductor; CL, conductor lobe; E, embolus; MA, median apophysis; P, paracymbium; Ra, radix; TA, terminal apophysis. Scale bars: A, B, 1 mm; C, D, 0.5 mm; E, 1.2 mm." figureDoi="10.3897/zse.97.67788.figure1" httpUri="https://binary.pensoft.net/fig/570631" pageId="0" pageNumber="393">1C</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Novakiella trituberculosa, male (WAM T 73571), expanded left pedipalp. A. Ventral view; B. Retrolateral view. Abbreviations: C, conductor; CL, conductor lobe; E, embolus; MA, median apophysis; P, paracymbium; Ra, radix; TA, terminal apophysis. Scale bars: 0.5 mm" figureDoi="10.3897/zse.97.67788.figure2" httpUri="https://binary.pensoft.net/fig/570632" pageId="0" pageNumber="393">2A, B</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Novakiella boletus sp. nov., male holotype (MV K 9867). A. Dorsal habitus; B. Ventral habitus; C. Left tibia, ventral view; D. Left pedipalp, mesal view; E. Left pedipalp, ventral view, F. left pedipalp, apical view. Abbreviations: C, conductor; CL, conductor lobe; E, embolus; MA, median apophysis; P, paracymbium; Ra, radix; TA, terminal apophysis. Scale bars: A, B, 2 mm; C, 1 mm; D-F, 0.5 mm." figureDoi="10.3897/zse.97.67788.figure4" httpUri="https://binary.pensoft.net/fig/570634" pageId="0" pageNumber="393">4D-F</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Novakiella boletus sp. nov., male holotype (MV K 9867), expanded left pedipalp. mesal view. Abbreviations: C, conductor; CL, conductor lobe; E, embolus; MA, median apophysis; Ra, radix; TA, terminal apophysis. Scale bar: 0.5 mm." figureDoi="10.3897/zse.97.67788.figure5" httpUri="https://binary.pensoft.net/fig/570635" pageId="0" pageNumber="393">5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/zse.97.67788.figure1" httpUri="https://binary.pensoft.net/fig/570631" pageId="0" pageNumber="393" start="Figure 1" startId="F1">
|
||
<paragraph pageId="0" pageNumber="393">
|
||
<emphasis bold="true" pageId="0" pageNumber="393">Figure 1.</emphasis>
|
||
<taxonomicName baseAuthorityName="Roewer" baseAuthorityYear="1942" class="Arachnida" family="Araneidae" genus="Novakiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Novakiella trituberculosa" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="species" species="trituberculosa">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Novakiella trituberculosa</emphasis>
|
||
</taxonomicName>
|
||
, male (WAM T73571).
|
||
<emphasis bold="true" pageId="0" pageNumber="393">A.</emphasis>
|
||
Dorsal habitus;
|
||
<emphasis bold="true" pageId="0" pageNumber="393">B.</emphasis>
|
||
ventral habitus;
|
||
<emphasis bold="true" pageId="0" pageNumber="393">C.</emphasis>
|
||
Left pedipalp, mesal view;
|
||
<emphasis bold="true" pageId="0" pageNumber="393">D.</emphasis>
|
||
Left pedipalp, ventral view. Abbreviations: C, conductor; CL, conductor lobe; E, embolus; MA, median apophysis; P, paracymbium; Ra, radix; TA, terminal apophysis. Scale bars:
|
||
<emphasis bold="true" pageId="0" pageNumber="393">A, B</emphasis>
|
||
, 1 mm;
|
||
<emphasis bold="true" pageId="0" pageNumber="393">C, D</emphasis>
|
||
, 0.5 mm;
|
||
<emphasis bold="true" pageId="0" pageNumber="393">E</emphasis>
|
||
, 1.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zse.97.67788.figure2" httpUri="https://binary.pensoft.net/fig/570632" pageId="0" pageNumber="393" start="Figure 2" startId="F2">
|
||
<paragraph pageId="0" pageNumber="393">
|
||
<emphasis bold="true" pageId="0" pageNumber="393">Figure 2.</emphasis>
|
||
<taxonomicName baseAuthorityName="Roewer" baseAuthorityYear="1942" class="Arachnida" family="Araneidae" genus="Novakiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Novakiella trituberculosa" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="species" species="trituberculosa">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Novakiella trituberculosa</emphasis>
|
||
</taxonomicName>
|
||
, male (WAM T73571), expanded left pedipalp.
|
||
<emphasis bold="true" pageId="0" pageNumber="393">A.</emphasis>
|
||
Ventral view;
|
||
<emphasis bold="true" pageId="0" pageNumber="393">B.</emphasis>
|
||
Retrolateral view. Abbreviations: C, conductor; CL, conductor lobe; E, embolus; MA, median apophysis; P, paracymbium; Ra, radix; TA, terminal apophysis. Scale bars: 0.5 mm
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="393">
|
||
Females of
|
||
<taxonomicName authorityName="Court & Forster, 1993, in Platnick" authorityYear="1993" class="Arachnida" family="Araneidae" genus="Novakiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Novakiella" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Novakiella</emphasis>
|
||
</taxonomicName>
|
||
have an elongated triangular scape without terminal pockets, as is typical for all backobourkiines above; however, these genera lack the subtriangular base plate with its transverse and lateral wrinkles (Figs
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Novakiella trituberculosa, female (WAM T 73556). A. Dorsal habitus; B. Ventral habitus; C. Epigyne, ventral view; D. Cleared epigyne, ventral view; E. Cleared epigyne, dorsal view. Abbreviations: CO, copulatory openings; S, scape; Sp, spermatheca. Scale bars: A, B, 2 mm; C-E, 0.1 mm." figureDoi="10.3897/zse.97.67788.figure3" httpUri="https://binary.pensoft.net/fig/570633" pageId="0" pageNumber="393">3C</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Novakiella boletus sp. nov., female (SAM). A. Dorsal habitus; B. Ventral habitus; C. Epigyne, ventral view. Abbreviations: CO, copulatory openings; L, lips; S, scape. Scale bars: A, B, 2 mm, C, 0.2 mm." figureDoi="10.3897/zse.97.67788.figure6" httpUri="https://binary.pensoft.net/fig/570636" pageId="0" pageNumber="393">6C</figureCitation>
|
||
;
|
||
<bibRefCitation author="Framenau, VW" journalOrPublisher="Arthropod Systematics & Phylogeny" pageId="0" pageNumber="393" pagination="79 - 111" refId="B5" refString="Framenau, VW, Duperre, N, Blackledge, TA, Vink, CJ, 2010. Systematics of the new Australasian orb-weaving spider genus Backobourkia (Araneae: Araneidae: Araneinae). Arthropod Systematics & Phylogeny 68: 79 - 111" title="Systematics of the new Australasian orb-weaving spider genus Backobourkia (Araneae: Araneidae: Araneinae)." volume="68" year="2010">Framenau et al. 2010</bibRefCitation>
