451 lines
56 KiB
XML
451 lines
56 KiB
XML
<document id="B11F2493BFCCF0760493E2AF5C0C0FFD" ID-DOI="10.5281/zenodo.179242" ID-GBIF-Dataset="9fb430d0-cd77-48f4-b277-8a7c3b316485" ID-ISSN="1175-5326" ID-Zenodo-Dep="179242" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459946210070" checkinUser="plazi" docAuthor="Ojanguren-Affilastro, Andrés A., Agusto, Pablo, Pizarro-Araya, Jaime & Mattoni, Camilo I." docDate="2007" docId="566287FEA0172B70048973A3FE8A7E54" docLanguage="en" docName="zt01623p068.pdf" docOrigin="Zootaxa 1623" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Brachistosternus (Leptosternus) coquimbo Ojanguren-Affilastro, Agusto, Pizarro-Araya & Mattoni, 2007, n. sp." docType="treatment" docVersion="10" lastPageNumber="65" masterDocId="AA5BFF86A0112B7A041E776CFFDB7F7E" masterDocTitle="Two new scorpion species of genus Brachistosternus (Scorpiones: Bothriuridae) from northern Chile" masterLastPageNumber="68" masterPageNumber="55" pageNumber="61" updateTime="1698230676319" updateUser="plazi">
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<mods:titleInfo id="4516AD981AA94B4149E815DAB51C23CA">
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<mods:title id="AEC956E2E04783765E5C7C2E84463D5D">Two new scorpion species of genus Brachistosternus (Scorpiones: Bothriuridae) from northern Chile</mods:title>
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</mods:titleInfo>
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<mods:name id="EA47A44196E03720B6E4FBB91A5B4383" type="personal">
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<mods:roleTerm id="02526D26CA445C8FD1A327BE75B75615">Author</mods:roleTerm>
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<mods:namePart id="36F9CDE7610AEC3DCAD741BC64A370BD">Ojanguren-Affilastro, Andrés A.</mods:namePart>
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<mods:roleTerm id="7551D39D633AB6210C186B513EFBE7CD">Author</mods:roleTerm>
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<mods:namePart id="5C876E9506C26B7D4DF7FCF506F01E77">Agusto, Pablo</mods:namePart>
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<mods:roleTerm id="153F94EB18ACC3783743B3B85DAAD69D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0FD5EEF6BDC6BC600FA33FC0B63B70A1">Pizarro-Araya, Jaime</mods:namePart>
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<mods:name id="54BCD90AD2399CDFC27A582D4CA60912" type="personal">
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<mods:roleTerm id="BB2539AC59C967FD2C29A88792A32A1B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7311A94D484CAF9B23225C488625A25F">Mattoni, Camilo I.</mods:namePart>
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<mods:typeOfResource id="E0DE8A7D99B5DBDC7752BDD718BF26B7">text</mods:typeOfResource>
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<mods:title id="970DE0E737D29991F794E4EF38F0EE28">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="1CA0D7F5C62217D009CCA37880C883B7">
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<mods:date id="F1243C759EBFAFF84C0F4B4B75AA7FF1">2007</mods:date>
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<mods:number id="AD1AD7F1E38B79728D6E0D41F289C50A">1623</mods:number>
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<mods:classification id="2DE13645DBC5595EF0F36BE3388ECEE7">journal article</mods:classification>
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<mods:identifier id="06DC63E2912278FDD2839C85E685B559" type="DOI">10.5281/zenodo.179242</mods:identifier>
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<mods:identifier id="F9D5CAD07D632086C87785D81EA0559E" type="ISSN">1175-5326</mods:identifier>
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<treatment id="566287FEA0172B70048973A3FE8A7E54" ID-DOI="http://doi.org/10.5281/zenodo.5670652" ID-GBIF-Taxon="119376228" ID-Zenodo-Dep="5670652" LSID="urn:lsid:plazi:treatment:566287FEA0172B70048973A3FE8A7E54" httpUri="http://treatment.plazi.org/id/566287FEA0172B70048973A3FE8A7E54" lastPageId="10" lastPageNumber="65" pageId="6" pageNumber="61">
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<subSubSection id="96D16563A0172B7C048973A3FE7F7A72" pageId="6" pageNumber="61" type="nomenclature">
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<paragraph id="DE7436E8A0172B7C048973A3FD387B97" blockId="6.[151,739,1231,1292]" box="[151,739,1231,1257]" pageId="6" pageNumber="61">
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<heading id="853C8184A0172B7C048973A3FD387B97" bold="true" box="[151,739,1231,1257]" fontSize="11" level="1" pageId="6" pageNumber="61" reason="1">
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<emphasis id="ECBFEAFAA0172B7C048973A3FD387B97" bold="true" box="[151,739,1231,1257]" pageId="6" pageNumber="61">
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||
<taxonomicName id="19CB4D6BA0172B7C048973A3FD427B97" ID-CoL="MTVM" box="[151,665,1231,1257]" class="Arachnida" family="Bothriuridae" genus="Brachistosternus" higherTaxonomySource="CoL" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="coquimbo" status="sp. nov." subGenus="Leptosternus">
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<emphasis id="ECBFEAFAA0172B7C048973A3FEBF7B97" bold="true" box="[151,356,1231,1257]" italics="true" pageId="6" pageNumber="61">Brachistosternus</emphasis>
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(
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<emphasis id="ECBFEAFAA0172B7C056B73A3FDCA7B97" bold="true" box="[373,529,1231,1257]" italics="true" pageId="6" pageNumber="61">Leptosternus</emphasis>
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)
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<emphasis id="ECBFEAFAA0172B7C063C73A3FD427B97" bold="true" box="[546,665,1231,1257]" italics="true" pageId="6" pageNumber="61">coquimbo</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="F78C5781A0172B7C06BF73A3FD387B97" box="[673,739,1231,1257]" pageId="6" pageNumber="61" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="DE7436E8A0172B7C0489739EFE7F7A72" blockId="6.[151,739,1231,1292]" box="[151,420,1266,1292]" pageId="6" pageNumber="61">
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<figureCitation id="46F02A6DA0172B7C0489739EFF3C7A72" box="[151,231,1266,1292]" captionStart="FIGURE 1" captionStartId="2.[151,255,1313,1337]" captionTargetBox="[275,1311,178,1278]" captionTargetId="figure@2.[225,1362,137,1307]" captionTargetPageId="2" captionText="FIGURE 1. Map of Coquimbo region, Chile, with localities of Brachistosternus (L.) cepedai n. sp. and Brachistosternus (L.) coquimbo n. sp." httpUri="https://zenodo.org/record/179243/files/figure.png" pageId="6" pageNumber="61">Figs. 1</figureCitation>
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,
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<figureCitation id="46F02A6DA0172B7C04EB739EFE9A7A72" box="[245,321,1266,1292]" captionStart-0="FIGURES 11 – 18" captionStart-1="FIGURES 19 – 22" captionStartId-0="7.[151,269,1715,1739]" captionStartId-1="9.[151,269,1454,1478]" captionTargetBox-0="[275,1322,207,1619]" captionTargetBox-1="[287,1307,651,1425]" captionTargetId-0="figure@7.[264,1323,194,1691]" captionTargetId-1="figure@9.[264,1322,634,1449]" captionTargetPageId-0="7" captionTargetPageId-1="9" captionText-0="FIGURES 11 – 18. Brachistosternus (L.) coquimbo n. sp. 11. left hemispermatophore, ventral aspect; 12. left hemispermatophore, dorsal aspect; 13. left hemispermatophore, detail of lobe region; 14. metasomal segment V, male, dorsal aspect; 15. metasomal segment V, male, ventral aspect; 16. telson, male, dorsal aspect; 17. telson, male, lateral aspect; 18. telson, female, lateral aspect. Scale bars = 1 mm." captionText-1="FIGURES 19 – 22. Brachistosternus (L.) coquimbo n. sp. 19. Left pedipalp chela, male, ventral aspect; 20. left pedipalp chela, male, internal aspect; 21. right pedipalp chela, male, external aspect; 22. right pedipalp chela, female, external aspect. Scale bars = 1 mm." httpUri-0="https://zenodo.org/record/179245/files/figure.png" httpUri-1="https://zenodo.org/record/179246/files/figure.png" pageId="6" pageNumber="61">11–22</figureCitation>
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;
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<tableCitation id="93490353A0172B7C0553739EFE7F7A72" box="[333,420,1266,1292]" captionStart="TABLE 1" captionStartId="3.[151,239,546,570]" captionTargetBox="[151,1436,754,1976]" captionTargetPageId="3" captionText="TABLE 1. Measurements of Brachistosternus (Leptosternus) cepedai n. sp. (Paratypes) and Brachistosternus (Leptosternus) coquimbo n. sp. (Holotype and Paratype). (In mm.)." httpUri="http://table.plazi.org/id/8AB46660A0122B790489754EFCEF7D1E" pageId="6" pageNumber="61" tableUuid="8AB46660A0122B790489754EFCEF7D1E">Table 1</tableCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="96D16563A0172B7C0489722EFDD67A82" pageId="6" pageNumber="61" type="materials_examined">
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<paragraph id="DE7436E8A0172B7C0489722EFDD67A82" blockId="6.[151,1436,1346,2012]" pageId="6" pageNumber="61">
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<emphasis id="ECBFEAFAA0172B7C0489722EFE8F7A22" bold="true" box="[151,340,1346,1372]" pageId="6" pageNumber="61">
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<typeStatus id="0170884AA0172B7C0489722EFF097A22" box="[151,210,1346,1372]" pageId="6" pageNumber="61">Type</typeStatus>
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material:
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</emphasis>
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<typeStatus id="0170884AA0172B7C057A722EFE0A7A22" box="[356,465,1346,1372]" pageId="6" pageNumber="61" type="holotype">Holotype</typeStatus>
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ɗ:
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<collectingCountry id="A6DC7678A0172B7C0614722EFD977A22" box="[522,588,1346,1372]" name="Chile" pageId="6" pageNumber="61">Chile</collectingCountry>
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, IV Región,
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<taxonomicName id="19CB4D6BA0172B7C06E8722EFCAE7A22" box="[758,885,1346,1372]" class="Arachnida" family="Bothriuridae" genus="Brachistosternus" higherTaxonomySource="CoL" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="65" phylum="Arthropoda" rank="species" species="coquimbo" status="sp. nov.">Coquimbo</taxonomicName>
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, between Juntas and Paso del Agua Negra,
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<geoCoordinate id="BBFF502FA0172B7C04897206FE9B7AFA" box="[151,320,1386,1412]" direction="south" orientation="latitude" pageId="6" pageNumber="61" precision="1" value="-30.270695">30°16'14.5'' S</geoCoordinate>
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,
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<geoCoordinate id="BBFF502FA0172B7C054D7206FDDD7AFA" box="[339,518,1386,1412]" direction="west" orientation="longitude" pageId="6" pageNumber="61" precision="1" value="-69.97442">69°58'27.9'' W</geoCoordinate>
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,
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<quantity id="19339B0DA0172B7C06097206FDAD7AFA" box="[535,630,1386,1412]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="61" unit="m" value="3000.0">3000 m</quantity>
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a.s.l,
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<date id="AA751028A0172B7C06D97206FCE37AFA" box="[711,824,1386,1412]" pageId="6" pageNumber="61" value="2006-02-06">6/II/2006</date>
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; A.A. Ojanguren, L. Compagnucci & C. Cuezzo (MACN-Ar).
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<typeStatus id="0170884AA0172B7C052772FEFE6B7AD2" box="[313,432,1426,1452]" pageId="6" pageNumber="61" type="paratype">Paratypes</typeStatus>
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: 4 ɗ,
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<specimenCount id="C8CDFD61A0172B7C05EC72FEFDB17AD2" box="[498,618,1426,1452]" pageId="6" pageNumber="61" type="juvenile">1 juvenile</specimenCount>
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; same data; 1 ɗ, Cancha Sky, El Indio mine,
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<geoCoordinate id="BBFF502FA0172B7C006472FEFACA7AD2" box="[1146,1297,1426,1452]" direction="south" orientation="latitude" pageId="6" pageNumber="61" precision="15" value="-29.85">29º51´00´´ S</geoCoordinate>
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,
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<geoCoordinate id="BBFF502FA0172B7C010072FEFF687AAA" direction="west" orientation="longitude" pageId="6" pageNumber="61" precision="15" value="-70.05">70º03´00´´ W</geoCoordinate>
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,
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<quantity id="19339B0DA0172B7C04DF72D6FEC67AAA" box="[193,285,1466,1492]" metricMagnitude="3" metricUnit="m" metricValue="3.3" pageId="6" pageNumber="61" unit="m" value="3300.0">3300 m</quantity>
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a.s.l;
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<date id="AA751028A0172B7C057772D6FE3C7AAA" box="[361,487,1466,1492]" pageId="6" pageNumber="61" value="1992-02-15">15/II/1992</date>
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, H. Vásquez; 1 Ψ, Sancarrón, El Indio mine,
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<geoCoordinate id="BBFF502FA0172B7C000A72D6FB757AAA" box="[1044,1198,1466,1492]" direction="south" orientation="latitude" pageId="6" pageNumber="61" precision="15" value="-29.55">29º33´00´´ S</geoCoordinate>
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,
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<geoCoordinate id="BBFF502FA0172B7C00A072D6FA807AAA" box="[1214,1371,1466,1492]" direction="west" orientation="longitude" pageId="6" pageNumber="61" precision="15" value="-70.23333">70º14´00´´W</geoCoordinate>
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;
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<date id="AA751028A0172B7C017772D6FF0E7A82" pageId="6" pageNumber="61" value="1993-02-05">5/II/ 1993</date>
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; H. Vásquez (MACN-Ar).
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</paragraph>
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</subSubSection>
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<subSubSection id="96D16563A0172B7C04D87166FACF795A" box="[198,1300,1546,1572]" pageId="6" pageNumber="61" type="etymology">
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<paragraph id="DE7436E8A0172B7C04D87166FACF795A" blockId="6.[151,1436,1346,2012]" box="[198,1300,1546,1572]" pageId="6" pageNumber="61">
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<emphasis id="ECBFEAFAA0172B7C04D87166FE8C795A" bold="true" box="[198,343,1546,1572]" pageId="6" pageNumber="61">Etymology:</emphasis>
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The name of this species refers to the Chilean region where it has been collected.
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</paragraph>
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</subSubSection>
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<subSubSection id="96D16563A0172B7C04D8715EFBE778A2" pageId="6" pageNumber="61" type="diagnosis">
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<paragraph id="DE7436E8A0172B7C04D8715EFBE778A2" blockId="6.[151,1436,1346,2012]" pageId="6" pageNumber="61">
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<emphasis id="ECBFEAFAA0172B7C04D8715EFE937932" bold="true" box="[198,328,1586,1612]" pageId="6" pageNumber="61">Diagnosis:</emphasis>
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<taxonomicName id="19CB4D6BA0172B7C054F715FFD1C7932" box="[337,711,1586,1612]" class="Arachnida" family="Bothriuridae" genus="Brachistosternus" higherTaxonomySource="CoL" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="coquimbo" status="sp. nov." subGenus="Leptosternus">
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<emphasis id="ECBFEAFAA0172B7C054F715FFDCC7932" box="[337,535,1587,1612]" italics="true" pageId="6" pageNumber="61">Brachistosternus</emphasis>
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(
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<emphasis id="ECBFEAFAA0172B7C0634715FFDE17932" box="[554,570,1587,1612]" italics="true" pageId="6" pageNumber="61">L</emphasis>
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.)
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<emphasis id="ECBFEAFAA0172B7C064A715FFD1C7932" box="[596,711,1587,1612]" italics="true" pageId="6" pageNumber="61">coquimbo</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="F78C5781A0172B7C06D1715EFCC87932" box="[719,787,1586,1612]" pageId="6" pageNumber="61" rank="species">
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||
<emphasis id="ECBFEAFAA0172B7C06D1715EFCC87932" bold="true" box="[719,787,1586,1612]" pageId="6" pageNumber="61">n. sp.</emphasis>
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</taxonomicNameLabel>
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is closely related with
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<taxonomicName id="19CB4D6BA0172B7C0039715FFEAF790A" authority="Roig Alsina, 1977" authorityName="Roig Alsina" authorityYear="1977" class="Arachnida" family="Bothriuridae" genus="Brachistosternus" higherTaxonomySource="CoL" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="montanus" subGenus="Leptosternus">
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<emphasis id="ECBFEAFAA0172B7C0039715FFB367932" box="[1063,1261,1587,1612]" italics="true" pageId="6" pageNumber="61">Brachistosternus</emphasis>
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(
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||
<emphasis id="ECBFEAFAA0172B7C011E715FFACB7932" box="[1280,1296,1587,1612]" italics="true" pageId="6" pageNumber="61">L</emphasis>
|
||
.)
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<emphasis id="ECBFEAFAA0172B7C0134715FFA477932" box="[1322,1436,1587,1612]" italics="true" pageId="6" pageNumber="61">montanus</emphasis>
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Roig Alsina, 1977
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</taxonomicName>
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. The two species can be distinguished by the following characters: (1)
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<taxonomicName id="19CB4D6BA0172B7C00D67137FA47790A" box="[1224,1436,1626,1652]" class="Arachnida" family="Bothriuridae" genus="Brachistosternus" higherTaxonomySource="CoL" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="coquimbo" subGenus="Leptosternus">
|
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<emphasis id="ECBFEAFAA0172B7C00D67137FB3F790A" box="[1224,1252,1627,1652]" italics="true" pageId="6" pageNumber="61">Br</emphasis>
|
||
. (
|
||
<emphasis id="ECBFEAFAA0172B7C00E07137FAD5790A" box="[1278,1294,1627,1652]" italics="true" pageId="6" pageNumber="61">L</emphasis>
|
||
.)
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<emphasis id="ECBFEAFAA0172B7C01367137FA47790A" box="[1320,1436,1627,1652]" italics="true" pageId="6" pageNumber="61">coquimbo</emphasis>
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</taxonomicName>
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has a narrower pedipalp chela. In males the chela length/width ratio varies from 4.7 to 5.09 (N = 6; mean = 4.85), and in the only studied female it is 4.33, whereas in the males of
|
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<taxonomicName id="19CB4D6BA0172B7C07CD71C7FB7A79BA" box="[979,1185,1706,1732]" class="Arachnida" family="Bothriuridae" genus="Brachistosternus" higherTaxonomySource="CoL" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="montanus" subGenus="Leptosternus">
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<emphasis id="ECBFEAFAA0172B7C07CD71C7FC3479BA" box="[979,1007,1707,1732]" italics="true" pageId="6" pageNumber="61">Br</emphasis>
|
||
. (
|
||
<emphasis id="ECBFEAFAA0172B7C001971C7FBCC79BA" box="[1031,1047,1707,1732]" italics="true" pageId="6" pageNumber="61">L</emphasis>
|
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.)
|
||
<emphasis id="ECBFEAFAA0172B7C003171C7FB7A79BA" box="[1071,1185,1707,1732]" italics="true" pageId="6" pageNumber="61">montanus</emphasis>
|
||
</taxonomicName>
|
||
it varies from 4.01 to 4.37 (N = 15; mean = 4.25) and in females it varies from 3.9 to 4.16 (N = 10; mean = 4.07); (2) in
|
||
<taxonomicName id="19CB4D6BA0172B7C015A71BFFED0786A" class="Arachnida" family="Bothriuridae" genus="Brachistosternus" higherTaxonomySource="CoL" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="coquimbo" subGenus="Leptosternus">
|
||
<emphasis id="ECBFEAFAA0172B7C015A71BFFABB7992" box="[1348,1376,1747,1772]" italics="true" pageId="6" pageNumber="61">Br</emphasis>
|
||
. (
|
||
<emphasis id="ECBFEAFAA0172B7C016571BFFA507992" box="[1403,1419,1747,1772]" italics="true" pageId="6" pageNumber="61">L</emphasis>
|
||
.)
|
||
<emphasis id="ECBFEAFAA0172B7C04897197FED0786A" box="[151,267,1787,1812]" italics="true" pageId="6" pageNumber="61">coquimbo</emphasis>
|
||
</taxonomicName>
|
||
males the chela length/height ratio varies from 3.63 to 3.93 (N = 6; mean = 3.81) and is
|
||
<quantity id="19339B0DA0172B7C01327196FAAB786A" box="[1324,1392,1786,1812]" metricMagnitude="-2" metricUnit="m" metricValue="8.89" pageId="6" pageNumber="61" unit="in" value="3.5">3.5 in</quantity>
|
||
the only studied female; whereas in
|
||
<taxonomicName id="19CB4D6BA0172B7C0605704FFD367842" box="[539,749,1826,1852]" class="Arachnida" family="Bothriuridae" genus="Brachistosternus" higherTaxonomySource="CoL" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="montanus" subGenus="Leptosternus">
|
||
<emphasis id="ECBFEAFAA0172B7C0605704FFDEC7842" box="[539,567,1827,1852]" italics="true" pageId="6" pageNumber="61">Br</emphasis>
|
||
. (
|
||
<emphasis id="ECBFEAFAA0172B7C064F704FFDBA7842" box="[593,609,1827,1852]" italics="true" pageId="6" pageNumber="61">L</emphasis>
|
||
.)
|
||
<emphasis id="ECBFEAFAA0172B7C0665704FFD367842" box="[635,749,1827,1852]" italics="true" pageId="6" pageNumber="61">montanus</emphasis>
|
||
</taxonomicName>
|
||
males it varies from 3.01 to 3.31 (N = 15; mean = 3.17), and in females it varies from 3.06 to 3.23 (N = 10; mean = 3.14).
|
||
<taxonomicName id="19CB4D6BA0172B7C07807027FAC8781A" box="[926,1299,1866,1892]" class="Arachnida" family="Bothriuridae" genus="Brachistosternus" higherTaxonomySource="CoL" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="coquimbo" status="sp. nov." subGenus="Leptosternus">
|
||
<emphasis id="ECBFEAFAA0172B7C07807027FBBF781A" box="[926,1124,1867,1892]" italics="true" pageId="6" pageNumber="61">Brachistosternus</emphasis>
|
||
(
|
||
<emphasis id="ECBFEAFAA0172B7C00687027FB5D781A" box="[1142,1158,1867,1892]" italics="true" pageId="6" pageNumber="61">L</emphasis>
|
||
.)
|
||
<emphasis id="ECBFEAFAA0172B7C00817027FAC8781A" box="[1183,1299,1867,1892]" italics="true" pageId="6" pageNumber="61">coquimbo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F78C5781A0172B7C01047026FA86781A" box="[1306,1373,1866,1892]" pageId="6" pageNumber="61" rank="species">
|
||
<emphasis id="ECBFEAFAA0172B7C01047026FA86781A" bold="true" box="[1306,1373,1866,1892]" pageId="6" pageNumber="61">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
lives in sympatry with
|
||
<taxonomicName id="19CB4D6BA0172B7C0570701FFB4778F2" authority="Ojanguren-Affilastro & Mattoni 2006" authorityName="Ojanguren-Affilastro & Mattoni" authorityYear="2006" box="[366,1180,1906,1932]" class="Arachnida" family="Bothriuridae" genus="Brachistosternus" higherTaxonomySource="CoL" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="perettii" subGenus="Leptosternus">
|
||
<emphasis id="ECBFEAFAA0172B7C0570701FFDEE78F2" box="[366,565,1907,1932]" italics="true" pageId="6" pageNumber="61">Brachistosternus</emphasis>
|
||
(
|
||
<emphasis id="ECBFEAFAA0172B7C0657701FFD8278F2" box="[585,601,1907,1932]" italics="true" pageId="6" pageNumber="61">L</emphasis>
|
||
.)
|
||
<emphasis id="ECBFEAFAA0172B7C066B701FFD1378F2" box="[629,712,1907,1932]" italics="true" pageId="6" pageNumber="61">perettii</emphasis>
|
||
<bibRefCitation id="BA5A4B19A0172B7C06CA701EFB4778F2" author="Ojanguren-Affilastro" box="[724,1180,1906,1932]" pageId="6" pageNumber="68" refString="Ojanguren-Affilastro, A. A. & C. I. Mattoni, (2006) A new species of Brachistosternus from Chilean central Andes (Scorpiones: Bothriuridae). Studies on Neotropical Fauna and Environment, 41, 79 - 85." type="journal article" year="2006">Ojanguren-Affilastro & Mattoni 2006</bibRefCitation>
|
||
</taxonomicName>
|
||
. The two species are easily distinguishable:
|
||
<taxonomicName id="19CB4D6BA0172B7C05BA70F7FD8D78CA" box="[420,598,1946,1972]" class="Arachnida" family="Bothriuridae" genus="Brachistosternus" higherTaxonomySource="CoL" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="perettii" subGenus="Leptosternus">
|
||
<emphasis id="ECBFEAFAA0172B7C05BA70F7FE1B78CA" box="[420,448,1947,1972]" italics="true" pageId="6" pageNumber="61">Br</emphasis>
|
||
. (
|
||
<emphasis id="ECBFEAFAA0172B7C05C770F7FE3278CA" box="[473,489,1947,1972]" italics="true" pageId="6" pageNumber="61">L</emphasis>
|
||
.)
|
||
<emphasis id="ECBFEAFAA0172B7C061D70F7FD8D78CA" box="[515,598,1947,1972]" italics="true" pageId="6" pageNumber="61">perettii</emphasis>
|
||
</taxonomicName>
|
||
do not posses a VM carina of metasomal segment V whereas it is well developed and extends on the entire length of the segment in
|
||
<taxonomicName id="19CB4D6BA0172B7C077B70AFFBED78A2" box="[869,1078,1986,2012]" class="Arachnida" family="Bothriuridae" genus="Brachistosternus" higherTaxonomySource="CoL" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="coquimbo" subGenus="Leptosternus">
|
||
<emphasis id="ECBFEAFAA0172B7C077B70AFFC5A78A2" box="[869,897,1987,2012]" italics="true" pageId="6" pageNumber="61">Br</emphasis>
|
||
. (
|
||
<emphasis id="ECBFEAFAA0172B7C078770AFFC7278A2" box="[921,937,1987,2012]" italics="true" pageId="6" pageNumber="61">L</emphasis>
|
||
.)
|
||
<emphasis id="ECBFEAFAA0172B7C07DC70AFFBED78A2" box="[962,1078,1987,2012]" italics="true" pageId="6" pageNumber="61">coquimbo</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="8AB46660A0162B7D048971DFFD12784C" httpUri="https://zenodo.org/record/179245/files/figure.png" pageId="7" pageNumber="62" targetBox="[275,1322,207,1619]" targetPageId="7">
|
||
<paragraph id="DE7436E8A0162B7D048971DFFD12784C" blockId="7.[151,1436,1715,1842]" pageId="7" pageNumber="62">
|
||
<emphasis id="ECBFEAFAA0162B7D048971DFFE8679B5" bold="true" box="[151,349,1715,1739]" pageId="7" pageNumber="62">FIGURES 11–18.</emphasis>
|
||
<taxonomicName id="19CB4D6BA0162B7D057A71DFFD6379B4" box="[356,696,1715,1738]" class="Arachnida" family="Bothriuridae" genus="Brachistosternus" higherTaxonomySource="CoL" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="coquimbo" status="sp. nov." subGenus="Leptosternus">
|
||
<emphasis id="ECBFEAFAA0162B7D057A71DFFDC379B4" box="[356,536,1715,1738]" italics="true" pageId="7" pageNumber="62">Brachistosternus</emphasis>
|
||
(
|
||
<emphasis id="ECBFEAFAA0162B7D063471DFFDE279B4" box="[554,569,1715,1738]" italics="true" pageId="7" pageNumber="62">L</emphasis>
|
||
.)
|
||
<emphasis id="ECBFEAFAA0162B7D065171DFFD6379B4" box="[591,696,1715,1738]" italics="true" pageId="7" pageNumber="62">coquimbo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F78C5781A0162B7D06DE71DFFD2679B5" box="[704,765,1715,1739]" pageId="7" pageNumber="62" rank="species">
|
||
<emphasis id="ECBFEAFAA0162B7D06DE71DFFD2679B5" bold="true" box="[704,765,1715,1739]" pageId="7" pageNumber="62">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
11. left hemispermatophore, ventral aspect; 12. left hemispermatophore, dorsal aspect; 13. left hemispermatophore, detail of lobe region; 14. metasomal segment V, male, dorsal aspect; 15. metasomal segment V, male, ventral aspect; 16. telson, male, dorsal aspect; 17. telson, male, lateral aspect; 18. telson, female, lateral aspect. Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="96D16563A0162B7304D870E4FB857837" lastPageId="9" lastPageNumber="64" pageId="7" pageNumber="62" type="description">
|
||
<paragraph id="DE7436E8A0162B7204D870E4FD1F7CD4" blockId="7.[151,1437,1928,2034]" lastBlockId="8.[151,1437,152,2018]" lastPageId="8" lastPageNumber="63" pageId="7" pageNumber="62">
|
||
<emphasis id="ECBFEAFAA0162B7D04D870E4FEBA78DC" bold="true" box="[198,353,1928,1954]" pageId="7" pageNumber="62">Description:</emphasis>
|
||
Measurements of a male specimen and a female specimen are recorded in
|
||
<tableCitation id="93490353A0162B7D00C470E4FAE878DC" box="[1242,1331,1928,1954]" captionStart="TABLE 1" captionStartId="3.[151,239,546,570]" captionTargetBox="[151,1436,754,1976]" captionTargetPageId="3" captionText="TABLE 1. Measurements of Brachistosternus (Leptosternus) cepedai n. sp. (Paratypes) and Brachistosternus (Leptosternus) coquimbo n. sp. (Holotype and Paratype). (In mm.)." httpUri="http://table.plazi.org/id/8AB46660A0122B790489754EFCEF7D1E" pageId="7" pageNumber="62" tableUuid="8AB46660A0122B790489754EFCEF7D1E">Table 1</tableCitation>
|
||
.
|
||
<emphasis id="ECBFEAFAA0162B7D015F70E4FA4778DC" bold="true" box="[1345,1436,1928,1954]" italics="true" pageId="7" pageNumber="62">Colour.</emphasis>
|
||
General colour dark yellow with a dense dusky pattern. Chelicerae: with a dark spot near the base of the fingers, movable finger with a dark spot on the external surface. Carapace: densely pigmented; ocular tubercle and lateral ocelli black; area around the ocular tubercle densely pigmented, with two lateral dark stripes from the lateral sulci to the lateral ocelli, and with two posterolateral dark spots; remaining tegument with reticulate pattern. Tergites I–VI: each tergite with three dark spots, two lateral and one median spot, connected by dense reticular pigmentation. Tergite VII with five dark spots, two antero-lateral, two postero-lateral, and an anteromedian spot, sometimes connected by a faint reticular pattern. Sternites: not pigmented. Metasoma: metasomal segments I–III: dorsal surface with an antero-median thin stripe, and two posterolateral dark spots; lateral surfaces depigmented; ventral surface with one thick VM and two thin VL stripes, joined distally; VM stripe barely visible in segment I, absent in some specimens, more noticeable in segment II, and well marked in segment III. Metasomal segment IV: dorsal surface with a faint small antero-median spot, and with two small posterolateral dark spots; lateral surfaces with reticulate pigment; ventral surface with one thick VM and two thin VL stripes joining in the posterior third of the segment,and connected by reticular pattern in the remaining surface. Metasomal segment V: dorsal surface with two DL faint stripes; lateral surface with slight reticular pattern; ventral surface with one thin, dark VM and two thick VL stripes that join in the posterior third of the segment; in the anterior half of the segment the VL stripes divided in four thin lines, two VL and two VSM, that join in the posterior half of the segment. Telson: vesicle with the ventral and dorsolateral surfaces densely pigmented; aculeus dark brawn. Pedipalps: femur densely pigmented near patella articulation, and in the anterior and posterior surfaces, with a dark stripe on the
|
||
<collectingCountry id="A6DC7678A0192B7207517474FCAD7C4C" box="[847,886,792,818]" name="South Georgia and the South Sandwich Islands" pageId="8" pageNumber="63">DE</collectingCountry>
|
||
margin; patella with one
|
||
<collectingCountry id="A6DC7678A0192B7200807474FB667C4C" box="[1182,1213,792,818]" name="Iceland" pageId="8" pageNumber="63">DI</collectingCountry>
|
||
and one
|
||
<collectingCountry id="A6DC7678A0192B7201387474FA967C4C" box="[1318,1357,792,818]" name="South Georgia and the South Sandwich Islands" pageId="8" pageNumber="63">DE</collectingCountry>
|
||
stripe, densely pigmented near chela articulation, remaining surfaces with dense reticular pattern; chela manus with longitudinal stripes joined by a faint reticular pigment, fingers not pigmented. Legs: femur and patella densely pigmented, remaining segments not pigmented.
|
||
</paragraph>
|
||
<paragraph id="DE7436E8A0192B7304D874DBFC557D3C" blockId="8.[151,1437,152,2018]" lastBlockId="9.[151,1437,152,578]" lastPageId="9" lastPageNumber="64" pageId="8" pageNumber="63">
|
||
<emphasis id="ECBFEAFAA0192B7204D874DBFEBC7CAF" bold="true" box="[198,359,951,977]" pageId="8" pageNumber="63">Morphology.</emphasis>
|
||
Carapace: Anterior margin with a slight median bulge; tegument smooth near the ocular tubercle, lateral surfaces densely granular; ocular tubercle slightly anterior to the middle of the carapace, interocular sulcus deeply marked, median ocelli two diameters apart, with a seta behind each eye; anterior, laterals and posterior longitudinal sulci well marked, postocular furrow deeply marked. Chelicerae with two subdistal teeth. Hemispermatophore: distal lamina thick and slightly curved, approximately of the same size as the basal portion (
|
||
<figureCitation id="46F02A6DA0192B72056873ECFDD07BE4" box="[374,523,1152,1178]" captionStart="FIGURES 11 – 18" captionStartId="7.[151,269,1715,1739]" captionTargetBox="[275,1322,207,1619]" captionTargetId="figure@7.[264,1323,194,1691]" captionTargetPageId="7" captionText="FIGURES 11 – 18. Brachistosternus (L.) coquimbo n. sp. 11. left hemispermatophore, ventral aspect; 12. left hemispermatophore, dorsal aspect; 13. left hemispermatophore, detail of lobe region; 14. metasomal segment V, male, dorsal aspect; 15. metasomal segment V, male, ventral aspect; 16. telson, male, dorsal aspect; 17. telson, male, lateral aspect; 18. telson, female, lateral aspect. Scale bars = 1 mm." httpUri="https://zenodo.org/record/179245/files/figure.png" pageId="8" pageNumber="63">Figs. 11, 12</figureCitation>
|
||
); cylindrical apophysis well developed, longer than the laminar apophysis; basal triangle well developed, formed by 2 or 3 crests; internal spines absent; basal spines and row of spines poorly developed, and aligned along the same axis (
|
||
<figureCitation id="46F02A6DA0192B7206E173BCFC8C7B94" box="[767,855,1232,1258]" captionStart="FIGURES 11 – 18" captionStartId="7.[151,269,1715,1739]" captionTargetBox="[275,1322,207,1619]" captionTargetId="figure@7.[264,1323,194,1691]" captionTargetPageId="7" captionText="FIGURES 11 – 18. Brachistosternus (L.) coquimbo n. sp. 11. left hemispermatophore, ventral aspect; 12. left hemispermatophore, dorsal aspect; 13. left hemispermatophore, detail of lobe region; 14. metasomal segment V, male, dorsal aspect; 15. metasomal segment V, male, ventral aspect; 16. telson, male, dorsal aspect; 17. telson, male, lateral aspect; 18. telson, female, lateral aspect. Scale bars = 1 mm." httpUri="https://zenodo.org/record/179245/files/figure.png" pageId="8" pageNumber="63">Fig. 13</figureCitation>
|
||
). Mesosoma: Tergites I–VI finely granular in the distal margin, remaining areas smooth. Tergite VII smooth in the median area, remaining areas densely granular, with two lateral carinae in the posterior half of the segment. Sternites: Sternite I smooth in the anterior half of the segment, coarsely granular in the posterior half; Sternites II–V densely granular. Spiracles narrow and well developed. Sternum
|
||
<typeStatus id="0170884AA0192B7205DE721CFE297AF4" box="[448,498,1392,1418]" pageId="8" pageNumber="63">type</typeStatus>
|
||
2 (
|
||
<bibRefCitation id="BA5A4B19A0192B720609721CFCD77AF4" author="Soleglad" box="[535,780,1392,1418]" pageId="8" pageNumber="68" refString="Soleglad, M. E. & V. Fet, (2003) The scorpion sternum: structure and phylogeny (Scorpiones: Orthosterni). Euscorpius, 5, 1 - 34." type="journal article" year="2003">Soleglad & Fet 2003</bibRefCitation>
|
||
) much wider than long, apex width almost equal to posterior width, posterior emargination quite well developed, with convex lateral lobes conspicuously separated. Pectines: pectinal teeth
|
||
<quantity id="19339B0DA0192B7205AF72ACFDC17AA4" box="[433,538,1472,1498]" metricMagnitude="-1" metricUnit="m" metricValue="8.001" metricValueMax="8.636" metricValueMin="7.366" pageId="8" pageNumber="63" unit="in" value="31.5" valueMax="34.0" valueMin="29.0">29–34 in</quantity>
|
||
males (N = 5; median = 32),
|
||
<quantity id="19339B0DA0192B72079F72ACFC317AA4" box="[897,1002,1472,1498]" metricMagnitude="-1" metricUnit="m" metricValue="7.493" metricValueMax="7.62" metricValueMin="7.366" pageId="8" pageNumber="63" unit="in" value="29.5" valueMax="30.0" valueMin="29.0">29–30 in</quantity>
|
||
the only studied female. Metasoma: Metasomal segments I–III: tegument densely granular; LSM and LIM carinae present on the distal 3/4 of the segment; DL carina weakly developed, extending on the entire length of the segment. Metasomal segment IV: dorsal and lateral surfaces granular; ventral surface smooth with a large number of scattered setae; DL carinae and LSM carinae extending on the entire length of the segment, but poorly developed; LIM carina only present in the last third of the segment. Metasomal segment V: slightly wider in its median part than near the anterior and posterior margins in males (
|
||
<figureCitation id="46F02A6DA0192B72066571DCFD0F79B4" box="[635,724,1712,1738]" captionStart="FIGURES 11 – 18" captionStartId="7.[151,269,1715,1739]" captionTargetBox="[275,1322,207,1619]" captionTargetId="figure@7.[264,1323,194,1691]" captionTargetPageId="7" captionText="FIGURES 11 – 18. Brachistosternus (L.) coquimbo n. sp. 11. left hemispermatophore, ventral aspect; 12. left hemispermatophore, dorsal aspect; 13. left hemispermatophore, detail of lobe region; 14. metasomal segment V, male, dorsal aspect; 15. metasomal segment V, male, ventral aspect; 16. telson, male, dorsal aspect; 17. telson, male, lateral aspect; 18. telson, female, lateral aspect. Scale bars = 1 mm." httpUri="https://zenodo.org/record/179245/files/figure.png" pageId="8" pageNumber="63">Fig. 14</figureCitation>
|
||
); dorsal surface smooth; androvestigia long and well developed, occupying almost 50 % of the dorsal surface (
|
||
<figureCitation id="46F02A6DA0192B72071771B4FCB9798C" box="[777,866,1752,1778]" captionStart="FIGURES 11 – 18" captionStartId="7.[151,269,1715,1739]" captionTargetBox="[275,1322,207,1619]" captionTargetId="figure@7.[264,1323,194,1691]" captionTargetPageId="7" captionText="FIGURES 11 – 18. Brachistosternus (L.) coquimbo n. sp. 11. left hemispermatophore, ventral aspect; 12. left hemispermatophore, dorsal aspect; 13. left hemispermatophore, detail of lobe region; 14. metasomal segment V, male, dorsal aspect; 15. metasomal segment V, male, ventral aspect; 16. telson, male, dorsal aspect; 17. telson, male, lateral aspect; 18. telson, female, lateral aspect. Scale bars = 1 mm." httpUri="https://zenodo.org/record/179245/files/figure.png" pageId="8" pageNumber="63">Fig. 15</figureCitation>
|
||
); lateral surface granular; LSM and DL carinae granular, extending the entire length of the segment but poorly developed; ventral surface irregularly granular, specially in the second half of the segment; VL and VM carinae extending on the entire length of the segment; four rows of ventral setae, two basal rows of four setae and two posterior rows of one or two setae each. Telson: dorsal gland in males almost triangular and well developed (
|
||
<figureCitation id="46F02A6DA0192B7207BD7014FC2778EC" box="[931,1020,1912,1938]" captionStart="FIGURES 11 – 18" captionStartId="7.[151,269,1715,1739]" captionTargetBox="[275,1322,207,1619]" captionTargetId="figure@7.[264,1323,194,1691]" captionTargetPageId="7" captionText="FIGURES 11 – 18. Brachistosternus (L.) coquimbo n. sp. 11. left hemispermatophore, ventral aspect; 12. left hemispermatophore, dorsal aspect; 13. left hemispermatophore, detail of lobe region; 14. metasomal segment V, male, dorsal aspect; 15. metasomal segment V, male, ventral aspect; 16. telson, male, dorsal aspect; 17. telson, male, lateral aspect; 18. telson, female, lateral aspect. Scale bars = 1 mm." httpUri="https://zenodo.org/record/179245/files/figure.png" pageId="8" pageNumber="63">Fig. 16</figureCitation>
|
||
); vesicle with rounded ventral surface (
|
||
<figureCitation id="46F02A6DA0192B7204C970CCFEBE78C4" box="[215,357,1952,1978]" captionStart="FIGURES 11 – 18" captionStartId="7.[151,269,1715,1739]" captionTargetBox="[275,1322,207,1619]" captionTargetId="figure@7.[264,1323,194,1691]" captionTargetPageId="7" captionText="FIGURES 11 – 18. Brachistosternus (L.) coquimbo n. sp. 11. left hemispermatophore, ventral aspect; 12. left hemispermatophore, dorsal aspect; 13. left hemispermatophore, detail of lobe region; 14. metasomal segment V, male, dorsal aspect; 15. metasomal segment V, male, ventral aspect; 16. telson, male, dorsal aspect; 17. telson, male, lateral aspect; 18. telson, female, lateral aspect. Scale bars = 1 mm." httpUri="https://zenodo.org/record/179245/files/figure.png" pageId="8" pageNumber="63">Figs. 17, 18</figureCitation>
|
||
), ventral surface slightly granular; aculeus slightly curved, of the same length as the vesicle. Pedipalps: Femur: internal surface slightly granular; VI,
|
||
<collectingCountry id="A6DC7678A0192B72072C70A4FC87789C" box="[818,860,1992,2018]" name="South Georgia and the South Sandwich Islands" pageId="8" pageNumber="63">DE</collectingCountry>
|
||
, and
|
||
<collectingCountry id="A6DC7678A0192B72078570A4FC61789C" box="[923,954,1992,2018]" name="Iceland" pageId="8" pageNumber="63">DI</collectingCountry>
|
||
carinae granular, extending on the entire length of the segment, but poorly developed; remaining surfaces smooth. Patella: internal surface with scattered granules;
|
||
<collectingCountry id="A6DC7678A0182B73054877ACFEAE7FA4" box="[342,373,192,218]" name="Iceland" pageId="9" pageNumber="64">DI</collectingCountry>
|
||
and VI carinae extending on the entire length of the segment, but poorly developed; remaining surfaces of the tegument smooth. Chela: manus very narrow, with smooth tegument; ventrointernal carina poorly developed (
|
||
<figureCitation id="46F02A6DA0182B7305CC767CFDF57E54" box="[466,558,272,298]" captionStart="FIGURES 19 – 22" captionStartId="9.[151,269,1454,1478]" captionTargetBox="[287,1307,651,1425]" captionTargetId="figure@9.[264,1322,634,1449]" captionTargetPageId="9" captionText="FIGURES 19 – 22. Brachistosternus (L.) coquimbo n. sp. 19. Left pedipalp chela, male, ventral aspect; 20. left pedipalp chela, male, internal aspect; 21. right pedipalp chela, male, external aspect; 22. right pedipalp chela, female, external aspect. Scale bars = 1 mm." httpUri="https://zenodo.org/record/179246/files/figure.png" pageId="9" pageNumber="64">Fig. 19</figureCitation>
|
||
); prolateral apophysis well developed in males; movable finger with a median row of granules and 5 or 6 internal and external granules that do not overlap with the median row of granules. Trichobothrial pattern of subgenus
|
||
<taxonomicName id="19CB4D6BA0182B7306BA760CFCE07E07" box="[676,827,352,377]" class="Arachnida" family="Bothriuridae" genus="Leptosternus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="64" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="ECBFEAFAA0182B7306BA760CFCE07E07" box="[676,827,352,377]" italics="true" pageId="9" pageNumber="64">Leptosternus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="46F02A6DA0182B730755760CFC077E04" box="[843,988,352,378]" captionStart="FIGURES 19 – 22" captionStartId="9.[151,269,1454,1478]" captionTargetBox="[287,1307,651,1425]" captionTargetId="figure@9.[264,1322,634,1449]" captionTargetPageId="9" captionText="FIGURES 19 – 22. Brachistosternus (L.) coquimbo n. sp. 19. Left pedipalp chela, male, ventral aspect; 20. left pedipalp chela, male, internal aspect; 21. right pedipalp chela, male, external aspect; 22. right pedipalp chela, female, external aspect. Scale bars = 1 mm." httpUri="https://zenodo.org/record/179246/files/figure.png" pageId="9" pageNumber="64">Figs. 19–22</figureCitation>
|
||
): Neobothriotaxic Major
|
||
<typeStatus id="0170884AA0182B73011A760CFAE57E04" box="[1284,1342,352,378]" pageId="9" pageNumber="64">Type</typeStatus>
|
||
C, with one accessory trichobothrium in
|
||
<emphasis id="ECBFEAFAA0182B73060B76E4FDFC7EDF" box="[533,551,392,417]" italics="true" pageId="9" pageNumber="64">V</emphasis>
|
||
series of chela; femur with 3 trichobothria (1
|
||
<emphasis id="ECBFEAFAA0182B73005E76E4FB947EDF" box="[1088,1103,392,417]" italics="true" pageId="9" pageNumber="64">d</emphasis>
|
||
,
|
||
<date id="AA751028A0182B73004376E4FBA17EDF" box="[1117,1146,392,418]" pageId="9" pageNumber="64">
|
||
1
|
||
<emphasis id="ECBFEAFAA0182B73006C76E4FBA17EDF" box="[1138,1146,392,417]" italics="true" pageId="9" pageNumber="64">i</emphasis>
|
||
</date>
|
||
and 1
|
||
<emphasis id="ECBFEAFAA0182B7300D676E4FB0E7EDF" box="[1224,1237,392,417]" italics="true" pageId="9" pageNumber="64">e</emphasis>
|
||
); patella with 19 trichobothria (
|
||
<date id="AA751028A0182B73055E76DCFEBD7EB7" box="[320,358,432,458]" pageId="9" pageNumber="64">
|
||
3
|
||
<emphasis id="ECBFEAFAA0182B73054676DCFEBD7EB7" box="[344,358,432,457]" italics="true" pageId="9" pageNumber="64">V</emphasis>
|
||
</date>
|
||
, 2
|
||
<emphasis id="ECBFEAFAA0182B73059376DCFE477EB7" box="[397,412,432,457]" italics="true" pageId="9" pageNumber="64">d</emphasis>
|
||
,
|
||
<date id="AA751028A0182B7305B276DCFE107EB7" box="[428,459,432,458]" pageId="9" pageNumber="64">
|
||
1
|
||
<emphasis id="ECBFEAFAA0182B7305DD76DCFE107EB7" box="[451,459,432,457]" italics="true" pageId="9" pageNumber="64">i</emphasis>
|
||
</date>
|
||
, 3
|
||
<emphasis id="ECBFEAFAA0182B7305EC76DCFDDC7EB7" box="[498,519,432,457]" italics="true" pageId="9" pageNumber="64">et</emphasis>
|
||
, 1
|
||
<emphasis id="ECBFEAFAA0182B73063176DCFD947EB7" box="[559,591,432,457]" italics="true" pageId="9" pageNumber="64">est</emphasis>
|
||
, 2
|
||
<emphasis id="ECBFEAFAA0182B73066876DCFD437EB7" box="[630,664,432,457]" italics="true" pageId="9" pageNumber="64">em</emphasis>
|
||
, 2
|
||
<emphasis id="ECBFEAFAA0182B7306DE76DCFD3C7EB7" box="[704,743,432,457]" italics="true" pageId="9" pageNumber="64">esb</emphasis>
|
||
, and 5
|
||
<emphasis id="ECBFEAFAA0182B73075F76DCFC867EB7" box="[833,861,432,457]" italics="true" pageId="9" pageNumber="64">eb</emphasis>
|
||
); chela with 27 trichobothria (1
|
||
<emphasis id="ECBFEAFAA0182B7300FF76DCFADD7EB7" box="[1249,1286,432,457]" italics="true" pageId="9" pageNumber="64">Est</emphasis>
|
||
, 5
|
||
<emphasis id="ECBFEAFAA0182B73013076DCFA937EB7" box="[1326,1352,432,457]" italics="true" pageId="9" pageNumber="64">Et</emphasis>
|
||
,
|
||
<date id="AA751028A0182B73014676DCFAA67EB7" box="[1368,1405,432,458]" pageId="9" pageNumber="64">
|
||
5
|
||
<emphasis id="ECBFEAFAA0182B73017176DCFAA67EB7" box="[1391,1405,432,457]" italics="true" pageId="9" pageNumber="64">V</emphasis>
|
||
</date>
|
||
, 1
|
||
<emphasis id="ECBFEAFAA0182B73048976B4FF187E8F" box="[151,195,472,497]" italics="true" pageId="9" pageNumber="64">Esb</emphasis>
|
||
, 3
|
||
<emphasis id="ECBFEAFAA0182B7304F876B4FEDC7E8F" box="[230,263,472,497]" italics="true" pageId="9" pageNumber="64">Eb</emphasis>
|
||
, 1
|
||
<emphasis id="ECBFEAFAA0182B73053576B4FE937E8F" box="[299,328,472,497]" italics="true" pageId="9" pageNumber="64">Dt</emphasis>
|
||
, 1
|
||
<emphasis id="ECBFEAFAA0182B73057276B4FE4B7E8F" box="[364,400,472,497]" italics="true" pageId="9" pageNumber="64">Db</emphasis>
|
||
, 1
|
||
<emphasis id="ECBFEAFAA0182B7305AD76B4FE137E8F" box="[435,456,472,497]" italics="true" pageId="9" pageNumber="64">et</emphasis>
|
||
, 1
|
||
<emphasis id="ECBFEAFAA0182B7305F276B4FDD67E8F" box="[492,525,472,497]" italics="true" pageId="9" pageNumber="64">est</emphasis>
|
||
, 1
|
||
<emphasis id="ECBFEAFAA0182B73062E76B4FD8C7E8F" box="[560,599,472,497]" italics="true" pageId="9" pageNumber="64">esb</emphasis>
|
||
, 1
|
||
<emphasis id="ECBFEAFAA0182B73066576B4FD4C7E8F" box="[635,663,472,497]" italics="true" pageId="9" pageNumber="64">eb</emphasis>
|
||
, 1
|
||
<emphasis id="ECBFEAFAA0182B7306A576B4FD097E8F" box="[699,722,472,497]" italics="true" pageId="9" pageNumber="64">dt</emphasis>
|
||
, 1
|
||
<emphasis id="ECBFEAFAA0182B7306EB76B4FCCC7E8F" box="[757,791,472,497]" italics="true" pageId="9" pageNumber="64">dst</emphasis>
|
||
, 1
|
||
<emphasis id="ECBFEAFAA0182B73072576B4FCBF7E8F" box="[827,868,472,497]" italics="true" pageId="9" pageNumber="64">dsb</emphasis>
|
||
, 1
|
||
<emphasis id="ECBFEAFAA0182B73079676B4FC7E7E8F" box="[904,933,472,497]" italics="true" pageId="9" pageNumber="64">db</emphasis>
|
||
, 1
|
||
<emphasis id="ECBFEAFAA0182B7307D776B4FC047E8F" box="[969,991,472,497]" italics="true" pageId="9" pageNumber="64">ib</emphasis>
|
||
, 1
|
||
<emphasis id="ECBFEAFAA0182B73001D76B4FBC87E8F" box="[1027,1043,472,497]" italics="true" pageId="9" pageNumber="64">it</emphasis>
|
||
). Legs: Tegument finely granular in femur; telotarsi I and II with the inner unguis 5 to 10 % shorter than the external one; internal and external basitarsal spurs well developed, of the same size in all segments.
|
||
</paragraph>
|
||
<caption id="8AB46660A0182B73048972C2FE6E7975" httpUri="https://zenodo.org/record/179246/files/figure.png" pageId="9" pageNumber="64" targetBox="[287,1307,651,1425]" targetPageId="9">
|
||
<paragraph id="DE7436E8A0182B73048972C2FE6E7975" blockId="9.[151,1436,1454,1547]" pageId="9" pageNumber="64">
|
||
<emphasis id="ECBFEAFAA0182B73048972C2FE857AB8" bold="true" box="[151,350,1454,1478]" pageId="9" pageNumber="64">FIGURES 19–22.</emphasis>
|
||
<taxonomicName id="19CB4D6BA0182B73057B72C3FD637AB8" box="[357,696,1455,1478]" class="Arachnida" family="Bothriuridae" genus="Brachistosternus" higherTaxonomySource="CoL" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="coquimbo" status="sp. nov." subGenus="Leptosternus">
|
||
<emphasis id="ECBFEAFAA0182B73057B72C3FDC17AB8" box="[357,538,1455,1478]" italics="true" pageId="9" pageNumber="64">Brachistosternus</emphasis>
|
||
(
|
||
<emphasis id="ECBFEAFAA0182B73063472C3FDE27AB8" box="[554,569,1455,1478]" italics="true" pageId="9" pageNumber="64">L</emphasis>
|
||
.)
|
||
<emphasis id="ECBFEAFAA0182B73065172C3FD637AB8" box="[591,696,1455,1478]" italics="true" pageId="9" pageNumber="64">coquimbo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F78C5781A0182B7306DE72C2FD277AB8" box="[704,764,1454,1478]" pageId="9" pageNumber="64" rank="species">
|
||
<emphasis id="ECBFEAFAA0182B7306DE72C2FD277AB8" bold="true" box="[704,764,1454,1478]" pageId="9" pageNumber="64">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
19. Left pedipalp chela, male, ventral aspect; 20. left pedipalp chela, male, internal aspect; 21. right pedipalp chela, male, external aspect; 22. right pedipalp chela, female, external aspect. Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="DE7436E8A0182B7304D87152FB857837" blockId="9.[151,1437,1598,2025]" pageId="9" pageNumber="64">
|
||
<emphasis id="ECBFEAFAA0182B7304D87152FEE27926" bold="true" box="[198,313,1598,1624]" pageId="9" pageNumber="64">Variation</emphasis>
|
||
: Pedipalp chela length/height ratio, male 3.63–3.93 (N = 6; mean = 3.81), female 3.5; pedipalp chela length/width ratio, male 4.7–5.09 (N = 6; mean = 4.85), female 4.33; metasomal segment V, length/ width ratio, male 1.5–1.7 (N = 6; mean = 1.61), female 1.4; metasomal segment V, ventral macroseta, 11–13 (N = 7; median = 12); metasomal segment V, ventrolateral macroseta, 8–9 (N = 7; median = 8); telotarsus III, dorsal macroseta, 10–12 (N = 7; median = 11); telotarsus III, ventrointernal macroseta, 5–8 (N = 7; median = 7); telotarsus III, ventroexternal macroseta, 6–8 (N = 7; median = 7); basitarsus III, dorsal macroseta, 5–9 (N = 7; median = 8); total length (mm), males, 49–57 (N = 6; mean = 53), female: 44.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="96D16563A0182B7004D8703AFE8A7E54" lastPageId="10" lastPageNumber="65" pageId="9" pageNumber="64" type="distribution">
|
||
<paragraph id="DE7436E8A0182B7304D8703AFD0F7897" blockId="9.[151,1437,1598,2025]" pageId="9" pageNumber="64">
|
||
<emphasis id="ECBFEAFAA0182B7304D8703AFDD8780E" bold="true" box="[198,515,1878,1904]" pageId="9" pageNumber="64">Distribution and ecology:</emphasis>
|
||
<taxonomicName id="19CB4D6BA0182B730614703BFCA6780E" box="[522,893,1879,1905]" class="Arachnida" family="Bothriuridae" genus="Brachistosternus" higherTaxonomySource="CoL" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="coquimbo" status="sp. nov." subGenus="Leptosternus">
|
||
<emphasis id="ECBFEAFAA0182B730614703BFD0A780E" box="[522,721,1879,1904]" italics="true" pageId="9" pageNumber="64">Brachistosternus</emphasis>
|
||
(
|
||
<emphasis id="ECBFEAFAA0182B7306FC703BFD29780E" box="[738,754,1879,1904]" italics="true" pageId="9" pageNumber="64">L</emphasis>
|
||
.)
|
||
<emphasis id="ECBFEAFAA0182B730714703BFCA6780E" box="[778,893,1879,1904]" italics="true" pageId="9" pageNumber="64">coquimbo</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="ECBFEAFAA0182B73079B703AFC1C780E" bold="true" box="[901,967,1878,1904]" pageId="9" pageNumber="64">
|
||
<taxonomicNameLabel id="F78C5781A0182B73079B703AFC1C780E" box="[901,967,1878,1904]" pageId="9" pageNumber="64" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has been collected in the Andean sector of
|
||
<taxonomicName id="19CB4D6BA0182B7304A87013FEE978E7" box="[182,306,1919,1945]" class="Arachnida" family="Bothriuridae" genus="Brachistosternus" higherTaxonomySource="CoL" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="65" phylum="Arthropoda" rank="species" species="coquimbo" status="sp. nov.">Coquimbo</taxonomicName>
|
||
Region, (
|
||
<collectingCountry id="A6DC7678A0182B7305BA7013FE3278E7" box="[420,489,1919,1945]" name="Chile" pageId="9" pageNumber="64">Chile</collectingCountry>
|
||
), between
|
||
<quantity id="19339B0DA0182B73067B7013FCC778E7" box="[613,796,1919,1945]" metricMagnitude="3" metricUnit="m" metricValue="2.925" metricValueMax="3.0" metricValueMin="2.85" pageId="9" pageNumber="64" unit="m" value="2925.0" valueMax="3000.0" valueMin="2850.0">2850 to 3000 m</quantity>
|
||
a.s.l (
|
||
<figureCitation id="46F02A6DA0182B73077C7013FC7D78E7" box="[866,934,1919,1945]" captionStart="FIGURE 1" captionStartId="2.[151,255,1313,1337]" captionTargetBox="[275,1311,178,1278]" captionTargetId="figure@2.[225,1362,137,1307]" captionTargetPageId="2" captionText="FIGURE 1. Map of Coquimbo region, Chile, with localities of Brachistosternus (L.) cepedai n. sp. and Brachistosternus (L.) coquimbo n. sp." httpUri="https://zenodo.org/record/179243/files/figure.png" pageId="9" pageNumber="64">Fig. 1</figureCitation>
|
||
). All localities are located within the high- Andean steppe of Doña Ana botanical subregion, of the Andean desert botanical region (Cepeda-Pizarro
|
||
<emphasis id="ECBFEAFAA0182B73017C70CBFA4778BE" box="[1378,1436,1959,1984]" italics="true" pageId="9" pageNumber="64">et al.</emphasis>
|
||
2006;
|
||
<bibRefCitation id="BA5A4B19A0182B7304FF70A3FE5F7897" author="Gajardo" box="[225,388,1999,2025]" pageId="9" pageNumber="67" refString="Gajardo, R. (1993) La vegetacion natural de Chile. Clasificacion y distribucion geografica. E ditorial Universitaria, Santiago, Chile, p. 165." type="book chapter" year="1993">Gajardo 1993</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="BA5A4B19A0182B73058F70A3FDB57897" author="Squeo" box="[401,622,1999,2025]" pageId="9" pageNumber="68" refString="Squeo, F. A., J. Cepeda-Pizarro, N. C. Olivares, & M. T. K. Arroyo, (2006 a) Interacciones ecologicas en la Alta Montana del Valle de Elqui. In: J. Cepeda-Pizarro (Ed.). Geoecologia de los Andes Deserticos: La Alta Montana del Valle del Elqui. Ediciones Universidad de La Serena, La Serena, Chile, pp. 69 - 103." type="book chapter" year="2006" yearSuffix="a">
|
||
Squeo
|
||
<emphasis id="ECBFEAFAA0182B7305FF70A3FDC77896" box="[481,540,1999,2024]" italics="true" pageId="9" pageNumber="64">et al.</emphasis>
|
||
2006a
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="BA5A4B19A0182B73066770A3FD137897" author="Squeo" box="[633,712,1999,2025]" pageId="9" pageNumber="68" refString="Squeo, F. A., E. Ibacache, B. Warner, D. Espinoza, R. Aravena, & J. R. Gutierrez, (2006 b) Productividad y diversidad floristica de la Vega Tambo, Cordillera de Dona Ana: variabilidad inter-anual, herbivoria y nivel freatico. In: J. Cepeda- Pizarro (Ed.). Geoecologia de los Andes Deserticos: La Alta Montana del Valle del Elqui. Ediciones Universidad de La Serena, La Serena, Chile, pp. 325 - 351." type="book chapter" year="2006" yearSuffix="b">2006b</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="DE7436E8A01B2B7004D877F4FE8A7E54" blockId="10.[151,1436,151,298]" pageId="10" pageNumber="65">
|
||
<taxonomicName id="19CB4D6BA01B2B7004D877F4FDE07FCF" box="[198,571,152,178]" class="Arachnida" family="Bothriuridae" genus="Brachistosternus" higherTaxonomySource="CoL" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="65" phylum="Arthropoda" rank="species" species="coquimbo" status="sp. nov." subGenus="Leptosternus">
|
||
<emphasis id="ECBFEAFAA01B2B7004D877F4FE567FCF" box="[198,397,152,177]" italics="true" pageId="10" pageNumber="65">Brachistosternus</emphasis>
|
||
(
|
||
<emphasis id="ECBFEAFAA01B2B70058077F4FE757FCF" box="[414,430,152,177]" italics="true" pageId="10" pageNumber="65">L</emphasis>
|
||
.)
|
||
<emphasis id="ECBFEAFAA01B2B7005D977F4FDE07FCF" box="[455,571,152,177]" italics="true" pageId="10" pageNumber="65">coquimbo</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F78C5781A01B2B70065C77FBFD5E7FCF" box="[578,645,151,177]" pageId="10" pageNumber="65" rank="species">
|
||
<emphasis id="ECBFEAFAA01B2B70065C77FBFD5E7FCF" bold="true" box="[578,645,151,177]" pageId="10" pageNumber="65">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
occurs in areas with fine clay substratum with scattered rocks. The vegetation of this area is composed mostly of small shrubs. This species has been collected in sympatry with
|
||
<taxonomicName id="19CB4D6BA01B2B7004897784FE9C7E7F" box="[151,327,232,258]" class="Arachnida" family="Bothriuridae" genus="Brachistosternus" higherTaxonomySource="CoL" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="65" phylum="Arthropoda" rank="species" species="perettii" subGenus="Leptosternus">
|
||
<emphasis id="ECBFEAFAA01B2B7004897784FF687E7F" box="[151,179,232,257]" italics="true" pageId="10" pageNumber="65">Br</emphasis>
|
||
. (
|
||
<emphasis id="ECBFEAFAA01B2B7004D57784FF007E7F" box="[203,219,232,257]" italics="true" pageId="10" pageNumber="65">L</emphasis>
|
||
.
|
||
<emphasis id="ECBFEAFAA01B2B7004FD7784FE9C7E7F" box="[227,327,232,257]" italics="true" pageId="10" pageNumber="65">) perettii</emphasis>
|
||
</taxonomicName>
|
||
, and with an undescribed species of
|
||
<taxonomicName id="19CB4D6BA01B2B7006E67784FB9A7E7C" authority="Maury, 1976" authorityName="Maury" authorityYear="1976" box="[760,1089,232,258]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="65" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="ECBFEAFAA01B2B7006E67784FC797E7F" box="[760,930,232,257]" italics="true" pageId="10" pageNumber="65">Orobothriurus</emphasis>
|
||
Maury, 1976
|
||
</taxonomicName>
|
||
, closely related to
|
||
<taxonomicName id="19CB4D6BA01B2B7001007784FE957E54" authority="Pocock, 1899" authorityName="Pocock" authorityYear="1899" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="65" phylum="Arthropoda" rank="species" species="alticola">
|
||
<collectingCountry id="A6DC7678A01B2B7001007784FAE87E7F" box="[1310,1331,232,257]" name="Iceland" pageId="10" pageNumber="65">
|
||
<emphasis id="ECBFEAFAA01B2B7001007784FAE87E7F" box="[1310,1331,232,257]" italics="true" pageId="10" pageNumber="65">O</emphasis>
|
||
</collectingCountry>
|
||
.
|
||
<emphasis id="ECBFEAFAA01B2B70015C7784FA477E7F" box="[1346,1436,232,257]" italics="true" pageId="10" pageNumber="65">alticola</emphasis>
|
||
(Pocock, 1899)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |