treatments-xml/data/56/62/87/566287FEA0132B7C04897290FD737B0C.xml

<document id="B54B0162AE443D7D2D8BE9BD62431C11" ID-DOI="10.5281/zenodo.179242" ID-GBIF-Dataset="9fb430d0-cd77-48f4-b277-8a7c3b316485" ID-ISSN="1175-5326" ID-Zenodo-Dep="179242" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459946210070" checkinUser="plazi" docAuthor="Ojanguren-Affilastro, Andrés A., Agusto, Pablo, Pizarro-Araya, Jaime &amp; Mattoni, Camilo I." docDate="2007" docId="566287FEA0132B7C04897290FD737B0C" docLanguage="en" docName="zt01623p068.pdf" docOrigin="Zootaxa 1623" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Brachistosternus (Leptosternus) cepedai Ojanguren-Affilastro, Agusto, Pizarro-Araya &amp; Mattoni, 2007, n. sp." docType="treatment" docVersion="10" lastPageNumber="61" masterDocId="AA5BFF86A0112B7A041E776CFFDB7F7E" masterDocTitle="Two new scorpion species of genus Brachistosternus (Scorpiones: Bothriuridae) from northern Chile" masterLastPageNumber="68" masterPageNumber="55" pageNumber="57" updateTime="1698230676319" updateUser="plazi">
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<mods:title id="61FB08AE8DAEF97C73744D80D5CEDFBF">Two new scorpion species of genus Brachistosternus (Scorpiones: Bothriuridae) from northern Chile</mods:title>
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<mods:namePart id="4293167D8108E89DA93A0569F136472F">Agusto, Pablo</mods:namePart>
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<mods:namePart id="BA5772ADE7BB22AA198D45C8371DEFC8">Mattoni, Camilo I.</mods:namePart>
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<treatment id="566287FEA0132B7C04897290FD737B0C" ID-DOI="http://doi.org/10.5281/zenodo.5670650" ID-GBIF-Taxon="119376229" ID-Zenodo-Dep="5670650" LSID="urn:lsid:plazi:treatment:566287FEA0132B7C04897290FD737B0C" httpUri="http://treatment.plazi.org/id/566287FEA0132B7C04897290FD737B0C" lastPageId="6" lastPageNumber="61" pageId="2" pageNumber="57">
<subSubSection id="96D16563A0132B7804897290FE0A7947" pageId="2" pageNumber="57" type="nomenclature">
<paragraph id="DE7436E8A0132B7804897290FD1C7968" blockId="2.[151,711,1532,1593]" box="[151,711,1532,1558]" pageId="2" pageNumber="57">
<heading id="853C8184A0132B7804897290FD1C7968" bold="true" box="[151,711,1532,1558]" fontSize="11" level="1" pageId="2" pageNumber="57" reason="1">
<emphasis id="ECBFEAFAA0132B7804897290FD1C7968" bold="true" box="[151,711,1532,1558]" pageId="2" pageNumber="57">
<taxonomicName id="19CB4D6BA0132B7804897290FDA67968" ID-CoL="MTVH" box="[151,637,1532,1558]" class="Arachnida" family="Bothriuridae" genus="Brachistosternus" higherTaxonomySource="CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="57" phylum="Arthropoda" rank="species" species="cepedai" status="sp. nov." subGenus="Leptosternus">
<emphasis id="ECBFEAFAA0132B7804897290FEBF7968" bold="true" box="[151,356,1532,1558]" italics="true" pageId="2" pageNumber="57">Brachistosternus</emphasis>
(
<emphasis id="ECBFEAFAA0132B78056B7290FDCA7968" bold="true" box="[373,529,1532,1558]" italics="true" pageId="2" pageNumber="57">Leptosternus</emphasis>
) 
<emphasis id="ECBFEAFAA0132B78063C7290FDA67968" bold="true" box="[546,637,1532,1558]" italics="true" pageId="2" pageNumber="57">cepedai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F78C5781A0132B78069B7290FD1C7968" box="[645,711,1532,1558]" pageId="2" pageNumber="57" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="DE7436E8A0132B7804897173FE0A7947" blockId="2.[151,711,1532,1593]" box="[151,465,1567,1593]" pageId="2" pageNumber="57">
<figureCitation id="46F02A6DA0132B7804897173FECE7947" box="[151,277,1567,1593]" captionStart-0="FIGURE 1" captionStart-1="FIGURES 2 – 10" captionStartId-0="2.[151,255,1313,1337]" captionStartId-1="4.[151,269,1714,1738]" captionTargetBox-0="[275,1311,178,1278]" captionTargetBox-1="[280,1321,250,1688]" captionTargetId-0="figure@2.[225,1362,137,1307]" captionTargetId-1="figure@4.[264,1322,194,1690]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionText-0="FIGURE 1. Map of Coquimbo region, Chile, with localities of Brachistosternus (L.) cepedai n. sp. and Brachistosternus (L.) coquimbo n. sp." captionText-1="FIGURES 2 – 10. Brachistosternus (L.) cepedai n. sp. 2. left hemispermatophore, ventral aspect; 3. left hemispermatophore, dorsal aspect; 4. metasomal segment V, male, dorsal aspect; 5. metasomal segment V, male, ventral aspect; 6. telson, male, lateral aspect; 7. telson, female, lateral aspect; 8. right pedipalp chela, male, ventral aspect; 9. right pedipalp chela, male, internal aspect; 10. left pedipalp chela, female, external aspect. Scale bars = 1 mm." httpUri-0="https://zenodo.org/record/179243/files/figure.png" httpUri-1="https://zenodo.org/record/179244/files/figure.png" pageId="2" pageNumber="57">Figs. 1–10</figureCitation>
, 
<figureCitation id="46F02A6DA0132B78053F7173FEB57947" box="[289,366,1567,1593]" captionStart="FIGURES 23 – 30" captionStartId="10.[151,269,1664,1688]" captionTargetBox="[327,1260,355,1634]" captionTargetId="figure@10.[319,1267,354,1640]" captionTargetPageId="10" captionText="FIGURES 23 – 30. Prolateral view of right leg telotarsi I and II of four Brachistosternus species. 23 – 24. Brachistosternus cepedai n. sp., 23. telotarsus I; 24. telotarsus II. 25 – 26 Brachistosternus sciosciae. 25. telotarsus I; 26. telotarsus II. 27 – 28 Brachistosternus roigalsinai. 27. telotarsus I; 28. telotarsus II. 29 – 30 Brachistosternus montanus. 29. telotarsus I; 30. telotarsus II. Scale bars 1 mm." httpUri="https://zenodo.org/record/179247/files/figure.png" pageId="2" pageNumber="57">23, 24</figureCitation>
; 
<tableCitation id="93490353A0132B7805647173FE0A7947" box="[378,465,1567,1593]" captionStart="TABLE 1" captionStartId="3.[151,239,546,570]" captionTargetBox="[151,1436,754,1976]" captionTargetPageId="3" captionText="TABLE 1. Measurements of Brachistosternus (Leptosternus) cepedai n. sp. (Paratypes) and Brachistosternus (Leptosternus) coquimbo n. sp. (Holotype and Paratype). (In mm.)." httpUri="http://table.plazi.org/id/8AB46660A0122B790489754EFCEF7D1E" pageId="2" pageNumber="57" tableUuid="8AB46660A0122B790489754EFCEF7D1E">Table 1</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="96D16563A0132B7804897102FB8F78DF" pageId="2" pageNumber="57" type="materials_examined">
<paragraph id="DE7436E8A0132B7804897102FBB67807" blockId="2.[151,1436,1646,2033]" pageId="2" pageNumber="57">
<emphasis id="ECBFEAFAA0132B7804897102FE8B79F6" bold="true" box="[151,336,1646,1672]" pageId="2" pageNumber="57">
<typeStatus id="0170884AA0132B7804897102FF0979F6" box="[151,210,1646,1672]" pageId="2" pageNumber="57">Type</typeStatus>
material:
</emphasis>
<typeStatus id="0170884AA0132B7805457103FE1379F7" box="[347,456,1647,1673]" pageId="2" pageNumber="57" type="holotype">Holotype</typeStatus>
ɗ: 
<collectingCountry id="A6DC7678A0132B7805E97103FDE279F7" box="[503,569,1647,1673]" name="Chile" pageId="2" pageNumber="57">Chile</collectingCountry>
, IV Región, Elqui Province, 
<quantity id="19339B0DA0132B7807877103FC0C79F7" box="[921,983,1647,1673]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="57" unit="km" value="5.0">5 km</quantity>
from Punta Choros, near the route to Choros, 
<geoCoordinate id="BBFF502FA0132B7804E371FBFE7979CF" box="[253,418,1687,1713]" direction="south" orientation="latitude" pageId="2" pageNumber="57" precision="1" value="-29.241388">29°14'29.0'' S</geoCoordinate>
, 
<geoCoordinate id="BBFF502FA0132B7805AF71FBFDBA79CF" box="[433,609,1687,1713]" direction="west" orientation="longitude" pageId="2" pageNumber="57" precision="1" value="-71.42941">71°25'45.9'' W</geoCoordinate>
, 
<quantity id="19339B0DA0132B78067171FBFD7679CF" box="[623,685,1687,1713]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="57" unit="m" value="12.0">12 m</quantity>
a.s.l, 
<date id="AA751028A0132B7806E971FBFCAD79CF" box="[759,886,1687,1713]" pageId="2" pageNumber="57" value="2006-02-22">22/II/2006</date>
, A.A. Ojanguren-Affilastro, L. Compagnucci, and A.C. Cuezzo, UV sampling in dunes (MACN-Ar 12246). 
<typeStatus id="0170884AA0132B78077771D3FC3B79A7" box="[873,992,1727,1753]" pageId="2" pageNumber="57" type="paratype">Paratypes</typeStatus>
: 2 ɗ, same data as 
<typeStatus id="0170884AA0132B7800A471D3FAFB79A7" box="[1210,1312,1727,1753]" pageId="2" pageNumber="57" type="holotype">holotype</typeStatus>
(
<collectionCode id="B8DAAE2DA0132B78013171D3FA4B79A7" LSID="urn:lsid:biocol.org:col:34925" box="[1327,1424,1727,1753]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="2" pageNumber="57">AMNH</collectionCode>
); 1 ɗ, 1 Ψ, same data as 
<typeStatus id="0170884AA0132B7805BA718BFDD0787F" box="[420,523,1767,1793]" pageId="2" pageNumber="57" type="holotype">holotype</typeStatus>
(
<collectionCode id="B8DAAE2DA0132B780605718BFD84787F" box="[539,607,1767,1793]" httpUri="http://grbio.org/cool/89m4-4i9r" name="Canadian Department of Agriculture" pageId="2" pageNumber="57">CDA</collectionCode>
); 20 ɗ, 5 Ψ, (same data as 
<typeStatus id="0170884AA0132B780784718BFBD9787F" box="[922,1026,1767,1793]" pageId="2" pageNumber="57" type="holotype">holotype</typeStatus>
) (MACN-Ar); 4 ɗ, 7 Ψ, same date as 
<typeStatus id="0170884AA0132B7804A87063FEC77857" box="[182,284,1807,1833]" pageId="2" pageNumber="57" type="holotype">holotype</typeStatus>
(
<collectionCode id="B8DAAE2DA0132B7805357063FE567857" box="[299,397,1807,1833]" pageId="2" pageNumber="57">LEULS</collectionCode>
); 3 ɗ, 10 Ψ, 
<quantity id="19339B0DA0132B7806007063FDB37857" box="[542,616,1807,1833]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="57" unit="km" value="20.0">20 km</quantity>
from Punta Choros, 
<geoCoordinate id="BBFF502FA0132B7807447063FC287857" box="[858,1011,1807,1833]" direction="south" orientation="latitude" pageId="2" pageNumber="57" precision="15" value="-27.638056">27°37´77´´ S</geoCoordinate>
, 67°53´126´´ W, 
<quantity id="19339B0DA0132B7800A37063FB237857" box="[1213,1272,1807,1833]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="57" unit="m" value="20.0">20 m</quantity>
a.s.l, 
<date id="AA751028A0132B7801227063FF0F782F" pageId="2" pageNumber="57" value="2005-08-26">26/VIII/ 2005</date>
, D. Valdivia, UV sampling in dunes (MZUC-UCCC). 3 ɗ, 10 Ψ, 
<quantity id="19339B0DA0132B7807C7705BFBFF782F" box="[985,1060,1847,1873]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="57" unit="km" value="15.0">15 km</quantity>
from Punta Choros, 
<geoCoordinate id="BBFF502FA0132B780105705BFF727807" direction="south" orientation="latitude" pageId="2" pageNumber="57" precision="15" value="-27.071667">27°03´78´´ S</geoCoordinate>
, 67°55´851´´ W, 
<date id="AA751028A0132B78056A7033FE2B7807" box="[372,496,1887,1913]" pageId="2" pageNumber="57" value="2005-11-09">9/XI/2005</date>
, J. Pizarro, 
<quantity id="19339B0DA0132B7806667033FD7E7807" box="[632,677,1887,1913]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="57" unit="m" value="2.0">2 m</quantity>
a.s.l, UV sampling in dunes (
<collectionCode id="B8DAAE2DA0132B78001F7033FBB97807" LSID="urn:lsid:biocol.org:col:34988" box="[1025,1122,1887,1913]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="57">MNHN</collectionCode>
).
</paragraph>
<paragraph id="DE7436E8A0132B7804D870EAFB8F78DF" blockId="2.[151,1436,1646,2033]" box="[198,1108,1926,1953]" pageId="2" pageNumber="57">
<emphasis id="ECBFEAFAA0132B7804D870EAFE1F78DE" bold="true" box="[198,452,1926,1952]" pageId="2" pageNumber="57">Additional material.</emphasis>
<specimenCount id="C8CDFD61A0132B7805D270EBFD9078DF" box="[460,587,1927,1953]" pageId="2" pageNumber="57" type="juvenile">2 juveniles</specimenCount>
(
<collectionCode id="B8DAAE2DA0132B78064470EBFD6978DF" box="[602,690,1927,1953]" httpUri="http://grbio.org/cool/m0b5-nwmg" name="Ambrose Monell Cryo Collection, American Museum of Natural History" pageId="2" pageNumber="57">AMCC</collectionCode>
[LP 5844]), same data as 
<typeStatus id="0170884AA0132B7807FB70EBFB9478DF" box="[997,1103,1927,1953]" pageId="2" pageNumber="57" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="96D16563A0132B7804D870C2FBB0788F" pageId="2" pageNumber="57" type="etymology">
<paragraph id="DE7436E8A0132B7804D870C2FBB0788F" blockId="2.[151,1436,1646,2033]" pageId="2" pageNumber="57">
<emphasis id="ECBFEAFAA0132B7804D870C2FE8878B6" bold="true" box="[198,339,1966,1992]" pageId="2" pageNumber="57">Etymology.</emphasis>
This species is named after the Chilean biologist Jorge Cepeda-Pizarro, who has carried out an extensive research on the epigean arthropods of the arid zones of northern 
<collectingCountry id="A6DC7678A0132B78003D70BBFBBC788F" box="[1059,1127,2007,2033]" name="Chile" pageId="2" pageNumber="57">Chile</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="96D16563A0122B7904D877FBFCD67EDC" pageId="3" pageNumber="58" type="diagnosis">
<paragraph id="DE7436E8A0122B7904D877FBFCD67EDC" blockId="3.[151,1436,151,498]" pageId="3" pageNumber="58">
<emphasis id="ECBFEAFAA0122B7904D877FBFE9D7FCF" bold="true" box="[198,326,151,177]" pageId="3" pageNumber="58">Diagnosis.</emphasis>
<taxonomicName id="19CB4D6BA0122B79055377F4FD727FCF" box="[333,681,152,178]" class="Arachnida" family="Bothriuridae" genus="Brachistosternus" higherTaxonomySource="CoL" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="cepedai" status="sp. nov." subGenus="Leptosternus">
<emphasis id="ECBFEAFAA0122B79055377F4FDCF7FCF" box="[333,532,152,177]" italics="true" pageId="3" pageNumber="58">Brachistosternus</emphasis>
(
<emphasis id="ECBFEAFAA0122B79063B77F4FDEE7FCF" box="[549,565,152,177]" italics="true" pageId="3" pageNumber="58">L</emphasis>
.) 
<emphasis id="ECBFEAFAA0122B79065077F4FD727FCF" box="[590,681,152,177]" italics="true" pageId="3" pageNumber="58">cepedai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F78C5781A0122B7906AF77FBFD287FCF" box="[689,755,151,177]" pageId="3" pageNumber="58" rank="species">
<emphasis id="ECBFEAFAA0122B7906AF77FBFD287FCF" bold="true" box="[689,755,151,177]" pageId="3" pageNumber="58">n. sp.</emphasis>
</taxonomicNameLabel>
is closely related to 
<taxonomicName id="19CB4D6BA0122B7907FB77F4FE417FA4" authority="Ojanguren-Affilastro 2002" authorityName="Ojanguren-Affilastro" authorityYear="2002" class="Arachnida" family="Bothriuridae" genus="Brachistosternus" higherTaxonomySource="CoL" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="sciosciae" subGenus="Leptosternus">
<emphasis id="ECBFEAFAA0122B7907FB77F4FB777FCF" box="[997,1196,152,177]" italics="true" pageId="3" pageNumber="58">Brachistosternus</emphasis>
(
<emphasis id="ECBFEAFAA0122B7900A377F4FB167FCF" box="[1213,1229,152,177]" italics="true" pageId="3" pageNumber="58">L</emphasis>
.) 
<emphasis id="ECBFEAFAA0122B7900F877F4FA8A7FCF" box="[1254,1361,152,177]" italics="true" pageId="3" pageNumber="58">sciosciae</emphasis>
Ojanguren-Affilastro 2002
</taxonomicName>
. These two species can be distinguished by the following characters: (1) androvestigiae are present but poorly developed in males of 
<taxonomicName id="19CB4D6BA0122B7906AB7784FCAB7E7F" box="[693,880,232,258]" class="Arachnida" family="Bothriuridae" genus="Brachistosternus" higherTaxonomySource="CoL" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="cepedai" status="sp. nov." subGenus="Leptosternus">
<emphasis id="ECBFEAFAA0122B7906AB7784FD0A7E7F" box="[693,721,232,257]" italics="true" pageId="3" pageNumber="58">Br</emphasis>
. (
<emphasis id="ECBFEAFAA0122B7906F57784FD207E7F" box="[747,763,232,257]" italics="true" pageId="3" pageNumber="58">L</emphasis>
.) 
<emphasis id="ECBFEAFAA0122B79070B7784FCAB7E7F" box="[789,880,232,257]" italics="true" pageId="3" pageNumber="58">cepedai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F78C5781A0122B790767778BFC667E7F" box="[889,957,231,257]" pageId="3" pageNumber="58" rank="species">
<emphasis id="ECBFEAFAA0122B790767778BFC667E7F" bold="true" box="[889,957,231,257]" pageId="3" pageNumber="58">n. sp.</emphasis>
</taxonomicNameLabel>
whereas they are absent in males of 
<taxonomicName id="19CB4D6BA0122B7901677784FEE37E57" class="Arachnida" family="Bothriuridae" genus="Brachistosternus" higherTaxonomySource="CoL" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="sciosciae" subGenus="Leptosternus">
<emphasis id="ECBFEAFAA0122B7901677784FA4E7E7F" box="[1401,1429,232,257]" italics="true" pageId="3" pageNumber="58">Br</emphasis>
. (
<emphasis id="ECBFEAFAA0122B7904BF767CFF6A7E57" box="[161,177,272,297]" italics="true" pageId="3" pageNumber="58">L</emphasis>
.) 
<emphasis id="ECBFEAFAA0122B7904D2767CFEE37E57" box="[204,312,272,297]" italics="true" pageId="3" pageNumber="58">sciosciae</emphasis>
</taxonomicName>
. (2) Hemispermatophores of these species are morphologically different: in 
<taxonomicName id="19CB4D6BA0122B7900D1767CFA477E57" box="[1231,1436,272,298]" class="Arachnida" family="Bothriuridae" genus="Brachistosternus" higherTaxonomySource="CoL" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="sciosciae" subGenus="Leptosternus">
<emphasis id="ECBFEAFAA0122B7900D1767CFB307E57" box="[1231,1259,272,297]" italics="true" pageId="3" pageNumber="58">Br</emphasis>
. (
<emphasis id="ECBFEAFAA0122B79011B767CFACE7E57" box="[1285,1301,272,297]" italics="true" pageId="3" pageNumber="58">L</emphasis>
.) 
<emphasis id="ECBFEAFAA0122B79012E767CFA477E57" box="[1328,1436,272,297]" italics="true" pageId="3" pageNumber="58">sciosciae</emphasis>
</taxonomicName>
the distal lamina is straight and longer than the basal portion, whereas in 
<taxonomicName id="19CB4D6BA0122B7907FB7654FB407E2F" box="[997,1179,312,338]" class="Arachnida" family="Bothriuridae" genus="Brachistosternus" higherTaxonomySource="CoL" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="cepedai" status="sp. nov." subGenus="Leptosternus">
<emphasis id="ECBFEAFAA0122B7907FB7654FBDA7E2F" box="[997,1025,312,337]" italics="true" pageId="3" pageNumber="58">Br</emphasis>
. (
<emphasis id="ECBFEAFAA0122B7900077654FBF27E2F" box="[1049,1065,312,337]" italics="true" pageId="3" pageNumber="58">L</emphasis>
.) 
<emphasis id="ECBFEAFAA0122B79005E7654FB407E2F" box="[1088,1179,312,337]" italics="true" pageId="3" pageNumber="58">cepedai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F78C5781A0122B7900BC765BFB387E2F" box="[1186,1251,311,337]" pageId="3" pageNumber="58" rank="species">
<emphasis id="ECBFEAFAA0122B7900BC765BFB387E2F" bold="true" box="[1186,1251,311,337]" pageId="3" pageNumber="58">n. sp.</emphasis>
</taxonomicNameLabel>
it is curved and of the same length as the basal portion; in 
<taxonomicName id="19CB4D6BA0122B790693760CFC8C7E07" box="[653,855,352,378]" class="Arachnida" family="Bothriuridae" genus="Brachistosternus" higherTaxonomySource="CoL" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="sciosciae" subGenus="Leptosternus">
<emphasis id="ECBFEAFAA0122B790693760CFD727E07" box="[653,681,352,377]" italics="true" pageId="3" pageNumber="58">Br</emphasis>
. (
<emphasis id="ECBFEAFAA0122B7906DC760CFD097E07" box="[706,722,352,377]" italics="true" pageId="3" pageNumber="58">L</emphasis>
.) 
<emphasis id="ECBFEAFAA0122B7906F5760CFC8C7E07" box="[747,855,352,377]" italics="true" pageId="3" pageNumber="58">sciosciae</emphasis>
</taxonomicName>
the basal triangle is vestigial, whereas in 
<taxonomicName id="19CB4D6BA0122B790158760CFF297EDF" class="Arachnida" family="Bothriuridae" genus="Brachistosternus" higherTaxonomySource="CoL" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="cepedai" status="sp. nov." subGenus="Leptosternus">
<emphasis id="ECBFEAFAA0122B790158760CFAB97E07" box="[1350,1378,352,377]" italics="true" pageId="3" pageNumber="58">Br</emphasis>
. (
<emphasis id="ECBFEAFAA0122B790165760CFA507E07" box="[1403,1419,352,377]" italics="true" pageId="3" pageNumber="58">L</emphasis>
.) 
<emphasis id="ECBFEAFAA0122B79048976E4FF297EDF" box="[151,242,392,417]" italics="true" pageId="3" pageNumber="58">cepedai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F78C5781A0122B7904E776EBFEE77EDF" box="[249,316,391,417]" pageId="3" pageNumber="58" rank="species">
<emphasis id="ECBFEAFAA0122B7904E776EBFEE77EDF" bold="true" box="[249,316,391,417]" pageId="3" pageNumber="58">n. sp.</emphasis>
</taxonomicNameLabel>
it is absent or reduced to a small bulge.
</paragraph>
</subSubSection>
<subSubSection id="96D16563A0122B7C04D876C3FAFF7D9C" lastPageId="6" lastPageNumber="61" pageId="3" pageNumber="58" type="description">
<paragraph id="DE7436E8A0122B7904D876C3FF767E8C" blockId="3.[151,1436,151,498]" pageId="3" pageNumber="58">
<emphasis id="ECBFEAFAA0122B7904D876C3FEBA7EB7" bold="true" box="[198,353,431,457]" pageId="3" pageNumber="58">Description:</emphasis>
Measurements of a male specimen and a female specimen (
<typeStatus id="0170884AA0122B79003F76DCFB4E7EB4" box="[1057,1173,432,458]" pageId="3" pageNumber="58" type="paratype">paratypes</typeStatus>
) are recorded in 
<tableCitation id="93490353A0122B79014576DCFF7C7E8C" captionStart="TABLE 1" captionStartId="3.[151,239,546,570]" captionTargetBox="[151,1436,754,1976]" captionTargetPageId="3" captionText="TABLE 1. Measurements of Brachistosternus (Leptosternus) cepedai n. sp. (Paratypes) and Brachistosternus (Leptosternus) coquimbo n. sp. (Holotype and Paratype). (In mm.)." httpUri="http://table.plazi.org/id/8AB46660A0122B790489754EFCEF7D1E" pageId="3" pageNumber="58" tableUuid="8AB46660A0122B790489754EFCEF7D1E">Table 1</tableCitation>
.
</paragraph>
<caption id="8AB46660A0122B790489754EFCEF7D1E" ID-Table-UUID="8AB46660A0122B790489754EFCEF7D1E" httpUri="http://table.plazi.org/id/8AB46660A0122B790489754EFCEF7D1E" pageId="3" pageNumber="58" targetBox="[151,1436,754,1976]" targetIsTable="true" targetPageId="3">
<paragraph id="DE7436E8A0122B790489754EFCEF7D1E" blockId="3.[151,1436,546,738]" pageId="3" pageNumber="58">
<emphasis id="ECBFEAFAA0122B790489754EFED67D44" bold="true" box="[151,269,546,570]" pageId="3" pageNumber="58">TABLE 1.</emphasis>
Measurements of 
<taxonomicName id="19CB4D6BA0122B7905FE754EFC437D47" box="[480,920,546,569]" class="Arachnida" family="Bothriuridae" genus="Brachistosternus" higherTaxonomySource="CoL" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="cepedai" status="sp. nov." subGenus="Leptosternus">
<emphasis id="ECBFEAFAA0122B7905FE754EFD4E7D47" box="[480,661,546,569]" italics="true" pageId="3" pageNumber="58">Brachistosternus</emphasis>
(
<emphasis id="ECBFEAFAA0122B7906B6754EFCE97D47" box="[680,818,546,569]" italics="true" pageId="3" pageNumber="58">Leptosternus</emphasis>
) 
<emphasis id="ECBFEAFAA0122B79075B754EFC437D47" box="[837,920,546,569]" italics="true" pageId="3" pageNumber="58">cepedai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F78C5781A0122B7907BD754EFC397D44" box="[931,994,546,570]" pageId="3" pageNumber="58" rank="species">
<emphasis id="ECBFEAFAA0122B7907BD754EFC397D44" bold="true" box="[931,994,546,570]" pageId="3" pageNumber="58">n. sp.</emphasis>
</taxonomicNameLabel>
(Paratypes) and 
<taxonomicName id="19CB4D6BA0122B7900BD754EFEAA7D1F" class="Arachnida" family="Bothriuridae" genus="Brachistosternus" higherTaxonomySource="CoL" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="coquimbo" status="sp. nov." subGenus="Leptosternus">
<emphasis id="ECBFEAFAA0122B7900BD754EFA837D47" box="[1187,1368,546,569]" italics="true" pageId="3" pageNumber="58">Brachistosternus</emphasis>
(
<emphasis id="ECBFEAFAA0122B790175754EFF227D1F" italics="true" pageId="3" pageNumber="58">Leptosternus</emphasis>
) 
<emphasis id="ECBFEAFAA0122B7905167526FEAA7D1F" box="[264,369,586,609]" italics="true" pageId="3" pageNumber="58">coquimbo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F78C5781A0122B7905667525FE687D1F" box="[376,435,585,609]" pageId="3" pageNumber="58" rank="species">
<emphasis id="ECBFEAFAA0122B7905667525FE687D1F" bold="true" box="[376,435,585,609]" pageId="3" pageNumber="58">n. sp.</emphasis>
</taxonomicNameLabel>
(Holotype and Paratype). (In mm.).
</paragraph>
</caption>
<paragraph id="DE7436E8A0122B79063D7511FB577DEA" blockId="3.[151,1436,546,738]" box="[547,1164,637,660]" pageId="3" pageNumber="58">
<taxonomicName id="19CB4D6BA0122B79063D7511FD117DEA" box="[547,714,637,660]" class="Arachnida" family="Bothriuridae" genus="Brachistosternus" higherTaxonomySource="CoL" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="cepedai" subGenus="Leptosternus">
<emphasis id="ECBFEAFAA0122B79063D7511FDE77DEA" box="[547,572,637,660]" italics="true" pageId="3" pageNumber="58">Br</emphasis>
. (
<emphasis id="ECBFEAFAA0122B79064C7511FDBA7DEA" box="[594,609,637,660]" italics="true" pageId="3" pageNumber="58">L</emphasis>
.) 
<emphasis id="ECBFEAFAA0122B7906697511FD117DEA" box="[631,714,637,660]" italics="true" pageId="3" pageNumber="58">cepedai</emphasis>
</taxonomicName>
<taxonomicName id="19CB4D6BA0122B7907D17511FB577DEA" box="[975,1164,637,660]" class="Arachnida" family="Bothriuridae" genus="Brachistosternus" higherTaxonomySource="CoL" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="coquimbo" subGenus="Leptosternus">
Br. (
<emphasis id="ECBFEAFAA0122B7907E07511FBD67DEA" box="[1022,1037,637,660]" italics="true" pageId="3" pageNumber="58">L</emphasis>
.) 
<emphasis id="ECBFEAFAA0122B79003D7511FB577DEA" box="[1059,1164,637,660]" italics="true" pageId="3" pageNumber="58">coquimbo</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="DE7436E8A0122B79063D75C9FAFB7DC9" blockId="3.[151,1436,546,738]" box="[547,1312,675,700]" pageId="3" pageNumber="58">
<typeStatus id="0170884AA0122B79063D75C9FD5B7DC2" box="[547,640,677,700]" pageId="3" pageNumber="58" type="paratype">Paratype</typeStatus>
ɗ 
<typeStatus id="0170884AA0122B7906E275C9FC827DC2" box="[764,857,677,700]" pageId="3" pageNumber="58" type="paratype">Paratype</typeStatus>
Ψ 
<typeStatus id="0170884AA0122B7907D175C9FBE97DC2" box="[975,1074,677,700]" pageId="3" pageNumber="58" type="holotype">Holotype</typeStatus>
ɗ 
<typeStatus id="0170884AA0122B7900B675C9FADE7DC2" box="[1192,1285,677,700]" pageId="3" pageNumber="58" type="paratype">Paratype</typeStatus>
Ψ
</paragraph>
<paragraph id="DE7436E8A0122B7F063475A7FC427D64" blockId="3.[151,1436,546,738]" lastBlockId="5.[151,1437,152,2018]" lastPageId="5" lastPageNumber="60" pageId="3" pageNumber="58">
LEULS LEULS MACN-Ar MACN-Ar 
<emphasis id="ECBFEAFAA0152B7E04D870EBFECC78DF" bold="true" box="[198,279,1927,1953]" pageId="4" pageNumber="59">Color:</emphasis>
Light yellow, with some faint spots on the carapace and the tergites. Chelicerae: not pigmented. Carapace: median ocular tubercle and area around lateral ocelli infuscated; without pigmentation except for a slight reticular pattern around the ocular tubercle. Tergites: with three faint spots, two laterals and one median; the median spot darker than the lateral ones, (in poorly pigmented specimens the only visible spot of the tergites). Sternites: not pigmented. Metasoma: segments I–III, dorsal surfaces each with an antero-median narrow stripe, and two posterolateral dark spots, lateral and ventral surfaces not pigmented, ventral surface with a narrow VM stripe, that is absent in poorly pigmented specimens; segment IV, dorsal surface with an anteromedian narrow stripe, lateral surfaces unpigmented, ventral surfaces with a narrow VM stripe; segment V, dorsal and lateral surfaces similar to segment IV, ventral surface with a narrow VM stripe and with two narrow VL stripes, that do not join with the VM stripe. Telson: vesicle not pigmented; aculeus dark brown. Pedipalps: femur slightly pigmented on the posterior margin, and near the articulation with the patella; patella slightly pigmented on anterior and posterior margins; chela not pigmented. Legs: femur slightly pigmented near articulation with patella; remaining segments not pigmented.
</paragraph>
<paragraph id="DE7436E8A0122B790481759EFB0C78C6" pageId="3" pageNumber="58">
<table id="ACCBC448A012D4850489759EFA4778C6" box="[151,1436,754,1976]" gridcols="4" gridrows="32" pageId="3" pageNumber="58">
<tr id="60FB34AAA012D4850489759EFA477C77" box="[151,1436,754,777]" gridrow="0" pageId="3" pageNumber="58">
<th id="232A5DD6A012D4850489759EFD847C77" box="[151,607,754,777]" gridcol="0" gridrow="0" pageId="3" pageNumber="58">Total length 42.2</th>
<th id="232A5DD6A012D48506E2759EFCE37C77" box="[764,824,754,777]" gridcol="1" gridrow="0" pageId="3" pageNumber="58">40.52</th>
<th id="232A5DD6A012D48507D1759EFBD07C77" box="[975,1035,754,777]" gridcol="2" gridrow="0" pageId="3" pageNumber="58">49.23</th>
<th id="232A5DD6A012D48500B6759EFA477C77" box="[1192,1436,754,777]" gridcol="3" gridrow="0" pageId="3" pageNumber="58">39.67</th>
</tr>
<tr id="60FB34AAA012D48504897475FA477C4E" box="[151,1436,793,816]" gridrow="1" pageId="3" pageNumber="58">
<th id="232A5DD6A012D48504897475FD847C4E" box="[151,607,793,816]" gridcol="0" gridrow="1" pageId="3" pageNumber="58">Carapace, length 5.25</th>
<td id="232A5DD6A012D48506E27475FCE37C4E" box="[764,824,793,816]" gridcol="1" gridrow="1" pageId="3" pageNumber="58">4.85</td>
<td id="232A5DD6A012D48507D17475FBD07C4E" box="[975,1035,793,816]" gridcol="2" gridrow="1" pageId="3" pageNumber="58">5.98</td>
<td id="232A5DD6A012D48500B67475FA477C4E" box="[1192,1436,793,816]" gridcol="3" gridrow="1" pageId="3" pageNumber="58">5.49</td>
</tr>
<tr id="60FB34AAA012D4850489742CFA477C29" box="[151,1436,832,855]" gridrow="2" pageId="3" pageNumber="58">
<th id="232A5DD6A012D4850489742CFD847C29" box="[151,607,832,855]" gridcol="0" gridrow="2" pageId="3" pageNumber="58">Carapace, anterior width 3.15</th>
<td id="232A5DD6A012D48506E2742CFCE37C29" box="[764,824,832,855]" gridcol="1" gridrow="2" pageId="3" pageNumber="58">3.23</td>
<td id="232A5DD6A012D48507D1742CFBD07C29" box="[975,1035,832,855]" gridcol="2" gridrow="2" pageId="3" pageNumber="58">4.04</td>
<td id="232A5DD6A012D48500B6742CFA477C29" box="[1192,1436,832,855]" gridcol="3" gridrow="2" pageId="3" pageNumber="58">4.04</td>
</tr>
<tr id="60FB34AAA012D4850489740AFA477C03" box="[151,1436,870,893]" gridrow="3" pageId="3" pageNumber="58">
<th id="232A5DD6A012D4850489740AFD847C03" box="[151,607,870,893]" gridcol="0" gridrow="3" pageId="3" pageNumber="58">Carapace, posterior width 4.93</th>
<td id="232A5DD6A012D48506E2740AFCE37C03" box="[764,824,870,893]" gridcol="1" gridrow="3" pageId="3" pageNumber="58">5.09</td>
<td id="232A5DD6A012D48507D1740AFBD07C03" box="[975,1035,870,893]" gridcol="2" gridrow="3" pageId="3" pageNumber="58">5.98</td>
<td id="232A5DD6A012D48500B6740AFA477C03" box="[1192,1436,870,893]" gridcol="3" gridrow="3" pageId="3" pageNumber="58">6.87</td>
</tr>
<tr id="60FB34AAA012D485048974E1FA477CDA" box="[151,1436,909,932]" gridrow="4" pageId="3" pageNumber="58">
<th id="232A5DD6A012D485048974E1FD847CDA" box="[151,607,909,932]" gridcol="0" gridrow="4" pageId="3" pageNumber="58">Mesosoma, total length 12.38</th>
<td id="232A5DD6A012D48506E274E1FCE37CDA" box="[764,824,909,932]" gridcol="1" gridrow="4" pageId="3" pageNumber="58">14.67</td>
<td id="232A5DD6A012D48507D174E1FBD07CDA" box="[975,1035,909,932]" gridcol="2" gridrow="4" pageId="3" pageNumber="58">12.23</td>
<td id="232A5DD6A012D48500B674E1FA477CDA" box="[1192,1436,909,932]" gridcol="3" gridrow="4" pageId="3" pageNumber="58">9.78</td>
</tr>
<tr id="60FB34AAA012D485048974D8FA477CB5" box="[151,1436,948,971]" gridrow="5" pageId="3" pageNumber="58">
<th id="232A5DD6A012D485048974D8FD847CB5" box="[151,607,948,971]" gridcol="0" gridrow="5" pageId="3" pageNumber="58">Metasoma, total length 19.47</th>
<td id="232A5DD6A012D48506E274D8FCE37CB5" box="[764,824,948,971]" gridcol="1" gridrow="5" pageId="3" pageNumber="58">16.15</td>
<td id="232A5DD6A012D48507D174D8FBD07CB5" box="[975,1035,948,971]" gridcol="2" gridrow="5" pageId="3" pageNumber="58">24.15</td>
<td id="232A5DD6A012D48500B674D8FA477CB5" box="[1192,1436,948,971]" gridcol="3" gridrow="5" pageId="3" pageNumber="58">18.74</td>
</tr>
<tr id="60FB34AAA012D485048974B6FA477C8F" box="[151,1436,986,1009]" gridrow="6" pageId="3" pageNumber="58">
<th id="232A5DD6A012D485048974B6FD847C8F" box="[151,607,986,1009]" gridcol="0" gridrow="6" pageId="3" pageNumber="58">Metasomal segment I, length 3.31</th>
<td id="232A5DD6A012D48506E274B6FCE37C8F" box="[764,824,986,1009]" gridcol="1" gridrow="6" pageId="3" pageNumber="58">2.83</td>
<td id="232A5DD6A012D48507D174B6FBD07C8F" box="[975,1035,986,1009]" gridcol="2" gridrow="6" pageId="3" pageNumber="58">4.04</td>
<td id="232A5DD6A012D48500B674B6FA477C8F" box="[1192,1436,986,1009]" gridcol="3" gridrow="6" pageId="3" pageNumber="58">2.99</td>
</tr>
<tr id="60FB34AAA012D4850489736DFA477B66" box="[151,1436,1025,1048]" gridrow="7" pageId="3" pageNumber="58">
<th id="232A5DD6A012D4850489736DFD847B66" box="[151,607,1025,1048]" gridcol="0" gridrow="7" pageId="3" pageNumber="58">Metasomal segment I, width 3.23</th>
<td id="232A5DD6A012D48506E2736DFCE37B66" box="[764,824,1025,1048]" gridcol="1" gridrow="7" pageId="3" pageNumber="58">2.75</td>
<td id="232A5DD6A012D48507D1736DFBD07B66" box="[975,1035,1025,1048]" gridcol="2" gridrow="7" pageId="3" pageNumber="58">3.79</td>
<td id="232A5DD6A012D48500B6736DFA477B66" box="[1192,1436,1025,1048]" gridcol="3" gridrow="7" pageId="3" pageNumber="58">3.34</td>
</tr>
<tr id="60FB34AAA012D48504897344FA477B41" box="[151,1436,1064,1087]" gridrow="8" pageId="3" pageNumber="58">
<th id="232A5DD6A012D48504897344FD847B41" box="[151,607,1064,1087]" gridcol="0" gridrow="8" pageId="3" pageNumber="58">Metasomal segment I, height 2.50</th>
<td id="232A5DD6A012D48506E27344FCE37B41" box="[764,824,1064,1087]" gridcol="1" gridrow="8" pageId="3" pageNumber="58">2.18</td>
<td id="232A5DD6A012D48507D17344FBD07B41" box="[975,1035,1064,1087]" gridcol="2" gridrow="8" pageId="3" pageNumber="58">2.99</td>
<td id="232A5DD6A012D48500B67344FA477B41" box="[1192,1436,1064,1087]" gridcol="3" gridrow="8" pageId="3" pageNumber="58">2.75</td>
</tr>
<tr id="60FB34AAA012D48504897322FA477B1B" box="[151,1436,1102,1125]" gridrow="9" pageId="3" pageNumber="58">
<th id="232A5DD6A012D48504897322FD847B1B" box="[151,607,1102,1125]" gridcol="0" gridrow="9" pageId="3" pageNumber="58">Metasomal segment II, length 3.64</th>
<td id="232A5DD6A012D48506E27322FCE37B1B" box="[764,824,1102,1125]" gridcol="1" gridrow="9" pageId="3" pageNumber="58">2.99</td>
<td id="232A5DD6A012D48507D17322FBD07B1B" box="[975,1035,1102,1125]" gridcol="2" gridrow="9" pageId="3" pageNumber="58">4.53</td>
<td id="232A5DD6A012D48500B67322FA477B1B" box="[1192,1436,1102,1125]" gridcol="3" gridrow="9" pageId="3" pageNumber="58">3.23</td>
</tr>
<tr id="60FB34AAA012D48504897319FA477BF2" box="[151,1436,1141,1164]" gridrow="10" pageId="3" pageNumber="58">
<th id="232A5DD6A012D48504897319FD847BF2" box="[151,607,1141,1164]" gridcol="0" gridrow="10" pageId="3" pageNumber="58">Metasomal segment II, width 3.07</th>
<td id="232A5DD6A012D48506E27319FCE37BF2" box="[764,824,1141,1164]" gridcol="1" gridrow="10" pageId="3" pageNumber="58">2.74</td>
<td id="232A5DD6A012D48507D17319FBD07BF2" box="[975,1035,1141,1164]" gridcol="2" gridrow="10" pageId="3" pageNumber="58">3.47</td>
<td id="232A5DD6A012D48500B67319FA477BF2" box="[1192,1436,1141,1164]" gridcol="3" gridrow="10" pageId="3" pageNumber="58">3.07</td>
</tr>
<tr id="60FB34AAA012D485048973F0FA477BCD" box="[151,1436,1180,1203]" gridrow="11" pageId="3" pageNumber="58">
<th id="232A5DD6A012D485048973F0FD847BCD" box="[151,607,1180,1203]" gridcol="0" gridrow="11" pageId="3" pageNumber="58">Metasomal segment II, height 2.51</th>
<td id="232A5DD6A012D48506E273F0FCE37BCD" box="[764,824,1180,1203]" gridcol="1" gridrow="11" pageId="3" pageNumber="58">2.18</td>
<td id="232A5DD6A012D48507D173F0FBD07BCD" box="[975,1035,1180,1203]" gridcol="2" gridrow="11" pageId="3" pageNumber="58">2.91</td>
<td id="232A5DD6A012D48500B673F0FA477BCD" box="[1192,1436,1180,1203]" gridcol="3" gridrow="11" pageId="3" pageNumber="58">2.67</td>
</tr>
<tr id="60FB34AAA012D485048973AEFA477BA7" box="[151,1436,1218,1241]" gridrow="12" pageId="3" pageNumber="58">
<th id="232A5DD6A012D485048973AEFD847BA7" box="[151,607,1218,1241]" gridcol="0" gridrow="12" pageId="3" pageNumber="58">Metasomal segment III, length 3.71</th>
<td id="232A5DD6A012D48506E273AEFCE37BA7" box="[764,824,1218,1241]" gridcol="1" gridrow="12" pageId="3" pageNumber="58">3.15</td>
<td id="232A5DD6A012D48507D173AEFBD07BA7" box="[975,1035,1218,1241]" gridcol="2" gridrow="12" pageId="3" pageNumber="58">4.68</td>
<td id="232A5DD6A012D48500B673AEFA477BA7" box="[1192,1436,1218,1241]" gridcol="3" gridrow="12" pageId="3" pageNumber="58">3.47</td>
</tr>
<tr id="60FB34AAA012D48504897385FA477A7E" box="[151,1436,1257,1280]" gridrow="13" pageId="3" pageNumber="58">
<th id="232A5DD6A012D48504897385FD847A7E" box="[151,607,1257,1280]" gridcol="0" gridrow="13" pageId="3" pageNumber="58">Metasomal segment III, width 2.91</th>
<td id="232A5DD6A012D48506E27385FCE37A7E" box="[764,824,1257,1280]" gridcol="1" gridrow="13" pageId="3" pageNumber="58">2.42</td>
<td id="232A5DD6A012D48507D17385FBD07A7E" box="[975,1035,1257,1280]" gridcol="2" gridrow="13" pageId="3" pageNumber="58">3.31</td>
<td id="232A5DD6A012D48500B67385FA477A7E" box="[1192,1436,1257,1280]" gridcol="3" gridrow="13" pageId="3" pageNumber="58">2.91</td>
</tr>
<tr id="60FB34AAA012D4850489727CFA477A59" box="[151,1436,1296,1319]" gridrow="14" pageId="3" pageNumber="58">
<th id="232A5DD6A012D4850489727CFD847A59" box="[151,607,1296,1319]" gridcol="0" gridrow="14" pageId="3" pageNumber="58">Metasomal segment III, height 2.50</th>
<td id="232A5DD6A012D48506E2727CFCE37A59" box="[764,824,1296,1319]" gridcol="1" gridrow="14" pageId="3" pageNumber="58">2.02</td>
<td id="232A5DD6A012D48507D1727CFBD07A59" box="[975,1035,1296,1319]" gridcol="2" gridrow="14" pageId="3" pageNumber="58">2.83</td>
<td id="232A5DD6A012D48500B6727CFA477A59" box="[1192,1436,1296,1319]" gridcol="3" gridrow="14" pageId="3" pageNumber="58">2.66</td>
</tr>
<tr id="60FB34AAA012D4850489725AFA477A33" box="[151,1436,1334,1357]" gridrow="15" pageId="3" pageNumber="58">
<th id="232A5DD6A012D4850489725AFD847A33" box="[151,607,1334,1357]" gridcol="0" gridrow="15" pageId="3" pageNumber="58">Metasomal segment IV, length 4.04</th>
<td id="232A5DD6A012D48506E2725AFCE37A33" box="[764,824,1334,1357]" gridcol="1" gridrow="15" pageId="3" pageNumber="58">3.39</td>
<td id="232A5DD6A012D48507D1725AFBD07A33" box="[975,1035,1334,1357]" gridcol="2" gridrow="15" pageId="3" pageNumber="58">5.25</td>
<td id="232A5DD6A012D48500B6725AFA477A33" box="[1192,1436,1334,1357]" gridcol="3" gridrow="15" pageId="3" pageNumber="58">4.04</td>
</tr>
<tr id="60FB34AAA012D48504897231FA477A0A" box="[151,1436,1373,1396]" gridrow="16" pageId="3" pageNumber="58">
<th id="232A5DD6A012D48504897231FD847A0A" box="[151,607,1373,1396]" gridcol="0" gridrow="16" pageId="3" pageNumber="58">Metasomal segment IV, width 2.58</th>
<td id="232A5DD6A012D48506E27231FCE37A0A" box="[764,824,1373,1396]" gridcol="1" gridrow="16" pageId="3" pageNumber="58">2.26</td>
<td id="232A5DD6A012D48507D17231FBD07A0A" box="[975,1035,1373,1396]" gridcol="2" gridrow="16" pageId="3" pageNumber="58">2.99</td>
<td id="232A5DD6A012D48500B67231FA477A0A" box="[1192,1436,1373,1396]" gridcol="3" gridrow="16" pageId="3" pageNumber="58">2.70</td>
</tr>
<tr id="60FB34AAA012D485048972E8FA477AE5" box="[151,1436,1412,1435]" gridrow="17" pageId="3" pageNumber="58">
<th id="232A5DD6A012D485048972E8FD847AE5" box="[151,607,1412,1435]" gridcol="0" gridrow="17" pageId="3" pageNumber="58">Metasomal segment IV, height 2.18</th>
<td id="232A5DD6A012D48506E272E8FCE37AE5" box="[764,824,1412,1435]" gridcol="1" gridrow="17" pageId="3" pageNumber="58">1.77</td>
<td id="232A5DD6A012D48507D172E8FBD07AE5" box="[975,1035,1412,1435]" gridcol="2" gridrow="17" pageId="3" pageNumber="58">2.67</td>
<td id="232A5DD6A012D48500B672E8FA477AE5" box="[1192,1436,1412,1435]" gridcol="3" gridrow="17" pageId="3" pageNumber="58">2.42</td>
</tr>
<tr id="60FB34AAA012D485048972C6FA477ABF" box="[151,1436,1450,1473]" gridrow="18" pageId="3" pageNumber="58">
<th id="232A5DD6A012D485048972C6FD847ABF" box="[151,607,1450,1473]" gridcol="0" gridrow="18" pageId="3" pageNumber="58">Metasomal segment V, length 4.77</th>
<td id="232A5DD6A012D48506E272C6FCE37ABF" box="[764,824,1450,1473]" gridcol="1" gridrow="18" pageId="3" pageNumber="58">3.79</td>
<td id="232A5DD6A012D48507D172C6FBD07ABF" box="[975,1035,1450,1473]" gridcol="2" gridrow="18" pageId="3" pageNumber="58">5.65</td>
<td id="232A5DD6A012D48500B672C6FA477ABF" box="[1192,1436,1450,1473]" gridcol="3" gridrow="18" pageId="3" pageNumber="58">5.01</td>
</tr>
<tr id="60FB34AAA012D485048972BDFA477A96" box="[151,1436,1489,1512]" gridrow="19" pageId="3" pageNumber="58">
<th id="232A5DD6A012D485048972BDFD847A96" box="[151,607,1489,1512]" gridcol="0" gridrow="19" pageId="3" pageNumber="58">Metasomal segment V, width 2.67</th>
<td id="232A5DD6A012D48506E272BDFCE37A96" box="[764,824,1489,1512]" gridcol="1" gridrow="19" pageId="3" pageNumber="58">2.26</td>
<td id="232A5DD6A012D48507D172BDFBD07A96" box="[975,1035,1489,1512]" gridcol="2" gridrow="19" pageId="3" pageNumber="58">3.07</td>
<td id="232A5DD6A012D48500B672BDFA477A96" box="[1192,1436,1489,1512]" gridcol="3" gridrow="19" pageId="3" pageNumber="58">2.70</td>
</tr>
<tr id="60FB34AAA012D48504897294FA477971" box="[151,1436,1528,1551]" gridrow="20" pageId="3" pageNumber="58">
<th id="232A5DD6A012D48504897294FD847971" box="[151,607,1528,1551]" gridcol="0" gridrow="20" pageId="3" pageNumber="58">Metasomal segment V, height 2.02</th>
<td id="232A5DD6A012D48506E27294FCE37971" box="[764,824,1528,1551]" gridcol="1" gridrow="20" pageId="3" pageNumber="58">1.61</td>
<td id="232A5DD6A012D48507D17294FBD07971" box="[975,1035,1528,1551]" gridcol="2" gridrow="20" pageId="3" pageNumber="58">2.50</td>
<td id="232A5DD6A012D48500B67294FA477971" box="[1192,1436,1528,1551]" gridcol="3" gridrow="20" pageId="3" pageNumber="58">2.26</td>
</tr>
<tr id="60FB34AAA012D48504897172FA47794B" box="[151,1436,1566,1589]" gridrow="21" pageId="3" pageNumber="58">
<th id="232A5DD6A012D48504897172FD84794B" box="[151,607,1566,1589]" gridcol="0" gridrow="21" pageId="3" pageNumber="58">Telson, length 5.01</th>
<td id="232A5DD6A012D48506E27172FCE3794B" box="[764,824,1566,1589]" gridcol="1" gridrow="21" pageId="3" pageNumber="58">4.85</td>
<td id="232A5DD6A012D48507D17172FBD0794B" box="[975,1035,1566,1589]" gridcol="2" gridrow="21" pageId="3" pageNumber="58">6.87</td>
<td id="232A5DD6A012D48500B67172FA47794B" box="[1192,1436,1566,1589]" gridcol="3" gridrow="21" pageId="3" pageNumber="58">5.66</td>
</tr>
<tr id="60FB34AAA012D48504897129FA477922" box="[151,1436,1605,1628]" gridrow="22" pageId="3" pageNumber="58">
<th id="232A5DD6A012D48504897129FD847922" box="[151,607,1605,1628]" gridcol="0" gridrow="22" pageId="3" pageNumber="58">Vesicle, length 2.42</th>
<td id="232A5DD6A012D48506E27129FCE37922" box="[764,824,1605,1628]" gridcol="1" gridrow="22" pageId="3" pageNumber="58">2.26</td>
<td id="232A5DD6A012D48507D17129FBD07922" box="[975,1035,1605,1628]" gridcol="2" gridrow="22" pageId="3" pageNumber="58">4.04</td>
<td id="232A5DD6A012D48500B67129FA477922" box="[1192,1436,1605,1628]" gridcol="3" gridrow="22" pageId="3" pageNumber="58">2.99</td>
</tr>
<tr id="60FB34AAA012D48504897100FA4779FD" box="[151,1436,1644,1667]" gridrow="23" pageId="3" pageNumber="58">
<th id="232A5DD6A012D48504897100FD8479FD" box="[151,607,1644,1667]" gridcol="0" gridrow="23" pageId="3" pageNumber="58">Vesicle, width 1.77</th>
<td id="232A5DD6A012D48506E27100FCE379FD" box="[764,824,1644,1667]" gridcol="1" gridrow="23" pageId="3" pageNumber="58">1.69</td>
<td id="232A5DD6A012D48507D17100FBD079FD" box="[975,1035,1644,1667]" gridcol="2" gridrow="23" pageId="3" pageNumber="58">2.30</td>
<td id="232A5DD6A012D48500B67100FA4779FD" box="[1192,1436,1644,1667]" gridcol="3" gridrow="23" pageId="3" pageNumber="58">2.10</td>
</tr>
<tr id="60FB34AAA012D485048971FEFA4779D7" box="[151,1436,1682,1705]" gridrow="24" pageId="3" pageNumber="58">
<th id="232A5DD6A012D485048971FEFD8479D7" box="[151,607,1682,1705]" gridcol="0" gridrow="24" pageId="3" pageNumber="58">Vesicle, height 1.61</th>
<td id="232A5DD6A012D48506E271FEFCE379D7" box="[764,824,1682,1705]" gridcol="1" gridrow="24" pageId="3" pageNumber="58">1.54</td>
<td id="232A5DD6A012D48507D171FEFBD079D7" box="[975,1035,1682,1705]" gridcol="2" gridrow="24" pageId="3" pageNumber="58">2.18</td>
<td id="232A5DD6A012D48500B671FEFA4779D7" box="[1192,1436,1682,1705]" gridcol="3" gridrow="24" pageId="3" pageNumber="58">1.78</td>
</tr>
<tr id="60FB34AAA012D485048971D5FA4779AE" box="[151,1436,1721,1744]" gridrow="25" pageId="3" pageNumber="58">
<th id="232A5DD6A012D485048971D5FD8479AE" box="[151,607,1721,1744]" gridcol="0" gridrow="25" pageId="3" pageNumber="58">Aculeus, length 2.58</th>
<td id="232A5DD6A012D48506E271D5FCE379AE" box="[764,824,1721,1744]" gridcol="1" gridrow="25" pageId="3" pageNumber="58">2.58</td>
<td id="232A5DD6A012D48507D171D5FBD079AE" box="[975,1035,1721,1744]" gridcol="2" gridrow="25" pageId="3" pageNumber="58">2.83</td>
<td id="232A5DD6A012D48500B671D5FA4779AE" box="[1192,1436,1721,1744]" gridcol="3" gridrow="25" pageId="3" pageNumber="58">2.66</td>
</tr>
<tr id="60FB34AAA012D4850489718CFA477989" box="[151,1436,1760,1783]" gridrow="26" pageId="3" pageNumber="58">
<th id="232A5DD6A012D4850489718CFD847989" box="[151,607,1760,1783]" gridcol="0" gridrow="26" pageId="3" pageNumber="58">Femur, length 3.64</th>
<td id="232A5DD6A012D48506E2718CFCE37989" box="[764,824,1760,1783]" gridcol="1" gridrow="26" pageId="3" pageNumber="58">3.07</td>
<td id="232A5DD6A012D48507D1718CFBD07989" box="[975,1035,1760,1783]" gridcol="2" gridrow="26" pageId="3" pageNumber="58">5.33</td>
<td id="232A5DD6A012D48500B6718CFA477989" box="[1192,1436,1760,1783]" gridcol="3" gridrow="26" pageId="3" pageNumber="58">4.00</td>
</tr>
<tr id="60FB34AAA012D4850489706AFA477863" box="[151,1436,1798,1821]" gridrow="27" pageId="3" pageNumber="58">
<th id="232A5DD6A012D4850489706AFD847863" box="[151,607,1798,1821]" gridcol="0" gridrow="27" pageId="3" pageNumber="58">Femur, width 1.13</th>
<td id="232A5DD6A012D48506E2706AFCE37863" box="[764,824,1798,1821]" gridcol="1" gridrow="27" pageId="3" pageNumber="58">1.21</td>
<td id="232A5DD6A012D48507D1706AFBD07863" box="[975,1035,1798,1821]" gridcol="2" gridrow="27" pageId="3" pageNumber="58">1.29</td>
<td id="232A5DD6A012D48500B6706AFA477863" box="[1192,1436,1798,1821]" gridcol="3" gridrow="27" pageId="3" pageNumber="58">1.37</td>
</tr>
<tr id="60FB34AAA012D48504897041FA47783A" box="[151,1436,1837,1860]" gridrow="28" pageId="3" pageNumber="58">
<th id="232A5DD6A012D48504897041FD84783A" box="[151,607,1837,1860]" gridcol="0" gridrow="28" pageId="3" pageNumber="58">Patella, length 3.23</th>
<td id="232A5DD6A012D48506E27041FCE3783A" box="[764,824,1837,1860]" gridcol="1" gridrow="28" pageId="3" pageNumber="58">3.23</td>
<td id="232A5DD6A012D48507D17041FBD0783A" box="[975,1035,1837,1860]" gridcol="2" gridrow="28" pageId="3" pageNumber="58">4.60</td>
<td id="232A5DD6A012D48500B67041FA47783A" box="[1192,1436,1837,1860]" gridcol="3" gridrow="28" pageId="3" pageNumber="58">3.88</td>
</tr>
<tr id="60FB34AAA012D48504897038FA477815" box="[151,1436,1876,1899]" gridrow="29" pageId="3" pageNumber="58">
<th id="232A5DD6A012D48504897038FD847815" box="[151,607,1876,1899]" gridcol="0" gridrow="29" pageId="3" pageNumber="58">Patella, width 1.45</th>
<td id="232A5DD6A012D48506E27038FCE37815" box="[764,824,1876,1899]" gridcol="1" gridrow="29" pageId="3" pageNumber="58">1.37</td>
<td id="232A5DD6A012D48507D17038FBD07815" box="[975,1035,1876,1899]" gridcol="2" gridrow="29" pageId="3" pageNumber="58">1.69</td>
<td id="232A5DD6A012D48500B67038FA477815" box="[1192,1436,1876,1899]" gridcol="3" gridrow="29" pageId="3" pageNumber="58">1.65</td>
</tr>
<tr id="60FB34AAA012D48504897016FA4778EF" box="[151,1436,1914,1937]" gridrow="30" pageId="3" pageNumber="58">
<th id="232A5DD6A012D48504897016FD8478EF" box="[151,607,1914,1937]" gridcol="0" gridrow="30" pageId="3" pageNumber="58">Chela, length 6.2</th>
<td id="232A5DD6A012D48506E27016FCE378EF" box="[764,824,1914,1937]" gridcol="1" gridrow="30" pageId="3" pageNumber="58">5.74</td>
<td id="232A5DD6A012D48507D17016FBD078EF" box="[975,1035,1914,1937]" gridcol="2" gridrow="30" pageId="3" pageNumber="58">8.16</td>
<td id="232A5DD6A012D48500B67016FA4778EF" box="[1192,1436,1914,1937]" gridcol="3" gridrow="30" pageId="3" pageNumber="58">14.8</td>
</tr>
<tr id="60FB34AAA012D485048970CDFA4778C6" box="[151,1436,1953,1976]" gridrow="31" pageId="3" pageNumber="58">
<th id="232A5DD6A012D485048970CDFD8478C6" box="[151,607,1953,1976]" gridcol="0" gridrow="31" pageId="3" pageNumber="58">Chela, width 1.65</th>
<td id="232A5DD6A012D48506E270CDFCE378C6" box="[764,824,1953,1976]" gridcol="1" gridrow="31" pageId="3" pageNumber="58">1.54</td>
<td id="232A5DD6A012D48507D170CDFBD078C6" box="[975,1035,1953,1976]" gridcol="2" gridrow="31" pageId="3" pageNumber="58">1.61</td>
<td id="232A5DD6A012D48500B670CDFA4778C6" box="[1192,1436,1953,1976]" gridcol="3" gridrow="31" pageId="3" pageNumber="58">3.42</td>
</tr>
</table>
</paragraph>
<caption id="8AB46660A0152B7E048971DEFB4A784F" httpUri="https://zenodo.org/record/179244/files/figure.png" pageId="4" pageNumber="59" targetBox="[280,1321,250,1688]" targetPageId="4">
<paragraph id="DE7436E8A0152B7E048971DEFB4A784F" blockId="4.[151,1436,1714,1841]" pageId="4" pageNumber="59">
<emphasis id="ECBFEAFAA0152B7E048971DEFE8A79B4" bold="true" box="[151,337,1714,1738]" pageId="4" pageNumber="59">FIGURES 2–10.</emphasis>
<taxonomicName id="19CB4D6BA0152B7E054771DEFD4379B7" box="[345,664,1714,1737]" class="Arachnida" family="Bothriuridae" genus="Brachistosternus" higherTaxonomySource="CoL" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="59" phylum="Arthropoda" rank="species" species="cepedai" status="sp. nov." subGenus="Leptosternus">
<emphasis id="ECBFEAFAA0152B7E054771DEFDD579B7" box="[345,526,1714,1737]" italics="true" pageId="4" pageNumber="59">Brachistosternus</emphasis>
(
<emphasis id="ECBFEAFAA0152B7E060171DEFDF579B7" box="[543,558,1714,1737]" italics="true" pageId="4" pageNumber="59">L</emphasis>
.) 
<emphasis id="ECBFEAFAA0152B7E065B71DEFD4379B7" box="[581,664,1714,1737]" italics="true" pageId="4" pageNumber="59">cepedai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F78C5781A0152B7E06BE71DEFD0579B4" box="[672,734,1714,1738]" pageId="4" pageNumber="59" rank="species">
<emphasis id="ECBFEAFAA0152B7E06BE71DEFD0579B4" bold="true" box="[672,734,1714,1738]" pageId="4" pageNumber="59">n. sp.</emphasis>
</taxonomicNameLabel>
2. left hemispermatophore, ventral aspect; 3. left hemispermatophore, dorsal aspect; 4. metasomal segment V, male, dorsal aspect; 5. metasomal segment V, male, ventral aspect; 6. telson, male, lateral aspect; 7. telson, female, lateral aspect; 8. right pedipalp chela, male, ventral aspect; 9. right pedipalp chela, male, internal aspect; 10. left pedipalp chela, female, external aspect. Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="DE7436E8A0142B7C04D8754BFCDC7EDC" blockId="5.[151,1437,152,2018]" lastBlockId="6.[151,1437,152,1138]" lastPageId="6" lastPageNumber="61" pageId="5" pageNumber="60">
<emphasis id="ECBFEAFAA0142B7F04D8754BFEBC7D3F" bold="true" box="[198,359,551,577]" pageId="5" pageNumber="60">Morphology.</emphasis>
Carapace: anterior margin convex with a slight median projection; anterior longitudinal sulcus well developed; ocular tubercle well developed, situated anteromedially, interocular sulcus weakly developed, median ocelli almost two diameters apart; three pairs of lateral ocelli, considerably smaller than the median ones; anterior and posterior longitudinal sulci, lateral sulcus and postocular furrow well developed; carapace lateral surfaces slightly granular, smooth near the ocular tubercle and the anterior longitudinal sulcus. Chelicerae with two subdistal teeth. Hemispermatophore: cylindrical apophysis vestigial (
<figureCitation id="46F02A6DA0142B7F00E4759CFA9F7C74" box="[1274,1348,752,778]" captionStart="FIGURES 2 – 10" captionStartId="4.[151,269,1714,1738]" captionTargetBox="[280,1321,250,1688]" captionTargetId="figure@4.[264,1322,194,1690]" captionTargetPageId="4" captionText="FIGURES 2 – 10. Brachistosternus (L.) cepedai n. sp. 2. left hemispermatophore, ventral aspect; 3. left hemispermatophore, dorsal aspect; 4. metasomal segment V, male, dorsal aspect; 5. metasomal segment V, male, ventral aspect; 6. telson, male, lateral aspect; 7. telson, female, lateral aspect; 8. right pedipalp chela, male, ventral aspect; 9. right pedipalp chela, male, internal aspect; 10. left pedipalp chela, female, external aspect. Scale bars = 1 mm." httpUri="https://zenodo.org/record/179244/files/figure.png" pageId="5" pageNumber="60">Fig. 2</figureCitation>
); distal lamina narrow, curved, similar in size to, or slightly longer than the basal portion (
<figureCitation id="46F02A6DA0142B7F008A7474FB047C4C" box="[1172,1247,792,818]" captionStart="FIGURES 2 – 10" captionStartId="4.[151,269,1714,1738]" captionTargetBox="[280,1321,250,1688]" captionTargetId="figure@4.[264,1322,194,1690]" captionTargetPageId="4" captionText="FIGURES 2 – 10. Brachistosternus (L.) cepedai n. sp. 2. left hemispermatophore, ventral aspect; 3. left hemispermatophore, dorsal aspect; 4. metasomal segment V, male, dorsal aspect; 5. metasomal segment V, male, ventral aspect; 6. telson, male, lateral aspect; 7. telson, female, lateral aspect; 8. right pedipalp chela, male, ventral aspect; 9. right pedipalp chela, male, internal aspect; 10. left pedipalp chela, female, external aspect. Scale bars = 1 mm." httpUri="https://zenodo.org/record/179244/files/figure.png" pageId="5" pageNumber="60">Fig. 3</figureCitation>
); basal triangle absent, or reduced to a small bulge; internal spines, basal spines and row of spines absent. Mesosoma: Tergites I–VI, entirely smooth in females, they are smooth in anterior two-thirds, and slightly granular in the posterior third in males; tergite VII, surface slightly granular in the posterior two-thirds, two PL and two VL carinae apparent in the second half. Sternites: granular in males, smooth in segments I–IV and coarsely granular in segment V in females; spiracles narrow, and medium-sized. Sternum 
<typeStatus id="0170884AA0142B7F07F8748CFBC37C84" box="[998,1048,992,1018]" pageId="5" pageNumber="60">type</typeStatus>
2 (
<bibRefCitation id="BA5A4B19A0142B7F005B748CFA9C7C84" author="Soleglad" box="[1093,1351,992,1018]" pageId="5" pageNumber="68" refString="Soleglad, M. E. &amp; V. Fet, (2003) The scorpion sternum: structure and phylogeny (Scorpiones: Orthosterni). Euscorpius, 5, 1 - 34." type="journal article" year="2003">Soleglad &amp; Fet 2003</bibRefCitation>
) much wider than long, apex width almost equal to posterior width, posterior emargination quite well developed, with convex lateral lobes conspicuously separated. Pectines: pectinal teeth, 
<quantity id="19339B0DA0142B7F0014735CFBA97B34" box="[1034,1138,1072,1098]" metricMagnitude="-1" metricUnit="m" metricValue="6.858" metricValueMax="7.366" metricValueMin="6.35" pageId="5" pageNumber="60" unit="in" value="27.0" valueMax="29.0" valueMin="25.0">25–29 in</quantity>
males (N = 20; median = 27), 
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females (N = 20; median = 21). Metasoma: segments I–IV each with a pair of dorsolateral macrosetae, segment V with one or two pairs of dorsal macrosetae; metasomal segments I–III: dorsal and ventral surfaces densely granular in males, smooth in females, lateral surface densely granular except the area between LSM and LIM carinae; LSM and LIM carina present only in the posterior two thirds of the segment, DL carina extending the entire length of segment, but almost indistinguishable because of the granules of the tegument; metasomal segment IV: ventral surface smooth, covered by scattered macrosetae; DL carina weakly developed, extending the entire length of segment; LIM carina poorly developed, only present in the distal quarter of the segment; LSM carina apparent only at the anterior and posterior margins of the segment; VL carina present on the entire length of the segment but poorly developed, only represented by a slight elevation of the tegument; surfaces between carinae smooth in females and slightly granular in males. Metasomal segment V: androvestigia small-sized, and very narrow, located submedially (
<figureCitation id="46F02A6DA0142B7F002B7284FB5B797C" box="[1077,1152,1512,1538]" captionStart="FIGURES 2 – 10" captionStartId="4.[151,269,1714,1738]" captionTargetBox="[280,1321,250,1688]" captionTargetId="figure@4.[264,1322,194,1690]" captionTargetPageId="4" captionText="FIGURES 2 – 10. Brachistosternus (L.) cepedai n. sp. 2. left hemispermatophore, ventral aspect; 3. left hemispermatophore, dorsal aspect; 4. metasomal segment V, male, dorsal aspect; 5. metasomal segment V, male, ventral aspect; 6. telson, male, lateral aspect; 7. telson, female, lateral aspect; 8. right pedipalp chela, male, ventral aspect; 9. right pedipalp chela, male, internal aspect; 10. left pedipalp chela, female, external aspect. Scale bars = 1 mm." httpUri="https://zenodo.org/record/179244/files/figure.png" pageId="5" pageNumber="60">Fig. 4</figureCitation>
); dorsal and lateral surfaces slightly granular (male) or smooth (female); ventral surface sparsely granular, more densely granular in the posterior third of the segment (
<figureCitation id="46F02A6DA0142B7F06357154FDA9792C" box="[555,626,1592,1618]" captionStart="FIGURES 2 – 10" captionStartId="4.[151,269,1714,1738]" captionTargetBox="[280,1321,250,1688]" captionTargetId="figure@4.[264,1322,194,1690]" captionTargetPageId="4" captionText="FIGURES 2 – 10. Brachistosternus (L.) cepedai n. sp. 2. left hemispermatophore, ventral aspect; 3. left hemispermatophore, dorsal aspect; 4. metasomal segment V, male, dorsal aspect; 5. metasomal segment V, male, ventral aspect; 6. telson, male, lateral aspect; 7. telson, female, lateral aspect; 8. right pedipalp chela, male, ventral aspect; 9. right pedipalp chela, male, internal aspect; 10. left pedipalp chela, female, external aspect. Scale bars = 1 mm." httpUri="https://zenodo.org/record/179244/files/figure.png" pageId="5" pageNumber="60">Fig. 5</figureCitation>
); ventral macrosetae usually comprising four rows, the first row with four macrosetae, the rest with one or two macrosetae; DL carinae weakly developed; VL carinae well developed, extending the entire length of segment; VM carina absent or weakly developed, represented only by some granules in the posterior half of segment. Telson: Vesicle with rounded ventral surface, lower in males; vesicle surface sparsely granular, with four granules slightly more developed than the rest and related with 4 macrosetae; telson gland absent or not evident; aculeus slightly curved, of the same length as the vesicle (
<figureCitation id="46F02A6DA0142B7F017F706CFF1D783C" captionStart="FIGURES 2 – 10" captionStartId="4.[151,269,1714,1738]" captionTargetBox="[280,1321,250,1688]" captionTargetId="figure@4.[264,1322,194,1690]" captionTargetPageId="4" captionText="FIGURES 2 – 10. Brachistosternus (L.) cepedai n. sp. 2. left hemispermatophore, ventral aspect; 3. left hemispermatophore, dorsal aspect; 4. metasomal segment V, male, dorsal aspect; 5. metasomal segment V, male, ventral aspect; 6. telson, male, lateral aspect; 7. telson, female, lateral aspect; 8. right pedipalp chela, male, ventral aspect; 9. right pedipalp chela, male, internal aspect; 10. left pedipalp chela, female, external aspect. Scale bars = 1 mm." httpUri="https://zenodo.org/record/179244/files/figure.png" pageId="5" pageNumber="60">Figs. 6, 7</figureCitation>
). Pedipalps: Femur: 
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and VI carinae granular and well developed, 
<collectingCountry id="A6DC7678A0142B7F07EA7044FBC0783C" box="[1012,1051,1832,1858]" name="South Georgia and the South Sandwich Islands" pageId="5" pageNumber="60">DE</collectingCountry>
carina only present in the proximal half of the segment, internal surface granular, remaining surfaces smooth, one macroseta (M1) associated with 
<emphasis id="ECBFEAFAA0142B7F04CA7014FF3878EF" box="[212,227,1912,1937]" italics="true" pageId="5" pageNumber="60">d</emphasis>
and 
<emphasis id="ECBFEAFAA0142B7F05007014FEF078EF" box="[286,299,1912,1937]" italics="true" pageId="5" pageNumber="60">e</emphasis>
trichobothria; patella: 
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and VI carinae granular and well developed in males, less granular in females, remaining surfaces smooth; chela: smooth, VM carina weakly developed (
<figureCitation id="46F02A6DA0142B7F007670CCFB7478C4" box="[1128,1199,1952,1978]" captionStart="FIGURES 2 – 10" captionStartId="4.[151,269,1714,1738]" captionTargetBox="[280,1321,250,1688]" captionTargetId="figure@4.[264,1322,194,1690]" captionTargetPageId="4" captionText="FIGURES 2 – 10. Brachistosternus (L.) cepedai n. sp. 2. left hemispermatophore, ventral aspect; 3. left hemispermatophore, dorsal aspect; 4. metasomal segment V, male, dorsal aspect; 5. metasomal segment V, male, ventral aspect; 6. telson, male, lateral aspect; 7. telson, female, lateral aspect; 8. right pedipalp chela, male, ventral aspect; 9. right pedipalp chela, male, internal aspect; 10. left pedipalp chela, female, external aspect. Scale bars = 1 mm." httpUri="https://zenodo.org/record/179244/files/figure.png" pageId="5" pageNumber="60">Fig. 8</figureCitation>
), internal apophysis of male well developed (
<figureCitation id="46F02A6DA0142B7F05A770A4FDDB789C" box="[441,512,1992,2018]" captionStart="FIGURES 2 – 10" captionStartId="4.[151,269,1714,1738]" captionTargetBox="[280,1321,250,1688]" captionTargetId="figure@4.[264,1322,194,1690]" captionTargetPageId="4" captionText="FIGURES 2 – 10. Brachistosternus (L.) cepedai n. sp. 2. left hemispermatophore, ventral aspect; 3. left hemispermatophore, dorsal aspect; 4. metasomal segment V, male, dorsal aspect; 5. metasomal segment V, male, ventral aspect; 6. telson, male, lateral aspect; 7. telson, female, lateral aspect; 8. right pedipalp chela, male, ventral aspect; 9. right pedipalp chela, male, internal aspect; 10. left pedipalp chela, female, external aspect. Scale bars = 1 mm." httpUri="https://zenodo.org/record/179244/files/figure.png" pageId="5" pageNumber="60">Fig. 9</figureCitation>
), fingers with a median row of denticles and five or six pairs of accessory denticles each, external denticles overlapping with median row near the base of the finger. Trichobothrial pattern of subgenus 
<taxonomicName id="19CB4D6BA0172B7C053277ACFE1F7FA7" box="[300,452,192,217]" class="Arachnida" family="Bothriuridae" genus="Leptosternus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="ECBFEAFAA0172B7C053277ACFE1F7FA7" box="[300,452,192,217]" italics="true" pageId="6" pageNumber="61">Leptosternus</emphasis>
</taxonomicName>
(
<figureCitation id="46F02A6DA0172B7C05CA77ACFD8E7FA4" box="[468,597,192,218]" captionStart="FIGURES 2 – 10" captionStartId="4.[151,269,1714,1738]" captionTargetBox="[280,1321,250,1688]" captionTargetId="figure@4.[264,1322,194,1690]" captionTargetPageId="4" captionText="FIGURES 2 – 10. Brachistosternus (L.) cepedai n. sp. 2. left hemispermatophore, ventral aspect; 3. left hemispermatophore, dorsal aspect; 4. metasomal segment V, male, dorsal aspect; 5. metasomal segment V, male, ventral aspect; 6. telson, male, lateral aspect; 7. telson, female, lateral aspect; 8. right pedipalp chela, male, ventral aspect; 9. right pedipalp chela, male, internal aspect; 10. left pedipalp chela, female, external aspect. Scale bars = 1 mm." httpUri="https://zenodo.org/record/179244/files/figure.png" pageId="6" pageNumber="61">Figs. 8–10</figureCitation>
): Neobothriotaxic Major 
<typeStatus id="0170884AA0172B7C079E77ACFC617FA4" box="[896,954,192,218]" pageId="6" pageNumber="61">Type</typeStatus>
C, with one accessory trichobothrium in 
<emphasis id="ECBFEAFAA0172B7C04897784FF727E7F" box="[151,169,232,257]" italics="true" pageId="6" pageNumber="61">V</emphasis>
series of chela; femur with 3 trichobothria (1 
<emphasis id="ECBFEAFAA0172B7C06DB7784FD0F7E7F" box="[709,724,232,257]" italics="true" pageId="6" pageNumber="61">d</emphasis>
, 
<date id="AA751028A0172B7C06FD7784FCDA7E7F" box="[739,769,232,258]" pageId="6" pageNumber="61">
1 
<emphasis id="ECBFEAFAA0172B7C06E77784FCDA7E7F" box="[761,769,232,257]" italics="true" pageId="6" pageNumber="61">i</emphasis>
</date>
and 1 
<emphasis id="ECBFEAFAA0172B7C074F7784FC857E7F" box="[849,862,232,257]" italics="true" pageId="6" pageNumber="61">e</emphasis>
); patella with 19 trichobothria (
<date id="AA751028A0172B7C00CB7784FB217E7F" box="[1237,1274,232,258]" pageId="6" pageNumber="61">
3 
<emphasis id="ECBFEAFAA0172B7C00F27784FB217E7F" box="[1260,1274,232,257]" italics="true" pageId="6" pageNumber="61">V</emphasis>
</date>
, 2 
<emphasis id="ECBFEAFAA0172B7C01017784FAF57E7F" box="[1311,1326,232,257]" italics="true" pageId="6" pageNumber="61">d</emphasis>
, 
<date id="AA751028A0172B7C01227784FA817E7F" box="[1340,1370,232,258]" pageId="6" pageNumber="61">
1 
<emphasis id="ECBFEAFAA0172B7C014C7784FA817E7F" box="[1362,1370,232,257]" italics="true" pageId="6" pageNumber="61">i</emphasis>
</date>
, 
<emphasis id="ECBFEAFAA0172B7C01777784FA4F7E7F" box="[1385,1428,232,257]" italics="true" pageId="6" pageNumber="61">3 et</emphasis>
, 1 
<emphasis id="ECBFEAFAA0172B7C04B2767CFF167E57" box="[172,205,272,297]" italics="true" pageId="6" pageNumber="61">est</emphasis>
, 2 
<emphasis id="ECBFEAFAA0172B7C04EF767CFEC87E57" box="[241,275,272,297]" italics="true" pageId="6" pageNumber="61">em</emphasis>
, 2 
<emphasis id="ECBFEAFAA0172B7C0526767CFEBD7E57" box="[312,358,272,297]" italics="true" pageId="6" pageNumber="61">esb,</emphasis>
and 5 
<emphasis id="ECBFEAFAA0172B7C05AA767CFE0B7E57" box="[436,464,272,297]" italics="true" pageId="6" pageNumber="61">eb</emphasis>
); chela with 27 trichobothria (1 
<emphasis id="ECBFEAFAA0172B7C0752767CFCAA7E57" box="[844,881,272,297]" italics="true" pageId="6" pageNumber="61">Est</emphasis>
, 5 
<emphasis id="ECBFEAFAA0172B7C078B767CFC747E57" box="[917,943,272,297]" italics="true" pageId="6" pageNumber="61">Et</emphasis>
, 
<date id="AA751028A0172B7C07A0767CFC397E57" box="[958,994,272,298]" pageId="6" pageNumber="61">
5 
<emphasis id="ECBFEAFAA0172B7C07CA767CFC397E57" box="[980,994,272,297]" italics="true" pageId="6" pageNumber="61">V</emphasis>
</date>
, 1 
<emphasis id="ECBFEAFAA0172B7C0018767CFBE97E57" box="[1030,1074,272,297]" italics="true" pageId="6" pageNumber="61">Esb</emphasis>
, 3 
<emphasis id="ECBFEAFAA0172B7C0048767CFBAC7E57" box="[1110,1143,272,297]" italics="true" pageId="6" pageNumber="61">Eb</emphasis>
, 1 
<emphasis id="ECBFEAFAA0172B7C0085767CFB637E57" box="[1179,1208,272,297]" italics="true" pageId="6" pageNumber="61">Dt</emphasis>
, 1 
<emphasis id="ECBFEAFAA0172B7C00C3767CFADA7E57" box="[1245,1281,272,297]" italics="true" pageId="6" pageNumber="61">Db</emphasis>
, 1 
<emphasis id="ECBFEAFAA0172B7C013B767CFAE17E57" box="[1317,1338,272,297]" italics="true" pageId="6" pageNumber="61">et</emphasis>
, 1 
<emphasis id="ECBFEAFAA0172B7C0140767CFAA47E57" box="[1374,1407,272,297]" italics="true" pageId="6" pageNumber="61">est</emphasis>
, 1 
<emphasis id="ECBFEAFAA0172B7C04897654FF657E2F" box="[151,190,312,337]" italics="true" pageId="6" pageNumber="61">esb</emphasis>
, 1 
<emphasis id="ECBFEAFAA0172B7C04FA7654FF247E2F" box="[228,255,312,337]" italics="true" pageId="6" pageNumber="61">eb</emphasis>
, 1 
<emphasis id="ECBFEAFAA0172B7C053B7654FEE77E2F" box="[293,316,312,337]" italics="true" pageId="6" pageNumber="61">dt</emphasis>
, 1 
<emphasis id="ECBFEAFAA0172B7C057C7654FE5F7E2F" box="[354,388,312,337]" italics="true" pageId="6" pageNumber="61">dst</emphasis>
, 1 
<emphasis id="ECBFEAFAA0172B7C05B47654FE087E2F" box="[426,467,312,337]" italics="true" pageId="6" pageNumber="61">dsb</emphasis>
, 1 
<emphasis id="ECBFEAFAA0172B7C05E67654FDCE7E2F" box="[504,533,312,337]" italics="true" pageId="6" pageNumber="61">db</emphasis>
, 1 
<emphasis id="ECBFEAFAA0172B7C06227654FD887E2F" box="[572,595,312,337]" italics="true" pageId="6" pageNumber="61">ib</emphasis>
, 1 
<emphasis id="ECBFEAFAA0172B7C06667654FD537E2F" box="[632,648,312,337]" italics="true" pageId="6" pageNumber="61">it</emphasis>
). Legs: ventral surface of the femur with two longitudinal carinae, the rest of the surface smooth; telotarsi I and II: each with inner unguis ca. 15% shorter than external, and with the inner pedal spur absent or vestigial (
<figureCitation id="46F02A6DA0172B7C067576E4FD217EDC" box="[619,762,392,418]" captionStart="FIGURES 23 – 30" captionStartId="10.[151,269,1664,1688]" captionTargetBox="[327,1260,355,1634]" captionTargetId="figure@10.[319,1267,354,1640]" captionTargetPageId="10" captionText="FIGURES 23 – 30. Prolateral view of right leg telotarsi I and II of four Brachistosternus species. 23 – 24. Brachistosternus cepedai n. sp., 23. telotarsus I; 24. telotarsus II. 25 – 26 Brachistosternus sciosciae. 25. telotarsus I; 26. telotarsus II. 27 – 28 Brachistosternus roigalsinai. 27. telotarsus I; 28. telotarsus II. 29 – 30 Brachistosternus montanus. 29. telotarsus I; 30. telotarsus II. Scale bars 1 mm." httpUri="https://zenodo.org/record/179247/files/figure.png" pageId="6" pageNumber="61">Figs. 23, 24</figureCitation>
).
</paragraph>
<paragraph id="DE7436E8A0172B7C04D876C3FAFF7D9C" blockId="6.[151,1437,152,1138]" pageId="6" pageNumber="61">
<emphasis id="ECBFEAFAA0172B7C04D876C3FEE27EB7" bold="true" box="[198,313,431,457]" pageId="6" pageNumber="61">Variation</emphasis>
: Pedipalp chela length/height ratio, male 2.77–3.26 (N = 20; mean = 3.09), female 2.25–3.44 (N = 20; mean = 2.88); pedipalp chela length/width ratio, male 3.85–4.54 (N = 20; mean = 4.18), female 3.78– 4.42 (N = 15; mean = 4.11); metasomal segment V, length/width ratio, male 1.8–1.95 (N = 20; mean = 1.86), female 1.78–1.97 (N = 20; mean = 1.89); metasomal segment V, ventral macroseta 7–10 (N = 20; median = 8); metasomal segment V, ventrolateral macroseta 11–17 (N = 20; median = 14); telotarsus III, dorsal macroseta 12–15 (N = 20; median = 14); telotarsus III, ventrointernal macroseta 10–12 (N = 20; median = 11); telotarsus III, ventroexternal macroseta 5–7 (N = 20; median = 7); basitarsus III, dorsal macroseta 6–8 (N = 20; median = 7); total length (mm), male 36.5–48 (N = 20; mean = 41.5), female 39–66 (N = 18; mean = 62.3).
</paragraph>
</subSubSection>
<subSubSection id="96D16563A0172B7C04D87583FD737B0C" pageId="6" pageNumber="61" type="distribution">
<paragraph id="DE7436E8A0172B7C04D87583FF387CD4" blockId="6.[151,1437,152,1138]" pageId="6" pageNumber="61">
<emphasis id="ECBFEAFAA0172B7C04D87583FDD17C77" bold="true" box="[198,522,751,777]" pageId="6" pageNumber="61">Distribution and ecology.</emphasis>
<taxonomicName id="19CB4D6BA0172B7C0609759CFCA77C77" box="[535,892,752,778]" class="Arachnida" family="Bothriuridae" genus="Brachistosternus" higherTaxonomySource="CoL" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="cepedai" status="sp. nov." subGenus="Leptosternus">
<emphasis id="ECBFEAFAA0172B7C0609759CFD067C77" box="[535,733,752,777]" italics="true" pageId="6" pageNumber="61">Brachistosternus</emphasis>
(
<emphasis id="ECBFEAFAA0172B7C06EA759CFCDF7C77" box="[756,772,752,777]" italics="true" pageId="6" pageNumber="61">L</emphasis>
.) 
<emphasis id="ECBFEAFAA0172B7C073F759CFCA77C77" box="[801,892,752,777]" italics="true" pageId="6" pageNumber="61">cepedai</emphasis>
</taxonomicName>
<emphasis id="ECBFEAFAA0172B7C07977583FC0B7C77" bold="true" box="[905,976,751,777]" pageId="6" pageNumber="61">
<taxonomicNameLabel id="F78C5781A0172B7C07977583FC0B7C77" box="[905,976,751,777]" pageId="6" pageNumber="61" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
inhabits in the 
<taxonomicName id="19CB4D6BA0172B7C0084759CFACD7C74" box="[1178,1302,752,778]" class="Arachnida" family="Bothriuridae" genus="Brachistosternus" higherTaxonomySource="CoL" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="65" phylum="Arthropoda" rank="species" species="coquimbo" status="sp. nov.">Coquimbo</taxonomicName>
Region of northern 
<collectingCountry id="A6DC7678A0172B7C051F7474FE9F7C4C" box="[257,324,792,818]" name="Chile" pageId="6" pageNumber="61">Chile</collectingCountry>
; all specimens have been collected in coastal dunes and in the coastal steppe near the localities of Choros and Punta Choros (
<figureCitation id="46F02A6DA0172B7C0616742CFD8A7C24" box="[520,593,832,858]" captionStart="FIGURE 1" captionStartId="2.[151,255,1313,1337]" captionTargetBox="[275,1311,178,1278]" captionTargetId="figure@2.[225,1362,137,1307]" captionTargetPageId="2" captionText="FIGURE 1. Map of Coquimbo region, Chile, with localities of Brachistosternus (L.) cepedai n. sp. and Brachistosternus (L.) coquimbo n. sp." httpUri="https://zenodo.org/record/179243/files/figure.png" pageId="6" pageNumber="61">Fig. 1</figureCitation>
). The localities where this species has been collected fall within the coastal desert of Huasco, pertaining to the transitional coastal desert of 
<collectingCountry id="A6DC7678A0172B7C07FA7404FBF87CFC" box="[996,1059,872,898]" name="Chile" pageId="6" pageNumber="61">Chile</collectingCountry>
(
<bibRefCitation id="BA5A4B19A0172B7C002D7404FB037CFC" author="Gajardo" box="[1075,1240,872,898]" pageId="6" pageNumber="67" refString="Gajardo, R. (1993) La vegetacion natural de Chile. Clasificacion y distribucion geografica. E ditorial Universitaria, Santiago, Chile, p. 165." type="book chapter" year="1993">Gajardo 1993</bibRefCitation>
, 
<bibRefCitation id="BA5A4B19A0172B7C00F87404FF0D7CD4" author="Cepeda-Pizarro" pageId="6" pageNumber="67" refString="Cepeda-Pizarro, J. (1995) Sintesis ecologica del desierto costero Peruano-Chileno. Investigacion y desarrollo, 1, 4 - 14." type="journal article" year="1995">Cepeda-Pizarro 1995</bibRefCitation>
).
</paragraph>
<paragraph id="DE7436E8A0172B7C04D874D4FD737B0C" blockId="6.[151,1437,152,1138]" pageId="6" pageNumber="61">
<taxonomicName id="19CB4D6BA0172B7C04D874D4FDF17CAF" box="[198,554,952,978]" class="Arachnida" family="Bothriuridae" genus="Brachistosternus" higherTaxonomySource="CoL" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="cepedai" status="sp. nov." subGenus="Leptosternus">
<emphasis id="ECBFEAFAA0172B7C04D874D4FE567CAF" box="[198,397,952,977]" italics="true" pageId="6" pageNumber="61">Brachistosternus</emphasis>
(
<emphasis id="ECBFEAFAA0172B7C05BC74D4FE697CAF" box="[418,434,952,977]" italics="true" pageId="6" pageNumber="61">L</emphasis>
.) 
<emphasis id="ECBFEAFAA0172B7C05D174D4FDF17CAF" box="[463,554,952,977]" italics="true" pageId="6" pageNumber="61">cepedai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F78C5781A0172B7C062874DBFDA67CAF" box="[566,637,951,977]" pageId="6" pageNumber="61" rank="species">
<emphasis id="ECBFEAFAA0172B7C062874DBFDA67CAF" bold="true" box="[566,637,951,977]" pageId="6" pageNumber="61">n. sp.</emphasis>
</taxonomicNameLabel>
inhabits in sandy areas with scarce vegetation, usually 
<emphasis id="ECBFEAFAA0172B7C013274D4FEF37C87" italics="true" pageId="6" pageNumber="61">Crystaria glaucophyla</emphasis>
Cav. (
<taxonomicName id="19CB4D6BA0172B7C0567748CFE277C84" box="[377,508,992,1018]" class="Magnoliopsida" family="Malvaceae" kingdom="Plantae" order="Malvales" pageId="6" pageNumber="61" phylum="Tracheophyta" rank="family">Malvaceae</taxonomicName>
) and 
<taxonomicName id="19CB4D6BA0172B7C065D748CFBC47C84" authority="Barn." authorityName="Barn." box="[579,1055,992,1018]" class="Magnoliopsida" family="Aizoaceae" genus="Tetragonia" kingdom="Animalia" order="Caryophyllales" pageId="6" pageNumber="61" phylum="Tracheophyta" rank="species" species="maritima">
<emphasis id="ECBFEAFAA0172B7C065D748CFCE27C87" box="[579,825,992,1017]" italics="true" pageId="6" pageNumber="61">Tetragonia maritima</emphasis>
Barn. (Aizoaceae)
</taxonomicName>
. This species lives in sympatry with the bothriurids 
<taxonomicName id="19CB4D6BA0172B7C05967364FCFE7B5C" authority="Pocock 1893" authorityName="Pocock" authorityYear="1893" box="[392,805,1032,1058]" class="Arachnida" family="Bothriuridae" genus="Bothriurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="coriaceus">
<emphasis id="ECBFEAFAA0172B7C05967364FD5A7B5F" box="[392,641,1032,1057]" italics="true" pageId="6" pageNumber="61">Bothriurus coriaceus</emphasis>
Pocock 1893
</taxonomicName>
, 
<taxonomicName id="19CB4D6BA0172B7C072C7364FF267B34" authority="Ojanguren-Affilastro 2002" authorityName="Ojanguren-Affilastro" authorityYear="2002" class="Arachnida" family="Bothriuridae" genus="Brachistosternus" higherTaxonomySource="CoL" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="roigalsinai" subGenus="Leptosternus">
<emphasis id="ECBFEAFAA0172B7C072C7364FC227B5F" box="[818,1017,1032,1057]" italics="true" pageId="6" pageNumber="61">Brachistosternus</emphasis>
(
<emphasis id="ECBFEAFAA0172B7C00157364FBC07B5F" box="[1035,1051,1032,1057]" italics="true" pageId="6" pageNumber="61">L</emphasis>
.) 
<emphasis id="ECBFEAFAA0172B7C002A7364FB687B5F" box="[1076,1203,1032,1057]" italics="true" pageId="6" pageNumber="61">roigalsinai</emphasis>
Ojanguren-Affilastro 2002
</taxonomicName>
, and an undescribed 
<taxonomicName id="19CB4D6BA0172B7C05EB735CFD677B37" box="[501,700,1072,1097]" class="Arachnida" family="Bothriuridae" genus="Brachistosternus" higherTaxonomySource="CoL" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="ECBFEAFAA0172B7C05EB735CFD677B37" box="[501,700,1072,1097]" italics="true" pageId="6" pageNumber="61">Brachistosternus</emphasis>
</taxonomicName>
species, and the iurid 
<taxonomicName id="19CB4D6BA0172B7C07DB735CFA4C7B34" authority="Pocock 1893" authorityName="Pocock" authorityYear="1893" box="[965,1431,1072,1098]" class="Arachnida" family="Caraboctonidae" genus="Caraboctonus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="keyserlingi">
<emphasis id="ECBFEAFAA0172B7C07DB735CFB287B37" box="[965,1267,1072,1097]" italics="true" pageId="6" pageNumber="61">Caraboctonus keyserlingi</emphasis>
Pocock 1893
</taxonomicName>
, being the later absent from the coastal dunes.
</paragraph>
</subSubSection>
</treatment>
</document>