|
||
, e.g. figs 6D, F;
|
||
<bibRefCitation DOI="https://doi.org/10.18195/issn.0312-3162.26(2).2011.191-201" author="Framenau, VW" journalOrPublisher="Records of the Western Australian Museum" pageId="0" pageNumber="393" pagination="191 - 201" refId="B4" refString="Framenau, VW, 2011. Lariniophora, a new monotypic orb-weaving spider genus from Australia (Araneae: Araneidae: Araneinae). Records of the Western Australian Museum 26 (2): 191 - 201, DOI: https://doi.org/10.18195/issn.0312-3162.26(2).2011.191-201" title="Lariniophora, a new monotypic orb-weaving spider genus from Australia (Araneae: Araneidae: Araneinae)." url="https://doi.org/10.18195/issn.0312-3162.26(2).2011.191-201" volume="26" year="2011">Framenau 2011</bibRefCitation>
|
||
, fig. 6;
|
||
<bibRefCitation DOI="https://doi.org/10.1111/j.1096-3642.2012.00845.x" author="Joseph, MM" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="393" pagination="279 - 341" refId="B7" refString="Joseph, MM, Framenau, VW, 2012. Systematic review of a new orb-weaving spider genus (Araneae: Araneidae), with special reference to the Australasian-Pacific and South-East Asian fauna. Zoological Journal of the Linnean Society 166: 279 - 341, DOI: https://doi.org/10.1111/j.1096-3642.2012.00845.x" title="Systematic review of a new orb-weaving spider genus (Araneae: Araneidae), with special reference to the Australasian-Pacific and South-East Asian fauna." url="https://doi.org/10.1111/j.1096-3642.2012.00845.x" volume="166" year="2012">Joseph and Framenau 2012</bibRefCitation>
|
||
, e.g. figs 4D, 8E).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="393">
|
||
<emphasis bold="true" pageId="0" pageNumber="393">Description</emphasis>
|
||
. Medium-sized (TL males ca. 5-9, females 8-12) orb-weaving spiders with males on average slightly smaller than females. Carapace longer than wide, pear-shaped; cephalic area similar in shape in both sexes (Figs
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Novakiella trituberculosa, male (WAM T 73571). A. Dorsal habitus; B. ventral habitus; C. Left pedipalp, mesal view; D. Left pedipalp, ventral view. Abbreviations: C, conductor; CL, conductor lobe; E, embolus; MA, median apophysis; P, paracymbium; Ra, radix; TA, terminal apophysis. Scale bars: A, B, 1 mm; C, D, 0.5 mm; E, 1.2 mm." figureDoi="10.3897/zse.97.67788.figure1" httpUri="https://binary.pensoft.net/fig/570631" pageId="0" pageNumber="393">1A</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Novakiella trituberculosa, female (WAM T 73556). A. Dorsal habitus; B. Ventral habitus; C. Epigyne, ventral view; D. Cleared epigyne, ventral view; E. Cleared epigyne, dorsal view. Abbreviations: CO, copulatory openings; S, scape; Sp, spermatheca. Scale bars: A, B, 2 mm; C-E, 0.1 mm." figureDoi="10.3897/zse.97.67788.figure3" httpUri="https://binary.pensoft.net/fig/570633" pageId="0" pageNumber="393">3A</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Novakiella boletus sp. nov., male holotype (MV K 9867). A. Dorsal habitus; B. Ventral habitus; C. Left tibia, ventral view; D. Left pedipalp, mesal view; E. Left pedipalp, ventral view, F. left pedipalp, apical view. Abbreviations: C, conductor; CL, conductor lobe; E, embolus; MA, median apophysis; P, paracymbium; Ra, radix; TA, terminal apophysis. Scale bars: A, B, 2 mm; C, 1 mm; D-F, 0.5 mm." figureDoi="10.3897/zse.97.67788.figure4" httpUri="https://binary.pensoft.net/fig/570634" pageId="0" pageNumber="393">4A</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Novakiella boletus sp. nov., female (SAM). A. Dorsal habitus; B. Ventral habitus; C. Epigyne, ventral view. Abbreviations: CO, copulatory openings; L, lips; S, scape. Scale bars: A, B, 2 mm, C, 0.2 mm." figureDoi="10.3897/zse.97.67788.figure6" httpUri="https://binary.pensoft.net/fig/570636" pageId="0" pageNumber="393">6A</figureCitation>
|
||
); fovea longer than wide in males and wider than long in females, and with a dark spot in both sexes (Figs
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Novakiella trituberculosa, male (WAM T 73571). A. Dorsal habitus; B. ventral habitus; C. Left pedipalp, mesal view; D. Left pedipalp, ventral view. Abbreviations: C, conductor; CL, conductor lobe; E, embolus; MA, median apophysis; P, paracymbium; Ra, radix; TA, terminal apophysis. Scale bars: A, B, 1 mm; C, D, 0.5 mm; E, 1.2 mm." figureDoi="10.3897/zse.97.67788.figure1" httpUri="https://binary.pensoft.net/fig/570631" pageId="0" pageNumber="393">1A</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Novakiella trituberculosa, female (WAM T 73556). A. Dorsal habitus; B. Ventral habitus; C. Epigyne, ventral view; D. Cleared epigyne, ventral view; E. Cleared epigyne, dorsal view. Abbreviations: CO, copulatory openings; S, scape; Sp, spermatheca. Scale bars: A, B, 2 mm; C-E, 0.1 mm." figureDoi="10.3897/zse.97.67788.figure3" httpUri="https://binary.pensoft.net/fig/570633" pageId="0" pageNumber="393">3A</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Novakiella boletus sp. nov., male holotype (MV K 9867). A. Dorsal habitus; B. Ventral habitus; C. Left tibia, ventral view; D. Left pedipalp, mesal view; E. Left pedipalp, ventral view, F. left pedipalp, apical view. Abbreviations: C, conductor; CL, conductor lobe; E, embolus; MA, median apophysis; P, paracymbium; Ra, radix; TA, terminal apophysis. Scale bars: A, B, 2 mm; C, 1 mm; D-F, 0.5 mm." figureDoi="10.3897/zse.97.67788.figure4" httpUri="https://binary.pensoft.net/fig/570634" pageId="0" pageNumber="393">4A</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Novakiella boletus sp. nov., female (SAM). A. Dorsal habitus; B. Ventral habitus; C. Epigyne, ventral view. Abbreviations: CO, copulatory openings; L, lips; S, scape. Scale bars: A, B, 2 mm, C, 0.2 mm." figureDoi="10.3897/zse.97.67788.figure6" httpUri="https://binary.pensoft.net/fig/570636" pageId="0" pageNumber="393">6A</figureCitation>
|
||
); colouration (of ethanol preserved specimens) varying from reddish-brown to yellowish-brown, with black patches along carapace borders (Figs
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Novakiella trituberculosa, male (WAM T 73571). A. Dorsal habitus; B. ventral habitus; C. Left pedipalp, mesal view; D. Left pedipalp, ventral view. Abbreviations: C, conductor; CL, conductor lobe; E, embolus; MA, median apophysis; P, paracymbium; Ra, radix; TA, terminal apophysis. Scale bars: A, B, 1 mm; C, D, 0.5 mm; E, 1.2 mm." figureDoi="10.3897/zse.97.67788.figure1" httpUri="https://binary.pensoft.net/fig/570631" pageId="0" pageNumber="393">1A</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Novakiella trituberculosa, female (WAM T 73556). A. Dorsal habitus; B. Ventral habitus; C. Epigyne, ventral view; D. Cleared epigyne, ventral view; E. Cleared epigyne, dorsal view. Abbreviations: CO, copulatory openings; S, scape; Sp, spermatheca. Scale bars: A, B, 2 mm; C-E, 0.1 mm." figureDoi="10.3897/zse.97.67788.figure3" httpUri="https://binary.pensoft.net/fig/570633" pageId="0" pageNumber="393">3A</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Novakiella boletus sp. nov., male holotype (MV K 9867). A. Dorsal habitus; B. Ventral habitus; C. Left tibia, ventral view; D. Left pedipalp, mesal view; E. Left pedipalp, ventral view, F. left pedipalp, apical view. Abbreviations: C, conductor; CL, conductor lobe; E, embolus; MA, median apophysis; P, paracymbium; Ra, radix; TA, terminal apophysis. Scale bars: A, B, 2 mm; C, 1 mm; D-F, 0.5 mm." figureDoi="10.3897/zse.97.67788.figure4" httpUri="https://binary.pensoft.net/fig/570634" pageId="0" pageNumber="393">4A</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Novakiella boletus sp. nov., female (SAM). A. Dorsal habitus; B. Ventral habitus; C. Epigyne, ventral view. Abbreviations: CO, copulatory openings; L, lips; S, scape. Scale bars: A, B, 2 mm, C, 0.2 mm." figureDoi="10.3897/zse.97.67788.figure6" httpUri="https://binary.pensoft.net/fig/570636" pageId="0" pageNumber="393">6A</figureCitation>
|
||
). Eyes ringed in black, anterior median eyes largest, posterior eye row slightly recurved, lateral eyes almost touching, posterior lateral eyes separated from posterior median eyes by more than their diameter and located on small tubercles at the clypeus border (Figs
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Novakiella trituberculosa, male (WAM T 73571). A. Dorsal habitus; B. ventral habitus; C. Left pedipalp, mesal view; D. Left pedipalp, ventral view. Abbreviations: C, conductor; CL, conductor lobe; E, embolus; MA, median apophysis; P, paracymbium; Ra, radix; TA, terminal apophysis. Scale bars: A, B, 1 mm; C, D, 0.5 mm; E, 1.2 mm." figureDoi="10.3897/zse.97.67788.figure1" httpUri="https://binary.pensoft.net/fig/570631" pageId="0" pageNumber="393">1A</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Novakiella trituberculosa, female (WAM T 73556). A. Dorsal habitus; B. Ventral habitus; C. Epigyne, ventral view; D. Cleared epigyne, ventral view; E. Cleared epigyne, dorsal view. Abbreviations: CO, copulatory openings; S, scape; Sp, spermatheca. Scale bars: A, B, 2 mm; C-E, 0.1 mm." figureDoi="10.3897/zse.97.67788.figure3" httpUri="https://binary.pensoft.net/fig/570633" pageId="0" pageNumber="393">3A</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Novakiella boletus sp. nov., male holotype (MV K 9867). A. Dorsal habitus; B. Ventral habitus; C. Left tibia, ventral view; D. Left pedipalp, mesal view; E. Left pedipalp, ventral view, F. left pedipalp, apical view. Abbreviations: C, conductor; CL, conductor lobe; E, embolus; MA, median apophysis; P, paracymbium; Ra, radix; TA, terminal apophysis. Scale bars: A, B, 2 mm; C, 1 mm; D-F, 0.5 mm." figureDoi="10.3897/zse.97.67788.figure4" httpUri="https://binary.pensoft.net/fig/570634" pageId="0" pageNumber="393">4A</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Novakiella boletus sp. nov., female (SAM). A. Dorsal habitus; B. Ventral habitus; C. Epigyne, ventral view. Abbreviations: CO, copulatory openings; L, lips; S, scape. Scale bars: A, B, 2 mm, C, 0.2 mm." figureDoi="10.3897/zse.97.67788.figure6" httpUri="https://binary.pensoft.net/fig/570636" pageId="0" pageNumber="393">6A</figureCitation>
|
||
). Chelicera paturon with dark hue, fangs reddish-brown. Labium wider than long, subtriangular, with front end bulging and beige (Figs
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Novakiella trituberculosa, male (WAM T 73571). A. Dorsal habitus; B. ventral habitus; C. Left pedipalp, mesal view; D. Left pedipalp, ventral view. Abbreviations: C, conductor; CL, conductor lobe; E, embolus; MA, median apophysis; P, paracymbium; Ra, radix; TA, terminal apophysis. Scale bars: A, B, 1 mm; C, D, 0.5 mm; E, 1.2 mm." figureDoi="10.3897/zse.97.67788.figure1" httpUri="https://binary.pensoft.net/fig/570631" pageId="0" pageNumber="393">1B</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Novakiella trituberculosa, female (WAM T 73556). A. Dorsal habitus; B. Ventral habitus; C. Epigyne, ventral view; D. Cleared epigyne, ventral view; E. Cleared epigyne, dorsal view. Abbreviations: CO, copulatory openings; S, scape; Sp, spermatheca. Scale bars: A, B, 2 mm; C-E, 0.1 mm." figureDoi="10.3897/zse.97.67788.figure3" httpUri="https://binary.pensoft.net/fig/570633" pageId="0" pageNumber="393">3B</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Novakiella boletus sp. nov., male holotype (MV K 9867). A. Dorsal habitus; B. Ventral habitus; C. Left tibia, ventral view; D. Left pedipalp, mesal view; E. Left pedipalp, ventral view, F. left pedipalp, apical view. Abbreviations: C, conductor; CL, conductor lobe; E, embolus; MA, median apophysis; P, paracymbium; Ra, radix; TA, terminal apophysis. Scale bars: A, B, 2 mm; C, 1 mm; D-F, 0.5 mm." figureDoi="10.3897/zse.97.67788.figure4" httpUri="https://binary.pensoft.net/fig/570634" pageId="0" pageNumber="393">4B</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Novakiella boletus sp. nov., female (SAM). A. Dorsal habitus; B. Ventral habitus; C. Epigyne, ventral view. Abbreviations: CO, copulatory openings; L, lips; S, scape. Scale bars: A, B, 2 mm, C, 0.2 mm." figureDoi="10.3897/zse.97.67788.figure6" httpUri="https://binary.pensoft.net/fig/570636" pageId="0" pageNumber="393">6B</figureCitation>
|
||
). Endites rounded, inner portion beige (Figs
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Novakiella trituberculosa, male (WAM T 73571). A. Dorsal habitus; B. ventral habitus; C. Left pedipalp, mesal view; D. Left pedipalp, ventral view. Abbreviations: C, conductor; CL, conductor lobe; E, embolus; MA, median apophysis; P, paracymbium; Ra, radix; TA, terminal apophysis. Scale bars: A, B, 1 mm; C, D, 0.5 mm; E, 1.2 mm." figureDoi="10.3897/zse.97.67788.figure1" httpUri="https://binary.pensoft.net/fig/570631" pageId="0" pageNumber="393">1B</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Novakiella trituberculosa, female (WAM T 73556). A. Dorsal habitus; B. Ventral habitus; C. Epigyne, ventral view; D. Cleared epigyne, ventral view; E. Cleared epigyne, dorsal view. Abbreviations: CO, copulatory openings; S, scape; Sp, spermatheca. Scale bars: A, B, 2 mm; C-E, 0.1 mm." figureDoi="10.3897/zse.97.67788.figure3" httpUri="https://binary.pensoft.net/fig/570633" pageId="0" pageNumber="393">3B</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Novakiella boletus sp. nov., male holotype (MV K 9867). A. Dorsal habitus; B. Ventral habitus; C. Left tibia, ventral view; D. Left pedipalp, mesal view; E. Left pedipalp, ventral view, F. left pedipalp, apical view. Abbreviations: C, conductor; CL, conductor lobe; E, embolus; MA, median apophysis; P, paracymbium; Ra, radix; TA, terminal apophysis. Scale bars: A, B, 2 mm; C, 1 mm; D-F, 0.5 mm." figureDoi="10.3897/zse.97.67788.figure4" httpUri="https://binary.pensoft.net/fig/570634" pageId="0" pageNumber="393">4B</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Novakiella boletus sp. nov., female (SAM). A. Dorsal habitus; B. Ventral habitus; C. Epigyne, ventral view. Abbreviations: CO, copulatory openings; L, lips; S, scape. Scale bars: A, B, 2 mm, C, 0.2 mm." figureDoi="10.3897/zse.97.67788.figure6" httpUri="https://binary.pensoft.net/fig/570636" pageId="0" pageNumber="393">6B</figureCitation>
|
||
). Sternum almost as long as wide with dark contour (Figs
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Novakiella trituberculosa, male (WAM T 73571). A. Dorsal habitus; B. ventral habitus; C. Left pedipalp, mesal view; D. Left pedipalp, ventral view. Abbreviations: C, conductor; CL, conductor lobe; E, embolus; MA, median apophysis; P, paracymbium; Ra, radix; TA, terminal apophysis. Scale bars: A, B, 1 mm; C, D, 0.5 mm; E, 1.2 mm." figureDoi="10.3897/zse.97.67788.figure1" httpUri="https://binary.pensoft.net/fig/570631" pageId="0" pageNumber="393">1B</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Novakiella trituberculosa, female (WAM T 73556). A. Dorsal habitus; B. Ventral habitus; C. Epigyne, ventral view; D. Cleared epigyne, ventral view; E. Cleared epigyne, dorsal view. Abbreviations: CO, copulatory openings; S, scape; Sp, spermatheca. Scale bars: A, B, 2 mm; C-E, 0.1 mm." figureDoi="10.3897/zse.97.67788.figure3" httpUri="https://binary.pensoft.net/fig/570633" pageId="0" pageNumber="393">3B</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Novakiella boletus sp. nov., male holotype (MV K 9867). A. Dorsal habitus; B. Ventral habitus; C. Left tibia, ventral view; D. Left pedipalp, mesal view; E. Left pedipalp, ventral view, F. left pedipalp, apical view. Abbreviations: C, conductor; CL, conductor lobe; E, embolus; MA, median apophysis; P, paracymbium; Ra, radix; TA, terminal apophysis. Scale bars: A, B, 2 mm; C, 1 mm; D-F, 0.5 mm." figureDoi="10.3897/zse.97.67788.figure4" httpUri="https://binary.pensoft.net/fig/570634" pageId="0" pageNumber="393">4B</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Novakiella boletus sp. nov., female (SAM). A. Dorsal habitus; B. Ventral habitus; C. Epigyne, ventral view. Abbreviations: CO, copulatory openings; L, lips; S, scape. Scale bars: A, B, 2 mm, C, 0.2 mm." figureDoi="10.3897/zse.97.67788.figure6" httpUri="https://binary.pensoft.net/fig/570636" pageId="0" pageNumber="393">6B</figureCitation>
|
||
). Legs (Figs
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Novakiella trituberculosa, male (WAM T 73571). A. Dorsal habitus; B. ventral habitus; C. Left pedipalp, mesal view; D. Left pedipalp, ventral view. Abbreviations: C, conductor; CL, conductor lobe; E, embolus; MA, median apophysis; P, paracymbium; Ra, radix; TA, terminal apophysis. Scale bars: A, B, 1 mm; C, D, 0.5 mm; E, 1.2 mm." figureDoi="10.3897/zse.97.67788.figure1" httpUri="https://binary.pensoft.net/fig/570631" pageId="0" pageNumber="393">1A, B, E</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Novakiella trituberculosa, female (WAM T 73556). A. Dorsal habitus; B. Ventral habitus; C. Epigyne, ventral view; D. Cleared epigyne, ventral view; E. Cleared epigyne, dorsal view. Abbreviations: CO, copulatory openings; S, scape; Sp, spermatheca. Scale bars: A, B, 2 mm; C-E, 0.1 mm." figureDoi="10.3897/zse.97.67788.figure3" httpUri="https://binary.pensoft.net/fig/570633" pageId="0" pageNumber="393">3A, B</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Novakiella boletus sp. nov., male holotype (MV K 9867). A. Dorsal habitus; B. Ventral habitus; C. Left tibia, ventral view; D. Left pedipalp, mesal view; E. Left pedipalp, ventral view, F. left pedipalp, apical view. Abbreviations: C, conductor; CL, conductor lobe; E, embolus; MA, median apophysis; P, paracymbium; Ra, radix; TA, terminal apophysis. Scale bars: A, B, 2 mm; C, 1 mm; D-F, 0.5 mm." figureDoi="10.3897/zse.97.67788.figure4" httpUri="https://binary.pensoft.net/fig/570634" pageId="0" pageNumber="393">4A-C</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Novakiella boletus sp. nov., female (SAM). A. Dorsal habitus; B. Ventral habitus; C. Epigyne, ventral view. Abbreviations: CO, copulatory openings; L, lips; S, scape. Scale bars: A, B, 2 mm, C, 0.2 mm." figureDoi="10.3897/zse.97.67788.figure6" httpUri="https://binary.pensoft.net/fig/570636" pageId="0" pageNumber="393">6A, B</figureCitation>
|
||
): Leg formula IV> I> II> III, all longer than
|
||
<normalizedToken originalValue="body’s">body's</normalizedToken>
|
||
length with dark spots on joints; tibia II of males with apico-prolateral spur bearing a thick macroseta or spine (less pronounced in
|
||
<taxonomicName class="Arachnida" genus="N." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="N. boletus" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="species" species="boletus">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">N. boletus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.). Abdomen subtriangular, longer than wide, with two distinct humeral humps and posterior tip reaching beyond spinnerets (Figs
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Novakiella trituberculosa, male (WAM T 73571). A. Dorsal habitus; B. ventral habitus; C. Left pedipalp, mesal view; D. Left pedipalp, ventral view. Abbreviations: C, conductor; CL, conductor lobe; E, embolus; MA, median apophysis; P, paracymbium; Ra, radix; TA, terminal apophysis. Scale bars: A, B, 1 mm; C, D, 0.5 mm; E, 1.2 mm." figureDoi="10.3897/zse.97.67788.figure1" httpUri="https://binary.pensoft.net/fig/570631" pageId="0" pageNumber="393">1A</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Novakiella trituberculosa, female (WAM T 73556). A. Dorsal habitus; B. Ventral habitus; C. Epigyne, ventral view; D. Cleared epigyne, ventral view; E. Cleared epigyne, dorsal view. Abbreviations: CO, copulatory openings; S, scape; Sp, spermatheca. Scale bars: A, B, 2 mm; C-E, 0.1 mm." figureDoi="10.3897/zse.97.67788.figure3" httpUri="https://binary.pensoft.net/fig/570633" pageId="0" pageNumber="393">3A</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Novakiella boletus sp. nov., male holotype (MV K 9867). A. Dorsal habitus; B. Ventral habitus; C. Left tibia, ventral view; D. Left pedipalp, mesal view; E. Left pedipalp, ventral view, F. left pedipalp, apical view. Abbreviations: C, conductor; CL, conductor lobe; E, embolus; MA, median apophysis; P, paracymbium; Ra, radix; TA, terminal apophysis. Scale bars: A, B, 2 mm; C, 1 mm; D-F, 0.5 mm." figureDoi="10.3897/zse.97.67788.figure4" httpUri="https://binary.pensoft.net/fig/570634" pageId="0" pageNumber="393">4A</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Novakiella boletus sp. nov., female (SAM). A. Dorsal habitus; B. Ventral habitus; C. Epigyne, ventral view. Abbreviations: CO, copulatory openings; L, lips; S, scape. Scale bars: A, B, 2 mm, C, 0.2 mm." figureDoi="10.3897/zse.97.67788.figure6" httpUri="https://binary.pensoft.net/fig/570636" pageId="0" pageNumber="393">6A</figureCitation>
|
||
); folium pattern distinct; sides varying in colour from yellowish-brown to black (Figs
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Novakiella trituberculosa, male (WAM T 73571). A. Dorsal habitus; B. ventral habitus; C. Left pedipalp, mesal view; D. Left pedipalp, ventral view. Abbreviations: C, conductor; CL, conductor lobe; E, embolus; MA, median apophysis; P, paracymbium; Ra, radix; TA, terminal apophysis. Scale bars: A, B, 1 mm; C, D, 0.5 mm; E, 1.2 mm." figureDoi="10.3897/zse.97.67788.figure1" httpUri="https://binary.pensoft.net/fig/570631" pageId="0" pageNumber="393">1A</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Novakiella trituberculosa, female (WAM T 73556). A. Dorsal habitus; B. Ventral habitus; C. Epigyne, ventral view; D. Cleared epigyne, ventral view; E. Cleared epigyne, dorsal view. Abbreviations: CO, copulatory openings; S, scape; Sp, spermatheca. Scale bars: A, B, 2 mm; C-E, 0.1 mm." figureDoi="10.3897/zse.97.67788.figure3" httpUri="https://binary.pensoft.net/fig/570633" pageId="0" pageNumber="393">3A</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Novakiella boletus sp. nov., male holotype (MV K 9867). A. Dorsal habitus; B. Ventral habitus; C. Left tibia, ventral view; D. Left pedipalp, mesal view; E. Left pedipalp, ventral view, F. left pedipalp, apical view. Abbreviations: C, conductor; CL, conductor lobe; E, embolus; MA, median apophysis; P, paracymbium; Ra, radix; TA, terminal apophysis. Scale bars: A, B, 2 mm; C, 1 mm; D-F, 0.5 mm." figureDoi="10.3897/zse.97.67788.figure4" httpUri="https://binary.pensoft.net/fig/570634" pageId="0" pageNumber="393">4A</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Novakiella boletus sp. nov., female (SAM). A. Dorsal habitus; B. Ventral habitus; C. Epigyne, ventral view. Abbreviations: CO, copulatory openings; L, lips; S, scape. Scale bars: A, B, 2 mm, C, 0.2 mm." figureDoi="10.3897/zse.97.67788.figure6" httpUri="https://binary.pensoft.net/fig/570636" pageId="0" pageNumber="393">6A</figureCitation>
|
||
), venter light coloured, generally mottled dark (Figs
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Novakiella trituberculosa, male (WAM T 73571). A. Dorsal habitus; B. ventral habitus; C. Left pedipalp, mesal view; D. Left pedipalp, ventral view. Abbreviations: C, conductor; CL, conductor lobe; E, embolus; MA, median apophysis; P, paracymbium; Ra, radix; TA, terminal apophysis. Scale bars: A, B, 1 mm; C, D, 0.5 mm; E, 1.2 mm." figureDoi="10.3897/zse.97.67788.figure1" httpUri="https://binary.pensoft.net/fig/570631" pageId="0" pageNumber="393">1B</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Novakiella trituberculosa, female (WAM T 73556). A. Dorsal habitus; B. Ventral habitus; C. Epigyne, ventral view; D. Cleared epigyne, ventral view; E. Cleared epigyne, dorsal view. Abbreviations: CO, copulatory openings; S, scape; Sp, spermatheca. Scale bars: A, B, 2 mm; C-E, 0.1 mm." figureDoi="10.3897/zse.97.67788.figure3" httpUri="https://binary.pensoft.net/fig/570633" pageId="0" pageNumber="393">3B</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Novakiella boletus sp. nov., male holotype (MV K 9867). A. Dorsal habitus; B. Ventral habitus; C. Left tibia, ventral view; D. Left pedipalp, mesal view; E. Left pedipalp, ventral view, F. left pedipalp, apical view. Abbreviations: C, conductor; CL, conductor lobe; E, embolus; MA, median apophysis; P, paracymbium; Ra, radix; TA, terminal apophysis. Scale bars: A, B, 2 mm; C, 1 mm; D-F, 0.5 mm." figureDoi="10.3897/zse.97.67788.figure4" httpUri="https://binary.pensoft.net/fig/570634" pageId="0" pageNumber="393">4B</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Novakiella boletus sp. nov., female (SAM). A. Dorsal habitus; B. Ventral habitus; C. Epigyne, ventral view. Abbreviations: CO, copulatory openings; L, lips; S, scape. Scale bars: A, B, 2 mm, C, 0.2 mm." figureDoi="10.3897/zse.97.67788.figure6" httpUri="https://binary.pensoft.net/fig/570636" pageId="0" pageNumber="393">6B</figureCitation>
|
||
). Male genitalia (Figs
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Novakiella trituberculosa, male (WAM T 73571). A. Dorsal habitus; B. ventral habitus; C. Left pedipalp, mesal view; D. Left pedipalp, ventral view. Abbreviations: C, conductor; CL, conductor lobe; E, embolus; MA, median apophysis; P, paracymbium; Ra, radix; TA, terminal apophysis. Scale bars: A, B, 1 mm; C, D, 0.5 mm; E, 1.2 mm." figureDoi="10.3897/zse.97.67788.figure1" httpUri="https://binary.pensoft.net/fig/570631" pageId="0" pageNumber="393">1C, D</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Novakiella trituberculosa, male (WAM T 73571), expanded left pedipalp. A. Ventral view; B. Retrolateral view. Abbreviations: C, conductor; CL, conductor lobe; E, embolus; MA, median apophysis; P, paracymbium; Ra, radix; TA, terminal apophysis. Scale bars: 0.5 mm" figureDoi="10.3897/zse.97.67788.figure2" httpUri="https://binary.pensoft.net/fig/570632" pageId="0" pageNumber="393">2A, B</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Novakiella boletus sp. nov., male holotype (MV K 9867). A. Dorsal habitus; B. Ventral habitus; C. Left tibia, ventral view; D. Left pedipalp, mesal view; E. Left pedipalp, ventral view, F. left pedipalp, apical view. Abbreviations: C, conductor; CL, conductor lobe; E, embolus; MA, median apophysis; P, paracymbium; Ra, radix; TA, terminal apophysis. Scale bars: A, B, 2 mm; C, 1 mm; D-F, 0.5 mm." figureDoi="10.3897/zse.97.67788.figure4" httpUri="https://binary.pensoft.net/fig/570634" pageId="0" pageNumber="393">4D-F</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Novakiella boletus sp. nov., male holotype (MV K 9867), expanded left pedipalp. mesal view. Abbreviations: C, conductor; CL, conductor lobe; E, embolus; MA, median apophysis; Ra, radix; TA, terminal apophysis. Scale bar: 0.5 mm." figureDoi="10.3897/zse.97.67788.figure5" httpUri="https://binary.pensoft.net/fig/570635" pageId="0" pageNumber="393">5</figureCitation>
|
||
): male pedipalp patella with a single strong macroseta; paracymbium well-developed and hook-like; cymbium longer than wide; radix thick and elongated, reaching from the base of median apophysis to near the cymbium tip; conductor lobe conspicuous and projected apically, being composed of two distinct lobes (
|
||
<taxonomicName class="Arachnida" genus="N." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="N. trituberculosa" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="species" species="trituberculosa">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">N. trituberculosa</emphasis>
|
||
</taxonomicName>
|
||
) or mushroom-shaped (
|
||
<taxonomicName class="Arachnida" genus="N." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="N. boletus" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="species" species="boletus">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">N. boletus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.); terminal apophysis wider than long, rounded and tapering terminally; conductor well-developed, subquadrate; embolus uncapped, elongated, pointed and almost straight; median apophysis stout, with an acute basally pointing tip. Female genitalia (Figs
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Novakiella trituberculosa, female (WAM T 73556). A. Dorsal habitus; B. Ventral habitus; C. Epigyne, ventral view; D. Cleared epigyne, ventral view; E. Cleared epigyne, dorsal view. Abbreviations: CO, copulatory openings; S, scape; Sp, spermatheca. Scale bars: A, B, 2 mm; C-E, 0.1 mm." figureDoi="10.3897/zse.97.67788.figure3" httpUri="https://binary.pensoft.net/fig/570633" pageId="0" pageNumber="393">3C-E</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Novakiella boletus sp. nov., female (SAM). A. Dorsal habitus; B. Ventral habitus; C. Epigyne, ventral view. Abbreviations: CO, copulatory openings; L, lips; S, scape. Scale bars: A, B, 2 mm, C, 0.2 mm." figureDoi="10.3897/zse.97.67788.figure6" httpUri="https://binary.pensoft.net/fig/570636" pageId="0" pageNumber="393">6C</figureCitation>
|
||
): epigyne plate wider than long, subtriangular; scape much longer than wide and extending posteriorly beyond plate (but length not known in
|
||
<taxonomicName class="Arachnida" genus="N." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="N. boletus" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="species" species="boletus">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">N. boletus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.), generally broken off. Spermathecae spherical and occupying most of genital area.
|
||
</paragraph>
|
||
<caption doi="10.3897/zse.97.67788.figure3" httpUri="https://binary.pensoft.net/fig/570633" pageId="0" pageNumber="393" start="Figure 3" startId="F3">
|
||
<paragraph pageId="0" pageNumber="393">
|
||
<emphasis bold="true" pageId="0" pageNumber="393">Figure 3.</emphasis>
|
||
<taxonomicName baseAuthorityName="Roewer" baseAuthorityYear="1942" class="Arachnida" family="Araneidae" genus="Novakiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Novakiella trituberculosa" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="species" species="trituberculosa">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Novakiella trituberculosa</emphasis>
|
||
</taxonomicName>
|
||
, female (WAM T73556).
|
||
<emphasis bold="true" pageId="0" pageNumber="393">A.</emphasis>
|
||
Dorsal habitus;
|
||
<emphasis bold="true" pageId="0" pageNumber="393">B.</emphasis>
|
||
Ventral habitus;
|
||
<emphasis bold="true" pageId="0" pageNumber="393">C.</emphasis>
|
||
Epigyne, ventral view;
|
||
<emphasis bold="true" pageId="0" pageNumber="393">D.</emphasis>
|
||
Cleared epigyne, ventral view;
|
||
<emphasis bold="true" pageId="0" pageNumber="393">E.</emphasis>
|
||
Cleared epigyne, dorsal view. Abbreviations: CO, copulatory openings; S, scape; Sp, spermatheca. Scale bars:
|
||
<emphasis bold="true" pageId="0" pageNumber="393">A, B</emphasis>
|
||
, 2 mm;
|
||
<emphasis bold="true" pageId="0" pageNumber="393">C-E</emphasis>
|
||
, 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="393" type="composition">
|
||
<paragraph pageId="0" pageNumber="393">Composition.</paragraph>
|
||
<paragraph pageId="0" pageNumber="393">
|
||
<taxonomicName baseAuthorityName="Roewer" baseAuthorityYear="1942" class="Arachnida" family="Araneidae" genus="Novakiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Novakiella trituberculosa" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="species" species="trituberculosa">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Novakiella trituberculosa</emphasis>
|
||
</taxonomicName>
|
||
(Roewer, 1942) and
|
||
<taxonomicName class="Arachnida" genus="N." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="N. boletus" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="species" species="boletus">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">N. boletus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
</paragraph>
|
||
<caption doi="10.3897/zse.97.67788.figure4" httpUri="https://binary.pensoft.net/fig/570634" pageId="0" pageNumber="393" start="Figure 4" startId="F4">
|
||
<paragraph pageId="0" pageNumber="393">
|
||
<emphasis bold="true" pageId="0" pageNumber="393">Figure 4.</emphasis>
|
||
<taxonomicName authorityName="Framenau & Vink & Scharff & Baptista & Castanheira" authorityYear="2021" class="Arachnida" family="Araneidae" genus="Novakiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Novakiella boletus" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="species" species="boletus">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Novakiella boletus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., male holotype (MV K9867).
|
||
<emphasis bold="true" pageId="0" pageNumber="393">A.</emphasis>
|
||
Dorsal habitus;
|
||
<emphasis bold="true" pageId="0" pageNumber="393">B.</emphasis>
|
||
Ventral habitus;
|
||
<emphasis bold="true" pageId="0" pageNumber="393">C.</emphasis>
|
||
Left tibia, ventral view;
|
||
<emphasis bold="true" pageId="0" pageNumber="393">D.</emphasis>
|
||
Left pedipalp, mesal view;
|
||
<emphasis bold="true" pageId="0" pageNumber="393">E.</emphasis>
|
||
Left pedipalp, ventral view,
|
||
<emphasis bold="true" pageId="0" pageNumber="393">F.</emphasis>
|
||
left pedipalp, apical view. Abbreviations: C, conductor; CL, conductor lobe; E, embolus; MA, median apophysis; P, paracymbium; Ra, radix; TA, terminal apophysis. Scale bars:
|
||
<emphasis bold="true" pageId="0" pageNumber="393">A, B</emphasis>
|
||
, 2 mm;
|
||
<emphasis bold="true" pageId="0" pageNumber="393">C</emphasis>
|
||
, 1 mm;
|
||
<emphasis bold="true" pageId="0" pageNumber="393">D-F</emphasis>
|
||
, 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="393" type="remarks">
|
||
<paragraph pageId="0" pageNumber="393">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="393">
|
||
The nomenclatural history of
|
||
<taxonomicName authorityName="Court & Forster, 1993, in Platnick" authorityYear="1993" class="Arachnida" family="Araneidae" genus="Novakiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Novakiella" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Novakiella</emphasis>
|
||
</taxonomicName>
|
||
is convoluted.
|
||
<taxonomicName baseAuthorityName="Roewer" baseAuthorityYear="1942" class="Arachnida" family="Araneidae" genus="Novakiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Novakiella trituberculosa" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="species" species="trituberculosa">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Novakiella trituberculosa</emphasis>
|
||
</taxonomicName>
|
||
was first described as
|
||
<taxonomicName authority="Walckenaer, 1805" authorityName="Walckenaer" authorityYear="1805" class="Arachnida" family="Araneidae" genus="Epeira" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Epeira tri-tuberculata" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="species" species="tri-tuberculata">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Epeira tri-tuberculata</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation author="Urquhart, AT" journalOrPublisher="Transactions of the New Zealand Institute" pageId="0" pageNumber="393" pagination="72 - 118" refId="B17" refString="Urquhart, AT, 1887. On new species of Araneida. Transactions of the New Zealand Institute 19: 72 - 118" title="On new species of Araneida." volume="19" year="1887">Urquhart (1887)</bibRefCitation>
|
||
, before
|
||
<bibRefCitation author="Roewer, CF" journalOrPublisher="Natura, Buchhandlung fuer Naturkunde und exakte Wissenschaften Paul Budy Bremen" pageId="0" pageNumber="393" refId="B14" refString="Roewer, CF, 1942. Katalog der Araneae von 1758 bis 1940. 1. Band (Mesothelae, Orthognatha, Labidognatha: Dysderaeformia, Scytodiformia, Pholciformia, Zodariiformia, Hersiliaeformia, Argyopiformia). Natura, Buchhandlung fuer Naturkunde und exakte Wissenschaften Paul Budy Bremen" title="Katalog der Araneae von 1758 bis 1940. 1. Band (Mesothelae, Orthognatha, Labidognatha: Dysderaeformia, Scytodiformia, Pholciformia, Zodariiformia, Hersiliaeformia, Argyopiformia)." year="1942">Roewer (1942)</bibRefCitation>
|
||
replaced the species-group name as it is a junior primary homonym of
|
||
<taxonomicName authority="Walckenaer, 1805" authorityName="Walckenaer" authorityYear="1805" class="Arachnida" family="Araneidae" genus="Epeira" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Epeira trituberculata" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="species" species="trituberculata">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Epeira trituberculata</emphasis>
|
||
</taxonomicName>
|
||
Lucas, 1846, currently listed as a junior synonym of
|
||
<taxonomicName baseAuthorityName="Costa" baseAuthorityYear="1834" class="Arachnida" family="Araneidae" genus="Cyclosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclosa insulana" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="species" species="insulana">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Cyclosa insulana</emphasis>
|
||
</taxonomicName>
|
||
(Costa, 1834).
|
||
<bibRefCitation author="Court, DJ" journalOrPublisher="Otago Museum Bulletin" pageId="0" pageNumber="393" pagination="68 - 124" refId="B3" refString="Court, DJ, Forster, RR, 1988. The spiders of New Zealand: Part VI. Family Araneidae. Otago Museum Bulletin 6: 68 - 124" title="The spiders of New Zealand: Part VI. Family Araneidae." volume="6" year="1988">Court and Forster (1988)</bibRefCitation>
|
||
described the genus
|
||
<taxonomicName class="Insecta" family="Araneidae" genus="Novakia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Novakia" order="Araneae" pageId="0" pageNumber="393" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Novakia</emphasis>
|
||
</taxonomicName>
|
||
to accommodate this species; however, this new genus-group name was also preoccupied, by
|
||
<taxonomicName class="Insecta" family="Araneidae" genus="Novakia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Novakia" order="Araneae" pageId="0" pageNumber="393" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Novakia</emphasis>
|
||
</taxonomicName>
|
||
Strobl, 1893 (
|
||
<taxonomicName class="Insecta" family="Araneidae" genus="Novakia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Diptera" pageId="0" pageNumber="393" phylum="Mollusca" rank="order">Diptera</taxonomicName>
|
||
) and
|
||
<taxonomicName class="Insecta" family="Araneidae" genus="Novakia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Novakia" order="Araneae" pageId="0" pageNumber="393" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Novakia</emphasis>
|
||
</taxonomicName>
|
||
Tolmachoff, 1926 (
|
||
<taxonomicName family="Araneidae" genus="Novakia" kingdom="Animalia" lsidName="" order="Araneae" pageId="0" pageNumber="393" phylum="Mollusca" rank="phylum">Mollusca</taxonomicName>
|
||
). Court and Forster, in
|
||
<bibRefCitation author="Platnick, NI" journalOrPublisher="Molecular Biology and Evolution" pageId="0" pageNumber="393" refId="B13" refString="Platnick, NI, 1993. Advances in spider taxonomy 1988-1991, with synonymies and transfers 1940-1980. The New York Entomological Society New York, 846 pp." title="Advances in spider taxonomy 1988 - 1991, with synonymies and transfers 1940 - 1980. The New York Entomological Society New York, 846 pp." year="1993">Platnick (1993)</bibRefCitation>
|
||
, proposed
|
||
<taxonomicName authorityName="Court & Forster, 1993, in Platnick" authorityYear="1993" class="Arachnida" family="Araneidae" genus="Novakiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Novakiella" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Novakiella</emphasis>
|
||
</taxonomicName>
|
||
Court & Forster, 1993 as a replacement name.
|
||
</paragraph>
|
||
<caption doi="10.3897/zse.97.67788.figure5" httpUri="https://binary.pensoft.net/fig/570635" pageId="0" pageNumber="393" start="Figure 5" startId="F5">
|
||
<paragraph pageId="0" pageNumber="393">
|
||
<emphasis bold="true" pageId="0" pageNumber="393">Figure 5.</emphasis>
|
||
<taxonomicName authorityName="Framenau & Vink & Scharff & Baptista & Castanheira" authorityYear="2021" class="Arachnida" family="Araneidae" genus="Novakiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Novakiella boletus" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="species" species="boletus">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Novakiella boletus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., male holotype (MV K9867), expanded left pedipalp. mesal view. Abbreviations: C, conductor; CL, conductor lobe; E, embolus; MA, median apophysis; Ra, radix; TA, terminal apophysis. Scale bar: 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zse.97.67788.figure6" httpUri="https://binary.pensoft.net/fig/570636" pageId="0" pageNumber="393" start="Figure 6" startId="F6">
|
||
<paragraph pageId="0" pageNumber="393">
|
||
<emphasis bold="true" pageId="0" pageNumber="393">Figure 6.</emphasis>
|
||
<taxonomicName authorityName="Framenau & Vink & Scharff & Baptista & Castanheira" authorityYear="2021" class="Arachnida" family="Araneidae" genus="Novakiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Novakiella boletus" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="species" species="boletus">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Novakiella boletus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., female (SAM).
|
||
<emphasis bold="true" pageId="0" pageNumber="393">A.</emphasis>
|
||
Dorsal habitus;
|
||
<emphasis bold="true" pageId="0" pageNumber="393">B.</emphasis>
|
||
Ventral habitus;
|
||
<emphasis bold="true" pageId="0" pageNumber="393">C.</emphasis>
|
||
Epigyne, ventral view. Abbreviations: CO, copulatory openings; L, lips; S, scape. Scale bars:
|
||
<emphasis bold="true" pageId="0" pageNumber="393">A, B</emphasis>
|
||
, 2 mm,
|
||
<emphasis bold="true" pageId="0" pageNumber="393">C</emphasis>
|
||
, 0.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="393" type="distribution">
|
||
<paragraph pageId="0" pageNumber="393">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="393">
|
||
Australia and New Zealand (Figs
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Distribution records of Novakiella trituberculosa (green circles) and Novakiella boletus sp. nov. (red stars) in Australia." figureDoi="10.3897/zse.97.67788.figure7" httpUri="https://binary.pensoft.net/fig/570637" pageId="0" pageNumber="393">7</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Distribution records of Novakiella trituberculosa in New Zealand; red circle points to material examined for this study and yellow diamonds point to records literature records (Court and Forster 1988)." figureDoi="10.3897/zse.97.67788.figure8" httpUri="https://binary.pensoft.net/fig/570638" pageId="0" pageNumber="393">8</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/zse.97.67788.figure7" httpUri="https://binary.pensoft.net/fig/570637" pageId="0" pageNumber="393" start="Figure 7" startId="F7">
|
||
<paragraph pageId="0" pageNumber="393">
|
||
<emphasis bold="true" pageId="0" pageNumber="393">Figure 7.</emphasis>
|
||
Distribution records of
|
||
<taxonomicName baseAuthorityName="Roewer" baseAuthorityYear="1942" class="Arachnida" family="Araneidae" genus="Novakiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Novakiella trituberculosa" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="species" species="trituberculosa">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Novakiella trituberculosa</emphasis>
|
||
</taxonomicName>
|
||
(green circles) and
|
||
<taxonomicName authorityName="Framenau & Vink & Scharff & Baptista & Castanheira" authorityYear="2021" class="Arachnida" family="Araneidae" genus="Novakiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Novakiella boletus" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="species" species="boletus">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Novakiella boletus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (red stars) in Australia.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zse.97.67788.figure8" httpUri="https://binary.pensoft.net/fig/570638" pageId="0" pageNumber="393" start="Figure 8" startId="F8">
|
||
<paragraph pageId="0" pageNumber="393">
|
||
<emphasis bold="true" pageId="0" pageNumber="393">Figure 8.</emphasis>
|
||
Distribution records of
|
||
<taxonomicName baseAuthorityName="Roewer" baseAuthorityYear="1942" class="Arachnida" family="Araneidae" genus="Novakiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Novakiella trituberculosa" order="Araneae" pageId="0" pageNumber="393" phylum="Arthropoda" rank="species" species="trituberculosa">
|
||
<emphasis italics="true" pageId="0" pageNumber="393">Novakiella trituberculosa</emphasis>
|
||
</taxonomicName>
|
||
in New Zealand; red circle points to material examined for this study and yellow diamonds point to records literature records (
|
||
<bibRefCitation author="Court, DJ" journalOrPublisher="Otago Museum Bulletin" pageId="0" pageNumber="393" pagination="68 - 124" refId="B3" refString="Court, DJ, Forster, RR, 1988. The spiders of New Zealand: Part VI. Family Araneidae. Otago Museum Bulletin 6: 68 - 124" title="The spiders of New Zealand: Part VI. Family Araneidae." volume="6" year="1988">Court and Forster 1988</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |