476 lines
73 KiB
XML
476 lines
73 KiB
XML
<document id="58162EEB0E82C0EFD94C3BB072ABFD0A" ID-CLB-Dataset="22057" ID-DOI="10.5852/ejt.2016.176" ID-GBIF-Dataset="1126ca50-610b-42ba-96e1-2b02949d6dba" ID-Zenodo-Dep="3832758" ID-ZooBank="C94C7774-5176-460C-804D-A38520284D4B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589831820681" checkinUser="valdenar" docAuthor="Yu, Daoyuan, Man, Le Cong & Deharveng, Louis" docDate="2016" docId="496BD62B7121F106FD8B3D28FAF29E5A" docLanguage="en" docName="ejt-2016-176.pdf.imf" docOrigin="European Journal of Taxonomy 176" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Tomocerus ocreatus Denis 1948" docType="treatment" docVersion="8" lastPageNumber="7" masterDocId="B552AE537123F101FFDD3E50FF859C65" masterDocTitle="Tomoceridae (Collembola, Entomobryomorpha) from the southern Annamitic cordillera: redescription of Tomocerus ocreatus Denis, 1948 and description of a new species of Tomocerina Yosii, 1955" masterLastPageNumber="14" masterPageNumber="1" pageNumber="3" updateTime="1698840175616" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
|
||
<mods:mods id="446553C6B26390BFB62D1D340FCB1439" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="8B5AE791FB2BB73C770E4FE4DA173C2F">
|
||
<mods:title id="867315D1405208093C1901775410261F">Tomoceridae (Collembola, Entomobryomorpha) from the southern Annamitic cordillera: redescription of Tomocerus ocreatus Denis, 1948 and description of a new species of Tomocerina Yosii, 1955</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="890DA5EB9C65130C56BD70B3BCF60556" type="personal">
|
||
<mods:role id="039EB19DC6FDE472C7DF99D0C2721C83">
|
||
<mods:roleTerm id="86FA038BEC016DF4696EAF793AAEF0FF">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="7720779D03BEFDE8C180546CCA11FFBA">Yu, Daoyuan</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="E4626D3848CB68F3586063BAC521D3D6" type="personal">
|
||
<mods:role id="50E7998A69B393726E40F821743AAB12">
|
||
<mods:roleTerm id="1FDE4E76FDD9D7E2186C4D1C1C1C6877">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="0AE598DA961F5FE22135067BD3847203">Man, Le Cong</mods:namePart>
|
||
<mods:nameIdentifier id="416C6F327B31E5A280F065C53B543A39" type="LSID">0D7A5067-F6EF-456F-8B32-D1B61DCC4CDD</mods:nameIdentifier>
|
||
<mods:nameIdentifier id="C52387DFCC917BC6E9C4F6A6A3F5A2D5" type="email">tranletamlinh@yahoo.com.vn</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="2DBB54903CB8EA1AAB24461D07FC1A01" type="personal">
|
||
<mods:role id="3D800E8A17E8E5A4251D4A4889414FD8">
|
||
<mods:roleTerm id="97A2B5E29C87EAE91EF23AEFCAEAA612">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="782363D2ED27C973979EBD32981D0816">Deharveng, Louis</mods:namePart>
|
||
<mods:nameIdentifier id="84CE56C10D83FFEB12326EF7C456ADE5" type="LSID">D8F5C679-C30C-442C-8621-D3B8EDB17EF7</mods:nameIdentifier>
|
||
<mods:nameIdentifier id="ADA4ACB96D1DE05FF334110017A3688C" type="email">deharven@mnhn.fr</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:typeOfResource id="9AA6D7457B4F147A3F66C8CA66297B09">text</mods:typeOfResource>
|
||
<mods:relatedItem id="53EB7201A1E481CDEFAA3593A7CDAEEA" type="host">
|
||
<mods:titleInfo id="A8624B66CC9ABE7218E15E2C01DE47FD">
|
||
<mods:title id="56BD1D79552E3EA059F2CF7A3A05B6D5">European Journal of Taxonomy</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="2353AA83C4B223FE5D6A9DC3F6FE6E15">
|
||
<mods:date id="D503AFF55C6D1FCBBF595D3388C20AA7">2016</mods:date>
|
||
<mods:detail id="8A616809C1772927534F902EAF291363" type="pubDate">
|
||
<mods:number id="F7BE20E777CCE11C99B97C5AFB3FF0C1">2016-02-19</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="E5C1C63C129C5E252EEBD185F4B761A4" type="volume">
|
||
<mods:number id="B7BA189F20FAC15721990FB9224A9E52">176</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="D7D8B42BE7406776699F243970EB8F29" unit="page">
|
||
<mods:start id="A6E8F62BE6CBC0882B3F47353B1AF8AA">1</mods:start>
|
||
<mods:end id="F45547C20ED7D8B90E46BE5A5EBC42D4">14</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="127651E032AD65A3B3BFCAAB65478E11">journal article</mods:classification>
|
||
<mods:identifier id="528125030A68BB4C4018B23CBF2F1C07" type="CLB-Dataset">22057</mods:identifier>
|
||
<mods:identifier id="12976D7F94F4646602F488AC78CDAF20" type="DOI">10.5852/ejt.2016.176</mods:identifier>
|
||
<mods:identifier id="8FC4635F2B2339AF80FD138C9C4867A4" type="GBIF-Dataset">1126ca50-610b-42ba-96e1-2b02949d6dba</mods:identifier>
|
||
<mods:identifier id="5CFC4308F4F0BC788773EA14A926D19F" type="Zenodo-Dep">3832758</mods:identifier>
|
||
<mods:identifier id="B753C23DBB4528AA7C5BB9B827904B0B" type="ZooBank">C94C7774-5176-460C-804D-A38520284D4B</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="496BD62B7121F106FD8B3D28FAF29E5A" ID-DOI="http://doi.org/10.5281/zenodo.3850237" ID-GBIF-Taxon="164167428" ID-Zenodo-Dep="3850237" LSID="urn:lsid:plazi:treatment:496BD62B7121F106FD8B3D28FAF29E5A" httpUri="http://treatment.plazi.org/id/496BD62B7121F106FD8B3D28FAF29E5A" lastPageId="7" lastPageNumber="7" pageId="2" pageNumber="3">
|
||
<subSubSection id="89D834B67121F103FD8B3D28FC5B9FF4" box="[598,990,887,914]" pageId="2" pageNumber="3" type="nomenclature">
|
||
<paragraph id="C17D673D7121F103FD8B3D28FC5B9FF4" blockId="2.[598,990,887,949]" box="[598,990,887,914]" pageId="2" pageNumber="3">
|
||
<heading id="9A35D0517121F103FD8B3D28FC5B9FF4" box="[598,990,887,914]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="3" reason="2">
|
||
<taxonomicName id="06C21CBE7121F103FD8B3D28FC5B9FF4" authority="Denis, 1948" authorityName="Denis" authorityYear="1948" box="[598,990,887,914]" class="Entognatha" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Collembola" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="ocreatus">
|
||
<emphasis id="F3B6BB2F7121F103FD8B3D28FCCC9FF7" bold="true" box="[598,841,888,914]" italics="true" pageId="2" pageNumber="3">Tomocerus ocreatus</emphasis>
|
||
<bibRefCitation id="A5531ACC7121F103FC8D3D27FC5B9FF4" author="Denis J. R." box="[848,990,887,913]" pageId="2" pageNumber="3" pagination="183 - 311" refId="ref6522" refString="Denis J. R. 1948. Collemboles d'Indochine. Notes d'Entomologie Chinoise 12: 183 - 311." type="journal article" year="1948">Denis, 1948</bibRefCitation>
|
||
</taxonomicName>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="89D834B67121F103FD133DCBFCE39FD0" box="[718,870,923,949]" pageId="2" pageNumber="3" type="description">
|
||
<paragraph id="C17D673D7121F103FD133DCBFCE39FD0" blockId="2.[598,990,887,949]" box="[718,870,923,949]" pageId="2" pageNumber="3">
|
||
<figureCitation id="59F97BB87121F103FD133DCBFCA89FD0" box="[718,813,923,949]" captionStart="Fig" captionStartId="3.[189,231,1965,1991]" captionTargetBox="[189,1399,1180,1925]" captionTargetId="figure@3.[189,1399,1180,1925]" captionTargetPageId="3" captionText="Fig. 1. Appearance of Tomocerus ocreatus Denis, 1948 and Tomocerina annamitica sp. nov. in alcohol. A. Tomocerus ocreatus (lateral view). B. Tomocerina annamitica sp. nov. (lateral view). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3832760" httpUri="https://zenodo.org/record/3832760/files/figure.png" pageId="2" pageNumber="3">Figs 1A</figureCitation>
|
||
,
|
||
<figureCitation id="59F97BB87121F103FCE73DCBFCCE9FD0" box="[826,843,923,949]" captionStart="Fig" captionStartId="4.[189,232,1824,1850]" captionTargetBox="[227,1302,491,1774]" captionTargetId="figure@4.[189,1398,473,1784]" captionTargetPageId="4" captionText="Fig. 2. Tomocerus ocreatus Denis, 1948. A. Labrum (dorsal view). B. Mandibular head (dorsal view). C. Lamella 5 of maxillary head (dorsal view). D. Maxillary outer lobe (ventral view). E. Cephalic dorsal chaetotaxy (circles = sockets of chaetae, same as below; 1 = anterior area; 2 = interocular area; 3 = postocular area; 4 = posterior area). F. Dorsal chaetotaxy of Th. II–Abd. VI (circle with a slash = pseudopore, same as below). G. Right tibiotarsi (lateral view). Scale bars: A, B, G = 100 μm; C = 10 μm; D = 50 μm; E, F = 500 μm." figureDoi="http://doi.org/10.5281/zenodo.3832762" httpUri="https://zenodo.org/record/3832762/files/figure.png" pageId="2" pageNumber="3">2</figureCitation>
|
||
,
|
||
<figureCitation id="59F97BB87121F103FC8A3DCBFCE39FD0" box="[855,870,923,949]" captionStart="Fig" captionStartId="5.[189,232,1860,1886]" captionTargetBox="[221,1369,611,1815]" captionTargetId="figure@5.[189,1398,598,1820]" captionTargetPageId="5" captionText="Fig. 3. Tomocerus ocreatus Denis, 1948. A. Diagram of tibiotarsal distal chaetae (distal view; t = tenent hair; a = accessory chaeta; g = guard chaeta; p = pretarsal chaeta, same as below). B. Hind claw (lateral view). C. Tenaculum (anterior view). D. Left side of manubrium (dorsal view). E. Disto-external corner of manubrium (dorsal view). F. Basal and middle subsegments of dens with dental spines (dorsal view). G. Mucro (dorso-inner view). Scale bars: B, C, E, G = 50 μm; D, F = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.3832764" httpUri="https://zenodo.org/record/3832764/files/figure.png" pageId="2" pageNumber="3">3</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="89D834B67121F103FF603D8DFC0899F7" pageId="2" pageNumber="3" type="diagnosis">
|
||
<paragraph id="C17D673D7121F103FF603D8DFEB39F92" blockId="2.[189,310,989,1015]" box="[189,310,989,1015]" pageId="2" pageNumber="3">
|
||
<heading id="9A35D0517121F103FF603D8DFEB39F92" bold="true" box="[189,310,989,1015]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
|
||
<emphasis id="F3B6BB2F7121F103FF603D8DFEB39F92" bold="true" box="[189,310,989,1015]" pageId="2" pageNumber="3">Diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="C17D673D7121F103FF603A5CFF7B98BD" blockId="2.[189,1399,1036,1240]" pageId="2" pageNumber="3">
|
||
Typical
|
||
<taxonomicName id="06C21CBE7121F103FEC33A5DFE1B9843" authorityName="Nicolet" authorityYear="1842" box="[286,414,1037,1062]" class="Entognatha" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Collembola" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F3B6BB2F7121F103FEC33A5DFE1B9843" box="[286,414,1037,1062]" italics="true" pageId="2" pageNumber="3">Tomocerus</emphasis>
|
||
</taxonomicName>
|
||
species with pale body colour, moderately long antennae and full set of 6+6 eyes. Head without distinct PAO; Th. II with relatively reduced number of macrochaetae; tenent hair clavate, moderately developed; unguis with 5–6 teeth; tenaculum unscaled, with numerous chaetae; manubrium without dorsal scales and blunt prominent chaetae; dental spines compound with numerous moderate sized denticles; no small dental spine between two distal large spines; mucro with 9–10 intermediate teeth.
|
||
</paragraph>
|
||
<paragraph id="C17D673D7121F103FF603B51FEA1997E" blockId="2.[189,292,1281,1307]" box="[189,292,1281,1307]" pageId="2" pageNumber="3">
|
||
<heading id="9A35D0517121F103FF603B51FEA1997E" bold="true" box="[189,292,1281,1307]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
|
||
<emphasis id="F3B6BB2F7121F103FF603B51FEA1997E" bold="true" box="[189,292,1281,1307]" pageId="2" pageNumber="3">
|
||
<typeStatus id="1E79D99F7121F103FF603B51FEA1997E" box="[189,292,1281,1307]" pageId="2" pageNumber="3" type="neotype">Neotype</typeStatus>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="C17D673D7121F103FF603B60FC0899F7" blockId="2.[189,1398,1327,1426]" pageId="2" pageNumber="3">
|
||
<materialsCitation id="71AA6D607121F103FF603B60FC0899F7" ID-GBIF-Occurrence="2620526302" collectingDate="2008-06-12" collectionCode="MNHN" collectorName="Louis Deharveng & Anne Bedos & Deposited" country="Vietnam" county="Hon Giao" elevation="1630" latitude="12.186389" location="Dalat" longLatPrecision="21" longitude="108.71472" municipality="Bi Doup" pageId="2" pageNumber="3" specimenCount="1" specimenCount-female="1" stateProvince="Lam Dong" typeStatus="neotype">
|
||
<collectingCountry id="B9D527AD7121F103FF603B60FECC992F" box="[189,329,1328,1354]" name="Vietnam" pageId="2" pageNumber="3">VIETNAM</collectingCountry>
|
||
:
|
||
<specimenCount id="D7C4ACB47121F103FE8F3B60FEED992F" box="[338,360,1328,1354]" pageId="2" pageNumber="3" type="female">♀</specimenCount>
|
||
, on slide. Collected in
|
||
<collectingCounty id="281C1FB17121F103FDB03B60FD65992C" box="[621,736,1327,1354]" pageId="2" pageNumber="3">Hon Giao</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="2119FD477121F103FD373B7FFCC9992F" box="[746,844,1327,1354]" pageId="2" pageNumber="3">Bi Doup</collectingMunicipality>
|
||
massif, northeast of
|
||
<location id="C41D31E67121F103FBE43B7FFBFE992C" LSID="urn:lsid:plazi:treatment:496BD62B7121F106FD8B3D28FAF29E5A:C41D31E67121F103FBE43B7FFBFE992C" box="[1081,1147,1327,1353]" country="Vietnam" county="Hon Giao" latitude="12.186389" longLatPrecision="21" longitude="108.71472" municipality="Bi Doup" name="Dalat" pageId="2" pageNumber="3" stateProvince="Lam Dong">Dalat</location>
|
||
,
|
||
<collectingRegion id="0306A9DF7121F103FB583B60FAF7992C" box="[1157,1394,1327,1354]" country="Vietnam" name="Lam Dong" pageId="2" pageNumber="3">Lam Dong Province</collectingRegion>
|
||
,
|
||
<geoCoordinate id="A4F601FA7121F103FF603B03FEDF990B" box="[189,346,1363,1390]" degrees="108" direction="east" minutes="42" orientation="longitude" pageId="2" pageNumber="3" precision="15" seconds="53" value="108.71472">108°42’53”E</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="A4F601FA7121F103FEBB3B03FE7C9908" box="[358,505,1363,1389]" degrees="12" direction="north" minutes="11" orientation="latitude" pageId="2" pageNumber="3" precision="15" seconds="11" value="12.186389">12°11’11’’N</geoCoordinate>
|
||
, alt.
|
||
<quantity id="063ACAD87121F103FDED3B03FD0F990B" box="[560,650,1363,1390]" metricMagnitude="3" metricUnit="m" metricValue="1.63" pageId="2" pageNumber="3" unit="m" value="1630.0">
|
||
<elevation id="4AEF800E7121F103FDED3B03FD0F990B" box="[560,650,1363,1390]" metricMagnitude="3" metricUnit="m" metricValue="1.63" pageId="2" pageNumber="3" unit="m" value="1630.0">1630 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="B57C41FD7121F103FD483B03FCA89908" box="[661,813,1363,1390]" pageId="2" pageNumber="3" value="2008-06-12">
|
||
<collectingDate id="A538B8157121F103FD483B03FCA89908" box="[661,813,1363,1390]" pageId="2" pageNumber="3" value="2008-06-12">12 Jun. 2008</collectingDate>
|
||
</date>
|
||
, by
|
||
<collectorName id="6C3702EB7121F103FC863B03FBA09908" box="[859,1061,1363,1389]" pageId="2" pageNumber="3">Louis Deharveng</collectorName>
|
||
&
|
||
<collectorName id="6C3702EB7121F103FB9A3B03FB509908" box="[1095,1237,1363,1389]" pageId="2" pageNumber="3">Anne Bedos</collectorName>
|
||
(sample code Vn08-150).
|
||
<collectorName id="6C3702EB7121F103FE913B27FE4699F4" box="[332,451,1399,1425]" pageId="2" pageNumber="3">Deposited</collectorName>
|
||
in
|
||
<collectionCode id="A7D3FFF87121F103FE343B27FDC699F4" box="[489,579,1399,1425]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="3">MNHN</collectionCode>
|
||
(specimen Vn08-150_To1).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="89D834B67121F107FF603BE9FAE89D9E" lastPageId="6" lastPageNumber="7" pageId="2" pageNumber="3" type="description">
|
||
<paragraph id="C17D673D7121F103FF603BE9FECB99B6" blockId="2.[189,334,1465,1491]" box="[189,334,1465,1491]" pageId="2" pageNumber="3">
|
||
<heading id="9A35D0517121F103FF603BE9FECB99B6" bold="true" box="[189,334,1465,1491]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
|
||
<emphasis id="F3B6BB2F7121F103FF603BE9FECB99B6" bold="true" box="[189,334,1465,1491]" pageId="2" pageNumber="3">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="C17D673D7121F103FF603BB8FB0E9A43" blockId="2.[189,1398,1512,1574]" pageId="2" pageNumber="3">
|
||
Body length 3.6 mm. Ground colour uniformly yellowish white. Ant. III+IV, antero-ventral part of head, coxae and tibiotarsi with dark pigment. Eye patches black. Scales brown (
|
||
<figureCitation id="59F97BB87121F103FBFD385BFBF89A43" box="[1056,1149,1547,1574]" captionStart="Fig" captionStartId="3.[189,231,1965,1991]" captionTargetBox="[189,1399,1180,1925]" captionTargetId="figure@3.[189,1399,1180,1925]" captionTargetPageId="3" captionText="Fig. 1. Appearance of Tomocerus ocreatus Denis, 1948 and Tomocerina annamitica sp. nov. in alcohol. A. Tomocerus ocreatus (lateral view). B. Tomocerina annamitica sp. nov. (lateral view). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3832760" httpUri="https://zenodo.org/record/3832760/files/figure.png" pageId="2" pageNumber="3">Fig. 1A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="C17D673D7121F103FF60381DFADF9BE0" blockId="2.[189,1399,1613,1926]" pageId="2" pageNumber="3">
|
||
Body densely clothed by scales and various
|
||
<typeStatus id="1E79D99F7121F103FD26381EFCBC9A0D" box="[763,825,1614,1640]" pageId="2" pageNumber="3">types</typeStatus>
|
||
of chaetae. Scales of typical morphology of
|
||
<taxonomicName id="06C21CBE7121F103FF603821FEDC9AEE" authorityName="Schaffer" authorityYear="1896" box="[189,345,1649,1675]" class="Entognatha" family="Tomoceridae" kingdom="Animalia" order="Collembola" pageId="2" pageNumber="3" phylum="Arthropoda" rank="subFamily" subFamily="Tomocerinae">Tomocerinae</taxonomicName>
|
||
, with continuous longitudinal ridges on surface (
|
||
<bibRefCitation id="A5531ACC7121F103FC6B3821FBEE9AEE" author="Lubbock J." box="[950,1131,1649,1675]" pageId="2" pageNumber="3" refId="ref6831" refString="Lubbock J. 1873. Monograph of the Collembola and Thysanura. Ray Society, London. http: // dx. doi. org / 10.5962 / bhl. title. 11583" type="book" year="1873">Lubbock 1873</bibRefCitation>
|
||
). Ordinary chaetae of different sizes. Microchaetae smooth and pointed. Macrochaetae and mesochaetae from slightly to strongly ciliated, some slightly ciliated mesochaetae appearing to be smooth under optical microscope. Most macrochaetae straight, rod-like and subcylindrical, some macrochaetae on posterior abdominal segments long, curved and acuminate. Mesochaetae acuminate, shorter and thinner than macrochaetae. S-chaetae subcylindrical, more hyaline than ordinary chaetae, as small as microchaetae except long ones on Abd. IV. Dorso-inner chaetae on dens modified as strong pointed spines. Pseudopores as small circular structures similar to chaetae sockets, distributed at least on Th. II to Abd. IV, coxae, and manubrium.
|
||
</paragraph>
|
||
<paragraph id="C17D673D7121F103FF6039FDFAF29B8E" blockId="2.[189,1399,1965,2027]" pageId="2" pageNumber="3">PAO not seen. Eyes 6+6. Antennae nearly as long as body. Antenna length ratio as I:II:III+IV= 1.0:1.7:15.4. Ant. I and Ant. II dorsally scaled, Ant. III+IV unscaled. Prelabral and labral chaetae (labral</paragraph>
|
||
<paragraph id="C17D673D7120F102FF603F5BFE569E5A" blockId="3.[189,1399,267,575]" pageId="3" pageNumber="4">
|
||
formula) 4/5, 5, 4, the distal 4 chaetae stronger. Distal edge of labrum with four curved spine-like papillae (
|
||
<figureCitation id="59F97BB87120F102FF1B3F7EFEA39D2C" box="[198,294,302,329]" captionStart="Fig" captionStartId="4.[189,232,1824,1850]" captionTargetBox="[227,1302,491,1774]" captionTargetId="figure@4.[189,1398,473,1784]" captionTargetPageId="4" captionText="Fig. 2. Tomocerus ocreatus Denis, 1948. A. Labrum (dorsal view). B. Mandibular head (dorsal view). C. Lamella 5 of maxillary head (dorsal view). D. Maxillary outer lobe (ventral view). E. Cephalic dorsal chaetotaxy (circles = sockets of chaetae, same as below; 1 = anterior area; 2 = interocular area; 3 = postocular area; 4 = posterior area). F. Dorsal chaetotaxy of Th. II–Abd. VI (circle with a slash = pseudopore, same as below). G. Right tibiotarsi (lateral view). Scale bars: A, B, G = 100 μm; C = 10 μm; D = 50 μm; E, F = 500 μm." figureDoi="http://doi.org/10.5281/zenodo.3832762" httpUri="https://zenodo.org/record/3832762/files/figure.png" pageId="3" pageNumber="4">Fig. 2A</figureCitation>
|
||
), and a well developed ventro-distal brush. Mandibular head asymmetrical, the left one with 4 teeth (including a basal rounded one) and the right one with 5, left molar plate distally with a tapered tooth (
|
||
<figureCitation id="59F97BB87120F102FED63F25FEED9DEA" box="[267,360,373,399]" captionStart="Fig" captionStartId="4.[189,232,1824,1850]" captionTargetBox="[227,1302,491,1774]" captionTargetId="figure@4.[189,1398,473,1784]" captionTargetPageId="4" captionText="Fig. 2. Tomocerus ocreatus Denis, 1948. A. Labrum (dorsal view). B. Mandibular head (dorsal view). C. Lamella 5 of maxillary head (dorsal view). D. Maxillary outer lobe (ventral view). E. Cephalic dorsal chaetotaxy (circles = sockets of chaetae, same as below; 1 = anterior area; 2 = interocular area; 3 = postocular area; 4 = posterior area). F. Dorsal chaetotaxy of Th. II–Abd. VI (circle with a slash = pseudopore, same as below). G. Right tibiotarsi (lateral view). Scale bars: A, B, G = 100 μm; C = 10 μm; D = 50 μm; E, F = 500 μm." figureDoi="http://doi.org/10.5281/zenodo.3832762" httpUri="https://zenodo.org/record/3832762/files/figure.png" pageId="3" pageNumber="4">Fig. 2B</figureCitation>
|
||
). Basal teeth of maxillary lamella 5 slightly longer than apical ones, without beard-like appendage (
|
||
<figureCitation id="59F97BB87120F102FE903FC8FE299DD7" box="[333,428,408,434]" captionStart="Fig" captionStartId="4.[189,232,1824,1850]" captionTargetBox="[227,1302,491,1774]" captionTargetId="figure@4.[189,1398,473,1784]" captionTargetPageId="4" captionText="Fig. 2. Tomocerus ocreatus Denis, 1948. A. Labrum (dorsal view). B. Mandibular head (dorsal view). C. Lamella 5 of maxillary head (dorsal view). D. Maxillary outer lobe (ventral view). E. Cephalic dorsal chaetotaxy (circles = sockets of chaetae, same as below; 1 = anterior area; 2 = interocular area; 3 = postocular area; 4 = posterior area). F. Dorsal chaetotaxy of Th. II–Abd. VI (circle with a slash = pseudopore, same as below). G. Right tibiotarsi (lateral view). Scale bars: A, B, G = 100 μm; C = 10 μm; D = 50 μm; E, F = 500 μm." figureDoi="http://doi.org/10.5281/zenodo.3832762" httpUri="https://zenodo.org/record/3832762/files/figure.png" pageId="3" pageNumber="4">Fig. 2C</figureCitation>
|
||
). Maxillary outer lobe with trifurcate palp, one basal chaeta and 4 sublobal hairs (
|
||
<figureCitation id="59F97BB87120F102FF1B3FEBFEA19DB0" box="[198,292,443,469]" captionStart="Fig" captionStartId="4.[189,232,1824,1850]" captionTargetBox="[227,1302,491,1774]" captionTargetId="figure@4.[189,1398,473,1784]" captionTargetPageId="4" captionText="Fig. 2. Tomocerus ocreatus Denis, 1948. A. Labrum (dorsal view). B. Mandibular head (dorsal view). C. Lamella 5 of maxillary head (dorsal view). D. Maxillary outer lobe (ventral view). E. Cephalic dorsal chaetotaxy (circles = sockets of chaetae, same as below; 1 = anterior area; 2 = interocular area; 3 = postocular area; 4 = posterior area). F. Dorsal chaetotaxy of Th. II–Abd. VI (circle with a slash = pseudopore, same as below). G. Right tibiotarsi (lateral view). Scale bars: A, B, G = 100 μm; C = 10 μm; D = 50 μm; E, F = 500 μm." figureDoi="http://doi.org/10.5281/zenodo.3832762" httpUri="https://zenodo.org/record/3832762/files/figure.png" pageId="3" pageNumber="4">Fig. 2D</figureCitation>
|
||
). Both dorsal and ventral sides of head scaled. Cephalic dorsal macrochaetotaxy: anterior area: 2, 2; interocular area: 2, 6, central uneven macrochaeta absent; postocular area: 2+2; posterior area: 2. Posterior margin of head with about 20+20 small chaetae (
|
||
<figureCitation id="59F97BB87120F102FCBF3C52FC399E79" box="[866,956,514,540]" captionStart="Fig" captionStartId="4.[189,232,1824,1850]" captionTargetBox="[227,1302,491,1774]" captionTargetId="figure@4.[189,1398,473,1784]" captionTargetPageId="4" captionText="Fig. 2. Tomocerus ocreatus Denis, 1948. A. Labrum (dorsal view). B. Mandibular head (dorsal view). C. Lamella 5 of maxillary head (dorsal view). D. Maxillary outer lobe (ventral view). E. Cephalic dorsal chaetotaxy (circles = sockets of chaetae, same as below; 1 = anterior area; 2 = interocular area; 3 = postocular area; 4 = posterior area). F. Dorsal chaetotaxy of Th. II–Abd. VI (circle with a slash = pseudopore, same as below). G. Right tibiotarsi (lateral view). Scale bars: A, B, G = 100 μm; C = 10 μm; D = 50 μm; E, F = 500 μm." figureDoi="http://doi.org/10.5281/zenodo.3832762" httpUri="https://zenodo.org/record/3832762/files/figure.png" pageId="3" pageNumber="4">Fig. 2E</figureCitation>
|
||
). Mentum with 5 chaetae, submentum with numerous chaetae.
|
||
</paragraph>
|
||
<paragraph id="C17D673D7120F102FF603C37FBC29FFF" blockId="3.[189,1399,614,922]" pageId="3" pageNumber="4">
|
||
Pattern of body chaetotaxy as in
|
||
<figureCitation id="59F97BB87120F102FDE93C36FD0C9EE5" box="[564,649,614,640]" captionStart="Fig" captionStartId="4.[189,232,1824,1850]" captionTargetBox="[227,1302,491,1774]" captionTargetId="figure@4.[189,1398,473,1784]" captionTargetPageId="4" captionText="Fig. 2. Tomocerus ocreatus Denis, 1948. A. Labrum (dorsal view). B. Mandibular head (dorsal view). C. Lamella 5 of maxillary head (dorsal view). D. Maxillary outer lobe (ventral view). E. Cephalic dorsal chaetotaxy (circles = sockets of chaetae, same as below; 1 = anterior area; 2 = interocular area; 3 = postocular area; 4 = posterior area). F. Dorsal chaetotaxy of Th. II–Abd. VI (circle with a slash = pseudopore, same as below). G. Right tibiotarsi (lateral view). Scale bars: A, B, G = 100 μm; C = 10 μm; D = 50 μm; E, F = 500 μm." figureDoi="http://doi.org/10.5281/zenodo.3832762" httpUri="https://zenodo.org/record/3832762/files/figure.png" pageId="3" pageNumber="4">
|
||
Fig.
|
||
<specimenCount id="D7C4ACB47120F102FDB53C36FD0C9EE5" box="[616,649,614,640]" pageId="3" pageNumber="4" type="generic">2F</specimenCount>
|
||
</figureCitation>
|
||
. Bothriotricha 2, 1/0, 0, 1, 2, 0, 0 on Th. II–Abd. VI, respectively, as typical in
|
||
<taxonomicName id="06C21CBE7120F102FE883CD9FE779EC6" authorityName="Schaffer" authorityYear="1896" box="[341,498,649,675]" class="Entognatha" family="Tomoceridae" kingdom="Animalia" order="Collembola" pageId="3" pageNumber="3" phylum="Arthropoda" rank="subFamily" subFamily="Tomocerinae">Tomocerinae</taxonomicName>
|
||
. Macrochaetae densely arranged along anterior margin of Th. II (not shown in figure). Th. II with a row of macrochaetae behind anterior margin. Number of macrochaetae or large mesochaetae in the posterior row as 3, 3/3, 3, 4, 2, 4 (3 dorsal+1 lateral) from Th. II to Abd. V. On Th. II no macrochaeta near the pseudopore contrary to most other tomocerids; on Abd. III s-microchaeta and accompanying microchaeta posterior to lateral macrochaeta; Abd. IV with one lateral macrochaeta and numerous long s-chaetae; on Abd. V inner macrochaeta smaller than others in posterior row; Abd. VI with numerous chaetae of moderate size. Most mesochaetae laterally and posteriorly on terga. Pseudopores near the axis of terga, 1, 1/1, 1, 1, 1, 0, 0 from Th. II to Abd. VI.
|
||
</paragraph>
|
||
<paragraph id="C17D673D7120F105FF603D92FD8D9DEA" blockId="3.[189,1399,961,1129]" lastBlockId="4.[189,1399,267,399]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
|
||
Legs with numerous ordinary chaetae. Trochantero-femoral organ with 1, 1 slender chaetae. Front, middle and hind tibiotarsus dorsally with 1, 1, 3 long prominent chaetae, ventrally with 4–5, 5, 6 blunt spine-like chaetae (
|
||
<figureCitation id="59F97BB87120F102FE793A58FD809847" box="[420,517,1032,1058]" captionStart="Fig" captionStartId="4.[189,232,1824,1850]" captionTargetBox="[227,1302,491,1774]" captionTargetId="figure@4.[189,1398,473,1784]" captionTargetPageId="4" captionText="Fig. 2. Tomocerus ocreatus Denis, 1948. A. Labrum (dorsal view). B. Mandibular head (dorsal view). C. Lamella 5 of maxillary head (dorsal view). D. Maxillary outer lobe (ventral view). E. Cephalic dorsal chaetotaxy (circles = sockets of chaetae, same as below; 1 = anterior area; 2 = interocular area; 3 = postocular area; 4 = posterior area). F. Dorsal chaetotaxy of Th. II–Abd. VI (circle with a slash = pseudopore, same as below). G. Right tibiotarsi (lateral view). Scale bars: A, B, G = 100 μm; C = 10 μm; D = 50 μm; E, F = 500 μm." figureDoi="http://doi.org/10.5281/zenodo.3832762" httpUri="https://zenodo.org/record/3832762/files/figure.png" pageId="3" pageNumber="4">Fig. 2G</figureCitation>
|
||
). Each tibiotarsus with a distal whorl of 11 chaetae, ventral 6 as ordinary chaetae, dorsal 5 modified: tenent hair clavate, as long as inner edge of unguis; 2 accessory chaetae extremely minute; 2 guard chaetae thin and long, slightly shorter than tenent hair (
|
||
<figureCitation id="59F97BB87120F102FB573A1EFA8E980C" box="[1162,1291,1102,1129]" captionStart="Fig" captionStartId="5.[189,232,1860,1886]" captionTargetBox="[221,1369,611,1815]" captionTargetId="figure@5.[189,1398,598,1820]" captionTargetPageId="5" captionText="Fig. 3. Tomocerus ocreatus Denis, 1948. A. Diagram of tibiotarsal distal chaetae (distal view; t = tenent hair; a = accessory chaeta; g = guard chaeta; p = pretarsal chaeta, same as below). B. Hind claw (lateral view). C. Tenaculum (anterior view). D. Left side of manubrium (dorsal view). E. Disto-external corner of manubrium (dorsal view). F. Basal and middle subsegments of dens with dental spines (dorsal view). G. Mucro (dorso-inner view). Scale bars: B, C, E, G = 50 μm; D, F = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.3832764" httpUri="https://zenodo.org/record/3832764/files/figure.png" pageId="3" pageNumber="4">Fig. 3A, B</figureCitation>
|
||
). Unguis slender, with baso-internal ridging visible in lateral view; lateral teeth pointed, of moderate size. Inner edge of unguis with 5–6 teeth, the basal tooth smaller, the other subequal in size. Unguiculus about half as long as unguis, its inner edge with 1 tooth. Pretarsal chaetae 1+1, much larger than tenent-hair accessory chaetae (
|
||
<figureCitation id="59F97BB87127F105FE433F25FE7F9DEA" box="[414,506,373,399]" captionStart="Fig" captionStartId="5.[189,232,1860,1886]" captionTargetBox="[221,1369,611,1815]" captionTargetId="figure@5.[189,1398,598,1820]" captionTargetPageId="5" captionText="Fig. 3. Tomocerus ocreatus Denis, 1948. A. Diagram of tibiotarsal distal chaetae (distal view; t = tenent hair; a = accessory chaeta; g = guard chaeta; p = pretarsal chaeta, same as below). B. Hind claw (lateral view). C. Tenaculum (anterior view). D. Left side of manubrium (dorsal view). E. Disto-external corner of manubrium (dorsal view). F. Basal and middle subsegments of dens with dental spines (dorsal view). G. Mucro (dorso-inner view). Scale bars: B, C, E, G = 50 μm; D, F = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.3832764" httpUri="https://zenodo.org/record/3832764/files/figure.png" pageId="4" pageNumber="5">Fig. 3B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="95BD37B57120F102FF6039FDFAF29B8E" ID-DOI="http://doi.org/10.5281/zenodo.3832760" ID-Zenodo-Dep="3832760" httpUri="https://zenodo.org/record/3832760/files/figure.png" pageId="3" pageNumber="4" startId="3.[189,231,1965,1991]" targetBox="[189,1399,1180,1925]" targetPageId="3">
|
||
<paragraph id="C17D673D7120F102FF6039FDFAF29B8E" blockId="3.[189,1399,1965,2028]" pageId="3" pageNumber="4">
|
||
<emphasis id="F3B6BB2F7120F102FF6039FDFE8E9BA2" bold="true" box="[189,267,1965,1991]" pageId="3" pageNumber="4">Fig. 1.</emphasis>
|
||
Appearance of
|
||
<taxonomicName id="06C21CBE7120F102FE1939FEFCC79BAD" authority="Denis, 1948" authorityName="Denis" authorityYear="1948" box="[452,834,1965,1992]" class="Entognatha" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Collembola" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="ocreatus">
|
||
<emphasis id="F3B6BB2F7120F102FE1939FEFD2B9BA2" box="[452,686,1966,1991]" italics="true" pageId="3" pageNumber="4">Tomocerus ocreatus</emphasis>
|
||
<bibRefCitation id="A5531ACC7120F102FD6839FDFCC79BAD" author="Denis J. R." box="[693,834,1965,1992]" pageId="3" pageNumber="4" pagination="183 - 311" refId="ref6522" refString="Denis J. R. 1948. Collemboles d'Indochine. Notes d'Entomologie Chinoise 12: 183 - 311." type="journal article" year="1948">Denis, 1948</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="06C21CBE7120F102FCA039FEFB179BA2" authority="Man & Deharveng, 2016" authorityName="Man & Deharveng" authorityYear="2016" box="[893,1170,1966,1991]" class="Entognatha" family="Tomoceridae" genus="Tomocerina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="annamitica" status="sp. nov.">
|
||
<emphasis id="F3B6BB2F7120F102FCA039FEFB179BA2" box="[893,1170,1966,1991]" italics="true" pageId="3" pageNumber="4">Tomocerina annamitica</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E88506547120F102FB4439FEFB719BAD" box="[1177,1268,1966,1992]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
in alcohol.
|
||
<emphasis id="F3B6BB2F7120F102FF603980FF579B8F" bold="true" box="[189,210,2000,2026]" pageId="3" pageNumber="4">A</emphasis>
|
||
.
|
||
<taxonomicName id="06C21CBE7120F102FF3D3982FE429B8E" authorityName="Denis" authorityYear="1948" box="[224,455,2002,2027]" class="Entognatha" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Collembola" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="ocreatus">
|
||
<emphasis id="F3B6BB2F7120F102FF3D3982FE429B8E" box="[224,455,2002,2027]" italics="true" pageId="3" pageNumber="4">Tomocerus ocreatus</emphasis>
|
||
</taxonomicName>
|
||
(lateral view).
|
||
<emphasis id="F3B6BB2F7120F102FDA83981FD0D9B8E" bold="true" box="[629,648,2001,2027]" pageId="3" pageNumber="4">B</emphasis>
|
||
.
|
||
<taxonomicName id="06C21CBE7120F102FD4B3982FC2C9B8E" authority="Man & Deharveng, 2016" authorityName="Man & Deharveng" authorityYear="2016" box="[662,937,2002,2027]" class="Entognatha" family="Tomoceridae" genus="Tomocerina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="annamitica" status="sp. nov.">
|
||
<emphasis id="F3B6BB2F7120F102FD4B3982FC2C9B8E" box="[662,937,2002,2027]" italics="true" pageId="3" pageNumber="4">Tomocerina annamitica</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E88506547120F102FC723981FB829B8E" box="[943,1031,2001,2027]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(lateral view). Scale bar: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="95BD37B57127F105FF603970FE7B9B8E" ID-DOI="http://doi.org/10.5281/zenodo.3832762" ID-Zenodo-Dep="3832762" httpUri="https://zenodo.org/record/3832762/files/figure.png" pageId="4" pageNumber="5" startId="4.[189,232,1824,1850]" targetBox="[227,1302,491,1774]" targetPageId="4">
|
||
<paragraph id="C17D673D7127F105FF603970FE7B9B8E" blockId="4.[189,1399,1824,2027]" pageId="4" pageNumber="5">
|
||
<emphasis id="F3B6BB2F7127F105FF603970FE899B5E" bold="true" box="[189,268,1824,1851]" pageId="4" pageNumber="5">Fig. 2.</emphasis>
|
||
<taxonomicName id="06C21CBE7127F105FEC93972FD1E9B5E" authority="Denis, 1948" authorityName="Denis" authorityYear="1948" box="[276,667,1825,1851]" class="Entognatha" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Collembola" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="ocreatus">
|
||
<emphasis id="F3B6BB2F7127F105FEC93972FD859B5E" box="[276,512,1826,1851]" italics="true" pageId="4" pageNumber="5">Tomocerus ocreatus</emphasis>
|
||
<bibRefCitation id="A5531ACC7127F105FDD53971FD1E9B5E" author="Denis J. R." box="[520,667,1825,1851]" pageId="4" pageNumber="5" pagination="183 - 311" refId="ref6522" refString="Denis J. R. 1948. Collemboles d'Indochine. Notes d'Entomologie Chinoise 12: 183 - 311." type="journal article" year="1948">Denis, 1948</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="F3B6BB2F7127F105FD7A3970FD399B5F" bold="true" box="[679,700,1824,1850]" pageId="4" pageNumber="5">A</emphasis>
|
||
. Labrum (dorsal view).
|
||
<emphasis id="F3B6BB2F7127F105FC033971FC779B5E" bold="true" box="[990,1010,1825,1851]" pageId="4" pageNumber="5">B</emphasis>
|
||
. Mandibular head (dorsal view).
|
||
<emphasis id="F3B6BB2F7127F105FF603914FF579B3B" bold="true" box="[189,210,1860,1886]" pageId="4" pageNumber="5">C</emphasis>
|
||
. Lamella 5 of maxillary head (dorsal view).
|
||
<emphasis id="F3B6BB2F7127F105FD223914FC919B3B" bold="true" box="[767,788,1860,1886]" pageId="4" pageNumber="5">D</emphasis>
|
||
. Maxillary outer lobe (ventral view).
|
||
<emphasis id="F3B6BB2F7127F105FB353914FB799B3B" bold="true" box="[1256,1276,1860,1886]" pageId="4" pageNumber="5">E</emphasis>
|
||
. Cephalic dorsal chaetotaxy (circles = sockets of chaetae, same as below; 1 = anterior area; 2 = interocular area; 3 = postocular area; 4 = posterior area).
|
||
<emphasis id="F3B6BB2F7127F105FD4339DAFD359BC1" bold="true" box="[670,688,1930,1956]" pageId="4" pageNumber="5">F</emphasis>
|
||
. Dorsal chaetotaxy of Th. II–Abd. VI (circle with a slash = pseudopore, same as below).
|
||
<emphasis id="F3B6BB2F7127F105FDCC39FDFDAD9BA2" bold="true" box="[529,552,1965,1991]" pageId="4" pageNumber="5">G</emphasis>
|
||
. Right tibiotarsi (lateral view). Scale bars: A, B, G = 100 μm; C = 10 μm; D = 50 μm; E, F = 500 μm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="C17D673D7126F107FF603F5BFAE89D9E" blockId="5.[189,1399,267,540]" lastBlockId="6.[189,1399,267,507]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
|
||
Ventral tube with scales on both anterior and posterior faces, lateral flap unscaled, anterior face with about 15 chaetae on each side, posterior face with about 55 chaetae, each lateral flap with 45–47 chaetae. Rami of tenaculum with 4+4 teeth, anterior face with 13 small chaetae and without scale (
|
||
<figureCitation id="59F97BB87126F104FAD43F02FAEC9D09" box="[1289,1385,338,364]" captionStart="Fig" captionStartId="5.[189,232,1860,1886]" captionTargetBox="[221,1369,611,1815]" captionTargetId="figure@5.[189,1398,598,1820]" captionTargetPageId="5" captionText="Fig. 3. Tomocerus ocreatus Denis, 1948. A. Diagram of tibiotarsal distal chaetae (distal view; t = tenent hair; a = accessory chaeta; g = guard chaeta; p = pretarsal chaeta, same as below). B. Hind claw (lateral view). C. Tenaculum (anterior view). D. Left side of manubrium (dorsal view). E. Disto-external corner of manubrium (dorsal view). F. Basal and middle subsegments of dens with dental spines (dorsal view). G. Mucro (dorso-inner view). Scale bars: B, C, E, G = 50 μm; D, F = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.3832764" httpUri="https://zenodo.org/record/3832764/files/figure.png" pageId="5" pageNumber="6">Fig. 3C</figureCitation>
|
||
). Furca ratio manubrium: dens: mucro=2.9–3.1:4.1–4.4:1.0. Manubrium ventrally scaled, without chaetae, laterally with large round scales and 11 chaetae, proximal 2 chaetae small and slender, distal 9 distinctly stronger and serrated; dorsal scales absent; each dorsal chaetal stripe with about 90 chaetae in different sizes, without blunt chaetae; pseudopores 11–12 on each side (
|
||
<figureCitation id="59F97BB87126F104FC403F8EFC799D9C" box="[925,1020,478,505]" captionStart="Fig" captionStartId="5.[189,232,1860,1886]" captionTargetBox="[221,1369,611,1815]" captionTargetId="figure@5.[189,1398,598,1820]" captionTargetPageId="5" captionText="Fig. 3. Tomocerus ocreatus Denis, 1948. A. Diagram of tibiotarsal distal chaetae (distal view; t = tenent hair; a = accessory chaeta; g = guard chaeta; p = pretarsal chaeta, same as below). B. Hind claw (lateral view). C. Tenaculum (anterior view). D. Left side of manubrium (dorsal view). E. Disto-external corner of manubrium (dorsal view). F. Basal and middle subsegments of dens with dental spines (dorsal view). G. Mucro (dorso-inner view). Scale bars: B, C, E, G = 50 μm; D, F = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.3832764" httpUri="https://zenodo.org/record/3832764/files/figure.png" pageId="5" pageNumber="6">Fig. 3D</figureCitation>
|
||
); external corner chaeta as large as small mesochaetae in chaetal stripe (
|
||
<figureCitation id="59F97BB87126F104FD413C52FD7F9E79" box="[668,762,514,540]" captionStart="Fig" captionStartId="5.[189,232,1860,1886]" captionTargetBox="[221,1369,611,1815]" captionTargetId="figure@5.[189,1398,598,1820]" captionTargetPageId="5" captionText="Fig. 3. Tomocerus ocreatus Denis, 1948. A. Diagram of tibiotarsal distal chaetae (distal view; t = tenent hair; a = accessory chaeta; g = guard chaeta; p = pretarsal chaeta, same as below). B. Hind claw (lateral view). C. Tenaculum (anterior view). D. Left side of manubrium (dorsal view). E. Disto-external corner of manubrium (dorsal view). F. Basal and middle subsegments of dens with dental spines (dorsal view). G. Mucro (dorso-inner view). Scale bars: B, C, E, G = 50 μm; D, F = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.3832764" httpUri="https://zenodo.org/record/3832764/files/figure.png" pageId="5" pageNumber="6">Fig. 3E</figureCitation>
|
||
). Dens basally without inner modified scale or outer strong chaetae. Dental spines formula as 3/4, II, distal spine strongest, about 0.1 times as long as dens; all spines with numerous denticles of moderate size (
|
||
<figureCitation id="59F97BB87125F107FCF03F7FFC039D2C" box="[813,902,303,329]" captionStart="Fig" captionStartId="5.[189,232,1860,1886]" captionTargetBox="[221,1369,611,1815]" captionTargetId="figure@5.[189,1398,598,1820]" captionTargetPageId="5" captionText="Fig. 3. Tomocerus ocreatus Denis, 1948. A. Diagram of tibiotarsal distal chaetae (distal view; t = tenent hair; a = accessory chaeta; g = guard chaeta; p = pretarsal chaeta, same as below). B. Hind claw (lateral view). C. Tenaculum (anterior view). D. Left side of manubrium (dorsal view). E. Disto-external corner of manubrium (dorsal view). F. Basal and middle subsegments of dens with dental spines (dorsal view). G. Mucro (dorso-inner view). Scale bars: B, C, E, G = 50 μm; D, F = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.3832764" httpUri="https://zenodo.org/record/3832764/files/figure.png" pageId="6" pageNumber="7">Fig. 3F</figureCitation>
|
||
). Dens dorsally with ordinary chaetae and feather-like chaetae as typical in
|
||
<taxonomicName id="06C21CBE7125F107FD923F02FD6E9D09" authorityName="Schaffer" authorityYear="1896" box="[591,747,338,364]" class="Entognatha" family="Tomoceridae" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="3" phylum="Arthropoda" rank="subFamily" subFamily="Tomocerinae">Tomocerinae</taxonomicName>
|
||
, ventrally with only scales. Mucro elongated, bearing numerous smooth chaetae with elongated sockets; both basal teeth with proximal lamellae, the outer tooth with a toothlet; apical and subapical tooth subequal; structure of dorsal lamellae of
|
||
<taxonomicName id="06C21CBE7125F107FB2A3FCBFAF29DD1" authorityName="Nicolet" authorityYear="1842" box="[1271,1399,411,436]" class="Entognatha" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F3B6BB2F7125F107FB2A3FCBFAF29DD1" box="[1271,1399,411,436]" italics="true" pageId="6" pageNumber="7">Tomocerus</emphasis>
|
||
</taxonomicName>
|
||
<typeStatus id="1E79D99F7125F107FF603FEEFF779DBD" box="[189,242,446,472]" pageId="6" pageNumber="7">type</typeStatus>
|
||
, two dorsal lamellae running from subapical tooth, outer lamella ending in inner basal tooth, inner lamella ending freely at base of mucro; outer lamella with 9–10 subequal intermediate teeth (
|
||
<figureCitation id="59F97BB87125F107FADC3FB1FADA9D9E" box="[1281,1375,481,507]" captionStart="Fig" captionStartId="5.[189,232,1860,1886]" captionTargetBox="[221,1369,611,1815]" captionTargetId="figure@5.[189,1398,598,1820]" captionTargetPageId="5" captionText="Fig. 3. Tomocerus ocreatus Denis, 1948. A. Diagram of tibiotarsal distal chaetae (distal view; t = tenent hair; a = accessory chaeta; g = guard chaeta; p = pretarsal chaeta, same as below). B. Hind claw (lateral view). C. Tenaculum (anterior view). D. Left side of manubrium (dorsal view). E. Disto-external corner of manubrium (dorsal view). F. Basal and middle subsegments of dens with dental spines (dorsal view). G. Mucro (dorso-inner view). Scale bars: B, C, E, G = 50 μm; D, F = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.3832764" httpUri="https://zenodo.org/record/3832764/files/figure.png" pageId="6" pageNumber="7">Fig. 3G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="95BD37B57126F104FF603914FBD79B8E" ID-DOI="http://doi.org/10.5281/zenodo.3832764" ID-Zenodo-Dep="3832764" httpUri="https://zenodo.org/record/3832764/files/figure.png" pageId="5" pageNumber="6" startId="5.[189,232,1860,1886]" targetBox="[221,1369,611,1815]" targetPageId="5">
|
||
<paragraph id="C17D673D7126F104FF603914FBD79B8E" blockId="5.[189,1399,1860,2028]" pageId="5" pageNumber="6">
|
||
<emphasis id="F3B6BB2F7126F104FF603914FE8F9B3B" bold="true" box="[189,266,1860,1886]" pageId="5" pageNumber="6">Fig. 3.</emphasis>
|
||
<taxonomicName id="06C21CBE7126F104FECC3915FD169B3B" authority="Denis, 1948" authorityName="Denis" authorityYear="1948" box="[273,659,1860,1886]" class="Entognatha" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Collembola" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="ocreatus">
|
||
<emphasis id="F3B6BB2F7126F104FECC3915FE7E9B3B" box="[273,507,1861,1886]" italics="true" pageId="5" pageNumber="6">Tomocerus ocreatus</emphasis>
|
||
<bibRefCitation id="A5531ACC7126F104FDDF3914FD169B3B" author="Denis J. R." box="[514,659,1860,1886]" pageId="5" pageNumber="6" pagination="183 - 311" refId="ref6522" refString="Denis J. R. 1948. Collemboles d'Indochine. Notes d'Entomologie Chinoise 12: 183 - 311." type="journal article" year="1948">Denis, 1948</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="F3B6BB2F7126F104FD433914FD369B3B" bold="true" box="[670,691,1860,1886]" pageId="5" pageNumber="6">A</emphasis>
|
||
. Diagram of tibiotarsal distal chaetae (distal view; t = tenent hair; a = accessory chaeta; g = guard chaeta; p = pretarsal chaeta, same as below).
|
||
<emphasis id="F3B6BB2F7126F104FB5F3937FB139BE4" bold="true" box="[1154,1174,1895,1921]" pageId="5" pageNumber="6">B</emphasis>
|
||
. Hind claw (lateral view).
|
||
<emphasis id="F3B6BB2F7126F104FED339DAFEA69BC1" bold="true" box="[270,291,1930,1956]" pageId="5" pageNumber="6">C</emphasis>
|
||
. Tenaculum (anterior view).
|
||
<emphasis id="F3B6BB2F7126F104FDA839DAFD0F9BC1" bold="true" box="[629,650,1930,1956]" pageId="5" pageNumber="6">D</emphasis>
|
||
. Left side of manubrium (dorsal view).
|
||
<emphasis id="F3B6BB2F7126F104FB8139DAFBF59BC1" bold="true" box="[1116,1136,1930,1956]" pageId="5" pageNumber="6">E</emphasis>
|
||
. Disto-external corner of manubrium (dorsal view).
|
||
<emphasis id="F3B6BB2F7126F104FDCE39FEFDA09BAD" bold="true" box="[531,549,1966,1992]" pageId="5" pageNumber="6">F</emphasis>
|
||
. Basal and middle subsegments of dens with dental spines (dorsal view).
|
||
<emphasis id="F3B6BB2F7126F104FF603980FF519B8F" bold="true" box="[189,212,2000,2026]" pageId="5" pageNumber="6">G</emphasis>
|
||
. Mucro (dorso-inner view). Scale bars: B, C, E, G = 50 μm; D, F = 100 μm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="89D834B67125F106FF603C73FAF29E5A" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="discussion">
|
||
<paragraph id="C17D673D7125F107FF603C73FEAA9E58" blockId="6.[189,303,547,573]" box="[189,303,547,573]" pageId="6" pageNumber="7">
|
||
<heading id="9A35D0517125F107FF603C73FEAA9E58" bold="true" box="[189,303,547,573]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
|
||
<emphasis id="F3B6BB2F7125F107FF603C73FEAA9E58" bold="true" box="[189,303,547,573]" pageId="6" pageNumber="7">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="C17D673D7125F107FF603C01FCF29FAA" blockId="6.[189,1399,593,976]" pageId="6" pageNumber="7">
|
||
<bibRefCitation id="A5531ACC7125F107FF603C01FEDD9E09" author="Denis J. R." box="[189,344,593,620]" pageId="6" pageNumber="7" pagination="183 - 311" refId="ref6522" refString="Denis J. R. 1948. Collemboles d'Indochine. Notes d'Entomologie Chinoise 12: 183 - 311." type="journal article" year="1948">Denis (1948)</bibRefCitation>
|
||
provided accurate description for
|
||
<taxonomicName id="06C21CBE7125F107FD323C02FC5E9E0E" authorityName="Denis" authorityYear="1948" box="[751,987,594,619]" class="Entognatha" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="ocreatus">
|
||
<emphasis id="F3B6BB2F7125F107FD323C02FC5E9E0E" box="[751,987,594,619]" italics="true" pageId="6" pageNumber="7">Tomocerus ocreatus</emphasis>
|
||
</taxonomicName>
|
||
. For instance, he described clearly the dental spines as “covered with secondary spines, more similar to scales than denticles”, and the figure showed that those scale-like denticles were of moderate size and covered at least basal half of each spine, on which account we rejected some records of non-type
|
||
<taxonomicName id="06C21CBE7125F107FC2D3CEDFB599EB3" authorityName="Denis" authorityYear="1948" box="[1008,1244,701,726]" class="Entognatha" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="ocreatus">
|
||
<emphasis id="F3B6BB2F7125F107FC2D3CEDFB599EB3" box="[1008,1244,701,726]" italics="true" pageId="6" pageNumber="7">Tomocerus ocreatus</emphasis>
|
||
</taxonomicName>
|
||
(see below). But naturally, some later revealed characters were not mentioned by him. For instance, chaetotaxy had not been used for taxonomy in
|
||
<taxonomicName id="06C21CBE7125F107FDEC3D53FD4F9F78" authorityName="Schaffer" authorityYear="1896" box="[561,714,771,797]" class="Entognatha" family="Tomoceridae" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="3" phylum="Arthropoda" rank="subFamily" subFamily="Tomocerinae">Tomocerinae</taxonomicName>
|
||
before
|
||
<bibRefCitation id="A5531ACC7125F107FCF83D53FC399F7B" author="Yosii R." box="[805,956,771,798]" pageId="6" pageNumber="7" pagination="1 - 109" refId="ref7076" refString="Yosii R. 1956. Monographie zur Hohlencollembolen Japans. Contributions from the Biological Laboratory Kyoto University 3: 1 - 109." type="journal article" year="1956">Yosii (1956)</bibRefCitation>
|
||
. The limit of original description is a main reason for the misidentification of
|
||
<taxonomicName id="06C21CBE7125F107FD7E3D78FC149F24" authorityName="Denis" authorityYear="1948" box="[675,913,808,833]" class="Entognatha" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="ocreatus">
|
||
<emphasis id="F3B6BB2F7125F107FD7E3D78FC149F24" box="[675,913,808,833]" italics="true" pageId="6" pageNumber="7">Tomocerus ocreatus</emphasis>
|
||
</taxonomicName>
|
||
in some subsequent records, and finally turned the species into a complex including a number of closely related forms from
|
||
<collectingCountry id="B9D527AD7125F107FB473D1AFB789F01" box="[1178,1277,842,868]" name="Vietnam" pageId="6" pageNumber="7">Vietnam</collectingCountry>
|
||
to eastern
|
||
<collectingCountry id="B9D527AD7125F107FF603D3EFE8A9FED" box="[189,271,878,904]" name="Russia" pageId="6" pageNumber="7">Russia</collectingCountry>
|
||
. To overcome the deep confusion about this species, a redescription of type specimen is necessary. Because the
|
||
<typeStatus id="1E79D99F7125F107FE8D3DC2FE329FC9" box="[336,439,914,940]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
|
||
and unique type specimen of
|
||
<taxonomicName id="06C21CBE7125F107FCC93DC3FC7A9FC9" authorityName="Denis" authorityYear="1948" box="[788,1023,915,940]" class="Entognatha" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="ocreatus">
|
||
<emphasis id="F3B6BB2F7125F107FCC93DC3FC7A9FC9" box="[788,1023,915,940]" italics="true" pageId="6" pageNumber="7">Tomocerus ocreatus</emphasis>
|
||
</taxonomicName>
|
||
was lost, we propose to set up a
|
||
<typeStatus id="1E79D99F7125F107FF603DE6FE9F9FB5" box="[189,282,950,976]" pageId="6" pageNumber="7" type="neotype">neotype</typeStatus>
|
||
for the species from specimens of the type locality.
|
||
</paragraph>
|
||
<paragraph id="C17D673D7125F107FF603DA7FBBF994B" blockId="6.[189,1399,1015,1326]" pageId="6" pageNumber="7">
|
||
<bibRefCitation id="A5531ACC7125F107FF603DA7FEDC9874" author="Denis J. R." box="[189,345,1015,1041]" pageId="6" pageNumber="7" pagination="183 - 311" refId="ref6522" refString="Denis J. R. 1948. Collemboles d'Indochine. Notes d'Entomologie Chinoise 12: 183 - 311." type="journal article" year="1948">Denis (1948)</bibRefCitation>
|
||
described
|
||
<taxonomicName id="06C21CBE7125F107FE073DA8FD429874" authorityName="Denis" authorityYear="1948" box="[474,711,1016,1041]" class="Entognatha" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="ocreatus">
|
||
<emphasis id="F3B6BB2F7125F107FE073DA8FD429874" box="[474,711,1016,1041]" italics="true" pageId="6" pageNumber="7">Tomocerus ocreatus</emphasis>
|
||
</taxonomicName>
|
||
on one specimen, from “plateau du Lang Biang,
|
||
<quantity id="063ACAD87125F107FAC83DA7FAF79877" box="[1301,1394,1015,1042]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="6" pageNumber="7" unit="m" value="2400.0">2400 m</quantity>
|
||
. alt., forêt tropicale”. It is not the Lang Bian peak itself that reaches
|
||
<quantity id="063ACAD87125F107FBD83A4BFBE79850" box="[1029,1122,1051,1077]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="6" pageNumber="7" unit="m" value="2400.0">2400 m</quantity>
|
||
but the Chu Yang Sin massif north of the Lang Bian (= Dalat) plateau, the Lang Bian peak being only
|
||
<quantity id="063ACAD87125F107FB4C3A6EFB68983C" box="[1169,1261,1086,1113]" metricMagnitude="3" metricUnit="m" metricValue="2.197" pageId="6" pageNumber="7" unit="m" value="2197.0">2197 m</quantity>
|
||
in altitude. The Chu Yang Sin massif was and still is of very difficult access, and was certainly not the place where the collector, Dawydoff, operated. There are two main patches of subtropical forests around Dalat: a small one on the Lang Bian peak, and a much larger one
|
||
<quantity id="063ACAD87125F107FC8B3AF9FC1A98A6" box="[854,927,1193,1220]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="6" pageNumber="7" unit="km" value="25.0">25 km</quantity>
|
||
northeast on the Bi Doup massif, which reaches
|
||
<quantity id="063ACAD87125F107FEC33A9DFEFC9882" box="[286,377,1229,1255]" metricMagnitude="3" metricUnit="m" metricValue="2.287" pageId="6" pageNumber="7" unit="m" value="2287.0">2287 m</quantity>
|
||
and is less easy to access. In any case, it is most likely on the slopes of one of these massifs that
|
||
<taxonomicName id="06C21CBE7125F107FE893AA1FDC5996F" authorityName="Denis" authorityYear="1948" box="[340,576,1265,1290]" class="Entognatha" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="ocreatus">
|
||
<emphasis id="F3B6BB2F7125F107FE893AA1FDC5996F" box="[340,576,1265,1290]" italics="true" pageId="6" pageNumber="7">Tomocerus ocreatus</emphasis>
|
||
</taxonomicName>
|
||
was collected. On this account, we propose to designate a specimen that we obtained from litter sample in the Bi Doup mountain as the
|
||
<typeStatus id="1E79D99F7125F107FC0B3B44FBB2994B" box="[982,1079,1300,1326]" pageId="6" pageNumber="7" type="neotype">neotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="C17D673D7125F107FF603B06FBA09A23" blockId="6.[189,1399,1366,1606]" pageId="6" pageNumber="7">
|
||
The
|
||
<typeStatus id="1E79D99F7125F107FF2C3B06FECB9915" box="[241,334,1366,1392]" pageId="6" pageNumber="7" type="neotype">neotype</typeStatus>
|
||
is highly identical with the original description in almost all described characters, including the body colour, the length of antennae, the structure of claws and the morphology and arrangement of dental spines. The only difference is that in the
|
||
<typeStatus id="1E79D99F7125F107FD283BCDFCD799D2" box="[757,850,1437,1463]" pageId="6" pageNumber="7" type="neotype">neotype</typeStatus>
|
||
specimen the numbers of ungual teeth, dental spines and mucronal teeth are slightly larger than in the original one. It could happen in
|
||
<taxonomicName id="06C21CBE7125F107FB033B90FAF299BF" authorityName="Borner" authorityYear="1913" box="[1246,1399,1472,1498]" class="Entognatha" family="Tomoceridae" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="7" phylum="Arthropoda" rank="family">Tomoceridae</taxonomicName>
|
||
that within the same species larger individuals have a few more teeth and dental spines than smaller ones, so the slight difference mentioned above can be treated as intraspecific considering the
|
||
<typeStatus id="1E79D99F7125F107FAC73858FAF29A47" box="[1306,1399,1544,1570]" pageId="6" pageNumber="7" type="neotype">neotype</typeStatus>
|
||
specimen is slightly larger than the lost
|
||
<typeStatus id="1E79D99F7125F107FD52387BFD739A20" box="[655,758,1579,1605]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
|
||
(3.6 mm versus 3.3 mm).
|
||
</paragraph>
|
||
<paragraph id="C17D673D7125F106FF60383DFAF29E5A" blockId="6.[189,1399,1645,2027]" lastBlockId="7.[189,1399,267,575]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
|
||
Regarding previous non-type records of
|
||
<taxonomicName id="06C21CBE7125F107FD55383EFBCE9AED" authority=", Stach (1964" authorityName=", Stach" authorityYear="1964" box="[648,1099,1645,1672]" class="Entognatha" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="ocreatus">
|
||
<emphasis id="F3B6BB2F7125F107FD55383EFCE89AE2" box="[648,877,1646,1671]" italics="true" pageId="6" pageNumber="7">Tomocerus ocreatus</emphasis>
|
||
,
|
||
<bibRefCitation id="A5531ACC7125F107FCA7383DFB809AE2" author="Stach J." box="[890,1029,1645,1671]" pageId="6" pageNumber="7" pagination="1 - 26" refId="ref6959" refString="Stach J. 1964. Materials to the knowledge of Chinese Collembolan Fauna. Acta Zoologica Cracoviensia 9: 1 - 26." type="journal article" year="1964">Stach (1964</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="A5531ACC7125F107FBCD383DFBCE9AED" author="Stach J." box="[1040,1099,1645,1672]" pageId="6" pageNumber="7" pagination="346 - 372" refId="ref6982" refString="Stach J. 1965. On some Collembola of North Vietnam. Acta Zoologica Cracoviensia 10: 346 - 372." type="journal article" year="1965">1965</bibRefCitation>
|
||
</taxonomicName>
|
||
) redescribed two different forms of
|
||
<taxonomicName id="06C21CBE7125F107FEFF38C1FD839ACF" authorityName=", Stach" authorityYear="1964" box="[290,518,1681,1706]" class="Entognatha" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="ocreatus">
|
||
<emphasis id="F3B6BB2F7125F107FEFF38C1FD839ACF" box="[290,518,1681,1706]" italics="true" pageId="6" pageNumber="7">Tomocerus ocreatus</emphasis>
|
||
</taxonomicName>
|
||
from southeastern
|
||
<collectingCountry id="B9D527AD7125F107FD0638C0FCA59ACF" box="[731,800,1680,1706]" name="China" pageId="6" pageNumber="7">China</collectingCountry>
|
||
and northern
|
||
<collectingCountry id="B9D527AD7125F107FC6438C0FB989ACF" box="[953,1053,1680,1706]" name="Vietnam" pageId="6" pageNumber="7">Vietnam</collectingCountry>
|
||
, respectively, and also claimed that the Japanese record of “
|
||
<taxonomicName id="06C21CBE7125F107FDDE38E5FD499AAB" baseAuthorityName="Lubbock" baseAuthorityYear="1862" box="[515,716,1717,1742]" class="Entognatha" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="minor">
|
||
<emphasis id="F3B6BB2F7125F107FDDE38E5FD499AAB" box="[515,716,1717,1742]" italics="true" pageId="6" pageNumber="7">Tomocerus minor</emphasis>
|
||
</taxonomicName>
|
||
” by
|
||
<bibRefCitation id="A5531ACC7125F107FCDE38E4FC2F9AAA" author="Uchida H." box="[771,938,1716,1743]" pageId="6" pageNumber="7" pagination="1 - 6" refId="ref7003" refString="Uchida H. 1953. The insect fauna of Mt. Ishizuchi and Omogo Valley, Iyo, Japan. The Collembola. Transactions of the Shikoku Entomological Society 4: 1 - 6." type="journal article" year="1953">Uchida (1953)</bibRefCitation>
|
||
was actually
|
||
<taxonomicName id="06C21CBE7125F107FB9A38E5FAA89AAB" authorityName=", Stach" authorityYear="1964" box="[1095,1325,1717,1742]" class="Entognatha" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="ocreatus">
|
||
<emphasis id="F3B6BB2F7125F107FB9A38E5FAA89AAB" box="[1095,1325,1717,1742]" italics="true" pageId="6" pageNumber="7">Tomocerus ocreatus</emphasis>
|
||
</taxonomicName>
|
||
; Yosii described a Japanese species
|
||
<taxonomicName id="06C21CBE7125F107FDD83888FC279A97" authority="(Yosii, 1954)" baseAuthorityName="Yosii" baseAuthorityYear="1954" box="[517,930,1751,1778]" class="Entognatha" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="kawamurai">
|
||
<emphasis id="F3B6BB2F7125F107FDD83888FC8C9A97" box="[517,777,1752,1778]" italics="true" pageId="6" pageNumber="7">Tomocerus kawamurai</emphasis>
|
||
(Yosii, 1954)
|
||
</taxonomicName>
|
||
and then synonymized it with
|
||
<taxonomicName id="06C21CBE7125F107FB243888FE9A9B70" authorityName=", Stach" authorityYear="1964" class="Entognatha" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="ocreatus">
|
||
<emphasis id="F3B6BB2F7125F107FB243888FE9A9B70" italics="true" pageId="6" pageNumber="7">Tomocerus ocreatus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="A5531ACC7125F107FEF238ABFE359B73" author="Yosii R." box="[303,432,1787,1814]" pageId="6" pageNumber="7" pagination="1 - 109" refId="ref7076" refString="Yosii R. 1956. Monographie zur Hohlencollembolen Japans. Contributions from the Biological Laboratory Kyoto University 3: 1 - 109." type="journal article" year="1956">Yosii 1956</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="A5531ACC7125F107FE6038ABFE7E9B73" author="Yosii R." box="[445,507,1787,1814]" pageId="6" pageNumber="7" pagination="1 - 54" refId="ref7162" refString="Yosii R. 1967. Studies on the Collembolan family Tomoceridae, with special reference to Japanese forms. Contributions from the Biological Laboratory Kyoto University 20: 1 - 54." type="journal article" year="1967">1967</bibRefCitation>
|
||
);
|
||
<bibRefCitation id="A5531ACC7125F107FDCC38ABFD2F9B73" author="Chiba S." box="[529,682,1787,1814]" pageId="6" pageNumber="7" pagination="24 - 26" refId="ref6362" refString="Chiba S. 1968. Collembola of the Mt. Hakkoda area. I. Family Tomoceridae. The Science Reports of the Hirosaki University 15: 24 - 26." type="journal article" year="1968">Chiba (1968)</bibRefCitation>
|
||
recognized three forms in Japanese
|
||
<taxonomicName id="06C21CBE7125F107FB8938ACFABE9B70" authorityName=", Stach" authorityYear="1964" box="[1108,1339,1788,1813]" class="Entognatha" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="ocreatus">
|
||
<emphasis id="F3B6BB2F7125F107FB8938ACFABE9B70" box="[1108,1339,1788,1813]" italics="true" pageId="6" pageNumber="7">Tomocerus ocreatus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="A5531ACC7125F107FA9638ACFE8C9B5F" author="Lee B. H." pageId="6" pageNumber="7" pagination="945 - 961" refId="ref6740" refString="Lee B. H. 1975. Etude de la faune coreenne des Insectes Collemboles. VI. Sur la famille des Tomoceridae, edaphiques, avec la description de quatre nouvelles especes et d'une nouvelle sous-espece. Bulletin du Museum national d'Histoire naturelle, serie 3 317: 945 - 961." type="journal article" year="1975">Lee (1975)</bibRefCitation>
|
||
made descriptive notes on Korean specimens;
|
||
<bibRefCitation id="A5531ACC7125F107FCF6394FFC789B5C" author="Martynova E. F." box="[811,1021,1823,1849]" pageId="6" pageNumber="7" pagination="3 - 16" refId="ref6899" refString="Martynova E. F. 1977. Springtails of the family Tomoceridae (Collembola) from the fauna of the Far East. Insect Fauna of the Far East 46: 3 - 16." type="journal article" year="1977">Martynova (1977)</bibRefCitation>
|
||
recorded
|
||
<taxonomicName id="06C21CBE7125F107FBAC3970FADD9B5C" authorityName=", Stach" authorityYear="1964" box="[1137,1368,1824,1849]" class="Entognatha" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="ocreatus">
|
||
<emphasis id="F3B6BB2F7125F107FBAC3970FADD9B5C" box="[1137,1368,1824,1849]" italics="true" pageId="6" pageNumber="7">Tomocerus ocreatus</emphasis>
|
||
</taxonomicName>
|
||
in Sakhalin and made some descriptive notes on it. These redescriptions had subtle to considerable differences to each other, and were never identical to the original description. The Chinese “
|
||
<taxonomicName id="06C21CBE7125F107FBA73937FB599BE5" authorityName=", Stach" authorityYear="1964" box="[1146,1244,1895,1920]" class="Entognatha" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="ocreatus">
|
||
<emphasis id="F3B6BB2F7125F107FBA73937FB599BE5" box="[1146,1244,1895,1920]" italics="true" pageId="6" pageNumber="7">ocreatus</emphasis>
|
||
</taxonomicName>
|
||
” recorded in Hangzhou (“Hangchow”),
|
||
<collectingRegion id="0306A9DF7125F107FE2A39D9FD489BC6" box="[503,717,1929,1955]" country="China" name="Zhejiang" pageId="6" pageNumber="7">Zhejiang Province</collectingRegion>
|
||
by
|
||
<bibRefCitation id="A5531ACC7125F107FD2039D9FC169BC1" author="Stach J." box="[765,915,1929,1956]" pageId="6" pageNumber="7" pagination="1 - 26" refId="ref6959" refString="Stach J. 1964. Materials to the knowledge of Chinese Collembolan Fauna. Acta Zoologica Cracoviensia 9: 1 - 26." type="journal article" year="1964">Stach (1964)</bibRefCitation>
|
||
has brownish body colour and distinctly short antenna about half the length of body; the northern Vietnamese record from
|
||
<collectingRegion id="0306A9DF7125F107FB5E39FEFAD59BA2" box="[1155,1360,1965,1992]" country="Vietnam" name="Lao Cai" pageId="6" pageNumber="7">Lao Cai Province</collectingRegion>
|
||
by the same author (
|
||
<bibRefCitation id="A5531ACC7125F107FE5F3981FD809B8E" author="Stach J." box="[386,517,2000,2027]" pageId="6" pageNumber="7" pagination="346 - 372" refId="ref6982" refString="Stach J. 1965. On some Collembola of North Vietnam. Acta Zoologica Cracoviensia 10: 346 - 372." type="journal article" year="1965">Stach 1965</bibRefCitation>
|
||
) bears a small dental spine between two large distal spines, and the denticles on spines are finer than those in the original description, therefore it is more similar to
|
||
<taxonomicName id="06C21CBE7124F106FB433F5CFECD9D2C" authority="Denis, 1929" authorityName="Denis" authorityYear="1929" class="Entognatha" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Collembola" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="folsomi">
|
||
<emphasis id="F3B6BB2F7124F106FB433F5CFAF39D40" box="[1182,1398,267,293]" italics="true" pageId="7" pageNumber="8">Tomocerus folsomi</emphasis>
|
||
<bibRefCitation id="A5531ACC7124F106FF603F7EFECD9D2C" author="Denis J. R." box="[189,328,302,329]" pageId="7" pageNumber="8" pagination="166 - 180" refId="ref6477" refString="Denis J. R. 1929. Notes sur les collemboles recoltes dans ses voyages par le Prof. F. Silvestri. Bollettino del Laboratorio di Zoologia generale e Agraria della R. Scuola Superiore d'Agricoltura in Portici 22: 166 - 180." type="journal article" year="1929">Denis, 1929</bibRefCitation>
|
||
</taxonomicName>
|
||
from
|
||
<collectingRegion id="0306A9DF7124F106FE533F7FFDD29D2D" box="[398,599,302,329]" country="China" name="Yunnan" pageId="7" pageNumber="8">Yunnan Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="B9D527AD7124F106FDBE3F7EFD2E9D2D" box="[611,683,302,328]" name="China" pageId="7" pageNumber="8">China</collectingCountry>
|
||
; the Japanese “
|
||
<taxonomicName id="06C21CBE7124F106FC813F7FFC3B9D2D" authorityName=", Stach" authorityYear="1964" box="[860,958,303,328]" class="Entognatha" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Collembola" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ocreatus">
|
||
<emphasis id="F3B6BB2F7124F106FC813F7FFC3B9D2D" box="[860,958,303,328]" italics="true" pageId="7" pageNumber="8">ocreatus</emphasis>
|
||
</taxonomicName>
|
||
” described by
|
||
<bibRefCitation id="A5531ACC7124F106FBB63F7EFB7F9D2C" author="Yosii R." box="[1131,1274,302,329]" pageId="7" pageNumber="8" pagination="1 - 54" refId="ref7162" refString="Yosii R. 1967. Studies on the Collembolan family Tomoceridae, with special reference to Japanese forms. Contributions from the Biological Laboratory Kyoto University 20: 1 - 54." type="journal article" year="1967">Yosii (1967)</bibRefCitation>
|
||
has dorsal scales and 2+2 blunt principal chaetae on manubrium which are absent in the true
|
||
<taxonomicName id="06C21CBE7124F106FB963F03FAAA9D09" authorityName=", Stach" authorityYear="1964" box="[1099,1327,339,364]" class="Entognatha" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Collembola" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ocreatus">
|
||
<emphasis id="F3B6BB2F7124F106FB963F03FAAA9D09" box="[1099,1327,339,364]" italics="true" pageId="7" pageNumber="8">Tomocerus ocreatus</emphasis>
|
||
</taxonomicName>
|
||
; other records (
|
||
<bibRefCitation id="A5531ACC7124F106FEFD3F25FE2F9DEA" author="Chiba S." box="[288,426,373,399]" pageId="7" pageNumber="8" pagination="24 - 26" refId="ref6362" refString="Chiba S. 1968. Collembola of the Mt. Hakkoda area. I. Family Tomoceridae. The Science Reports of the Hirosaki University 15: 24 - 26." type="journal article" year="1968">Chiba 1968</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="A5531ACC7124F106FE683F25FDA69DF5" author="Lee B. H." box="[437,547,373,400]" pageId="7" pageNumber="8" pagination="945 - 961" refId="ref6740" refString="Lee B. H. 1975. Etude de la faune coreenne des Insectes Collemboles. VI. Sur la famille des Tomoceridae, edaphiques, avec la description de quatre nouvelles especes et d'une nouvelle sous-espece. Bulletin du Museum national d'Histoire naturelle, serie 3 317: 945 - 961." type="journal article" year="1975">Lee 1975</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="A5531ACC7124F106FDF33F25FD6B9DEA" author="Martynova E. F." box="[558,750,373,399]" pageId="7" pageNumber="8" pagination="3 - 16" refId="ref6899" refString="Martynova E. F. 1977. Springtails of the family Tomoceridae (Collembola) from the fauna of the Far East. Insect Fauna of the Far East 46: 3 - 16." type="journal article" year="1977">Martynova 1977</bibRefCitation>
|
||
) all have dental spines with one or two whorls of crownlike denticles near the base which is not the morphology of the true
|
||
<taxonomicName id="06C21CBE7124F106FC0C3FC9FBB69DD7" authorityName=", Stach" authorityYear="1964" box="[977,1075,409,434]" class="Entognatha" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Collembola" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ocreatus">
|
||
<emphasis id="F3B6BB2F7124F106FC0C3FC9FBB69DD7" box="[977,1075,409,434]" italics="true" pageId="7" pageNumber="8">ocreatus</emphasis>
|
||
</taxonomicName>
|
||
form. But these remarkable differences were treated as intraspecific variations when other characters showed high similarity. Inferred from the present morphological review and the molecular study on Chinese
|
||
<taxonomicName id="06C21CBE7124F106FBCB3F8FFBFD9D9D" authorityName=", Stach" authorityYear="1964" box="[1046,1144,479,504]" class="Entognatha" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Collembola" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ocreatus">
|
||
<emphasis id="F3B6BB2F7124F106FBCB3F8FFBFD9D9D" box="[1046,1144,479,504]" italics="true" pageId="7" pageNumber="8">ocreatus</emphasis>
|
||
</taxonomicName>
|
||
complex (
|
||
<bibRefCitation id="A5531ACC7124F106FB333F8EFF7F9E79" author="Zhang F. & Yu D. Y. & Luo Y. Z. & Ho S. Y. W. & Wang B. X. & Zhu C. D." pageId="7" pageNumber="8" pagination="393 - 404" refId="ref7375" refString="Zhang F., Yu D. Y., Luo Y. Z., Ho S. Y. W., Wang B. X. & Zhu C. D. 2014. Cryptic diversity, diversification and vicariance in two species complexes of Tomocerus (Collembola, Tomoceridae) from China. Zoologica Scripta 43 (4): 393 - 404. http: // dx. doi. org / 10.1111 / zsc. 12056" type="journal article" year="2014">
|
||
Zhang
|
||
<emphasis id="F3B6BB2F7124F106FAE33F8FFAF39D9C" box="[1342,1398,479,505]" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
|
||
2014
|
||
</bibRefCitation>
|
||
), the aforementioned non-type “
|
||
<taxonomicName id="06C21CBE7124F106FDB53C53FD4F9E79" authorityName=", Stach" authorityYear="1964" box="[616,714,515,540]" class="Entognatha" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Collembola" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ocreatus">
|
||
<emphasis id="F3B6BB2F7124F106FDB53C53FD4F9E79" box="[616,714,515,540]" italics="true" pageId="7" pageNumber="8">ocreatus</emphasis>
|
||
</taxonomicName>
|
||
” may each represent an independent species in the complex, and so far the true
|
||
<taxonomicName id="06C21CBE7124F106FE523C76FDF19E5A" authorityName=", Stach" authorityYear="1964" box="[399,628,550,575]" class="Entognatha" family="Tomoceridae" genus="Tomocerus" kingdom="Animalia" order="Collembola" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ocreatus">
|
||
<emphasis id="F3B6BB2F7124F106FE523C76FDF19E5A" box="[399,628,550,575]" italics="true" pageId="7" pageNumber="8">Tomocerus ocreatus</emphasis>
|
||
</taxonomicName>
|
||
is known as only endemic to southern Annamitic range in
|
||
<collectingCountry id="B9D527AD7124F106FAD33C75FAF69E5A" box="[1294,1395,549,575]" name="Vietnam" pageId="7" pageNumber="8">Vietnam</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |