treatments-xml/data/A9/98/B6/A998B605B61B555F968D6793005DCAD1.xml

<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.48.4446" ID-PMC="PMC4408729" ID-Pensoft-Pub="1314-2003-48-21" ID-Pensoft-UUID="FFEAFFE2DA1FFF8F69507C75FF9CFF83" ID-PubMed="25931971" ID-Zenodo-Dep="576257" ModsDocID="1314-2003-48-21" checkinTime="1451251555515" checkinUser="pensoft" docAuthor="Hamzaoglu, Ergin, Koc, Murat &amp; Aksoy, Ahmet" docDate="2015" docId="A998B605B61B555F968D6793005DCAD1" docLanguage="en" docName="PhytoKeys 48: 21-28" docOrigin="PhytoKeys 48" docPubDate="2015-04-02" docSource="http://dx.doi.org/10.3897/phytokeys.48.4446" docTitle="Dianthus aticii Hamzaoglu 2015, sp. nov." docType="treatment" docVersion="3" id="FFEAFFE2DA1FFF8F69507C75FF9CFF83" lastPageNumber="26" masterDocId="FFEAFFE2DA1FFF8F69507C75FF9CFF83" masterDocTitle="Dianthus aticii, a new species from Turkey (Caryophyllaceae)" masterLastPageNumber="28" masterPageNumber="21" pageNumber="22" updateTime="1668141338959" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Dianthus aticii, a new species from Turkey (Caryophyllaceae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hamzaoglu, Ergin</mods:namePart>
<mods:affiliation>Department of Elementary Education, Gazi Faculty of Education, Gazi University, 06500, Ankara, Turkey</mods:affiliation>
<mods:nameIdentifier type="email">erginhamzaoglu@yahoo.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Koc, Murat</mods:namePart>
<mods:affiliation>Animal Production High School, Bozok University, 66900, Yozgat, Turkey</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Aksoy, Ahmet</mods:namePart>
<mods:affiliation>Department of Biology, Science Faculty, Akdeniz University, 07058, Antalya, Turkey</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2015</mods:date>
<mods:detail type="pubDate">
<mods:number>2015-04-02</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>48</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>21</mods:start>
<mods:end>28</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/phytokeys.48.4446</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.48.4446</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-48-21</mods:identifier>
<mods:identifier type="Pensoft-UUID">FFEAFFE2DA1FFF8F69507C75FF9CFF83</mods:identifier>
<mods:identifier type="Zenodo-Dep">576257</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="182228351" LSID="urn:lsid:plazi:treatment:A998B605B61B555F968D6793005DCAD1" httpUri="http://treatment.plazi.org/id/A998B605B61B555F968D6793005DCAD1" lastPageId="5" lastPageNumber="26" pageId="1" pageNumber="22">
<subSubSection pageId="1" pageNumber="22" type="nomenclature">
<paragraph pageId="1" pageNumber="22">
<taxonomicName LSID="A998B605-B61B-555F-968D-6793005DCAD1" authority="Hamzaoğlu" authorityName="Hamzaoglu" authorityYear="2015" class="Magnoliopsida" family="Caryophyllaceae" genus="Dianthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dianthus aticii" order="Caryophyllales" pageId="1" pageNumber="22" phylum="Tracheophyta" rank="species" species="aticii" status="sp. nov.">
Dianthus aticii 
<normalizedToken originalValue="Hamzaoğlu">Hamzaoglu</normalizedToken>
</taxonomicName>
<taxonomicNameLabel pageId="1" pageNumber="22">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Photographs of plant habit and flowers of Dianthus aticii and Dianthus zonatus. Dianthus aticii - A 1 Habit B 1 Flower; Dianthus zonatus - A 2 Habit B 2 Flower." figureDoi="10.3897/phytokeys.48.4446.figure1" httpUri="https://binary.pensoft.net/fig/41398" pageId="1" pageNumber="22">Figs 1</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. SEM photographs of the seed coat. A Dianthus aticii B Dianthus zonatus 1 - 3 dorsal surface 4 - 6 ventral surface (scale bars: 1 and 4: 1 mm, 2 and 5: 20 μm, 3 and 6: 10 μm)." figureDoi="10.3897/phytokeys.48.4446.figure2" httpUri="https://binary.pensoft.net/fig/41399" pageId="1" pageNumber="22">, 2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection lastPageId="2" lastPageNumber="23" pageId="1" pageNumber="22" type="diagnosis">
<paragraph pageId="1" pageNumber="22">Diagnosis.</paragraph>
<paragraph lastPageId="2" lastPageNumber="23" pageId="1" pageNumber="22">
Stems suffruticose (not herbaceous); sterile shoot leaves subulate, subcanaliculate (not linear, flattened or absent); inner epicalyx scales with scarious margin 
<pageBreakToken pageId="2" pageNumber="23" start="start">0.2</pageBreakToken>
-0.4 mm wide, arista 1/10-1/7 as long as scale (not with scarious margin 0.3-0.8 mm wide, arista 1/7-1/3 as long as scale).
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="23" type="type">
<paragraph pageId="2" pageNumber="23">Type.</paragraph>
<paragraph pageId="2" pageNumber="23">
<emphasis bold="true" pageId="2" pageNumber="23">TURKEY. Bilecik:</emphasis>
Bilecik highway exit towards 
<normalizedToken originalValue="Eskişehir">Eskisehir</normalizedToken>
, 
<geoCoordinate degrees="40" direction="north" minutes="06" orientation="latitude" precision="15" seconds="27" value="40.1075">40°06'27&quot;N</geoCoordinate>
, 29°59'47'E, 330 m, stony slopes and steppes, 16 June 2013 (fl, fr), 
<emphasis italics="true" pageId="2" pageNumber="23">
E. 
<normalizedToken originalValue="Hamzaoğlu">Hamzaoglu</normalizedToken>
et al. 6743
</emphasis>
(holotype: GAZI; isotypes: GAZI, ANK); 
<emphasis bold="true" pageId="2" pageNumber="23">Bolu:</emphasis>
Seben, between Bozyer and Korucuk villages, 1025 m, forest clearings, flowing slopes, 19 July 2013, 
<emphasis italics="true" pageId="2" pageNumber="23">
M. 
<normalizedToken originalValue="Koç">Koc</normalizedToken>
&amp; E. 
<normalizedToken originalValue="Hamzaoğlu">Hamzaoglu</normalizedToken>
6868
</emphasis>
(paratypes: GAZI, ANK); 
<emphasis bold="true" pageId="2" pageNumber="23">Ankara:</emphasis>
<normalizedToken originalValue="Nallıhan">Nallihan</normalizedToken>
, 
<normalizedToken originalValue="Gökçeöz">Goekceoez</normalizedToken>
village, road of forest watchtower, 820 m, forest clearings, stony slopes, 19 July 2013, 
<emphasis italics="true" pageId="2" pageNumber="23">
M. 
<normalizedToken originalValue="Koç">Koc</normalizedToken>
&amp; E. 
<normalizedToken originalValue="Hamzaoğlu">Hamzaoglu</normalizedToken>
6869
</emphasis>
(paratypes: GAZI, ANK).
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="23" type="specimens examined">
<paragraph pageId="2" pageNumber="23">Specimens examined.</paragraph>
<paragraph pageId="2" pageNumber="23">
<emphasis bold="true" pageId="2" pageNumber="23">
<taxonomicName authorityName="Hamzaoglu" authorityYear="2015" class="Magnoliopsida" family="Caryophyllaceae" genus="Dianthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dianthus aticii" order="Caryophyllales" pageId="2" pageNumber="23" phylum="Tracheophyta" rank="species" species="aticii">
<emphasis italics="true" pageId="2" pageNumber="23">Dianthus aticii</emphasis>
</taxonomicName>
<normalizedToken originalValue="Hamzaoğlu">Hamzaoglu</normalizedToken>
sp. nov.
</emphasis>
- 
<emphasis bold="true" pageId="2" pageNumber="23">TURKEY. Bilecik:</emphasis>
Bilecik highway exit towards 
<normalizedToken originalValue="Eskişehir">Eskisehir</normalizedToken>
, 
<geoCoordinate degrees="40" direction="north" minutes="06" orientation="latitude" precision="15" seconds="27" value="40.1075">40°06'27&quot;N</geoCoordinate>
, 29°59'47'E, 330 m, stony slopes and steppes, 16 June 2013, 
<emphasis italics="true" pageId="2" pageNumber="23">
E. 
<normalizedToken originalValue="Hamzaoğlu">Hamzaoglu</normalizedToken>
et al. 6743
</emphasis>
(holotype: GAZI; isotypes: GAZI, ANK); 
<emphasis bold="true" pageId="2" pageNumber="23">Bolu:</emphasis>
Seben, between Bozyer and Korucuk villages, 1025 m, forest clearings, flowing slopes, 19 July 2013, 
<emphasis italics="true" pageId="2" pageNumber="23">
M. 
<normalizedToken originalValue="Koç">Koc</normalizedToken>
&amp; E. 
<normalizedToken originalValue="Hamzaoğlu">Hamzaoglu</normalizedToken>
6868
</emphasis>
(paratypes: GAZI, ANK); 
<emphasis bold="true" pageId="2" pageNumber="23">Ankara:</emphasis>
<normalizedToken originalValue="Nallıhan">Nallihan</normalizedToken>
, 
<normalizedToken originalValue="Gökçeöz">Goekceoez</normalizedToken>
village, road of forest watchtower, 820 m, forest clearings, stony slopes, 19 July 2013, 
<emphasis italics="true" pageId="2" pageNumber="23">
M. 
<normalizedToken originalValue="Koç">Koc</normalizedToken>
&amp; E. 
<normalizedToken originalValue="Hamzaoğlu">Hamzaoglu</normalizedToken>
6869
</emphasis>
(paratypes: GAZI, ANK); 
<emphasis bold="true" pageId="2" pageNumber="23">
<taxonomicName authorityName="Fenzl" authorityYear="1842" class="Magnoliopsida" family="Caryophyllaceae" genus="Dianthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dianthus zonatus" order="Caryophyllales" pageId="2" pageNumber="23" phylum="Tracheophyta" rank="species" species="zonatus">
<emphasis italics="true" pageId="2" pageNumber="23">Dianthus zonatus</emphasis>
</taxonomicName>
Fenzl
</emphasis>
- 
<emphasis bold="true" pageId="2" pageNumber="23">TURKEY. Manisa:</emphasis>
Spil 
<normalizedToken originalValue="Dağı">Dagi</normalizedToken>
National Park, road of 
<normalizedToken originalValue="Atalanı">Atalani</normalizedToken>
resting area, 1320 m, calcerous rocks, 2 July 2011, 
<emphasis italics="true" pageId="2" pageNumber="23">
M. 
<normalizedToken originalValue="Koç">Koc</normalizedToken>
&amp; E. 
<normalizedToken originalValue="Hamzaoğlu">Hamzaoglu</normalizedToken>
6106
</emphasis>
(GAZI); 
<emphasis bold="true" pageId="2" pageNumber="23">
<normalizedToken originalValue="Kütahya">Kuetahya</normalizedToken>
:
</emphasis>
<normalizedToken originalValue="İscehisar">Iscehisar</normalizedToken>
, around Seydiler, 1150 m, rocks, 5 August 2012, 
<emphasis italics="true" pageId="2" pageNumber="23">
E. 
<normalizedToken originalValue="Hamzaoğlu">Hamzaoglu</normalizedToken>
et al. 6584
</emphasis>
(GAZI); 
<emphasis bold="true" pageId="2" pageNumber="23">
<normalizedToken originalValue="Eskişehir">Eskisehir</normalizedToken>
:
</emphasis>
Around Sivrihisar, 1115 m, rocks, 24 June 2012, 
<emphasis italics="true" pageId="2" pageNumber="23">
E. 
<normalizedToken originalValue="Hamzaoğlu">Hamzaoglu</normalizedToken>
et al. 6339
</emphasis>
(GAZI); 
<emphasis bold="true" pageId="2" pageNumber="23">Konya:</emphasis>
Between Kulu and Cihanbeyli, Kulu exit, 1130 m, steppe, 13 July 2011, 
<emphasis italics="true" pageId="2" pageNumber="23">
E. 
<normalizedToken originalValue="Hamzaoğlu">Hamzaoglu</normalizedToken>
et al. 6122
</emphasis>
(GAZI); 
<emphasis bold="true" pageId="2" pageNumber="23">Ankara:</emphasis>
<normalizedToken originalValue="Polatlı">Polatli</normalizedToken>
, above 
<normalizedToken originalValue="Babayokuş">Babayokus</normalizedToken>
village, 900 m, stony places, 2 July 2010, 
<emphasis italics="true" pageId="2" pageNumber="23">
M. 
<normalizedToken originalValue="Koç">Koc</normalizedToken>
et al. 1205
</emphasis>
(GAZI); 
<emphasis bold="true" pageId="2" pageNumber="23">
<normalizedToken originalValue="Aydın">Aydin</normalizedToken>
:
</emphasis>
Between 
<normalizedToken originalValue="Söke">Soeke</normalizedToken>
and Didim, after 4 km from 
<normalizedToken originalValue="Güllübahçe">Guelluebahce</normalizedToken>
exit, 820 m, 25 June 2006, 
<emphasis italics="true" pageId="2" pageNumber="23">
E. 
<normalizedToken originalValue="Hamzaoğlu">Hamzaoglu</normalizedToken>
et al. 4071
</emphasis>
(GAZI); 
<emphasis bold="true" pageId="2" pageNumber="23">
<normalizedToken originalValue="Muğla">Mugla</normalizedToken>
:
</emphasis>
<normalizedToken originalValue="Köyceğiz">Koeycegiz</normalizedToken>
, above Yayla village, from 
<normalizedToken originalValue="Gökçeova">Goekceova</normalizedToken>
Lake to Sandras Mountain summit, 1950 m, serpentine rocks, 15 July 2011, 
<emphasis italics="true" pageId="2" pageNumber="23">
E. 
<normalizedToken originalValue="Hamzaoğlu">Hamzaoglu</normalizedToken>
et al. 6198
</emphasis>
(GAZI); 
<emphasis bold="true" pageId="2" pageNumber="23">Antalya:</emphasis>
<normalizedToken originalValue="Elmalı">Elmali</normalizedToken>
, N of Vahhabi 
<normalizedToken originalValue="Ümmi">Uemmi</normalizedToken>
<normalizedToken originalValue="Türbesi">Tuerbesi</normalizedToken>
, 1480 m, rocks, 12 June 2007, 
<emphasis italics="true" pageId="2" pageNumber="23">
M. 
<normalizedToken originalValue="Koç">Koc</normalizedToken>
&amp; 
<normalizedToken originalValue="Ü">Ue</normalizedToken>
. Budak 2152
</emphasis>
(GAZI); 
<emphasis bold="true" pageId="2" pageNumber="23">Karaman:</emphasis>
Between Ermenek and Karaman, 16 km, 1670 m, Pine forest openings, stony places, 18 July 2005, 
<emphasis italics="true" pageId="2" pageNumber="23">
<normalizedToken originalValue="Ü">Ue</normalizedToken>
. Budak et al. 1743
</emphasis>
(GAZI); 
<emphasis bold="true" pageId="2" pageNumber="23">
<normalizedToken originalValue="Niğde">Nigde</normalizedToken>
:
</emphasis>
<normalizedToken originalValue="Çamardı">Camardi</normalizedToken>
, above 
<normalizedToken originalValue="Demirkazık">Demirkazik</normalizedToken>
village, 1475 m, rocks, 11.7.2012, 
<emphasis italics="true" pageId="2" pageNumber="23">
E. 
<normalizedToken originalValue="Hamzaoğlu">Hamzaoglu</normalizedToken>
et al. 6449
</emphasis>
(GAZI).
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="24" pageId="2" pageNumber="23" type="description">
<paragraph pageId="2" pageNumber="23">Description.</paragraph>
<paragraph lastPageId="3" lastPageNumber="24" pageId="2" pageNumber="23">
Suffruticose, several-stemmed, subpruinose herbs. Stems erect, fragile, 20-35 cm tall, branching from upper nodes, 6-10-nodes, glabrous or puberulent. Leaves subcanaliculate, thick, glabrous or puberulent, margins scabrous, ciliate and scarious at base, apex acuminate; sterile shoot leaves subulate, equal or longer than cauline leaves; cauline leaves subulate to linear-filiform, 11-22 
<normalizedToken originalValue="×">x</normalizedToken>
0.6-1.2 mm, appressed to stem, obviously shorter than internodes, rigid, 3-veined, sheaths equal or slightly longer than wide; upper similar but smaller. Flowers solitary or few in racemes; branches angled at 5-15°, glabrous or sparsely puberulent, up to 3 cm long; pedicels 5-15 mm, glabrous or sparsely puberulent, greenish. Epicalyx scales (4-)6-8(-12), cartilaginous, greenish or straw-coloured, glabrous or puberulent, appressed to calyx, apex acute to acuminate except arista; outer linear-lanceolate, veinless below, indistinctly 5-9-veined above, 1/5-2/5 as long as calyx, 4-8 
<normalizedToken originalValue="×">x</normalizedToken>
0.8-1.2 mm, with narrowly scari 
<pageBreakToken pageId="3" pageNumber="24" start="start">ous</pageBreakToken>
(c. 0.2 mm) margins, arista 1/2-2/3 as long as scale; inner oblong-oblanceolate, veinless below, indistinctly 7-9-veined above, 2/5-1/2 as long as calyx, 6-9 
<normalizedToken originalValue="×">x</normalizedToken>
2.5-3.5 mm, with scarious (0.2-0.4 mm) margins, arista 1/10-1/7 as long as scale. Calyx cylindric-lanceolate, 16-22 
<normalizedToken originalValue="×">x</normalizedToken>
3-4.5 mm, distinctly 36-40-veined above, glabrous or puberulent, pale green or sometimes purplish; teeth triangular-lanceolate, 4-5.5 
<normalizedToken originalValue="×">x</normalizedToken>
1.2-2 mm, 7-veined, with ciliate and scarious margins, apex acute to acuminate, sometimes short mucronate. Petals 20-23 mm long; limb broadly cuneate, 7-8 
<normalizedToken originalValue="×">x</normalizedToken>
6-7 mm, c. 1/3 as long as petal, completely exserted from calyx, usually spotted, barbulate, pink, yellowish-green beneath, 7-11-toothed to apex, teeth triangular, up to 1/6 as long as limb; claw 12-15 
<normalizedToken originalValue="×">x</normalizedToken>
1.5 mm, collar almost as wide as claw. Capsule equal in length to calyx. Seeds elliptical, 2-3 
<normalizedToken originalValue="×">x</normalizedToken>
1.4-2 mm, blackish.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="24" type="distinction from other taxa">
<paragraph pageId="3" pageNumber="24">Distinction from other taxa.</paragraph>
<paragraph pageId="3" pageNumber="24">
<taxonomicName authorityName="Hamzaoglu" authorityYear="2015" class="Magnoliopsida" family="Caryophyllaceae" genus="Dianthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dianthus aticii" order="Caryophyllales" pageId="3" pageNumber="24" phylum="Tracheophyta" rank="species" species="aticii">
<emphasis italics="true" pageId="3" pageNumber="24">Dianthus aticii</emphasis>
</taxonomicName>
shows close similarities to 
<taxonomicName authorityName="Fenzl" authorityYear="1842" class="Magnoliopsida" family="Caryophyllaceae" genus="Dianthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dianthus zonatus" order="Caryophyllales" pageId="3" pageNumber="24" phylum="Tracheophyta" rank="species" species="zonatus">
<emphasis italics="true" pageId="3" pageNumber="24">Dianthus zonatus</emphasis>
</taxonomicName>
Fenzl because of toothed and barbulate petals, solitary or double flowers, and epicalyx scales that reach up to half of its calyx length (
<bibRefCitation author="Fenzl, E" journalOrPublisher="Vienne" pageId="6" pageNumber="27" refId="B7" refString="Fenzl, E, 1842. Pugillus Plantarum Novarum Syriae et Tauri Occidentalis Primus. apud F. Beck. Vienne, 11 pp." title="Pugillus Plantarum Novarum Syriae et Tauri Occidentalis Primus. apud F. Beck" year="1842">Fenzl 1842</bibRefCitation>
, 
<bibRefCitation author="Boissier, E" journalOrPublisher="Typis Marci Ducloux et Cons, Paris" pageId="5" pageNumber="26" pagination="67 - 68" refId="B3" refString="Boissier, E, 1849. Diagnoses Plantarum Orientalium novarum. Typis Marci Ducloux et Cons, Paris: 67 - 68" title="Diagnoses Plantarum Orientalium novarum" year="1849">Boissier 1849</bibRefCitation>
, 
<bibRefCitation author="Tchihatcheff, P" journalOrPublisher="Gide [et] Baudry, Paris" pageId="6" pageNumber="27" refId="B15" refString="Tchihatcheff, P, 1860. Asie Mineure: description physique, statistique et archeologique de cette contree. Gide [et] Baudry, Paris" title="Asie Mineure: description physique, statistique et archeologique de cette contree" year="1860">Tchihatcheff 1860</bibRefCitation>
, 
<bibRefCitation author="Reeve, H" journalOrPublisher="Flora of Turkey and the East Aegean Islands" pageId="6" pageNumber="27" pagination="99 - 131" refId="B14" refString="Reeve, H, 1967. Dianthus L. Flora of Turkey and the East Aegean Islands Vol. 2: 99 - 131" title="Dianthus L." volume="Vol. 2" year="1967">Reeve 1967</bibRefCitation>
). Despite these similarities, there are very distinctive differences between 
<taxonomicName authorityName="Hamzaoglu" authorityYear="2015" class="Magnoliopsida" family="Caryophyllaceae" genus="Dianthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dianthus aticii" order="Caryophyllales" pageId="3" pageNumber="24" phylum="Tracheophyta" rank="species" species="aticii">
<emphasis italics="true" pageId="3" pageNumber="24">Dianthus aticii</emphasis>
</taxonomicName>
and 
<taxonomicName authorityName="Fenzl" authorityYear="1842" class="Magnoliopsida" family="Caryophyllaceae" genus="Dianthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dianthus zonatus" order="Caryophyllales" pageId="3" pageNumber="24" phylum="Tracheophyta" rank="species" species="zonatus">
<emphasis italics="true" pageId="3" pageNumber="24">Dianthus zonatus</emphasis>
</taxonomicName>
such as stem morphology, leaf shape, and size of epicalyx scales and petals (Table 
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Diagnostic characters between Dianthus aticii and Dianthus zonatus." httpUri="http://table.plazi.org/id/AB3FCB63F7013DE8DB74F83666F1C16A" pageId="3" pageNumber="24" tableUuid="AB3FCB63F7013DE8DB74F83666F1C16A">1</tableCitation>
, Figure 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. SEM photographs of the seed coat. A Dianthus aticii B Dianthus zonatus 1 - 3 dorsal surface 4 - 6 ventral surface (scale bars: 1 and 4: 1 mm, 2 and 5: 20 μm, 3 and 6: 10 μm)." figureDoi="10.3897/phytokeys.48.4446.figure2" httpUri="https://binary.pensoft.net/fig/41399" pageId="3" pageNumber="24">2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="24" type="key to the two closely related dianthus species">
<paragraph pageId="3" pageNumber="24">
Key to the two closely related 
<taxonomicName class="Magnoliopsida" family="Caryophyllaceae" genus="Dianthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dianthus" order="Caryophyllales" pageId="3" pageNumber="24" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="3" pageNumber="24">Dianthus</emphasis>
</taxonomicName>
species
</paragraph>
<paragraph pageId="3" pageNumber="24">
<table inLine="true" pageId="3" pageNumber="24">
<tr pageId="3" pageNumber="24">
<td colspan="1" pageId="3" pageNumber="24" rowspan="1">1</td>
<td colspan="1" pageId="3" pageNumber="24" rowspan="1">Stems suffruticose; sterile shoot leaves subulate and subcanaliculate; inner epicalyx scales with scarious margin 0.2-0.4 mm wide; petals 20-23 mm long</td>
<td colspan="1" pageId="3" pageNumber="24" rowspan="1">
<emphasis bold="true" pageId="3" pageNumber="24">
<taxonomicName authorityName="Hamzaoglu" authorityYear="2015" class="Magnoliopsida" family="Caryophyllaceae" genus="Dianthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dianthus aticii" order="Caryophyllales" pageId="3" pageNumber="24" phylum="Tracheophyta" rank="species" species="aticii">
<emphasis italics="true" pageId="3" pageNumber="24">Dianthus aticii</emphasis>
</taxonomicName>
sp. nov.
</emphasis>
</td>
</tr>
<tr pageId="3" pageNumber="24">
<td colspan="1" pageId="3" pageNumber="24" rowspan="1">-</td>
<td colspan="1" pageId="3" pageNumber="24" rowspan="1">Stems herbaceous; sterile shoot leaves linear and flattened or absent; inner epicalyx scales with scarious margin 0.3-0.8 mm wide; petals 24-31 mm long</td>
<td colspan="1" pageId="3" pageNumber="24" rowspan="1">
<taxonomicName authorityName="Fenzl" authorityYear="1842" class="Magnoliopsida" family="Caryophyllaceae" genus="Dianthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dianthus zonatus" order="Caryophyllales" pageId="3" pageNumber="24" phylum="Tracheophyta" rank="species" species="zonatus">
<emphasis bold="true" italics="true" pageId="3" pageNumber="24">Dianthus zonatus</emphasis>
</taxonomicName>
</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="25" type="seed morphology">
<paragraph pageId="4" pageNumber="25">
<pageBreakToken pageId="4" pageNumber="25" start="start">Seed</pageBreakToken>
morphology.
</paragraph>
<paragraph pageId="4" pageNumber="25">
Seeds of 
<taxonomicName authorityName="Hamzaoglu" authorityYear="2015" class="Magnoliopsida" family="Caryophyllaceae" genus="Dianthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dianthus aticii" order="Caryophyllales" pageId="4" pageNumber="25" phylum="Tracheophyta" rank="species" species="aticii">
<emphasis italics="true" pageId="4" pageNumber="25">Dianthus aticii</emphasis>
</taxonomicName>
are elliptical, 2-3 
<normalizedToken originalValue="×">x</normalizedToken>
1.4-2 mm, black, granular; dorsal surface convex, with regular rectangular cells, tuberculate, with 4-7 teeth on each margin, teeth V-undulate, apparent; ventral surface flat, with irregular rectangular cells, tuberculate, with 4-7 teeth on each margin, teeth S-undulate, not apparent; apex beaked. The seeds of 
<taxonomicName authorityName="Hamzaoglu" authorityYear="2015" class="Magnoliopsida" family="Caryophyllaceae" genus="Dianthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dianthus aticii" order="Caryophyllales" pageId="4" pageNumber="25" phylum="Tracheophyta" rank="species" species="aticii">
<emphasis italics="true" pageId="4" pageNumber="25">Dianthus aticii</emphasis>
</taxonomicName>
are different from the seeds of 
<taxonomicName authorityName="Fenzl" authorityYear="1842" class="Magnoliopsida" family="Caryophyllaceae" genus="Dianthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dianthus zonatus" order="Caryophyllales" pageId="4" pageNumber="25" phylum="Tracheophyta" rank="species" species="zonatus">
<emphasis italics="true" pageId="4" pageNumber="25">Dianthus zonatus</emphasis>
</taxonomicName>
in terms of shape and cell edges of both the dorsal and ventral surfaces (Table 
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Diagnostic characters between Dianthus aticii and Dianthus zonatus." httpUri="http://table.plazi.org/id/AB3FCB63F7013DE8DB74F83666F1C16A" pageId="4" pageNumber="25" tableUuid="AB3FCB63F7013DE8DB74F83666F1C16A">1</tableCitation>
, Figure 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. SEM photographs of the seed coat. A Dianthus aticii B Dianthus zonatus 1 - 3 dorsal surface 4 - 6 ventral surface (scale bars: 1 and 4: 1 mm, 2 and 5: 20 μm, 3 and 6: 10 μm)." figureDoi="10.3897/phytokeys.48.4446.figure2" httpUri="https://binary.pensoft.net/fig/41399" pageId="4" pageNumber="25">2</figureCitation>
).
</paragraph>
<caption ID-Zenodo-Dep="961045" doi="10.3897/phytokeys.48.4446.figure1" httpUri="https://binary.pensoft.net/fig/41398" pageId="4" pageNumber="25" start="Figure 1" startId="F1">
<paragraph pageId="4" pageNumber="25">
<emphasis bold="true" pageId="4" pageNumber="25">Figure 1.</emphasis>
Photographs of plant habit and flowers of 
<taxonomicName authorityName="Hamzaoglu" authorityYear="2015" class="Magnoliopsida" family="Caryophyllaceae" genus="Dianthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dianthus aticii" order="Caryophyllales" pageId="4" pageNumber="25" phylum="Tracheophyta" rank="species" species="aticii">
<emphasis italics="true" pageId="4" pageNumber="25">Dianthus aticii</emphasis>
</taxonomicName>
and 
<taxonomicName authorityName="Fenzl" authorityYear="1842" class="Magnoliopsida" family="Caryophyllaceae" genus="Dianthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dianthus zonatus" order="Caryophyllales" pageId="4" pageNumber="25" phylum="Tracheophyta" rank="species" species="zonatus">
<emphasis italics="true" pageId="4" pageNumber="25">Dianthus zonatus</emphasis>
</taxonomicName>
. 
<taxonomicName authorityName="Hamzaoglu" authorityYear="2015" class="Magnoliopsida" family="Caryophyllaceae" genus="Dianthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dianthus aticii" order="Caryophyllales" pageId="4" pageNumber="25" phylum="Tracheophyta" rank="species" species="aticii">
<emphasis italics="true" pageId="4" pageNumber="25">Dianthus aticii</emphasis>
</taxonomicName>
- 
<emphasis bold="true" pageId="4" pageNumber="25">A1</emphasis>
Habit 
<emphasis bold="true" pageId="4" pageNumber="25">B1</emphasis>
Flower; 
<taxonomicName authorityName="Fenzl" authorityYear="1842" class="Magnoliopsida" family="Caryophyllaceae" genus="Dianthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dianthus zonatus" order="Caryophyllales" pageId="4" pageNumber="25" phylum="Tracheophyta" rank="species" species="zonatus">
<emphasis italics="true" pageId="4" pageNumber="25">Dianthus zonatus</emphasis>
</taxonomicName>
- 
<emphasis bold="true" pageId="4" pageNumber="25">A2</emphasis>
Habit 
<emphasis bold="true" pageId="4" pageNumber="25">B2</emphasis>
Flower.
</paragraph>
</caption>
<caption ID-Zenodo-Dep="961047" doi="10.3897/phytokeys.48.4446.figure2" httpUri="https://binary.pensoft.net/fig/41399" pageId="4" pageNumber="25" start="Figure 2" startId="F2">
<paragraph pageId="4" pageNumber="25">
<emphasis bold="true" pageId="4" pageNumber="25">Figure 2.</emphasis>
SEM photographs of the seed coat. 
<emphasis bold="true" pageId="4" pageNumber="25">A</emphasis>
<taxonomicName authorityName="Hamzaoglu" authorityYear="2015" class="Magnoliopsida" family="Caryophyllaceae" genus="Dianthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dianthus aticii" order="Caryophyllales" pageId="4" pageNumber="25" phylum="Tracheophyta" rank="species" species="aticii">
<emphasis italics="true" pageId="4" pageNumber="25">Dianthus aticii</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="4" pageNumber="25">B</emphasis>
<taxonomicName authorityName="Fenzl" authorityYear="1842" class="Magnoliopsida" family="Caryophyllaceae" genus="Dianthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dianthus zonatus" order="Caryophyllales" pageId="4" pageNumber="25" phylum="Tracheophyta" rank="species" species="zonatus">
<emphasis italics="true" pageId="4" pageNumber="25">Dianthus zonatus</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="4" pageNumber="25">1-3</emphasis>
dorsal surface 
<emphasis bold="true" pageId="4" pageNumber="25">4-6</emphasis>
ventral surface (scale bars: 
<emphasis bold="true" pageId="4" pageNumber="25">1</emphasis>
and 
<emphasis bold="true" pageId="4" pageNumber="25">4:</emphasis>
1 mm, 
<emphasis bold="true" pageId="4" pageNumber="25">2</emphasis>
and 
<emphasis bold="true" pageId="4" pageNumber="25">5:</emphasis>
20 
<normalizedToken originalValue="μm">μm</normalizedToken>
, 
<emphasis bold="true" pageId="4" pageNumber="25">3</emphasis>
and 
<emphasis bold="true" pageId="4" pageNumber="25">6:</emphasis>
10 
<normalizedToken originalValue="μm">μm</normalizedToken>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="4" pageNumber="25" type="phenology">
<paragraph pageId="4" pageNumber="25">Phenology.</paragraph>
<paragraph pageId="4" pageNumber="25">The new species was observed flowering in June and July, in stony slopes and steppes, between 330 and 1025 m.</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="26" pageId="4" pageNumber="25" type="chorology and ecology">
<paragraph pageId="4" pageNumber="25">Chorology and ecology.</paragraph>
<paragraph pageId="5" pageNumber="26">
<taxonomicName authorityName="Hamzaoglu" authorityYear="2015" class="Magnoliopsida" family="Caryophyllaceae" genus="Dianthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dianthus aticii" order="Caryophyllales" pageId="5" pageNumber="26" phylum="Tracheophyta" rank="species" species="aticii">
<emphasis italics="true" pageId="5" pageNumber="26">
<pageBreakToken pageId="5" pageNumber="26" start="start">Dianthus</pageBreakToken>
aticii
</emphasis>
</taxonomicName>
grows in relatively sub-arid forest clearings in Bilecik, Seben (Bolu), and 
<normalizedToken originalValue="Nallıhan">Nallihan</normalizedToken>
(Ankara); 
<emphasis italics="true" pageId="5" pageNumber="26">it</emphasis>
grows in moist areas where the Euro-Siberian and Irano-Turanian phytogeographic regions coincide in the northwest part of Turkey (
<bibRefCitation author="Davis, PH" journalOrPublisher="Flora of Turkey and the East Aegean Islands" pageId="5" pageNumber="26" pagination="1 - 25" refId="B5" refString="Davis, PH, 1965. . Flora of Turkey and the East Aegean Islands Vol. 1: 1 - 25" volume="Vol. 1" year="1965">Davis 1965</bibRefCitation>
). The forest clearings of these areas that are sub-arid, compared with the oceanic climate zone, were occupied by some semi-xeric species. These areas where the forest and steppe formations co-exist are the ideal habitats for 
<taxonomicName authorityName="Hamzaoglu" authorityYear="2015" class="Magnoliopsida" family="Caryophyllaceae" genus="Dianthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dianthus aticii" order="Caryophyllales" pageId="5" pageNumber="26" phylum="Tracheophyta" rank="species" species="aticii">
<emphasis italics="true" pageId="5" pageNumber="26">Dianthus aticii</emphasis>
</taxonomicName>
. The species grows on stony slopes within forest openings together with 
<taxonomicName class="Dicotyledoneae" family="Fagaceae" genus="Quercus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Quercus pubescens" order="Fagales" pageId="5" pageNumber="26" phylum="Angiospermae" rank="species" species="pubescens">
<emphasis italics="true" pageId="5" pageNumber="26">Quercus pubescens</emphasis>
</taxonomicName>
Willd., 
<taxonomicName class="Pinopsida" family="Cupressaceae" genus="Juniperus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Juniperus oxycedrus" order="Pinales" pageId="5" pageNumber="26" phylum="Coniferophyta" rank="species" species="oxycedrus">
<emphasis italics="true" pageId="5" pageNumber="26">Juniperus oxycedrus</emphasis>
</taxonomicName>
L., 
<taxonomicName authority="Jacq. subsp. monogyna" class="Dicotyledoneae" family="Rosaceae" genus="Crataegus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Crataegus monogyna subsp. monogyna" order="Rosales" pageId="5" pageNumber="26" phylum="Angiospermae" rank="subSpecies" species="monogyna" subSpecies="monogyna">Crataegus monogyna Jacq. subsp. monogyna</taxonomicName>
, 
<taxonomicName class="Eudicots" family="Cistaceae" genus="Cistus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Cistus creticus" order="Malvales" pageId="5" pageNumber="26" phylum="Magnoliophyta" rank="species" species="creticus">
<emphasis italics="true" pageId="5" pageNumber="26">Cistus creticus</emphasis>
</taxonomicName>
L., 
<taxonomicName class="Dicotyledoneae" family="Oleaceae" genus="Jasminum" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Jasminum fruticans" order="Oleales" pageId="5" pageNumber="26" phylum="Angiospermae" rank="species" species="fruticans">
<emphasis italics="true" pageId="5" pageNumber="26">Jasminum fruticans</emphasis>
</taxonomicName>
L., 
<taxonomicName class="Dicotyledoneae" family="Cistaceae" genus="Helianthemum" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Helianthemum nummularium" order="Violales" pageId="5" pageNumber="26" phylum="Angiospermae" rank="species" species="nummularium">
<emphasis italics="true" pageId="5" pageNumber="26">Helianthemum nummularium</emphasis>
</taxonomicName>
(L.) Miller, 
<taxonomicName class="Eudicots" family="Cistaceae" genus="Fumana" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Fumana thymifolia" order="Malvales" pageId="5" pageNumber="26" phylum="Magnoliophyta" rank="species" species="thymifolia">
<emphasis italics="true" pageId="5" pageNumber="26">Fumana thymifolia</emphasis>
</taxonomicName>
(L.) Verlot, 
<taxonomicName class="Dicotyledoneae" family="Brassicaceae" genus="Alyssum" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Alyssum sibiricum" order="Papaverales" pageId="5" pageNumber="26" phylum="Angiospermae" rank="species" species="sibiricum">
<emphasis italics="true" pageId="5" pageNumber="26">Alyssum sibiricum</emphasis>
</taxonomicName>
Willd., 
<taxonomicName class="Eudicots" family="Caryophyllaceae" genus="Silene" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Silene italica" order="Caryophyllales" pageId="5" pageNumber="26" phylum="Magnoliophyta" rank="species" species="italica">
<emphasis italics="true" pageId="5" pageNumber="26">Silene italica</emphasis>
</taxonomicName>
(L.) Pers., 
<taxonomicName class="Eudicots" family="Asteraceae" genus="Pilosella" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Pilosella piloselloides" order="Asterales" pageId="5" pageNumber="26" phylum="Magnoliophyta" rank="species" species="piloselloides">
<emphasis italics="true" pageId="5" pageNumber="26">Pilosella piloselloides</emphasis>
</taxonomicName>
(Vill.) Sojak, 
<taxonomicName class="Dicotyledoneae" family="Boraginaceae" genus="Onosma" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Onosma tauricum" order="Tubiflorae" pageId="5" pageNumber="26" phylum="Angiospermae" rank="species" species="tauricum">
<emphasis italics="true" pageId="5" pageNumber="26">Onosma tauricum</emphasis>
</taxonomicName>
Pallas ex Willd., 
<taxonomicName class="Dicotyledoneae" family="Scrophulariaceae" genus="Veronica" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Veronica multifida" order="Tubiflorae" pageId="5" pageNumber="26" phylum="Angiospermae" rank="species" species="multifida">
<emphasis italics="true" pageId="5" pageNumber="26">Veronica multifida</emphasis>
</taxonomicName>
L., 
<taxonomicName class="Dicotyledoneae" family="Lamiaceae" genus="Teucrium" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Teucrium polium" order="Tubiflorae" pageId="5" pageNumber="26" phylum="Angiospermae" rank="species" species="polium">
<emphasis italics="true" pageId="5" pageNumber="26">Teucrium polium</emphasis>
</taxonomicName>
L., 
<taxonomicName class="Magnoliopsida" family="Plumbaginaceae" genus="Acantholimon" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="Acantholimon acerosum" order="Caryophyllales" pageId="5" pageNumber="26" phylum="Tracheophyta" rank="species" species="acerosum">
<emphasis italics="true" pageId="5" pageNumber="26">Acantholimon acerosum</emphasis>
</taxonomicName>
(Willd.) Boiss., 
<taxonomicName class="Dicotyledoneae" family="Clusiaceae" genus="Hypericum" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Hypericum perforatum" order="Guttiferales" pageId="5" pageNumber="26" phylum="Angiospermae" rank="species" species="perforatum">
<emphasis italics="true" pageId="5" pageNumber="26">Hypericum perforatum</emphasis>
</taxonomicName>
L., 
<taxonomicName class="Dicotyledoneae" family="Fabaceae" genus="Genista" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Genista tinctoria" order="Rosales" pageId="5" pageNumber="26" phylum="Angiospermae" rank="species" species="tinctoria">
<emphasis italics="true" pageId="5" pageNumber="26">Genista tinctoria</emphasis>
</taxonomicName>
L., 
<taxonomicName class="Dicotyledoneae" family="Fabaceae" genus="Vicia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Vicia cracca subsp. stenophylla" order="Rosales" pageId="5" pageNumber="26" phylum="Angiospermae" rank="subSpecies" species="cracca" subSpecies="stenophylla">Vicia cracca L. subsp. stenophylla</taxonomicName>
Vel., 
<taxonomicName class="Dicotyledoneae" family="Fabaceae" genus="Astragalus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Astragalus vulnerariae" order="Rosales" pageId="5" pageNumber="26" phylum="Angiospermae" rank="species" species="vulnerariae">
<emphasis italics="true" pageId="5" pageNumber="26">Astragalus vulnerariae</emphasis>
</taxonomicName>
DC., 
<taxonomicName class="Dicotyledoneae" family="Fabaceae" genus="Astragalus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Astragalus microcephalus" order="Rosales" pageId="5" pageNumber="26" phylum="Angiospermae" rank="species" species="microcephalus">
<emphasis italics="true" pageId="5" pageNumber="26">Astragalus microcephalus</emphasis>
</taxonomicName>
Willd., 
<taxonomicName class="Dicotyledoneae" family="Rosaceae" genus="Rosa" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Rosa canina" order="Rosales" pageId="5" pageNumber="26" phylum="Angiospermae" rank="species" species="canina">
<emphasis italics="true" pageId="5" pageNumber="26">Rosa canina</emphasis>
</taxonomicName>
L., and 
<taxonomicName class="Dicotyledoneae" family="Asteraceae" genus="Centaurea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Centaurea urvillei" order="Campanulales" pageId="5" pageNumber="26" phylum="Angiospermae" rank="species" species="urvillei">
<emphasis italics="true" pageId="5" pageNumber="26">Centaurea urvillei</emphasis>
</taxonomicName>
DC.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="26" type="conservation status">
<paragraph pageId="5" pageNumber="26">Conservation status.</paragraph>
<paragraph pageId="5" pageNumber="26">
According to the current data 
<taxonomicName authorityName="Hamzaoglu" authorityYear="2015" class="Magnoliopsida" family="Caryophyllaceae" genus="Dianthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dianthus aticii" order="Caryophyllales" pageId="5" pageNumber="26" phylum="Tracheophyta" rank="species" species="aticii">
<emphasis italics="true" pageId="5" pageNumber="26">Dianthus aticii</emphasis>
</taxonomicName>
grows in the Bilecik, Seben (Bolu), and 
<normalizedToken originalValue="Nallıhan">Nallihan</normalizedToken>
(Ankara) districts, which have an area of approximately 7000 km2. This has a discontinuous distribution due to dense forests, settlement, and farming areas. The open areas, which this species prefers, have the potential of possible settlements and agricultural activities. Therefore, the habitat of this species is under danger of being decreased and disturbed/destroyed in the future. Therefore, it is proposed that the species should be classified as 
<taxonomicName genus="Vulnerable" lsidName="Vulnerable" pageId="5" pageNumber="26" rank="genus">
<emphasis italics="true" pageId="5" pageNumber="26">Vulnerable</emphasis>
</taxonomicName>
[VU (B1b-iii) according to the International Union for Conservation of Nature (IUCN) categories (
<bibRefCitation author="IUCN" journalOrPublisher="Turkish Journal of Botany" pageId="6" pageNumber="27" publicationUrl="http://jr.iucnredlist.org/documents/RedListGuidelines.pdf" refId="B11" refString="IUCN, 2014. http://jr.iucnredlist.org/documents/RedListGuidelines.pdf" url="http://jr.iucnredlist.org/documents/RedListGuidelines.pdf" year="2014">IUCN 2014</bibRefCitation>
)].
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="26" type="etymology">
<paragraph pageId="5" pageNumber="26">Etymology.</paragraph>
<paragraph pageId="5" pageNumber="26">
The species is named in honour of the eminent Turkish hydrobiologist Prof Dr Tahir 
<normalizedToken originalValue="Atıcı">Atici</normalizedToken>
(Gazi Faculty of Education, Gazi University, Ankara).
</paragraph>
</subSubSection>
</treatment>
</document>