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<document id="930261299D233C3D1282D47BF744305A" ID-CLB-Dataset="8807" ID-DOI="10.11646/zootaxa.4822.2.1" ID-GBIF-Dataset="73a6f164-2334-466a-aee0-d39129bdf7e2" ID-ISSN="1175-5326" ID-Zenodo-Dep="4401369" ID-ZooBank="E3B5713F-ADE8-4075-9ABF-8F6DE9D3A88E" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609335057997" checkinUser="plazi" docAuthor="Miranda, Gil Felipe Gonçalves, Skevington, Jeffrey H. &amp; Marshall, Stephen A." docDate="2020" docId="A51D5F67FFED3A6DFF50FEF6FDF49B76" docLanguage="en" docName="zootaxa.4822.2.1.pdf" docOrigin="Zootaxa 4822 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Fragosa rugosifrons Miranda &amp; Skevington &amp; Marshall 2020" docType="treatment" docVersion="8" lastPageNumber="154" masterDocId="5924271FFFEE3A6EFFC7FFBDFFE39F0B" masterDocTitle="New generic concepts for orphaned lineages formerly treated as part of the genus Ocyptamus Macquart, 1834 (Diptera, Syrphidae)" masterLastPageNumber="174" masterPageNumber="151" pageNumber="154" updateTime="1698849646355" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="FA9E12C03B6C2F2F92C49A9B91378C9E">New generic concepts for orphaned lineages formerly treated as part of the genus Ocyptamus Macquart, 1834 (Diptera, Syrphidae)</mods:title>
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<mods:namePart id="D2A6D35A7B45C36CCF6F9CB3BBD1D71C">Miranda, Gil Felipe Gonçalves</mods:namePart>
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<mods:nameIdentifier id="01387D1D5CAD6457E961FB85D709C56B" type="email">gilfgm@gmail.com</mods:nameIdentifier>
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<mods:namePart id="43CBD64F54F488DA7EA48163103B4E4B">Skevington, Jeffrey H.</mods:namePart>
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<mods:affiliation id="FC34744234A8E11B261B7D63E84D9C60">jhskevington @ gmail. com; https: // orcid. org / 0000 - 0002 - 1445 - 9870</mods:affiliation>
<mods:nameIdentifier id="6D2F578E0E5A409959B5FF09A25AB5AF" type="email">gilfgm@gmail.com</mods:nameIdentifier>
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<mods:namePart id="0EEDBC398A7E48E1A7F0257B0C23E819">Marshall, Stephen A.</mods:namePart>
<mods:nameIdentifier id="F2D251A317651BBB6542E54AFBF55B6F" type="ORCID">0000-0001-7919-2639</mods:nameIdentifier>
<mods:affiliation id="9F9A198557C612FD10DE07754906A153">University of Guelph Insect Collection and Insect Systematics Laboratory - School of Environmental Sciences - University of Guelph. Guelph, Ontario, Canada.</mods:affiliation>
<mods:nameIdentifier id="F70C3ECF899C97D270B8FD4CEF8F6444" type="email">gilfgm@gmail.com</mods:nameIdentifier>
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<mods:title id="0E3EBAD5953AF9EEB9AEFE8E7535AD91">Zootaxa</mods:title>
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<mods:date id="4395468559F0A05AEF9A32484C32F3B7">2020</mods:date>
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<treatment id="A51D5F67FFED3A6DFF50FEF6FDF49B76" ID-DOI="http://doi.org/10.5281/zenodo.4449973" ID-GBIF-Taxon="177008124" ID-Zenodo-Dep="4449973" LSID="urn:lsid:plazi:treatment:A51D5F67FFED3A6DFF50FEF6FDF49B76" httpUri="http://treatment.plazi.org/id/A51D5F67FFED3A6DFF50FEF6FDF49B76" lastPageNumber="154" pageId="3" pageNumber="154">
<subSubSection id="65AEBDFAFFED3A6DFF50FEF6FCBD9E6F" box="[151,862,330,357]" pageId="3" pageNumber="154" type="nomenclature">
<paragraph id="2D0BEE71FFED3A6DFF50FEF6FCBD9E6F" blockId="3.[151,862,330,357]" box="[151,862,330,357]" pageId="3" pageNumber="154">
<heading id="7643591DFFED3A6DFF50FEF6FCBD9E6F" bold="true" box="[151,862,330,357]" fontSize="11" level="1" pageId="3" pageNumber="154" reason="1">
<emphasis id="1FC03263FFED3A6DFF50FEF6FCBD9E6F" bold="true" box="[151,862,330,357]" pageId="3" pageNumber="154">
<taxonomicName id="EAB495F2FFED3A6DFF50FEF6FE739E6F" ID-CoL="8C7C9" authorityName="Miranda &amp; Skevington &amp; Marshall" authorityYear="2020" baseAuthorityName="Schiner" baseAuthorityYear="1868" box="[151,400,330,357]" class="Insecta" family="Syrphidae" genus="Fragosa" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="rugosifrons">
<emphasis id="1FC03263FFED3A6DFF50FEF6FE739E6F" bold="true" box="[151,400,330,357]" italics="true" pageId="3" pageNumber="154">Fragosa rugosifrons</emphasis>
</taxonomicName>
species group (
<emphasis id="1FC03263FFED3A6DFD97FEF6FCB79E6E" bold="true" box="[592,852,331,357]" italics="true" pageId="3" pageNumber="154">Fragosa sensu stricto</emphasis>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="65AEBDFAFFED3A6DFF50FE2FFBEA9CC2" pageId="3" pageNumber="154" type="description">
<paragraph id="2D0BEE71FFED3A6DFF50FE2FFD8D9C1E" blockId="3.[151,1437,402,1149]" pageId="3" pageNumber="154">
<emphasis id="1FC03263FFED3A6DFF50FE2FFE8C9EA6" bold="true" box="[151,367,402,429]" pageId="3" pageNumber="154">
Description.
<emphasis id="1FC03263FFED3A6DFEF6FE2FFE8C9EA6" bold="true" box="[305,367,402,429]" italics="true" pageId="3" pageNumber="154">Head</emphasis>
</emphasis>
. Face usually dark dorsal to tubercle; tubercle pointed and medially positioned (
<figureCitation id="B58FF2F4FFED3A6DFAC6FE2EFAA59EA6" box="[1281,1350,402,429]" captionStart="FIGURES 111" captionStartId="4.[151,264,1631,1655]" captionTargetBox="[177,1414,185,1601]" captionTargetId="figure-19@4.[165,1423,181,1608]" captionTargetPageId="4" captionText="FIGURES 111. Fragosa Miranda gen. nov. 1: Fragosa sp. (INPA-DIP001362), female, habitus, dorsal view, photo previously published in Mengual et al. (2018). 2: Fragosa aff. titania (CNC), female, abdomen, dorsal view, photo previously published in Mengual et al. (2018). 3: Fragosa argentina (Curran) (CNC_Diptera196006), female, wing. 4: Fragosa stenogaster (Williston) (debu01088839), female, head, frontal view, photo previously published in Mengual et al. (2018). 5: Fragosa stenogaster (Williston) (debu01088840), male, head, lateral view. 6: Fragosa sp., female, habitus, dorsal view, by Steve Marshall. 7: Fragosa stenogaster (Williston) (debu01088840), male, epandrium, lateral view, photo previously published in Mengual et al. (2018). 8: Fragosa stenogaster (Williston) (debu01088839), female, apex of abdomen, dorsal view, photo previously published in Mengual et al. (2018). 911. Fragosa stenogaster (Williston) (debu01088840), male. 9: epandrium, ventral view. 10: hypandrium, lateral view. 11: hypandrium, ventral view. ap: apodeme; b: basiphallus; bm: cell bm; br: cell br; c: cell c; d: distiphallus; e: epiproct; f: frons; h: hypandrium; oc: occiput; p: postgonite; s: surstylus; se: subepandrial sclerite; t: facial tubercle; t2, t3, t7 and t8: terga 2, 3, 7 and 8." figureDoi="http://doi.org/10.5281/zenodo.4401371" httpUri="https://zenodo.org/record/4401371/files/figure.png" pageId="3" pageNumber="154">Fig. 5</figureCitation>
). Frons dark. Frons/frontal triangle rugose (
<figureCitation id="B58FF2F4FFED3A6DFDE0FE0AFD8F9EDA" box="[551,620,439,465]" captionStart="FIGURES 111" captionStartId="4.[151,264,1631,1655]" captionTargetBox="[177,1414,185,1601]" captionTargetId="figure-19@4.[165,1423,181,1608]" captionTargetPageId="4" captionText="FIGURES 111. Fragosa Miranda gen. nov. 1: Fragosa sp. (INPA-DIP001362), female, habitus, dorsal view, photo previously published in Mengual et al. (2018). 2: Fragosa aff. titania (CNC), female, abdomen, dorsal view, photo previously published in Mengual et al. (2018). 3: Fragosa argentina (Curran) (CNC_Diptera196006), female, wing. 4: Fragosa stenogaster (Williston) (debu01088839), female, head, frontal view, photo previously published in Mengual et al. (2018). 5: Fragosa stenogaster (Williston) (debu01088840), male, head, lateral view. 6: Fragosa sp., female, habitus, dorsal view, by Steve Marshall. 7: Fragosa stenogaster (Williston) (debu01088840), male, epandrium, lateral view, photo previously published in Mengual et al. (2018). 8: Fragosa stenogaster (Williston) (debu01088839), female, apex of abdomen, dorsal view, photo previously published in Mengual et al. (2018). 911. Fragosa stenogaster (Williston) (debu01088840), male. 9: epandrium, ventral view. 10: hypandrium, lateral view. 11: hypandrium, ventral view. ap: apodeme; b: basiphallus; bm: cell bm; br: cell br; c: cell c; d: distiphallus; e: epiproct; f: frons; h: hypandrium; oc: occiput; p: postgonite; s: surstylus; se: subepandrial sclerite; t: facial tubercle; t2, t3, t7 and t8: terga 2, 3, 7 and 8." figureDoi="http://doi.org/10.5281/zenodo.4401371" httpUri="https://zenodo.org/record/4401371/files/figure.png" pageId="3" pageNumber="154">Fig. 4</figureCitation>
). Frontal triangle with white microtrichosity concentrated laterally along eye margin (
<figureCitation id="B58FF2F4FFED3A6DFEE1FE66FE999EFE" box="[294,378,475,501]" captionStart="FIGURES 7888. 78" captionStartId="17.[151,264,1665,1689]" captionTargetBox="[301,1287,192,1634]" captionTargetId="figure-18@17.[294,1294,181,1642]" captionTargetPageId="17" captionText="FIGURES 7888. 78: Fragosa harlequina (Hull) (holotype Baccha harlequina Hull, AMNH), male, thorax, lateral, by AMNH (line drawing represents outline of the alula). 79: Fragosa argentina (Curran) (CNC_Diptera174232), thorax, postero-ventral (line drawing represents outline of the incomplete post-metacoxal bridge). 8081: Fragosa virgilio (Hull) (holotype Baccha virgilio Hull, CNC, CNC_Diptera189124), male. 80: habitus, lateral, by CNC. 81: head, oblique frontal, by CNC. 82: Fragosa titania (Hull) (JSS25229), female, head, dorsal. 83: Fragosa stenogaster (Williston) (debu01088840), male, head, dorsal. 8485: Hypocritanus fascipennis (Wiedemann), female, head. 84: frontal, by Ximo Mengual (http://syrphidae.myspecies.info/ taxonomy/term/879). 85: lateral (CNC_Diptera501). 8687: Nuntianus cecrops (Hull) (INPA-DIP000246), female, head. 86: dorsal. 87: lateral. 88: Victoriana aff. melanorrhina (DEBU), female, head, lateral. a: alula; ai: antennal insertion; f: frons; ft: frontal triangle; lem: lateral eye margin; mt: metafemur; o: ocellar triangle; oc: occiput; pmt: post-metacoxal bridge; t: facial tubercle." figureDoi="http://doi.org/10.5281/zenodo.4401387" httpUri="https://zenodo.org/record/4401387/files/figure.png" pageId="3" pageNumber="154">Fig. 83</figureCitation>
). Female ocellar triangle separated by less than its length from posterior eye margin (
<figureCitation id="B58FF2F4FFED3A6DFA8FFE66FA689EFE" box="[1352,1419,475,501]" captionStart="FIGURES 111" captionStartId="4.[151,264,1631,1655]" captionTargetBox="[177,1414,185,1601]" captionTargetId="figure-19@4.[165,1423,181,1608]" captionTargetPageId="4" captionText="FIGURES 111. Fragosa Miranda gen. nov. 1: Fragosa sp. (INPA-DIP001362), female, habitus, dorsal view, photo previously published in Mengual et al. (2018). 2: Fragosa aff. titania (CNC), female, abdomen, dorsal view, photo previously published in Mengual et al. (2018). 3: Fragosa argentina (Curran) (CNC_Diptera196006), female, wing. 4: Fragosa stenogaster (Williston) (debu01088839), female, head, frontal view, photo previously published in Mengual et al. (2018). 5: Fragosa stenogaster (Williston) (debu01088840), male, head, lateral view. 6: Fragosa sp., female, habitus, dorsal view, by Steve Marshall. 7: Fragosa stenogaster (Williston) (debu01088840), male, epandrium, lateral view, photo previously published in Mengual et al. (2018). 8: Fragosa stenogaster (Williston) (debu01088839), female, apex of abdomen, dorsal view, photo previously published in Mengual et al. (2018). 911. Fragosa stenogaster (Williston) (debu01088840), male. 9: epandrium, ventral view. 10: hypandrium, lateral view. 11: hypandrium, ventral view. ap: apodeme; b: basiphallus; bm: cell bm; br: cell br; c: cell c; d: distiphallus; e: epiproct; f: frons; h: hypandrium; oc: occiput; p: postgonite; s: surstylus; se: subepandrial sclerite; t: facial tubercle; t2, t3, t7 and t8: terga 2, 3, 7 and 8." figureDoi="http://doi.org/10.5281/zenodo.4401371" httpUri="https://zenodo.org/record/4401371/files/figure.png" pageId="3" pageNumber="154">Fig. 1</figureCitation>
). Occiput microtrichia homogeneously distributed (
<figureCitation id="B58FF2F4FFED3A6DFD05FE42FCE59D12" box="[706,774,510,537]" captionStart="FIGURES 111" captionStartId="4.[151,264,1631,1655]" captionTargetBox="[177,1414,185,1601]" captionTargetId="figure-19@4.[165,1423,181,1608]" captionTargetPageId="4" captionText="FIGURES 111. Fragosa Miranda gen. nov. 1: Fragosa sp. (INPA-DIP001362), female, habitus, dorsal view, photo previously published in Mengual et al. (2018). 2: Fragosa aff. titania (CNC), female, abdomen, dorsal view, photo previously published in Mengual et al. (2018). 3: Fragosa argentina (Curran) (CNC_Diptera196006), female, wing. 4: Fragosa stenogaster (Williston) (debu01088839), female, head, frontal view, photo previously published in Mengual et al. (2018). 5: Fragosa stenogaster (Williston) (debu01088840), male, head, lateral view. 6: Fragosa sp., female, habitus, dorsal view, by Steve Marshall. 7: Fragosa stenogaster (Williston) (debu01088840), male, epandrium, lateral view, photo previously published in Mengual et al. (2018). 8: Fragosa stenogaster (Williston) (debu01088839), female, apex of abdomen, dorsal view, photo previously published in Mengual et al. (2018). 911. Fragosa stenogaster (Williston) (debu01088840), male. 9: epandrium, ventral view. 10: hypandrium, lateral view. 11: hypandrium, ventral view. ap: apodeme; b: basiphallus; bm: cell bm; br: cell br; c: cell c; d: distiphallus; e: epiproct; f: frons; h: hypandrium; oc: occiput; p: postgonite; s: surstylus; se: subepandrial sclerite; t: facial tubercle; t2, t3, t7 and t8: terga 2, 3, 7 and 8." figureDoi="http://doi.org/10.5281/zenodo.4401371" httpUri="https://zenodo.org/record/4401371/files/figure.png" pageId="3" pageNumber="154">Fig. 5</figureCitation>
).
<emphasis id="1FC03263FFED3A6DFCDDFE43FC939D12" bold="true" box="[794,880,510,537]" italics="true" pageId="3" pageNumber="154">Thorax</emphasis>
. Scutum dull pale-microtrichose with sub-median pair of weak white microtrichose patches on anterior region, sometimes with weak stripes in place of patches (
<figureCitation id="B58FF2F4FFED3A6DFAA8FD9EFF479D6A" captionStart="FIGURES 111" captionStartId="4.[151,264,1631,1655]" captionTargetBox="[177,1414,185,1601]" captionTargetId="figure-19@4.[165,1423,181,1608]" captionTargetPageId="4" captionText="FIGURES 111. Fragosa Miranda gen. nov. 1: Fragosa sp. (INPA-DIP001362), female, habitus, dorsal view, photo previously published in Mengual et al. (2018). 2: Fragosa aff. titania (CNC), female, abdomen, dorsal view, photo previously published in Mengual et al. (2018). 3: Fragosa argentina (Curran) (CNC_Diptera196006), female, wing. 4: Fragosa stenogaster (Williston) (debu01088839), female, head, frontal view, photo previously published in Mengual et al. (2018). 5: Fragosa stenogaster (Williston) (debu01088840), male, head, lateral view. 6: Fragosa sp., female, habitus, dorsal view, by Steve Marshall. 7: Fragosa stenogaster (Williston) (debu01088840), male, epandrium, lateral view, photo previously published in Mengual et al. (2018). 8: Fragosa stenogaster (Williston) (debu01088839), female, apex of abdomen, dorsal view, photo previously published in Mengual et al. (2018). 911. Fragosa stenogaster (Williston) (debu01088840), male. 9: epandrium, ventral view. 10: hypandrium, lateral view. 11: hypandrium, ventral view. ap: apodeme; b: basiphallus; bm: cell bm; br: cell br; c: cell c; d: distiphallus; e: epiproct; f: frons; h: hypandrium; oc: occiput; p: postgonite; s: surstylus; se: subepandrial sclerite; t: facial tubercle; t2, t3, t7 and t8: terga 2, 3, 7 and 8." figureDoi="http://doi.org/10.5281/zenodo.4401371" httpUri="https://zenodo.org/record/4401371/files/figure.png" pageId="3" pageNumber="154">Fig. 1</figureCitation>
). Scutellum pale. Post-metacoxal bridge almost complete (
<figureCitation id="B58FF2F4FFED3A6DFCFEFDFAFC6E9D6A" box="[825,909,583,609]" captionStart="FIGURES 7888. 78" captionStartId="17.[151,264,1665,1689]" captionTargetBox="[301,1287,192,1634]" captionTargetId="figure-18@17.[294,1294,181,1642]" captionTargetPageId="17" captionText="FIGURES 7888. 78: Fragosa harlequina (Hull) (holotype Baccha harlequina Hull, AMNH), male, thorax, lateral, by AMNH (line drawing represents outline of the alula). 79: Fragosa argentina (Curran) (CNC_Diptera174232), thorax, postero-ventral (line drawing represents outline of the incomplete post-metacoxal bridge). 8081: Fragosa virgilio (Hull) (holotype Baccha virgilio Hull, CNC, CNC_Diptera189124), male. 80: habitus, lateral, by CNC. 81: head, oblique frontal, by CNC. 82: Fragosa titania (Hull) (JSS25229), female, head, dorsal. 83: Fragosa stenogaster (Williston) (debu01088840), male, head, dorsal. 8485: Hypocritanus fascipennis (Wiedemann), female, head. 84: frontal, by Ximo Mengual (http://syrphidae.myspecies.info/ taxonomy/term/879). 85: lateral (CNC_Diptera501). 8687: Nuntianus cecrops (Hull) (INPA-DIP000246), female, head. 86: dorsal. 87: lateral. 88: Victoriana aff. melanorrhina (DEBU), female, head, lateral. a: alula; ai: antennal insertion; f: frons; ft: frontal triangle; lem: lateral eye margin; mt: metafemur; o: ocellar triangle; oc: occiput; pmt: post-metacoxal bridge; t: facial tubercle." figureDoi="http://doi.org/10.5281/zenodo.4401387" httpUri="https://zenodo.org/record/4401387/files/figure.png" pageId="3" pageNumber="154">Fig. 79</figureCitation>
). Metafemur pale.
<emphasis id="1FC03263FFED3A6DFB98FDFAFB7F9D6A" bold="true" box="[1119,1180,583,609]" italics="true" pageId="3" pageNumber="154">Wing</emphasis>
. Female wing hyaline; female wing bare on basal 1/4 (or almost the whole cell) of cell c, basal 1/3 to 1/2 of br, and baso-anterior margin of bm.
<emphasis id="1FC03263FFED3A6DFF2CFD33FE899DA2" bold="true" box="[235,362,654,681]" italics="true" pageId="3" pageNumber="154">Terminalia</emphasis>
. Male epandrium enlarged; hypandrium reduced; *surstylus quadrangular with filiform apical extension (
<figureCitation id="B58FF2F4FFED3A6DFED3FD0EFEB99DC6" box="[276,346,691,717]" captionStart="FIGURES 111" captionStartId="4.[151,264,1631,1655]" captionTargetBox="[177,1414,185,1601]" captionTargetId="figure-19@4.[165,1423,181,1608]" captionTargetPageId="4" captionText="FIGURES 111. Fragosa Miranda gen. nov. 1: Fragosa sp. (INPA-DIP001362), female, habitus, dorsal view, photo previously published in Mengual et al. (2018). 2: Fragosa aff. titania (CNC), female, abdomen, dorsal view, photo previously published in Mengual et al. (2018). 3: Fragosa argentina (Curran) (CNC_Diptera196006), female, wing. 4: Fragosa stenogaster (Williston) (debu01088839), female, head, frontal view, photo previously published in Mengual et al. (2018). 5: Fragosa stenogaster (Williston) (debu01088840), male, head, lateral view. 6: Fragosa sp., female, habitus, dorsal view, by Steve Marshall. 7: Fragosa stenogaster (Williston) (debu01088840), male, epandrium, lateral view, photo previously published in Mengual et al. (2018). 8: Fragosa stenogaster (Williston) (debu01088839), female, apex of abdomen, dorsal view, photo previously published in Mengual et al. (2018). 911. Fragosa stenogaster (Williston) (debu01088840), male. 9: epandrium, ventral view. 10: hypandrium, lateral view. 11: hypandrium, ventral view. ap: apodeme; b: basiphallus; bm: cell bm; br: cell br; c: cell c; d: distiphallus; e: epiproct; f: frons; h: hypandrium; oc: occiput; p: postgonite; s: surstylus; se: subepandrial sclerite; t: facial tubercle; t2, t3, t7 and t8: terga 2, 3, 7 and 8." figureDoi="http://doi.org/10.5281/zenodo.4401371" httpUri="https://zenodo.org/record/4401371/files/figure.png" pageId="3" pageNumber="154">Fig. 7</figureCitation>
); *subepandrial sclerite with strongly sclerotized rectangular middle area and wide, but weakly sclerotized, ventrally directed lateral expansions (
<figureCitation id="B58FF2F4FFED3A6DFD16FD6AFCFB9DFA" box="[721,792,727,753]" captionStart="FIGURES 111" captionStartId="4.[151,264,1631,1655]" captionTargetBox="[177,1414,185,1601]" captionTargetId="figure-19@4.[165,1423,181,1608]" captionTargetPageId="4" captionText="FIGURES 111. Fragosa Miranda gen. nov. 1: Fragosa sp. (INPA-DIP001362), female, habitus, dorsal view, photo previously published in Mengual et al. (2018). 2: Fragosa aff. titania (CNC), female, abdomen, dorsal view, photo previously published in Mengual et al. (2018). 3: Fragosa argentina (Curran) (CNC_Diptera196006), female, wing. 4: Fragosa stenogaster (Williston) (debu01088839), female, head, frontal view, photo previously published in Mengual et al. (2018). 5: Fragosa stenogaster (Williston) (debu01088840), male, head, lateral view. 6: Fragosa sp., female, habitus, dorsal view, by Steve Marshall. 7: Fragosa stenogaster (Williston) (debu01088840), male, epandrium, lateral view, photo previously published in Mengual et al. (2018). 8: Fragosa stenogaster (Williston) (debu01088839), female, apex of abdomen, dorsal view, photo previously published in Mengual et al. (2018). 911. Fragosa stenogaster (Williston) (debu01088840), male. 9: epandrium, ventral view. 10: hypandrium, lateral view. 11: hypandrium, ventral view. ap: apodeme; b: basiphallus; bm: cell bm; br: cell br; c: cell c; d: distiphallus; e: epiproct; f: frons; h: hypandrium; oc: occiput; p: postgonite; s: surstylus; se: subepandrial sclerite; t: facial tubercle; t2, t3, t7 and t8: terga 2, 3, 7 and 8." figureDoi="http://doi.org/10.5281/zenodo.4401371" httpUri="https://zenodo.org/record/4401371/files/figure.png" pageId="3" pageNumber="154">Fig. 9</figureCitation>
); *hypandrium oval (
<figureCitation id="B58FF2F4FFED3A6DFBD4FD6AFB849DFA" box="[1043,1127,727,753]" captionStart="FIGURES 111" captionStartId="4.[151,264,1631,1655]" captionTargetBox="[177,1414,185,1601]" captionTargetId="figure-19@4.[165,1423,181,1608]" captionTargetPageId="4" captionText="FIGURES 111. Fragosa Miranda gen. nov. 1: Fragosa sp. (INPA-DIP001362), female, habitus, dorsal view, photo previously published in Mengual et al. (2018). 2: Fragosa aff. titania (CNC), female, abdomen, dorsal view, photo previously published in Mengual et al. (2018). 3: Fragosa argentina (Curran) (CNC_Diptera196006), female, wing. 4: Fragosa stenogaster (Williston) (debu01088839), female, head, frontal view, photo previously published in Mengual et al. (2018). 5: Fragosa stenogaster (Williston) (debu01088840), male, head, lateral view. 6: Fragosa sp., female, habitus, dorsal view, by Steve Marshall. 7: Fragosa stenogaster (Williston) (debu01088840), male, epandrium, lateral view, photo previously published in Mengual et al. (2018). 8: Fragosa stenogaster (Williston) (debu01088839), female, apex of abdomen, dorsal view, photo previously published in Mengual et al. (2018). 911. Fragosa stenogaster (Williston) (debu01088840), male. 9: epandrium, ventral view. 10: hypandrium, lateral view. 11: hypandrium, ventral view. ap: apodeme; b: basiphallus; bm: cell bm; br: cell br; c: cell c; d: distiphallus; e: epiproct; f: frons; h: hypandrium; oc: occiput; p: postgonite; s: surstylus; se: subepandrial sclerite; t: facial tubercle; t2, t3, t7 and t8: terga 2, 3, 7 and 8." figureDoi="http://doi.org/10.5281/zenodo.4401371" httpUri="https://zenodo.org/record/4401371/files/figure.png" pageId="3" pageNumber="154">Fig. 11</figureCitation>
), much smaller than epandrium; postgonite elongated (
<figureCitation id="B58FF2F4FFED3A6DFE27FD46FD839C1E" box="[480,608,763,789]" captionStart="FIGURES 111" captionStartId="4.[151,264,1631,1655]" captionTargetBox="[177,1414,185,1601]" captionTargetId="figure-19@4.[165,1423,181,1608]" captionTargetPageId="4" captionText="FIGURES 111. Fragosa Miranda gen. nov. 1: Fragosa sp. (INPA-DIP001362), female, habitus, dorsal view, photo previously published in Mengual et al. (2018). 2: Fragosa aff. titania (CNC), female, abdomen, dorsal view, photo previously published in Mengual et al. (2018). 3: Fragosa argentina (Curran) (CNC_Diptera196006), female, wing. 4: Fragosa stenogaster (Williston) (debu01088839), female, head, frontal view, photo previously published in Mengual et al. (2018). 5: Fragosa stenogaster (Williston) (debu01088840), male, head, lateral view. 6: Fragosa sp., female, habitus, dorsal view, by Steve Marshall. 7: Fragosa stenogaster (Williston) (debu01088840), male, epandrium, lateral view, photo previously published in Mengual et al. (2018). 8: Fragosa stenogaster (Williston) (debu01088839), female, apex of abdomen, dorsal view, photo previously published in Mengual et al. (2018). 911. Fragosa stenogaster (Williston) (debu01088840), male. 9: epandrium, ventral view. 10: hypandrium, lateral view. 11: hypandrium, ventral view. ap: apodeme; b: basiphallus; bm: cell bm; br: cell br; c: cell c; d: distiphallus; e: epiproct; f: frons; h: hypandrium; oc: occiput; p: postgonite; s: surstylus; se: subepandrial sclerite; t: facial tubercle; t2, t3, t7 and t8: terga 2, 3, 7 and 8." figureDoi="http://doi.org/10.5281/zenodo.4401371" httpUri="https://zenodo.org/record/4401371/files/figure.png" pageId="3" pageNumber="154">Figs 10, 11</figureCitation>
).
</paragraph>
<paragraph id="2D0BEE71FFED3A6DFF00FCA2FBEA9CC2" blockId="3.[151,1437,402,1149]" pageId="3" pageNumber="154">
<emphasis id="1FC03263FFED3A6DFF00FCA2FE2A9C33" bold="true" box="[199,457,798,825]" pageId="3" pageNumber="154">Included species (11):</emphasis>
<taxonomicName id="EAB495F2FFED3A6DFE17FCA2FCE99C32" authority="(Curran, 1939)" authorityName="Miranda &amp; Skevington &amp; Marshall" authorityYear="2020" baseAuthorityName="Curran" baseAuthorityYear="1939" box="[464,778,799,825]" class="Insecta" family="Syrphidae" genus="Fragosa" kingdom="Plantae" order="Diptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="argentina" status="comb. nov.">
<emphasis id="1FC03263FFED3A6DFE17FCA2FE029C33" box="[464,481,799,824]" italics="true" pageId="3" pageNumber="154">F</emphasis>
.
<emphasis id="1FC03263FFED3A6DFE37FCA2FDBF9C33" box="[496,604,799,824]" italics="true" pageId="3" pageNumber="154">argentina</emphasis>
(
<bibRefCitation id="49259380FFED3A6DFDACFCA2FCE19C32" author="Curran, C. H." box="[619,770,799,825]" pageId="3" pageNumber="154" pagination="1 - 12" refId="ref15834" refString="Curran, C. H. (1939) New Neotropical Baccha Fabricius (Syrphidae: Diptera). American Museum Novitates, 1041, 1 - 12." type="journal article" year="1939">Curran, 1939</bibRefCitation>
)
</taxonomicName>
<emphasis id="1FC03263FFED3A6DFCD5FCA2FC709C32" bold="true" box="[786,915,799,825]" pageId="3" pageNumber="154">
<taxonomicNameLabel id="04F38F18FFED3A6DFCD5FCA2FC709C32" box="[786,915,799,825]" pageId="3" pageNumber="154" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
[1b],
<taxonomicName id="EAB495F2FFED3A6DFC10FCA2FAF79C32" authority="(Curran, 1930 a)" authorityName="Miranda &amp; Skevington &amp; Marshall" authorityYear="2020" baseAuthorityName="Curran" baseAuthorityYear="1930" box="[983,1300,799,825]" class="Insecta" family="Syrphidae" genus="Fragosa" kingdom="Plantae" order="Diptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="deceptor" status="comb. nov.">
<emphasis id="1FC03263FFED3A6DFC10FCA2FC0B9C33" box="[983,1000,799,824]" italics="true" pageId="3" pageNumber="154">F</emphasis>
.
<emphasis id="1FC03263FFED3A6DFC30FCA2FBBA9C32" box="[1015,1113,799,825]" italics="true" pageId="3" pageNumber="154">deceptor</emphasis>
(
<bibRefCitation id="49259380FFED3A6DFBAFFCA2FAE89C32" author="Curran, C. H." box="[1128,1291,799,825]" pageId="3" pageNumber="154" pagination="1 - 16" refId="ref15673" refString="Curran, C. H. (1930 a) New species of Diptera belonging to the genus Baccha Fabricius (Syrphidae). American Museum Novitates, 403, 1 - 16." type="journal article" year="1930">Curran, 1930a</bibRefCitation>
)
</taxonomicName>
<emphasis id="1FC03263FFED3A6DFADCFCA2FA7F9C32" bold="true" box="[1307,1436,799,825]" pageId="3" pageNumber="154">
<taxonomicNameLabel id="04F38F18FFED3A6DFADCFCA2FA7F9C32" box="[1307,1436,799,825]" pageId="3" pageNumber="154" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
[1b],
<taxonomicName id="EAB495F2FFED3A6DFF14FCFEFE109C56" authority="(Shannon, 1927)" authorityName="Miranda &amp; Skevington &amp; Marshall" authorityYear="2020" baseAuthorityName="Shannon" baseAuthorityYear="1927" box="[211,499,835,861]" class="Insecta" family="Syrphidae" genus="Fragosa" kingdom="Plantae" order="Diptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="filiola" status="comb. nov.">
<emphasis id="1FC03263FFED3A6DFF14FCFEFF079C57" box="[211,228,835,860]" italics="true" pageId="3" pageNumber="154">F</emphasis>
.
<emphasis id="1FC03263FFED3A6DFF36FCFEFED79C56" box="[241,308,835,861]" italics="true" pageId="3" pageNumber="154">filiola</emphasis>
(
<bibRefCitation id="49259380FFED3A6DFE86FCFEFE099C56" author="Shannon, R. C." box="[321,490,835,861]" pageId="3" pageNumber="154" pagination="1 - 34" refId="ref17964" refString="Shannon, R. C. (1927) A review of the South American two-winged flies of the family Syrphidae. Proceedings of the United States National Museum, 70 (9), No. 2658, 1 - 34. https: // doi. org / 10.5479 / si. 00963801.70 - 2658.1" type="journal article" year="1927">Shannon, 1927</bibRefCitation>
)
</taxonomicName>
<emphasis id="1FC03263FFED3A6DFE3EFCFEFD9B9C56" bold="true" box="[505,632,835,861]" pageId="3" pageNumber="154">
<taxonomicNameLabel id="04F38F18FFED3A6DFE3EFCFEFD9B9C56" box="[505,632,835,861]" pageId="3" pageNumber="154" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
[2],
<taxonomicName id="EAB495F2FFED3A6DFD6CFCFEFC309C56" authority="(Hull, 1948)" authorityName="Miranda &amp; Skevington &amp; Marshall" authorityYear="2020" baseAuthorityName="Hull" baseAuthorityYear="1948" box="[683,979,835,861]" class="Insecta" family="Syrphidae" genus="Fragosa" kingdom="Plantae" order="Diptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="harlequina" status="comb. nov.">
<emphasis id="1FC03263FFED3A6DFD6CFCFEFD5F9C57" box="[683,700,835,860]" italics="true" pageId="3" pageNumber="154">F</emphasis>
.
<emphasis id="1FC03263FFED3A6DFD0EFCFEFCA79C56" box="[713,836,835,861]" italics="true" pageId="3" pageNumber="154">harlequina</emphasis>
(
<bibRefCitation id="49259380FFED3A6DFC95FCFEFC299C56" author="Hull, F. M." box="[850,970,835,861]" pageId="3" pageNumber="154" pagination="1 - 12" refId="ref16645" refString="Hull, F. M. (1948) Some Neotropical species of syrphids. Entomological News, 59, 1 - 12." type="journal article" year="1948">Hull, 1948</bibRefCitation>
)
</taxonomicName>
<emphasis id="1FC03263FFED3A6DFC1EFCFEFBBB9C56" bold="true" box="[985,1112,835,861]" pageId="3" pageNumber="154">
<taxonomicNameLabel id="04F38F18FFED3A6DFC1EFCFEFBBB9C56" box="[985,1112,835,861]" pageId="3" pageNumber="154" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
[1b, 1c],
<taxonomicName id="EAB495F2FFED3A6DFB07FCFEFF069C8A" authority="(Hull, 1947 a)" authorityName="Miranda &amp; Skevington &amp; Marshall" authorityYear="2020" baseAuthorityName="Hull" baseAuthorityYear="1947" class="Insecta" family="Syrphidae" genus="Fragosa" kingdom="Plantae" order="Diptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="hyacinthia" status="comb. nov.">
<emphasis id="1FC03263FFED3A6DFB07FCFEFB329C57" box="[1216,1233,835,860]" italics="true" pageId="3" pageNumber="154">F</emphasis>
.
<emphasis id="1FC03263FFED3A6DFB19FCFEFAB79C56" box="[1246,1364,835,861]" italics="true" pageId="3" pageNumber="154">hyacinthia</emphasis>
(
<bibRefCitation id="49259380FFED3A6DFAA5FCFEFF3E9C8A" author="Hull, F. M." pageId="3" pageNumber="154" pagination="395 - 410" refId="ref16574" refString="Hull, F. M. (1947 a) More flies of the genus Baccha (Diptera, Syrphidae). Revista de Entomologia, Rio de Janeiro, 18, 395 - 410." type="journal article" year="1947">Hull, 1947a</bibRefCitation>
)
</taxonomicName>
<emphasis id="1FC03263FFED3A6DFF2CFCDAFE889C8A" bold="true" box="[235,363,871,897]" pageId="3" pageNumber="154">
<taxonomicNameLabel id="04F38F18FFED3A6DFF2CFCDAFE889C8A" box="[235,363,871,897]" pageId="3" pageNumber="154" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
[2],
<taxonomicName id="EAB495F2FFED3A6DFE67FCDAFD459C8A" authority="(Curran, 1941)" authorityName="Miranda &amp; Skevington &amp; Marshall" authorityYear="2020" baseAuthorityName="Curran" baseAuthorityYear="1941" box="[416,678,871,897]" class="Insecta" family="Syrphidae" genus="Fragosa" kingdom="Plantae" order="Diptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="mara" status="comb. nov.">
<emphasis id="1FC03263FFED3A6DFE67FCDAFE529C8B" box="[416,433,871,896]" italics="true" pageId="3" pageNumber="154">F</emphasis>
.
<emphasis id="1FC03263FFED3A6DFE79FCDAFE1A9C8B" box="[446,505,871,896]" italics="true" pageId="3" pageNumber="154">mara</emphasis>
(
<bibRefCitation id="49259380FFED3A6DFDCFFCDAFD7D9C8A" author="Curran, C. H." box="[520,670,871,897]" pageId="3" pageNumber="154" pagination="243 - 304" refId="ref15863" refString="Curran, C. H. (1941) New American Syrphidae. Bulletin of the American Museum of Natural History, 78, 243 - 304." type="journal article" year="1941">Curran, 1941</bibRefCitation>
)
</taxonomicName>
<emphasis id="1FC03263FFED3A6DFD6BFCDAFCCF9C8A" bold="true" box="[684,812,871,897]" pageId="3" pageNumber="154">
<taxonomicNameLabel id="04F38F18FFED3A6DFD6BFCDAFCCF9C8A" box="[684,812,871,897]" pageId="3" pageNumber="154" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
[1b],
<taxonomicName id="EAB495F2FFED3A6DFCA8FCDAFB989C8A" authority="(Hull, 1949 a)" authorityName="Miranda &amp; Skevington &amp; Marshall" authorityYear="2020" baseAuthorityName="Hull" baseAuthorityYear="1949" box="[879,1147,871,897]" class="Insecta" family="Syrphidae" genus="Fragosa" kingdom="Plantae" order="Diptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="oenone" status="comb. nov.">
<emphasis id="1FC03263FFED3A6DFCA8FCDAFC639C8B" box="[879,896,871,896]" italics="true" pageId="3" pageNumber="154">F</emphasis>
.
<emphasis id="1FC03263FFED3A6DFC4AFCDAFC3D9C8B" box="[909,990,871,896]" italics="true" pageId="3" pageNumber="154">oenone</emphasis>
(
<bibRefCitation id="49259380FFED3A6DFC2AFCDAFB979C8A" author="Hull, F. M." box="[1005,1140,871,897]" pageId="3" pageNumber="154" pagination="89 - 285" refId="ref16669" refString="Hull, F. M. (1949 a) The genus Baccha from the New World. Entomologica Americana, 27, 89 - 285, 47 pls. [1947]" type="journal article" year="1949">Hull, 1949a</bibRefCitation>
)
</taxonomicName>
<emphasis id="1FC03263FFED3A6DFB45FCDAFAE19C8A" bold="true" box="[1154,1282,871,897]" pageId="3" pageNumber="154">
<taxonomicNameLabel id="04F38F18FFED3A6DFB45FCDAFAE19C8A" box="[1154,1282,871,897]" pageId="3" pageNumber="154" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
[2,
<typeStatus id="F20F50D3FFED3A6DFAEAFCDAFABE9C8A" box="[1325,1373,871,897]" pageId="3" pageNumber="154">type</typeStatus>
lost],
<taxonomicName id="EAB495F2FFED3A6DFF50FC36FE3A9CAE" authority="(Curran, 1939)" authorityName="Miranda &amp; Skevington &amp; Marshall" authorityYear="2020" baseAuthorityName="Curran" baseAuthorityYear="1939" box="[151,473,907,933]" class="Insecta" family="Syrphidae" genus="Fragosa" kingdom="Plantae" order="Diptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="provocans" status="comb. nov.">
<emphasis id="1FC03263FFED3A6DFF50FC36FF4B9CAF" box="[151,168,907,932]" italics="true" pageId="3" pageNumber="154">F</emphasis>
.
<emphasis id="1FC03263FFED3A6DFF70FC36FEC89CAF" box="[183,299,907,932]" italics="true" pageId="3" pageNumber="154">provocans</emphasis>
(
<bibRefCitation id="49259380FFED3A6DFEFDFC36FE329CAE" author="Curran, C. H." box="[314,465,907,933]" pageId="3" pageNumber="154" pagination="1 - 12" refId="ref15834" refString="Curran, C. H. (1939) New Neotropical Baccha Fabricius (Syrphidae: Diptera). American Museum Novitates, 1041, 1 - 12." type="journal article" year="1939">Curran, 1939</bibRefCitation>
)
</taxonomicName>
<emphasis id="1FC03263FFED3A6DFE26FC36FD819CAE" bold="true" box="[481,610,907,933]" pageId="3" pageNumber="154">
<taxonomicNameLabel id="04F38F18FFED3A6DFE26FC36FD819CAE" box="[481,610,907,933]" pageId="3" pageNumber="154" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
[1b],
<taxonomicName id="EAB495F2FFED3A6DFD60FC36FC1D9CAD" authority="(Schiner, 1868)" authorityName="Miranda &amp; Skevington &amp; Marshall" authorityYear="2020" baseAuthorityName="Schiner" baseAuthorityYear="1868" box="[679,1022,907,934]" class="Insecta" family="Syrphidae" genus="Fragosa" kingdom="Plantae" order="Diptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="rugosifrons" status="comb. nov.">
<emphasis id="1FC03263FFED3A6DFD60FC36FD5B9CAF" box="[679,696,907,932]" italics="true" pageId="3" pageNumber="154">F</emphasis>
.
<emphasis id="1FC03263FFED3A6DFD01FC36FCA59CAE" box="[710,838,907,933]" italics="true" pageId="3" pageNumber="154">rugosifrons</emphasis>
(
<bibRefCitation id="49259380FFED3A6DFC91FC36FC169CAD" author="Schiner, I. R." box="[854,1013,907,934]" pageId="3" pageNumber="154" refId="ref17881" refString="Schiner, I. R. (1868) Diptera. In: Wullerstorf-Urbair, B. von (in charge), Reise der osterreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859 unter den Befehlen des Commodore B. von Wullerstorf-Urbair. Vol. 2. Band 1. Abtheilung B. Kaiserlich-konigliche Hof- und Staatsdruckerei in Commission bei Karl Gerold's Sohn, Wien, pp. i-vi + 1 - 388, 4 pls." type="journal volume" year="1868">Schiner, 1868</bibRefCitation>
)
</taxonomicName>
,
<emphasis id="1FC03263FFED3A6DFBCDFC36FB689CAE" bold="true" box="[1034,1163,907,933]" pageId="3" pageNumber="154">
<taxonomicNameLabel id="04F38F18FFED3A6DFBCDFC36FB689CAE" box="[1034,1163,907,933]" pageId="3" pageNumber="154" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
[2, 3, 4],
<taxonomicName id="EAB495F2FFED3A6DFB3CFC36FEB49CC2" authority="(Williston, 1888)" authorityName="Miranda &amp; Skevington &amp; Marshall" authorityYear="2020" baseAuthorityName="Williston" baseAuthorityYear="1888" class="Insecta" family="Syrphidae" genus="Ocyptamus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="stenogaster" status="comb. nov.">
<emphasis id="1FC03263FFED3A6DFB3CFC36FAEF9CAF" box="[1275,1292,907,932]" italics="true" pageId="3" pageNumber="154">F</emphasis>
.
<emphasis id="1FC03263FFED3A6DFADCFC36FA7F9CAF" box="[1307,1436,907,932]" italics="true" pageId="3" pageNumber="154">stenogaster</emphasis>
(
<bibRefCitation id="49259380FFED3A6DFF58FC12FEAC9CC2" author="Williston, S. W." box="[159,335,943,969]" pageId="3" pageNumber="154" pagination="243 - 292" refId="ref18565" refString="Williston, S. W. (1888) Diptera Brasiliana, ab H. H. Smith collecta. Part I, Stratiomyidae, Syrphidae. Transactions of the American Entomological Society, 15, 243 - 292. https: // doi. org / 10.2307 / 25076505" type="journal article" year="1888">Williston, 1888</bibRefCitation>
)
</taxonomicName>
<emphasis id="1FC03263FFED3A6DFE99FC12FE3D9CC2" bold="true" box="[350,478,943,969]" pageId="3" pageNumber="154">
<taxonomicNameLabel id="04F38F18FFED3A6DFE99FC12FE3D9CC2" box="[350,478,943,969]" pageId="3" pageNumber="154" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
[1b],
<taxonomicName id="EAB495F2FFED3A6DFDE5FC12FCA69CC1" authority="(Hull, 1947 b)" authorityName="Miranda &amp; Skevington &amp; Marshall" authorityYear="2020" baseAuthorityName="Hull" baseAuthorityYear="1947" box="[546,837,943,970]" class="Insecta" family="Syrphidae" genus="Fragosa" kingdom="Plantae" order="Diptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="zephyrea" status="comb. nov.">
<emphasis id="1FC03263FFED3A6DFDE5FC12FDD09CC3" box="[546,563,943,968]" italics="true" pageId="3" pageNumber="154">F</emphasis>
.
<emphasis id="1FC03263FFED3A6DFD86FC12FD469CC2" box="[577,677,943,969]" italics="true" pageId="3" pageNumber="154">zephyrea</emphasis>
(
<bibRefCitation id="49259380FFED3A6DFD73FC12FCDF9CC1" author="Hull, F. M." box="[692,828,943,970]" pageId="3" pageNumber="154" pagination="230 - 240" refId="ref16610" refString="Hull, F. M. (1947 b) Some American syrphid flies. Psyche, 54, 230 - 240. https: // doi. org / 10.1155 / 1947 / 51282" type="journal article" year="1947">Hull, 1947b</bibRefCitation>
)
</taxonomicName>
<emphasis id="1FC03263FFED3A6DFC8CFC12FC2F9CC2" bold="true" box="[843,972,943,969]" pageId="3" pageNumber="154">
<taxonomicNameLabel id="04F38F18FFED3A6DFC8CFC12FC2F9CC2" box="[843,972,943,969]" pageId="3" pageNumber="154" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
[1b].
</paragraph>
</subSubSection>
<subSubSection id="65AEBDFAFFED3A6DFF00FC6FFDF49B76" pageId="3" pageNumber="154" type="discussion">
<paragraph id="2D0BEE71FFED3A6DFF00FC6FFDF49B76" blockId="3.[151,1437,402,1149]" pageId="3" pageNumber="154">
<emphasis id="1FC03263FFED3A6DFF00FC6FFEAC9CE7" bold="true" box="[199,335,978,1004]" pageId="3" pageNumber="154">Comments.</emphasis>
The rugose frons/frontal triangle is a distinct character of this group (
<figureCitation id="B58FF2F4FFED3A6DFB98FC6EFB479CE6" box="[1119,1188,979,1005]" captionStart="FIGURES 111" captionStartId="4.[151,264,1631,1655]" captionTargetBox="[177,1414,185,1601]" captionTargetId="figure-19@4.[165,1423,181,1608]" captionTargetPageId="4" captionText="FIGURES 111. Fragosa Miranda gen. nov. 1: Fragosa sp. (INPA-DIP001362), female, habitus, dorsal view, photo previously published in Mengual et al. (2018). 2: Fragosa aff. titania (CNC), female, abdomen, dorsal view, photo previously published in Mengual et al. (2018). 3: Fragosa argentina (Curran) (CNC_Diptera196006), female, wing. 4: Fragosa stenogaster (Williston) (debu01088839), female, head, frontal view, photo previously published in Mengual et al. (2018). 5: Fragosa stenogaster (Williston) (debu01088840), male, head, lateral view. 6: Fragosa sp., female, habitus, dorsal view, by Steve Marshall. 7: Fragosa stenogaster (Williston) (debu01088840), male, epandrium, lateral view, photo previously published in Mengual et al. (2018). 8: Fragosa stenogaster (Williston) (debu01088839), female, apex of abdomen, dorsal view, photo previously published in Mengual et al. (2018). 911. Fragosa stenogaster (Williston) (debu01088840), male. 9: epandrium, ventral view. 10: hypandrium, lateral view. 11: hypandrium, ventral view. ap: apodeme; b: basiphallus; bm: cell bm; br: cell br; c: cell c; d: distiphallus; e: epiproct; f: frons; h: hypandrium; oc: occiput; p: postgonite; s: surstylus; se: subepandrial sclerite; t: facial tubercle; t2, t3, t7 and t8: terga 2, 3, 7 and 8." figureDoi="http://doi.org/10.5281/zenodo.4401371" httpUri="https://zenodo.org/record/4401371/files/figure.png" pageId="3" pageNumber="154">Fig. 4</figureCitation>
), but not unique since it occurs in the
<taxonomicName id="EAB495F2FFED3A6DFE83FC4AFD639B1A" authorityName="Schiner" authorityYear="1868" box="[324,640,1015,1041]" class="Insecta" family="Syrphidae" genus="Pelecinobaccha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="brevipennis">
<emphasis id="1FC03263FFED3A6DFE83FC4AFD639B1A" box="[324,640,1015,1041]" italics="true" pageId="3" pageNumber="154">Pelecinobaccha brevipennis</emphasis>
</taxonomicName>
species group as well. Further distinct characters for the
<taxonomicName id="EAB495F2FFED3A6DFAC6FC4AFA7F9B1A" authorityName="Miranda &amp; Skevington &amp; Marshall" authorityYear="2020" baseAuthorityName="Schiner" baseAuthorityYear="1868" box="[1281,1436,1015,1041]" class="Insecta" family="Syrphidae" genus="Fragosa" kingdom="Plantae" order="Diptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="rugosifrons">
<emphasis id="1FC03263FFED3A6DFAC6FC4AFA7F9B1A" box="[1281,1436,1015,1041]" italics="true" pageId="3" pageNumber="154">F. rugosifrons</emphasis>
</taxonomicName>
group include the face that is dark dorsal to tubercle, the pointed and medially positioned facial tubercle (
<figureCitation id="B58FF2F4FFED3A6DFAE5FBA6FA869B3E" box="[1314,1381,1050,1077]" captionStart="FIGURES 111" captionStartId="4.[151,264,1631,1655]" captionTargetBox="[177,1414,185,1601]" captionTargetId="figure-19@4.[165,1423,181,1608]" captionTargetPageId="4" captionText="FIGURES 111. Fragosa Miranda gen. nov. 1: Fragosa sp. (INPA-DIP001362), female, habitus, dorsal view, photo previously published in Mengual et al. (2018). 2: Fragosa aff. titania (CNC), female, abdomen, dorsal view, photo previously published in Mengual et al. (2018). 3: Fragosa argentina (Curran) (CNC_Diptera196006), female, wing. 4: Fragosa stenogaster (Williston) (debu01088839), female, head, frontal view, photo previously published in Mengual et al. (2018). 5: Fragosa stenogaster (Williston) (debu01088840), male, head, lateral view. 6: Fragosa sp., female, habitus, dorsal view, by Steve Marshall. 7: Fragosa stenogaster (Williston) (debu01088840), male, epandrium, lateral view, photo previously published in Mengual et al. (2018). 8: Fragosa stenogaster (Williston) (debu01088839), female, apex of abdomen, dorsal view, photo previously published in Mengual et al. (2018). 911. Fragosa stenogaster (Williston) (debu01088840), male. 9: epandrium, ventral view. 10: hypandrium, lateral view. 11: hypandrium, ventral view. ap: apodeme; b: basiphallus; bm: cell bm; br: cell br; c: cell c; d: distiphallus; e: epiproct; f: frons; h: hypandrium; oc: occiput; p: postgonite; s: surstylus; se: subepandrial sclerite; t: facial tubercle; t2, t3, t7 and t8: terga 2, 3, 7 and 8." figureDoi="http://doi.org/10.5281/zenodo.4401371" httpUri="https://zenodo.org/record/4401371/files/figure.png" pageId="3" pageNumber="154">Fig. 5</figureCitation>
), the entirely pale scutellum, the almost complete post-metacoxal bridge (
<figureCitation id="B58FF2F4FFED3A6DFC5EFB82FC0F9B52" box="[921,1004,1087,1113]" captionStart="FIGURES 7888. 78" captionStartId="17.[151,264,1665,1689]" captionTargetBox="[301,1287,192,1634]" captionTargetId="figure-18@17.[294,1294,181,1642]" captionTargetPageId="17" captionText="FIGURES 7888. 78: Fragosa harlequina (Hull) (holotype Baccha harlequina Hull, AMNH), male, thorax, lateral, by AMNH (line drawing represents outline of the alula). 79: Fragosa argentina (Curran) (CNC_Diptera174232), thorax, postero-ventral (line drawing represents outline of the incomplete post-metacoxal bridge). 8081: Fragosa virgilio (Hull) (holotype Baccha virgilio Hull, CNC, CNC_Diptera189124), male. 80: habitus, lateral, by CNC. 81: head, oblique frontal, by CNC. 82: Fragosa titania (Hull) (JSS25229), female, head, dorsal. 83: Fragosa stenogaster (Williston) (debu01088840), male, head, dorsal. 8485: Hypocritanus fascipennis (Wiedemann), female, head. 84: frontal, by Ximo Mengual (http://syrphidae.myspecies.info/ taxonomy/term/879). 85: lateral (CNC_Diptera501). 8687: Nuntianus cecrops (Hull) (INPA-DIP000246), female, head. 86: dorsal. 87: lateral. 88: Victoriana aff. melanorrhina (DEBU), female, head, lateral. a: alula; ai: antennal insertion; f: frons; ft: frontal triangle; lem: lateral eye margin; mt: metafemur; o: ocellar triangle; oc: occiput; pmt: post-metacoxal bridge; t: facial tubercle." figureDoi="http://doi.org/10.5281/zenodo.4401387" httpUri="https://zenodo.org/record/4401387/files/figure.png" pageId="3" pageNumber="154">Fig. 79</figureCitation>
), the enlarged epandrium (
<figureCitation id="B58FF2F4FFED3A6DFADEFB82FABD9B52" box="[1305,1374,1087,1113]" captionStart="FIGURES 111" captionStartId="4.[151,264,1631,1655]" captionTargetBox="[177,1414,185,1601]" captionTargetId="figure-19@4.[165,1423,181,1608]" captionTargetPageId="4" captionText="FIGURES 111. Fragosa Miranda gen. nov. 1: Fragosa sp. (INPA-DIP001362), female, habitus, dorsal view, photo previously published in Mengual et al. (2018). 2: Fragosa aff. titania (CNC), female, abdomen, dorsal view, photo previously published in Mengual et al. (2018). 3: Fragosa argentina (Curran) (CNC_Diptera196006), female, wing. 4: Fragosa stenogaster (Williston) (debu01088839), female, head, frontal view, photo previously published in Mengual et al. (2018). 5: Fragosa stenogaster (Williston) (debu01088840), male, head, lateral view. 6: Fragosa sp., female, habitus, dorsal view, by Steve Marshall. 7: Fragosa stenogaster (Williston) (debu01088840), male, epandrium, lateral view, photo previously published in Mengual et al. (2018). 8: Fragosa stenogaster (Williston) (debu01088839), female, apex of abdomen, dorsal view, photo previously published in Mengual et al. (2018). 911. Fragosa stenogaster (Williston) (debu01088840), male. 9: epandrium, ventral view. 10: hypandrium, lateral view. 11: hypandrium, ventral view. ap: apodeme; b: basiphallus; bm: cell bm; br: cell br; c: cell c; d: distiphallus; e: epiproct; f: frons; h: hypandrium; oc: occiput; p: postgonite; s: surstylus; se: subepandrial sclerite; t: facial tubercle; t2, t3, t7 and t8: terga 2, 3, 7 and 8." figureDoi="http://doi.org/10.5281/zenodo.4401371" httpUri="https://zenodo.org/record/4401371/files/figure.png" pageId="3" pageNumber="154">Fig. 7</figureCitation>
), and the reduced hypandrium (
<figureCitation id="B58FF2F4FFED3A6DFE70FBDEFDE99B76" box="[439,522,1123,1149]" captionStart="FIGURES 111" captionStartId="4.[151,264,1631,1655]" captionTargetBox="[177,1414,185,1601]" captionTargetId="figure-19@4.[165,1423,181,1608]" captionTargetPageId="4" captionText="FIGURES 111. Fragosa Miranda gen. nov. 1: Fragosa sp. (INPA-DIP001362), female, habitus, dorsal view, photo previously published in Mengual et al. (2018). 2: Fragosa aff. titania (CNC), female, abdomen, dorsal view, photo previously published in Mengual et al. (2018). 3: Fragosa argentina (Curran) (CNC_Diptera196006), female, wing. 4: Fragosa stenogaster (Williston) (debu01088839), female, head, frontal view, photo previously published in Mengual et al. (2018). 5: Fragosa stenogaster (Williston) (debu01088840), male, head, lateral view. 6: Fragosa sp., female, habitus, dorsal view, by Steve Marshall. 7: Fragosa stenogaster (Williston) (debu01088840), male, epandrium, lateral view, photo previously published in Mengual et al. (2018). 8: Fragosa stenogaster (Williston) (debu01088839), female, apex of abdomen, dorsal view, photo previously published in Mengual et al. (2018). 911. Fragosa stenogaster (Williston) (debu01088840), male. 9: epandrium, ventral view. 10: hypandrium, lateral view. 11: hypandrium, ventral view. ap: apodeme; b: basiphallus; bm: cell bm; br: cell br; c: cell c; d: distiphallus; e: epiproct; f: frons; h: hypandrium; oc: occiput; p: postgonite; s: surstylus; se: subepandrial sclerite; t: facial tubercle; t2, t3, t7 and t8: terga 2, 3, 7 and 8." figureDoi="http://doi.org/10.5281/zenodo.4401371" httpUri="https://zenodo.org/record/4401371/files/figure.png" pageId="3" pageNumber="154">Fig. 10</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>