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<document id="36CC029373BCEAB34D5A260B159C6667" ID-DOI="10.5281/zenodo.199681" ID-GBIF-Dataset="62e1623e-d85e-46ea-8edf-6cb01d7c88eb" ID-ISSN="1175-5326" ID-Zenodo-Dep="199681" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460343126729" checkinUser="plazi" docAuthor="Blaimer, Bonnie B." docDate="2010" docId="6516CC460D0CFFF7FF7A8D7360C7659F" docLanguage="en" docName="zt02714p039.pdf" docOrigin="Zootaxa 2714" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Crematogaster grevei Forel 1891" docType="treatment" docVersion="8" lastPageNumber="17" masterDocId="992FB43E0D01FFE7FFED8A216475640D" masterDocTitle="Taxonomy and Natural History of the Crematogaster (Decacrema) - group (Hymenoptera: Formicidae) in Madagascar" masterLastPageNumber="39" masterPageNumber="1" pageNumber="14" updateTime="1698261518362" updateUser="plazi">
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<mods:title id="4C4991E9044AB3C3F875E51CEF07807F">Taxonomy and Natural History of the Crematogaster (Decacrema) - group (Hymenoptera: Formicidae) in Madagascar</mods:title>
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<mods:namePart id="A7A6B70B189904390843D9A85100CDF2">Blaimer, Bonnie B.</mods:namePart>
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<mods:date id="2CEA884198692302CE36498864871E21">2010</mods:date>
<mods:detail id="4BF3932CDFF97A470E2E0756B24B983F" type="volume">
<mods:number id="24AE29527960C3B84A5133753E0CEF2E">2714</mods:number>
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<mods:identifier id="5634B837937D3BA25FF08460D77837BE" type="DOI">10.5281/zenodo.199681</mods:identifier>
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<mods:identifier id="539AA5BCEB094CE4AAB1C8350F892CB7" type="Zenodo-Dep">199681</mods:identifier>
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<treatment id="6516CC460D0CFFF7FF7A8D7360C7659F" ID-DOI="http://doi.org/10.5281/zenodo.6199370" ID-GBIF-Taxon="119406764" ID-Zenodo-Dep="6199370" LSID="urn:lsid:plazi:treatment:6516CC460D0CFFF7FF7A8D7360C7659F" httpUri="http://treatment.plazi.org/id/6516CC460D0CFFF7FF7A8D7360C7659F" lastPageId="16" lastPageNumber="17" pageId="13" pageNumber="14">
<subSubSection id="A5A52EDB0D0CFFEAFF7A8D7365346382" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="ED007D500D0CFFEAFF7A8D7366506361" blockId="13.[151,549,1874,1935]" box="[151,549,1874,1900]" pageId="13" pageNumber="14">
<heading id="B648CA3C0D0CFFEAFF7A8D7366506361" bold="true" box="[151,549,1874,1900]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
<taxonomicName id="2ABF06D30D0CFFEAFF7A8D7366506361" ID-CoL="6BDC5" authority="Forel 1891" authorityName="Forel" authorityYear="1891" box="[151,549,1874,1900]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="grevei">
<emphasis id="DFCBA1420D0CFFEAFF7A8D7366506361" bold="true" box="[151,549,1874,1900]" pageId="13" pageNumber="14">
<emphasis id="DFCBA1420D0CFFEAFF7A8D7365EC6361" bold="true" box="[151,409,1874,1900]" italics="true" pageId="13" pageNumber="14">Crematogaster grevei</emphasis>
<bibRefCitation id="892E00A10D0CFFEAFE4D8D7366506361" author="Forel" box="[416,549,1874,1900]" pageId="13" pageNumber="14" refString="Forel, A. (1891) Les Formicides. [part]. In: A. Grandidier (Ed), Histoire physique, naturelle, et politique de Madagascar. Volume XX. Histoire naturelle des Hymenopteres. Deuxieme partie (28 e fascicule). Hachette et Cie, Paris, p. 1 - 280." type="book chapter" year="1891">Forel 1891</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="ED007D500D0CFFEAFF7A8D5465346382" blockId="13.[151,549,1874,1935]" box="[151,321,1909,1935]" pageId="13" pageNumber="14">
(
<figureCitation id="758461D50D0CFFEAFF728D5464866382" box="[159,243,1909,1935]" captionStart="FIGURES 32 – 37" captionStartId="29.[151,270,1974,1998]" captionTargetBox="[190,1397,196,1939]" captionTargetId="figure@29.[179,1407,166,1953]" captionTargetPageId="29" captionText="FIGURES 32 – 37. Malagasy Crematogaster (Decacrema) species; a: full-face view, b: lateral view, c: dorsal view. 32: C. nosibeensis; 33: C. sisa; 34: C. sabatra; 35: C. mahery; 36: C. malala; 37: C. grevei." httpUri="https://zenodo.org/record/199685/files/figure.png" pageId="13" pageNumber="14">Fig. 37</figureCitation>
&amp;
<figureCitation id="758461D50D0CFFEAFEF48D54654D6382" box="[281,312,1909,1935]" captionStart="FIGURES 38 – 43" captionStartId="30.[151,270,1894,1918]" captionTargetBox="[221,1364,193,1874]" captionTargetId="figure@30.[200,1385,164,1882]" captionTargetPageId="30" captionText="FIGURES 38 – 43. Malagasy Crematogaster (Decacrema) species; a: petiole &amp; postpetiole in dorsal view, b: petiole &amp; postpetiole in lateral view. 38: C. nosibeensis; 39: C. sisa, 40: C. sabatra; 41: C. mahery; 42: C. malala; 43: C. grevei." httpUri="https://zenodo.org/record/199686/files/figure.png" pageId="13" pageNumber="14">43</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="A5A52EDB0D0CFFE9FF7A8D9B609364E2" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="reference_group">
<paragraph id="ED007D500D0CFFE9FF7A8D9B674B64A2" blockId="13.[151,1436,1978,2033]" lastBlockId="14.[151,1266,151,239]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<treatmentCitationGroup id="CDAF5A7E0D0CFFE9FF7A8D9B674B64A2" infered="true" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<taxonomicName id="2ABF06D30D0CFFEAFF7A8D9B661663DC" ID-CoL="6BDC5" authority="Forel, A. 1891:183" authorityName="Forel, A." authorityPageNumber="183" authorityYear="1891" box="[151,611,1978,2001]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="grevei">
<emphasis id="DFCBA1420D0CFFEAFF7A8D9B65F763DC" box="[151,386,1978,2001]" italics="true" pageId="13" pageNumber="14">Crematogaster grevei</emphasis>
<treatmentCitation id="6C1E5B410D0CFFEAFE618D9B661663DC" author="Forel" box="[396,611,1978,2001]" page="183" pageId="13" pageNumber="14" year="1891">
<bibRefCitation id="892E00A10D0CFFEAFE618D9B664163DC" author="Forel" box="[396,564,1978,2001]" pageId="13" pageNumber="14" refString="Forel, A. (1891) Les Formicides. [part]. In: A. Grandidier (Ed), Histoire physique, naturelle, et politique de Madagascar. Volume XX. Histoire naturelle des Hymenopteres. Deuxieme partie (28 e fascicule). Hachette et Cie, Paris, p. 1 - 280." type="book chapter" year="1891">Forel, A. 1891</bibRefCitation>
:183
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="3204C3F20D0CFFEAFD9D8D9B66A163DC" box="[624,724,1978,2001]" pageId="13" pageNumber="14" type="syntype">Syntypes</typeStatus>
,
<specimenCount id="FBB9B6D90D0CFFEAFD0E8D9B672163DC" box="[739,852,1978,2001]" pageId="13" pageNumber="14" type="worker">2 workers</specimenCount>
, Morondava (Grevé) (MHNG) [examined] [The two MHNG
<typeStatus id="3204C3F20D0CFFEAFEDC8DFB65FB63FC" box="[305,398,2010,2033]" pageId="13" pageNumber="14" type="syntype">syntypes</typeStatus>
are also imaged on AntWeb: CASENT0101572, CASENT0101558]. One
<typeStatus id="3204C3F20D0CFFEAFAA48DFB61E963FC" box="[1353,1436,2010,2033]" pageId="13" pageNumber="14" type="syntype">syntype</typeStatus>
(CASENT0101572) hereby designated the
<typeStatus id="3204C3F20D0FFFE9FD7E8AB6674264A2" box="[659,823,151,175]" pageId="14" pageNumber="15" type="lectotype">
<emphasis id="DFCBA1420D0FFFE9FD7E8AB6674264A2" bold="true" box="[659,823,151,175]" pageId="14" pageNumber="15">LECTOTYPE</emphasis>
</typeStatus>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="ED007D500D0FFFE9FF7A8A99609364E2" blockId="14.[151,1266,151,239]" pageId="14" pageNumber="15">
<treatmentCitationGroup id="CDAF5A7E0D0FFFE9FF7A8A99609364E2" pageId="14" pageNumber="15">
<taxonomicName id="2ABF06D30D0FFFE9FF7A8A99677764C2" ID-CoL="6BDC5" authority="Forel, Forel 1910a:18" authorityName="Forel, Forel" authorityPageNumber="18" authorityYear="1910" box="[151,770,184,207]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="grevei" subGenus="Decacrema">
<emphasis id="DFCBA1420D0FFFE9FF7A8A99654364C2" box="[151,310,184,207]" italics="true" pageId="14" pageNumber="15">Crematogaster</emphasis>
(
<emphasis id="DFCBA1420D0FFFE9FEA88A9965CA64C2" box="[325,447,184,207]" italics="true" pageId="14" pageNumber="15">Decacrema</emphasis>
)
<emphasis id="DFCBA1420D0FFFE9FE228A99666464C2" box="[463,529,184,207]" italics="true" pageId="14" pageNumber="15">grevei</emphasis>
<treatmentCitation id="6C1E5B410D0FFFE9FDFA8A99677764C2" author="Forel" box="[535,770,184,207]" page="18" pageId="14" pageNumber="15" year="1910">
<bibRefCitation id="892E00A10D0FFFE9FDFA8A99669164C2" author="Forel" box="[535,740,184,207]" pageId="14" pageNumber="15" refString="Forel, A. (1910 a) Glanures myrmecologiques. Annales de la Societe Entomologique de Belgique, 54, 6 - 32." type="journal article" year="1910" yearSuffix="a">Forel, Forel 1910a</bibRefCitation>
:18
</treatmentCitation>
</taxonomicName>
. Combination in
<emphasis id="DFCBA1420D0FFFE9FC548A99608764C2" box="[953,1266,184,207]" italics="true" pageId="14" pageNumber="15">
<taxonomicName id="2ABF06D30D0FFFE9FC548A99609B64C2" box="[953,1262,184,207]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="10" phylum="Arthropoda" rank="subGenus" subGenus="Decacrema">Crematogaster (Decacrema)</taxonomicName>
.
</emphasis>
<taxonomicName id="2ABF06D30D0FFFE9FF7A8AF965BD64E2" ID-CoL="6BDC5" authority="(Decacrema)" authorityName="Forel, Forel" authorityPageNumber="9" authorityYear="1910" baseAuthorityName="Decacrema" box="[151,456,216,239]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="DFCBA1420D0FFFE9FF7A8AF9654364E2" box="[151,310,216,239]" italics="true" pageId="14" pageNumber="15">Crematogaster</emphasis>
(
<emphasis id="DFCBA1420D0FFFE9FEA88AF965CA64E2" box="[325,447,216,239]" italics="true" pageId="14" pageNumber="15">Decacrema</emphasis>
)
</taxonomicName>
<emphasis id="DFCBA1420D0FFFE9FE228AF9666464E2" box="[463,529,216,239]" italics="true" pageId="14" pageNumber="15">grevei</emphasis>
<treatmentCitation id="6C1E5B410D0FFFE9FDFA8AF9668264E2" author="Forel" box="[535,759,216,239]" page="9" pageId="14" pageNumber="15" year="1910">
<bibRefCitation id="892E00A10D0FFFE9FDFA8AF9669064E2" author="Forel" box="[535,741,216,239]" pageId="14" pageNumber="15" refString="Forel, A. (1910 b) Zoologische und anthropologische Ergebnisse einer Forschungsreise im westlichen und zentralen Sudafrika ausgefuhrt in den Jahren 1903 - 1905 von Dr. Leonhard Schultze. Vierter Band. Systematik und Tiergeographie. D) Formicidae. Denkschriften der Medizinsch-Naturwissenschaftlichen Gesellschaft Jena, 16, 1 - 30." type="journal article" year="1910" yearSuffix="b">Forel, Forel 1910b</bibRefCitation>
:9
</treatmentCitation>
. Combination in
<taxonomicName id="2ABF06D30D0FFFE9FC408AF960AA64E2" box="[941,1247,216,239]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="10" phylum="Arthropoda" rank="subGenus" subGenus="Decacrema">
<emphasis id="DFCBA1420D0FFFE9FC408AF9603964E2" box="[941,1100,216,239]" italics="true" pageId="14" pageNumber="15">Crematogaster</emphasis>
(
<emphasis id="DFCBA1420D0FFFE9FBB18AF960A364E2" box="[1116,1238,216,239]" italics="true" pageId="14" pageNumber="15">Decacrema</emphasis>
)
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="A5A52EDB0D0FFFE9FF7A8B3C615860BA" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="ED007D500D0FFFE9FF7A8B3C615860BA" blockId="14.[151,1437,285,2028]" pageId="14" pageNumber="15">
<emphasis id="DFCBA1420D0FFFE9FF7A8B3C65D7653A" bold="true" box="[151,418,285,311]" pageId="14" pageNumber="15">Material examined.</emphasis>
(
<collectionCode id="8BAEE5950D0FFFE9FE598B3C6678653A" box="[436,525,285,311]" pageId="14" pageNumber="15">BBBC</collectionCode>
,
<collectionCode id="8BAEE5950D0FFFE9FDF68B3C6605653A" box="[539,624,285,311]" pageId="14" pageNumber="15">CASC</collectionCode>
,
<collectionCode id="8BAEE5950D0FFFE9FD938B3C66A2653A" box="[638,727,285,311]" pageId="14" pageNumber="15">PSWC</collectionCode>
,
<collectionCode id="8BAEE5950D0FFFE9FD088B3C6733653A" box="[741,838,285,311]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="14" pageNumber="15">MHNG</collectionCode>
)
<collectingCountry id="95A83DC00D0FFFE9FCB58B3C6040653A" box="[856,1077,285,311]" name="Madagascar" pageId="14" pageNumber="15">MADAGASCAR</collectingCountry>
<emphasis id="DFCBA1420D0FFFE9FBD38B3C608D653B" box="[1086,1272,285,310]" italics="true" pageId="14" pageNumber="15">Fianarantsoa:</emphasis>
P.N. Isalo: - 22.29833, 45.
<quantity id="2A47D0B50D0FFFE9FEBC8B6365D46551" box="[337,417,322,348]" metricMagnitude="7" metricUnit="m" metricValue="3.516799" pageId="14" pageNumber="15" unit="m" value="3.516799E7">35167</quantity>
, 990m (B.L.Fisher
<emphasis id="DFCBA1420D0FFFE9FD5C8B63668D6556" box="[689,760,322,347]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
); near P.N Isalo: -22.62667, 45.35817,
<quantity id="2A47D0B50D0FFFE9FAFB8B6361E96551" box="[1302,1436,322,348]" metricMagnitude="2" metricUnit="m" metricValue="7.625" metricValueMax="8.25" metricValueMin="7.0" pageId="14" pageNumber="15" unit="m" value="762.5" valueMax="825.0" valueMin="700.0">700-825m</quantity>
(R.Harin’Hala);
<emphasis id="DFCBA1420D0FFFE9FE8F8B49658C658C" box="[354,505,360,385]" italics="true" pageId="14" pageNumber="15">Mahajanga:</emphasis>
P.N. Namoroka: -16.37667, 45.32667,
<quantity id="2A47D0B50D0FFFE9FC338B496051658F" box="[990,1060,360,386]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="15" unit="m" value="100.0">100m</quantity>
(B.L.Fisher
<emphasis id="DFCBA1420D0FFFE9FB2F8B496175658C" box="[1218,1280,360,385]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
); Mahavavy River: -16.05167, 45.90833,
<quantity id="2A47D0B50D0FFFE9FDE98BAC664F65AA" box="[516,570,397,423]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="15" unit="m" value="20.0">20m</quantity>
(B.L.Fisher
<emphasis id="DFCBA1420D0FFFE9FD318BAC676965AB" box="[732,796,397,422]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
); Rés. forest. Beanka: -18.02649, 44.05051,
<quantity id="2A47D0B50D0FFFE9FABA8BAC61E865AA" box="[1367,1437,397,423]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="14" pageNumber="15" unit="m" value="250.0">250m</quantity>
(B.L.Fisher
<emphasis id="DFCBA1420D0FFFE9FECB8B93651765C6" box="[294,354,434,459]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
); Morondava: -20.28333, 44.28333 (M. Grevé);
<quantity id="2A47D0B50D0FFFE9FC4B8B93679C65C1" box="[934,1001,434,460]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="14" pageNumber="15" unit="km" value="50.0">50km</quantity>
N Morondava: -20.06670, 44.58330 (A.Pauly),
<quantity id="2A47D0B50D0FFFE9FEF58BF9652E65FF" box="[280,347,472,498]" metricMagnitude="4" metricUnit="m" metricValue="4.8" pageId="14" pageNumber="15" unit="km" value="48.0">48km</quantity>
ENE Morondava: -20.29650, 44.28150,
<quantity id="2A47D0B50D0FFFE9FCD18BF9671A65FF" box="[828,879,472,498]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="15" unit="m" value="30.0">30m</quantity>
(D.M.Olson);
<quantity id="2A47D0B50D0FFFE9FBF38BF9601C65FF" box="[1054,1129,472,498]" metricMagnitude="3" metricUnit="m" metricValue="3.7" pageId="14" pageNumber="15" unit="km" value="3.7">3.7km</quantity>
102º ESE Belo-sur-mer: - 20.73472, 44.04000,
<quantity id="2A47D0B50D0FFFE9FE7D8BDC65B1661A" box="[400,452,509,535]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="15" unit="m" value="10.0">10m</quantity>
(B.L.Fisher
<emphasis id="DFCBA1420D0FFFE9FDB68BDC66E3661B" box="[603,662,509,534]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
); Bemeraha: -18.65000, 44.71670 (D.C.Lees); P.N. Baie de Baly: -16.01000, 45.26500,
<quantity id="2A47D0B50D0FFFE9FE70880365A76631" box="[413,466,546,572]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="15" unit="m" value="10.0">10m</quantity>
(B.L.Fisher
<emphasis id="DFCBA1420D0FFFE9FD87880366D36636" box="[618,678,546,571]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
); F Tsimembo-19.02139, 44.44067,
<quantity id="2A47D0B50D0FFFE9FBB7880360E76631" box="[1114,1170,546,572]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="15" unit="m" value="20.0">20m</quantity>
, -18.99528, 44.44350,
<quantity id="2A47D0B50D0FFFE9FF7A886964BE666F" box="[151,203,584,610]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="14" pageNumber="15" unit="m" value="50.0">50m</quantity>
(B.L.Fisher
<emphasis id="DFCBA1420D0FFFE9FE8F886965E8666C" box="[354,413,584,609]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
);
<emphasis id="DFCBA1420D0FFFE9FE5A88696666666C" box="[439,531,584,609]" italics="true" pageId="14" pageNumber="15">Toliara:</emphasis>
P.N. Kirindy-Mite: -20.79528, 44.14700,
<quantity id="2A47D0B50D0FFFE9FBEA8869604E666F" box="[1031,1083,584,610]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="14" pageNumber="15" unit="m" value="80.0">80m</quantity>
(B.L.Fisher
<emphasis id="DFCBA1420D0FFFE9FB3F88696178666C" box="[1234,1293,584,609]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
); Kirindy: - 20.04500, 44.66222,
<quantity id="2A47D0B50D0FFFE9FE7D884C65A6668A" box="[400,467,621,647]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="15" unit="m" value="100.0">100m</quantity>
(B.L.Fisher
<emphasis id="DFCBA1420D0FFFE9FD84884C66D6668B" box="[617,675,621,646]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
); P.N. Zombitse: -22.84330, 44.71000,
<quantity id="2A47D0B50D0FFFE9FB9B884C60CC668A" box="[1142,1209,621,647]" metricMagnitude="2" metricUnit="m" metricValue="7.7" pageId="14" pageNumber="15" unit="m" value="770.0">770m</quantity>
(B.L.Fisher
<emphasis id="DFCBA1420D0FFFE9FAA2884C61FF668B" box="[1359,1418,621,646]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
); Isoky-Vohimena: -22.68330, 44.83330,
<quantity id="2A47D0B50D0FFFE9FD9A88B366CE66A1" box="[631,699,658,684]" metricMagnitude="2" metricUnit="m" metricValue="7.3" pageId="14" pageNumber="15" unit="m" value="730.0">730m</quantity>
(B.L.Fisher
<emphasis id="DFCBA1420D0FFFE9FCBB88B367E666A6" box="[854,915,658,683]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
); F Vohibasia: -22.46670, 44.85000,
<quantity id="2A47D0B50D0FFFE9FAB588B361E966A1" box="[1368,1436,658,684]" metricMagnitude="2" metricUnit="m" metricValue="7.8" pageId="14" pageNumber="15" unit="m" value="780.0">780m</quantity>
(B.L.Fisher
<emphasis id="DFCBA1420D0FFFE9FECB8899651466DC" box="[294,353,696,721]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
);
<quantity id="2A47D0B50D0FFFE9FE97889965C866DF" box="[378,445,696,722]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="15" unit="km" value="15.0">15km</quantity>
E Sakaraha: -22.90000, 44.68333,
<quantity id="2A47D0B50D0FFFE9FCB3889967D466DF" box="[862,929,696,722]" metricMagnitude="2" metricUnit="m" metricValue="7.6" pageId="14" pageNumber="15" unit="m" value="760.0">760m</quantity>
(P.S.Ward); Ranobe: -23.03975, 43.61090,
<quantity id="2A47D0B50D0FFFE9FF7A88FC64B966FA" box="[151,204,733,759]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="15" unit="m" value="30.0">30m</quantity>
(Frontier Project,
<collectionCode id="8BAEE5950D0FFFE9FE4A88FC659966FA" box="[423,492,733,759]" httpUri="http://grbio.org/cool/nsep-8f7n" name="Museum George Frey" pageId="14" pageNumber="15">MGF</collectionCode>
); Rés. Berenty (F Bealoka): -24.95694, 46.27150,
<quantity id="2A47D0B50D0FFFE9FBAB88FC600F66FA" box="[1094,1146,733,759]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="14" pageNumber="15" unit="m" value="35.0">35m</quantity>
(B.L.Fisher
<emphasis id="DFCBA1420D0FFFE9FAFF88FC613B66FB" box="[1298,1358,733,758]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
); Rés. Berenty (F Malaza): -25.00778, 46.30600,
<quantity id="2A47D0B50D0FFFE9FD78892366BF6711" box="[661,714,770,796]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="14" pageNumber="15" unit="m" value="40.0">40m</quantity>
(B.L.Fisher
<emphasis id="DFCBA1420D0FFFE9FC8F892367EB6716" box="[866,926,770,795]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
); Rés. Berenty: -25.02100, 46.30550,
<quantity id="2A47D0B50D0FFFE9FA85892361E96711" box="[1384,1436,770,796]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="14" pageNumber="15" unit="m" value="35.0">35m</quantity>
(R.Harin’Hala), -25.01670, 46.30000 (P.S.Ward); F Mahavelo: -24.75830, 46.15717,
<quantity id="2A47D0B50D0FFFE9FB2C8909617C674F" box="[1217,1289,808,834]" metricMagnitude="2" metricUnit="m" metricValue="1.1" pageId="14" pageNumber="15" unit="m" value="110.0">110m</quantity>
, -24.75000, 46.16670,
<quantity id="2A47D0B50D0FFFE9FEFA896C652C676A" box="[279,345,845,871]" metricMagnitude="2" metricUnit="m" metricValue="1.15" pageId="14" pageNumber="15" unit="m" value="115.0">115m</quantity>
(B.L.Fisher
<emphasis id="DFCBA1420D0FFFE9FE18896C6647676B" box="[501,562,845,870]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
); P.N. Andohahela: -24.93000, 46.64550,
<quantity id="2A47D0B50D0FFFE9FBDE896C600D676A" box="[1075,1144,845,871]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="15" unit="m" value="300.0">300m</quantity>
(B.L.Fisher
<emphasis id="DFCBA1420D0FFFE9FAFF896C613A676B" box="[1298,1359,845,870]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
); P.N. Andohahela: -25.01366, 46.64650,
<quantity id="2A47D0B50D0FFFE9FDD58953660E6781" box="[568,635,882,908]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="14" pageNumber="15" unit="m" value="160.0">160m</quantity>
(B.B.Blaimer); F Beroboka: -22.23306, 43.36633,
<quantity id="2A47D0B50D0FFFE9FB34895361786781" box="[1241,1293,882,908]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="14" pageNumber="15" unit="m" value="80.0">80m</quantity>
(B.L.Fisher
<emphasis id="DFCBA1420D0FFFE9FF7A89B964A067BC" box="[151,213,920,945]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
); Mahafaly Plateau: -24.65361, 43.99667,
<quantity id="2A47D0B50D0FFFE9FD0989B9676F67BF" box="[740,794,920,946]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="14" pageNumber="15" unit="m" value="80.0">80m</quantity>
(B.L.Fisher
<emphasis id="DFCBA1420D0FFFE9FC5B89B9678167BC" box="[950,1012,920,945]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
); F Tsimanampetsotsa: -24.10056, 43.76000,
<quantity id="2A47D0B50D0FFFE9FEFB899C653E67DA" box="[278,331,957,983]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="14" pageNumber="15" unit="m" value="25.0">25m</quantity>
(B.L.Fisher
<emphasis id="DFCBA1420D0FFFE9FE09899C665467DB" box="[484,545,957,982]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
); Cap St. Marie: -25.59444, 45.14683,
<quantity id="2A47D0B50D0FFFE9FC11899C604A67DA" box="[1020,1087,957,983]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="14" pageNumber="15" unit="m" value="160.0">160m</quantity>
(B.L.Fisher
<emphasis id="DFCBA1420D0FFFE9FB34899C616367DB" box="[1241,1302,957,982]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
); F Mite: - 23.52417, 44.12133,
<quantity id="2A47D0B50D0FFFE9FE6289C365B667F1" box="[399,451,994,1020]" metricMagnitude="1" metricUnit="m" metricValue="7.5" pageId="14" pageNumber="15" unit="m" value="75.0">75m</quantity>
(B.L.Fisher
<emphasis id="DFCBA1420D0FFFE9FDBA89C366E467F6" box="[599,657,994,1019]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
); Manderano: -23.52417, 44.09278,
<quantity id="2A47D0B50D0FFFE9FBAD89C3600167F1" box="[1088,1140,994,1020]" metricMagnitude="1" metricUnit="m" metricValue="7.5" pageId="14" pageNumber="15" unit="m" value="75.0">75m</quantity>
(Frontier Project,
<collectionCode id="8BAEE5950D0FFFE9FAA789C361FA67F1" box="[1354,1423,994,1020]" httpUri="http://grbio.org/cool/nsep-8f7n" name="Museum George Frey" pageId="14" pageNumber="15">MGF</collectionCode>
); Fiheranana: -23.23528, 43.87083,
<quantity id="2A47D0B50D0FFFE9FDBC8E2966FD602F" box="[593,648,1032,1058]" metricMagnitude="1" metricUnit="m" metricValue="6.5" pageId="14" pageNumber="15" unit="m" value="65.0">65m</quantity>
(B.L.Fisher
<emphasis id="DFCBA1420D0FFFE9FCC38E29671A602C" box="[814,879,1032,1057]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
); F Tsinjoriaky: -22.80222, 43.42067,
<quantity id="2A47D0B50D0FFFE9FA8B8E2961E9602F" box="[1382,1436,1032,1058]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="14" pageNumber="15" unit="m" value="70.0">70m</quantity>
(B.L.Fisher
<emphasis id="DFCBA1420D0FFFE9FEDE8E0C6501604B" box="[307,372,1069,1094]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
); Rés. Beza-Mahafaly: -23.65000, 44.63333,
<quantity id="2A47D0B50D0FFFE9FC2C8E0C607D604A" box="[961,1032,1069,1095]" metricMagnitude="2" metricUnit="m" metricValue="1.35" pageId="14" pageNumber="15" unit="m" value="135.0">135m</quantity>
(B.L.Fisher
<emphasis id="DFCBA1420D0FFFE9FB408E0C609B604B" box="[1197,1262,1069,1094]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
.), -23.68650, 44.59100,
<quantity id="2A47D0B50D0FFFE9FEF48E73652B6061" box="[281,350,1106,1132]" metricMagnitude="2" metricUnit="m" metricValue="1.65" pageId="14" pageNumber="15" unit="m" value="165.0">165m</quantity>
(R. Harin’Hala); Rés. Beza-Mahafaly, parcel 1: -23.65000, 44.63330,
<quantity id="2A47D0B50D0FFFE9FB2E8E73617D6061" box="[1219,1288,1106,1132]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="14" pageNumber="15" unit="m" value="130.0">130m</quantity>
(P.S.Ward); Manombo: -22.81092, 43.73440,
<quantity id="2A47D0B50D0FFFE9FDCA8E59661F609F" box="[551,618,1144,1170]" metricMagnitude="2" metricUnit="m" metricValue="1.65" pageId="14" pageNumber="15" unit="m" value="165.0">165m</quantity>
(Frontier Project,
<collectionCode id="8BAEE5950D0FFFE9FCAA8E5967F9609F" box="[839,908,1144,1170]" httpUri="http://grbio.org/cool/nsep-8f7n" name="Museum George Frey" pageId="14" pageNumber="15">MGF</collectionCode>
); F Vohidava: -24.24070, 46.28780,
<quantity id="2A47D0B50D0FFFE9FAD28E5961F0609F" box="[1343,1413,1144,1170]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="14" pageNumber="15" unit="m" value="500.0">500m</quantity>
, - 24.23900, 46.28230,
<quantity id="2A47D0B50D0FFFE9FE7D8EBC65A760BA" box="[400,466,1181,1207]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="14" pageNumber="15" unit="m" value="850.0">850m</quantity>
(B.L.Fisher
<emphasis id="DFCBA1420D0FFFE9FD858EBC66D660BB" box="[616,675,1181,1206]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
); F Mikea: -22.90367, 43.47550,
<quantity id="2A47D0B50D0FFFE9FBDC8EBC601060BA" box="[1073,1125,1181,1207]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="14" pageNumber="15" unit="m" value="35.0">35m</quantity>
(R. Harin’Hala).
</paragraph>
</subSubSection>
<subSubSection id="A5A52EDB0D0FFFE8FF288EE36033605A" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="description">
<paragraph id="ED007D500D0FFFE9FF288EE365BA6141" blockId="14.[151,1437,285,2028]" pageId="14" pageNumber="15">
<emphasis id="DFCBA1420D0FFFE9FF288EE3663060D1" bold="true" box="[197,581,1218,1244]" pageId="14" pageNumber="15">Worker measurements (n=12).</emphasis>
HW 0.82–0.89; HL 0.72–0.78; EL 0.16–0.21; SL 0.66–0.74; WL 0.79– 1.07; SPL 0.13–0.18; PTH 0.18–0.22; PTL 0.24–0.29; PTW 0.26–0.31; PPL 0.15–0.20; PPW 0.27–0.32; LHT 0.57–0.63; CI 1.11–1.18; OI 0.21–0.27; SI 0.92–0.98; SPI 0.13–0.22; PTHI 0.74–0.79; PTWI 1.01–1.20; PPI 1.43–1.84; LBI 1.29–1.71.
</paragraph>
<paragraph id="ED007D500D0FFFE9FF2B8F766786617F" blockId="14.[151,1437,285,2028]" box="[198,1011,1367,1394]" pageId="14" pageNumber="15">
<emphasis id="DFCBA1420D0FFFE9FF2B8F7665B6617C" bold="true" box="[198,451,1367,1393]" pageId="14" pageNumber="15">Worker description.</emphasis>
Small species (HW 0.82–0.89, WL 0.79–1.07).
</paragraph>
<paragraph id="ED007D500D0FFFE9FF288F5C672A61EF" blockId="14.[151,1437,285,2028]" pageId="14" pageNumber="15">Masticatory margin of mandibles with 4 teeth; posterior margin of head in full-face view with subangular corners, sometimes medially depressed; antennal scapes surpassing posterior margin of head easily; midline of eyes situated well above midline of head in full face view.</paragraph>
<paragraph id="ED007D500D0FFFE9FF288FCC669C6301" blockId="14.[151,1437,285,2028]" pageId="14" pageNumber="15">Promesonotum medially depressed and laterally with raised shoulders, these transitioning into lateral carinae and terminating in small denticles that abruptly demarcate promesonotum from propodeum; in lateral view outline of promesonotum flat; promesonotal suture absent; promesonotum without a distinct posterior face medially; propodeal spines small-medium sized (SPI 0.13–0.22), length roughly a third of the width between their bases, quickly tapering, in lateral view more or less straight, in dorsal view almost parallel and barely diverging; petiole in dorsal view suboval, with dorso-lateral margins carinate, terminating in small postero-lateral tubercules; subpetiolar process developed as broad, rounded protuberance; postpetiole distinctly bilobed and broadly medially impressed.</paragraph>
<paragraph id="ED007D500D0FFFE8FF288D39654064F1" blockId="14.[151,1437,285,2028]" lastBlockId="15.[151,1438,152,2007]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">Head sculpture reduced aciculate to aciculate-reticulate, more pronounced in darker coloured form than in orange-red variety; mesosoma with promesonotum dorsally irregularly carinulate-reticulate; meso- and metapleuron carinulate-areolate; propodeum with dorsal face carinulate and first half of posterior face reticulate, otherwise shiny; legs shiny to reticulate; dorsal face of petiole mostly reticulate, centrally shiny; postpetiole dorsally feebly reticulate; lateral and ventral face of petiole and postpetiole coarsely reticulate; helcium dorsally shiny or reticulate, at most feebly carinulate; promesonotum with 4-8 erect humeral setae, rarely with 2 additional setae on lateral carinae; petiole with single stiff, long erect setae on each posterolateral tubercle; postpetiole with a pair of erect dorso-posterior setae; abdominal tergites 4–7 with scattered erect pilosity.</paragraph>
<paragraph id="ED007D500D0EFFE8FF288B296075659F" blockId="15.[151,1438,152,2007]" pageId="15" pageNumber="16">
Two distinct colour variants: first variant distinctly bi-coloured, with pale to bright orange-red head, mesosoma, legs, petiole and postpetiole that contrast with a brown to almost black gaster (as shown in
<figureCitation id="758461D50D0EFFE8FAAA8B0C61E9654A" box="[1351,1436,301,327]" captionStart="FIGURES 32 – 37" captionStartId="29.[151,270,1974,1998]" captionTargetBox="[190,1397,196,1939]" captionTargetId="figure@29.[179,1407,166,1953]" captionTargetPageId="29" captionText="FIGURES 32 – 37. Malagasy Crematogaster (Decacrema) species; a: full-face view, b: lateral view, c: dorsal view. 32: C. nosibeensis; 33: C. sisa; 34: C. sabatra; 35: C. mahery; 36: C. malala; 37: C. grevei." httpUri="https://zenodo.org/record/199685/files/figure.png" pageId="15" pageNumber="16">
<emphasis id="DFCBA1420D0EFFE8FAAA8B0C61E9654A" bold="true" box="[1351,1436,301,327]" pageId="15" pageNumber="16">Fig. 37</emphasis>
</figureCitation>
and
<figureCitation id="758461D50D0EFFE8FF248B7364936561" box="[201,230,338,364]" captionStart="FIGURES 38 – 43" captionStartId="30.[151,270,1894,1918]" captionTargetBox="[221,1364,193,1874]" captionTargetId="figure@30.[200,1385,164,1882]" captionTargetPageId="30" captionText="FIGURES 38 – 43. Malagasy Crematogaster (Decacrema) species; a: petiole &amp; postpetiole in dorsal view, b: petiole &amp; postpetiole in lateral view. 38: C. nosibeensis; 39: C. sisa, 40: C. sabatra; 41: C. mahery; 42: C. malala; 43: C. grevei." httpUri="https://zenodo.org/record/199686/files/figure.png" pageId="15" pageNumber="16">
<emphasis id="DFCBA1420D0EFFE8FF248B7364936561" bold="true" box="[201,230,338,364]" pageId="15" pageNumber="16">43</emphasis>
</figureCitation>
); second variant (to which the
<typeStatus id="3204C3F20D0EFFE8FDB78B7366B36561" box="[602,710,338,364]" pageId="15" pageNumber="16" type="lectotype">lectotype</typeStatus>
belongs) uni-coloured, from reddish brown to brown to dark brown, to entirely black (specimen image on Antweb: CASENT0454184).
</paragraph>
<paragraph id="ED007D500D0EFFE8FF2B8BBC6021662A" blockId="15.[151,1438,152,2007]" pageId="15" pageNumber="16">
<emphasis id="DFCBA1420D0EFFE8FF2B8BBC665065BA" bold="true" box="[198,549,413,439]" pageId="15" pageNumber="16">Queen measurements (n=3).</emphasis>
HW 1.26–1.33, HL 1.06–1.15, EL 0.36–0.39, SL 0.87–0.93, MSNW 0.92– 1.05, MSNL 1.71–1.93, PTH 0.31–0.36, PTL 0.41–0.42, PTW 0.48–0.53, PPL 0.30–0.34, PPW 0.46–0.53, SPL 0.03–0.08, WL 2.06–2.23, LHT 0.88–0.96, CI 1.11–1.26, OI 0.32–0.37, SI 0.76–0.83, MSNI 0.51–0.55, PTHI 0.74–0.87, PTWI 1.15–1.28, PPI 1.54–1.58, SPI 0.01–0.04, LBI 2.33–2.38.
</paragraph>
<paragraph id="ED007D500D0EFFE8FF2B88136612669A" blockId="15.[151,1438,152,2007]" pageId="15" pageNumber="16">
<emphasis id="DFCBA1420D0EFFE8FF2B881365C76641" bold="true" box="[198,434,562,588]" pageId="15" pageNumber="16">Queen description.</emphasis>
Small (HW 1.26–1.33, WL 2.06–2.23). With worker characters, except as follows. Antennal scapes just barely surpassing posterior margin of head; eyes situated at midline of head in full face view; posterior margin of head straight.
</paragraph>
<paragraph id="ED007D500D0EFFE8FF28888361136791" blockId="15.[151,1438,152,2007]" pageId="15" pageNumber="16">Mesosoma slender (MSNI 0.51–0.55, WL 2.06–2.23); mesoscutum in dorsal view oval, about half as wide as long; mesopleuron with episternal groove carinulate to broadly carinate; in lateral view mesepisternum meeting pronotum at an oblique angle; postscutellum with posterior face rounded to flattened; propodeal suture deep and laterally reaching until level of propodeal spiracle; dorsal face of propodeum short, about one third of the size of posterior face; propodeal spines reduced, ranging from tubercule to denticles to very sharp points; petiole and postpetiole as in worker, but less prominently shaped and dorso-posterior tubercule on petiole lacking; antero-ventral subpetiolar tooth present, but reduced with respect to worker.</paragraph>
<paragraph id="ED007D500D0EFFE8FF288989656A603F" blockId="15.[151,1438,152,2007]" pageId="15" pageNumber="16">Propodeum with vertical carinulae on dorsal and posterior face; petiole reticulate throughout; postpetiole feebly reticulate throughout; face with 10–12 erect setae and abundant appressed pilosity; mesonotum with 20–26 erect setae; petiole and postpetiole with dorso-posterior setae and further with appressed pilosity throughout.</paragraph>
<paragraph id="ED007D500D0EFFE8FF288E1C6033605A" blockId="15.[151,1438,152,2007]" box="[197,1094,1085,1111]" pageId="15" pageNumber="16">Colour brown to dark brown, or orange-red with gaster dark brown or black.</paragraph>
</subSubSection>
<subSubSection id="A5A52EDB0D0EFFE8FF2B8E43601A61C1" pageId="15" pageNumber="16" type="discussion">
<paragraph id="ED007D500D0EFFE8FF2B8E4360466151" blockId="15.[151,1438,152,2007]" pageId="15" pageNumber="16">
<emphasis id="DFCBA1420D0EFFE8FF2B8E4365206071" bold="true" box="[198,341,1122,1148]" pageId="15" pageNumber="16">Comments.</emphasis>
This species is recognizable by a combination of the following characters. The lateral portions of the promesonotum are raised with respect to the median portion and end in postero-lateral carinate denticles. Lateral carinae border the metanotal groove. The petiole has postero-lateral tubercles dorsally that each bear a single, stiff and long erect seta; the postpetiole is distinctly bilobed and broadly medially impressed, and also bears a pair of dorso-posterior stiff, long erect setae. Further noteworthy are the short and straight spines, the erect pilosity on abdominal tergites 4–7, and the mandibles that possess only 4 teeth.
<taxonomicName id="2ABF06D30D0EFFE8FA6C8F3C64AA6151" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="grevei">
<emphasis id="DFCBA1420D0EFFE8FA6C8F3C64AA6151" italics="true" pageId="15" pageNumber="16">C. grevei</emphasis>
</taxonomicName>
is strongly supported by both nuclear and mitochondrial molecular data.
</paragraph>
<paragraph id="ED007D500D0EFFE8FF7A8F49601A61C1" blockId="15.[151,1438,152,2007]" pageId="15" pageNumber="16">
Both
<typeStatus id="3204C3F20D0EFFE8FF378F49654D618F" box="[218,312,1384,1410]" pageId="15" pageNumber="16" type="syntype">syntype</typeStatus>
specimens in the MHNG correspond well with the brown and black coloured variants in the CASC material, and with the
<typeStatus id="3204C3F20D0EFFE8FE1E8FAC665061AA" box="[499,549,1421,1447]" pageId="15" pageNumber="16">type</typeStatus>
description. One of the MHNG
<typeStatus id="3204C3F20D0EFFE8FC4F8FAC607C61AA" box="[930,1033,1421,1447]" pageId="15" pageNumber="16" type="syntype">syntypes</typeStatus>
carries matching locality data and collector information with the
<typeStatus id="3204C3F20D0EFFE8FE108F93664561C1" box="[509,560,1458,1484]" pageId="15" pageNumber="16">type</typeStatus>
description and is hence chosen as the
<typeStatus id="3204C3F20D0EFFE8FC118F93601961C1" box="[1020,1132,1458,1484]" pageId="15" pageNumber="16" type="lectotype">lectotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="A5A52EDB0D0EFFE8FF2B8FF66488626F" pageId="15" pageNumber="16" type="description">
<paragraph id="ED007D500D0EFFE8FF2B8FF66488626F" blockId="15.[151,1438,152,2007]" pageId="15" pageNumber="16">
<emphasis id="DFCBA1420D0EFFE8FF2B8FF6653461FC" bold="true" box="[198,321,1495,1521]" pageId="15" pageNumber="16">Variation.</emphasis>
Apart from colouration, the two variants of
<taxonomicName id="2ABF06D30D0EFFE8FCA58FF967D961FC" box="[840,940,1496,1521]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="grevei">
<emphasis id="DFCBA1420D0EFFE8FCA58FF967D961FC" box="[840,940,1496,1521]" italics="true" pageId="15" pageNumber="16">C.grevei</emphasis>
</taxonomicName>
exhibit slight variations in head sculpture. Some size polymorphism is exhibited in the workers of this species. Larger workers that have a disproportionately larger mesosoma than head are often found among colony samples of normal-sized workers.
</paragraph>
</subSubSection>
<subSubSection id="A5A52EDB0D0EFFF7FF2B8C4C60C7659F" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="distribution">
<paragraph id="ED007D500D0EFFF7FF2B8C4C665864DA" blockId="15.[151,1438,152,2007]" lastBlockId="16.[151,1436,152,402]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
<emphasis id="DFCBA1420D0EFFE8FF2B8C4C6674628A" bold="true" box="[198,513,1645,1671]" pageId="15" pageNumber="16">Distribution and biology.</emphasis>
<taxonomicName id="2ABF06D30D0EFFE8FDE78C4C677F628B" box="[522,778,1645,1670]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="grevei">
<emphasis id="DFCBA1420D0EFFE8FDE78C4C677F628B" box="[522,778,1645,1670]" italics="true" pageId="15" pageNumber="16">Crematogaster grevei</emphasis>
</taxonomicName>
occurs throughout the seasonally dry forests and spiny forests of western and southwestern
<collectingCountry id="95A83DC00D0EFFE8FDBC8CB3669762A1" box="[593,738,1682,1708]" name="Madagascar" pageId="15" pageNumber="16">Madagascar</collectingCountry>
(
<figureCitation id="758461D50D0EFFE8FD198CB3673962A1" box="[756,844,1682,1708]" captionStart="FIGURES 54 – 57" captionStartId="33.[151,270,1948,1972]" captionTargetBox="[294,1298,205,1923]" captionTargetId="figure@33.[285,1302,194,1927]" captionTargetPageId="33" captionText="FIGURES 54 – 57. Distributions of Malagasy Crematogaster (Decacrema). 54: C. grevei; 55: C. mahery; 56: C. malala; 57: C. nosibeensis." httpUri="https://zenodo.org/record/199689/files/figure.png" pageId="15" pageNumber="16">
<emphasis id="DFCBA1420D0EFFE8FD198CB3673962A1" bold="true" box="[756,844,1682,1708]" pageId="15" pageNumber="16">Fig. 54</emphasis>
</figureCitation>
). Among members of the
<collectingCountry id="95A83DC00D0EFFE8FB618CB3617562A1" box="[1164,1280,1682,1708]" name="Madagascar" pageId="15" pageNumber="16">Malagasy</collectingCountry>
<taxonomicName id="2ABF06D30D0EFFE8FAE48CB261E762A1" box="[1289,1426,1683,1708]" class="Insecta" family="Formicidae" genus="Decacrema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="DFCBA1420D0EFFE8FAE48CB261E762A1" box="[1289,1426,1683,1708]" italics="true" pageId="15" pageNumber="16">Decacrema</emphasis>
</taxonomicName>
- group, these habitats are almost exclusively inhabited by
<taxonomicName id="2ABF06D30D0EFFE8FCBA8C9967B362DC" box="[855,966,1720,1745]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="grevei">
<emphasis id="DFCBA1420D0EFFE8FCBA8C9967B362DC" box="[855,966,1720,1745]" italics="true" pageId="15" pageNumber="16">C. grevei</emphasis>
</taxonomicName>
, its range is parapatric or allopatric in regard to most other species in the group. Narrow sympatry exists with the
<taxonomicName id="2ABF06D30D0EFFE8FBD58CFC60E262FB" box="[1080,1175,1757,1782]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="hova">
<emphasis id="DFCBA1420D0EFFE8FBD58CFC60E262FB" box="[1080,1175,1757,1782]" italics="true" pageId="15" pageNumber="16">C. hova</emphasis>
</taxonomicName>
-complex in the Isalo Range, where isolated pockets of dry forest occur at higher elevations, in remnants of western subhumid forest in the P.N. Zombitse, and in the “tsingy” of P.N. Namoroka. In the Andohahela Range
<taxonomicName id="2ABF06D30D0EFFE8FB308D09613D634C" box="[1245,1352,1832,1857]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="grevei">
<emphasis id="DFCBA1420D0EFFE8FB308D09613D634C" box="[1245,1352,1832,1857]" italics="true" pageId="15" pageNumber="16">C. grevei</emphasis>
</taxonomicName>
occurs at low elevation in drier habitats, and the
<emphasis id="DFCBA1420D0EFFE8FD938D6C6696636B" box="[638,739,1869,1894]" italics="true" pageId="15" pageNumber="16">C. hova-</emphasis>
complex and
<taxonomicName id="2ABF06D30D0EFFE8FC6F8D6C6789636B" box="[898,1020,1869,1894]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="mahery">
<emphasis id="DFCBA1420D0EFFE8FC6F8D6C6789636B" box="[898,1020,1869,1894]" italics="true" pageId="15" pageNumber="16">C. mahery</emphasis>
</taxonomicName>
take over at higher elevation in the humid forest. Fisher
<emphasis id="DFCBA1420D0EFFE8FE618D5265BD6381" box="[396,456,1907,1932]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
frequently collected this species nesting in dead twigs on and above the ground, in rotten logs, from leaf litter samples, and also found it occasionally nesting under stones or in stems of live plants. Carton nesting has not been recorded for this species. The absence of this behaviour is probably related to the drier habitat of this species, as rotting plant material here may simply lose moisture too quickly to be implemented in carton fabrication.
</paragraph>
<paragraph id="ED007D500D11FFF7FF288AC360C7659F" blockId="16.[151,1436,152,402]" pageId="16" pageNumber="17">
The two colour variants of
<taxonomicName id="2ABF06D30D11FFF7FDEA8AC3661E64F6" box="[519,619,226,251]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="grevei">
<emphasis id="DFCBA1420D11FFF7FDEA8AC3661E64F6" box="[519,619,226,251]" italics="true" pageId="16" pageNumber="17">C.grevei</emphasis>
</taxonomicName>
have nearly allopatric distributions. The nature of these distributions suggests that the orange variant is associated with the spiny forests of the far south and southwest parts of
<collectingCountry id="95A83DC00D11FFF7FF7A8B0C6552654A" box="[151,295,301,327]" name="Madagascar" pageId="16" pageNumber="17">Madagascar</collectingCountry>
, while the brown or black variant is adapted to western dry and western subhumid forest habitats. Very likely some environmental variables play a role in stabilizing this colour dimorphism, at the moment however too little is known about the ecology of the species to make further assumptions.
</paragraph>
</subSubSection>
</treatment>
</document>
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