treatments-xml/data/49/32/87/4932878EFF9BFFA266A0FD27FA09FA55.xml

<document id="FD15CDA97FD98D3858C795F0AD5FA8E5" ID-CLB-Dataset="46195" ID-DOI="10.11646/zootaxa.3779.5.6" ID-GBIF-Dataset="b798f275-02d9-456e-98d0-f116607218b2" ID-ISSN="1175-5326" ID-Zenodo-Dep="226678" ID-ZooBank="491EB308-B6CD-411C-AF74-78B37D8C526F" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460398479612" checkinUser="plazi" docAuthor="Archangelsky, Miguel &amp; Brand, Cecilia" docDate="2014" docId="4932878EFF9BFFA266A0FD27FA09FA55" docLanguage="en" docName="zt03779p572.pdf" docOrigin="Zootaxa 3779 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Luchoelmis kapenkemkensis Archangelsky &amp; Brand, 2014, sp. nov." docType="treatment" docVersion="9" lastPageNumber="567" masterDocId="B50BFFF6FF9AFFA66637FF94FF95FFB7" masterDocTitle="A new species of Luchoelmis Spangler &amp; Staines (Coleoptera: Elmidae) from Argentina and its probable larva" masterLastPageNumber="572" masterPageNumber="563" pageNumber="564" updateTime="1698298959420" updateUser="ExternalLinkService">
<mods:mods id="7D5DB450D5B11518766D77E51B74777A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="00859C8DD16A3D7C8D0EDC0DDCE8E776">
<mods:title id="56F4048F459AFC0DA1C75303B068A0CD">A new species of Luchoelmis Spangler &amp; Staines (Coleoptera: Elmidae) from Argentina and its probable larva</mods:title>
</mods:titleInfo>
<mods:name id="0B62425E48D5DA119BF15FBD497A3E53" type="personal">
<mods:role id="7A7587896BEED94C320A0511824258C0">
<mods:roleTerm id="AF80B504033CD1374C4A97B285E142C1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="ACF44AD3DBF3F786134C0757754D3513">Archangelsky, Miguel</mods:namePart>
</mods:name>
<mods:name id="4069DF48F2110113AE1EB5FBB79FC7F4" type="personal">
<mods:role id="E35371F21E820A8FE88D206B7E1F3D66">
<mods:roleTerm id="72BD9E81405B42A4979C0F8C03A34E02">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B9AA2C43176C21609760C619B604E064">Brand, Cecilia</mods:namePart>
</mods:name>
<mods:typeOfResource id="E2EDB6F9E179D313AABBA2BBC7AD5176">text</mods:typeOfResource>
<mods:relatedItem id="E8963625CED75EFE5AB3FAC09E9FBA07" type="host">
<mods:titleInfo id="004FF81C8E54FC71305F85CA3561EC8E">
<mods:title id="D5473E6C8387FB46499F8F43C9B8BC03">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="197A096FB6089ECE561E4673241A7B18">
<mods:date id="9976C9162B214C9B7F57F2735A44895B">2014</mods:date>
<mods:detail id="8701994326FD7AC19B9283752E6431F2" type="volume">
<mods:number id="20226840C561DB7A5121F70113A4160E">3779</mods:number>
</mods:detail>
<mods:detail id="317FE62E75785E467C40492C7035DB32" type="issue">
<mods:number id="342164682742C0EF3F6C38F3EC09DBF7">5</mods:number>
</mods:detail>
<mods:extent id="EFF1DC4F649E950EA315FA267AD427F0" unit="page">
<mods:start id="9D922CE9231BD0F58EF40BA0572035C3">563</mods:start>
<mods:end id="7B6E3F4AD6EB3D263476CCBB537548B1">572</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="AB1129BF32B451AEB6FB192BBDE5B112">journal article</mods:classification>
<mods:identifier id="F5299A160B1C610DFBE126B59189BD80" type="CLB-Dataset">46195</mods:identifier>
<mods:identifier id="C3CBC7960845408D11961C84D8AC68BF" type="DOI">10.11646/zootaxa.3779.5.6</mods:identifier>
<mods:identifier id="3BD0ED7F2A93F8268A295FECF1C28E9C" type="GBIF-Dataset">b798f275-02d9-456e-98d0-f116607218b2</mods:identifier>
<mods:identifier id="DF66D710AB7766577B155EF2E05FB843" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="8D38567C6E222E919141B4F1F2CD5ED7" type="Zenodo-Dep">226678</mods:identifier>
<mods:identifier id="00B498560749BB14CD70E218CBF4D944" type="ZooBank">491EB308-B6CD-411C-AF74-78B37D8C526F</mods:identifier>
</mods:mods>
<treatment id="4932878EFF9BFFA266A0FD27FA09FA55" ID-DOI="http://doi.org/10.5281/zenodo.6134049" ID-GBIF-Taxon="119418913" ID-Zenodo-Dep="6134049" LSID="urn:lsid:plazi:treatment:4932878EFF9BFFA266A0FD27FA09FA55" httpUri="http://treatment.plazi.org/id/4932878EFF9BFFA266A0FD27FA09FA55" lastPageId="4" lastPageNumber="567" pageId="1" pageNumber="564">
<subSubSection id="89816513FF9BFFA766A0FD27FDC2FD7A" box="[151,599,691,717]" pageId="1" pageNumber="564" type="nomenclature">
<paragraph id="C1243698FF9BFFA766A0FD27FDC2FD7A" blockId="1.[151,599,691,717]" box="[151,599,691,717]" pageId="1" pageNumber="564">
<heading id="9A6C81F4FF9BFFA766A0FD27FDC2FD7A" bold="true" box="[151,599,691,717]" fontSize="11" level="1" pageId="1" pageNumber="564" reason="1">
<emphasis id="F3EFEA8AFF9BFFA766A0FD27FDC2FD7A" bold="true" box="[151,599,691,717]" pageId="1" pageNumber="564">
<taxonomicName id="069B4D1BFF9BFFA766A0FD27FE67FD7A" ID-CoL="3WBCC" box="[151,498,691,717]" class="Insecta" family="Elmidae" genus="Luchoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="564" phylum="Arthropoda" rank="species" species="kapenkemkensis" status="sp. nov.">
<emphasis id="F3EFEA8AFF9BFFA766A0FD27FE67FD7A" bold="true" box="[151,498,691,717]" italics="true" pageId="1" pageNumber="564">Luchoelmis kapenkemkensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8DC57F1FF9BFFA767CDFD27FDC2FD7A" box="[506,599,691,717]" pageId="1" pageNumber="564" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="89816513FF9BFFA766A0FD6EFEFEFC14" pageId="1" pageNumber="564" type="materials_examined">
<paragraph id="C1243698FF9BFFA766A0FD6EFE8FFC80" blockId="1.[151,1436,762,2011]" pageId="1" pageNumber="564">
<emphasis id="F3EFEA8AFF9BFFA766A0FD6EFED5FCA4" bold="true" box="[151,320,762,787]" pageId="1" pageNumber="564">
<typeStatus id="1E20883AFF9BFFA766A0FD6EFF47FCA4" box="[151,210,762,787]" pageId="1" pageNumber="564">Type</typeStatus>
material
</emphasis>
. 
<typeStatus id="1E20883AFF9BFFA76778FD6EFE29FCA4" box="[335,444,762,787]" pageId="1" pageNumber="564" type="holotype">
<emphasis id="F3EFEA8AFF9BFFA76778FD6EFE29FCA4" bold="true" box="[335,444,762,787]" pageId="1" pageNumber="564">Holotype</emphasis>
</typeStatus>
(
<emphasis id="F3EFEA8AFF9BFFA767F8FD6EFD9CFCA4" bold="true" box="[463,521,762,787]" pageId="1" pageNumber="564">male</emphasis>
): 
<collectingCountry id="B98C7608FF9BFFA76414FD6EFD00FCA4" box="[547,661,762,787]" name="Argentina" pageId="1" pageNumber="564">Argentina</collectingCountry>
, Chubut Province, Arroyo Nant y Fall int. Rt. prov. 17; 
<date id="B5251058FF9BFFA76329FD6EFAE0FCA4" box="[1310,1397,762,787]" pageId="1" pageNumber="564" value="2008-04">iv.2008</date>
; C. Brand Coll.
</paragraph>
<paragraph id="C1243698FF9BFFA766F0FCD6FEFEFC14" blockId="1.[151,1436,762,2011]" pageId="1" pageNumber="564">
<emphasis id="F3EFEA8AFF9BFFA766F0FCD6FEDDFCEC" bold="true" box="[199,328,834,859]" pageId="1" pageNumber="564">
<typeStatus id="1E20883AFF9BFFA766F0FCD6FED6FCEC" box="[199,323,834,859]" pageId="1" pageNumber="564" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<collectingCountry id="B98C7608FF9BFFA76778FCD6FE54FCEC" box="[335,449,834,859]" name="Argentina" pageId="1" pageNumber="564">Argentina</collectingCountry>
, Chubut Province, Arroyo Nant y Fall int. Rt. prov. 17: 3 ex 
<date id="B5251058FF9BFFA76250FCD6FB46FCEC" box="[1127,1235,834,859]" pageId="1" pageNumber="564" value="2008-03-08">8.iii.2008</date>
M. Archangelsky coll.; 2 ex 
<date id="B5251058FF9BFFA76722FCF2FEF9FCC8" box="[277,364,870,895]" pageId="1" pageNumber="564" value="2008-03">iii.2008</date>
C. Brand coll.; 10 ex 
<date id="B5251058FF9BFFA76445FCF2FD5AFCC8" box="[626,719,870,895]" pageId="1" pageNumber="564" value="2008-04">iv. 2008</date>
C. Brand coll.; 1 ex 
<date id="B5251058FF9BFFA765FFFCF2FB83FCC8" box="[968,1046,870,895]" pageId="1" pageNumber="564" value="2009-02">ii.2009</date>
C. Brand coll.; 2 ex 
<date id="B5251058FF9BFFA76327FCF2FAE7FCC8" box="[1296,1394,870,895]" pageId="1" pageNumber="564" value="2011-02-02">2.ii.2011</date>
M. Archangelsky coll.
</paragraph>
</subSubSection>
<subSubSection id="89816513FF9BFFA766F0FC3AFC83FB84" pageId="1" pageNumber="564" type="diagnosis">
<paragraph id="C1243698FF9BFFA766F0FC3AFC83FB84" blockId="1.[151,1436,762,2011]" pageId="1" pageNumber="564">
<emphasis id="F3EFEA8AFF9BFFA766F0FC3AFEAFFC70" bold="true" box="[199,314,942,967]" pageId="1" pageNumber="564">Diagnosis</emphasis>
. The new species can be easily recognized from other known species of the genus by the following combination of characters: 1) color pattern with two reddish-brown areas on each elytron (
<figureCitation id="59A02A1DFF9BFFA762AAFC46FB75FC5C" box="[1181,1248,978,1003]" captionStart="FIGURES 1 – 5" captionStartId="2.[151,264,1717,1740]" captionTargetBox="[327,1245,222,1679]" captionTargetId="figure@2.[281,1306,193,1696]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Adults of Luchoelmis: 1) habitus of L. kapenkemkensis; 2) abdominal ventrites of L. kapenkemkensis; 3) lateral view of L. kapenkemkensis; 4) habitus of L. cekalovici; 5) abdominal ventrites of L. cekalovici. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/226679/files/figure.png" pageId="1" pageNumber="564">Fig. 1</figureCitation>
); 2) presence of plastron on margins of abdominal ventrites I–III (
<figureCitation id="59A02A1DFF9BFFA764F5FC62FCB2FBB8" box="[706,807,1014,1039]" captionStart="FIGURES 1 – 5" captionStartId="2.[151,264,1717,1740]" captionTargetBox="[327,1245,222,1679]" captionTargetId="figure@2.[281,1306,193,1696]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Adults of Luchoelmis: 1) habitus of L. kapenkemkensis; 2) abdominal ventrites of L. kapenkemkensis; 3) lateral view of L. kapenkemkensis; 4) habitus of L. cekalovici; 5) abdominal ventrites of L. cekalovici. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/226679/files/figure.png" pageId="1" pageNumber="564">Figs 2, 3</figureCitation>
); 3) presence of cleaning fringes on distal half of meso- and metatibiae; 4) shape of the male genitalia (
<figureCitation id="59A02A1DFF9BFFA76494FB8EFC9DFB84" box="[675,776,1050,1075]" captionStart="FIGURES 6 – 8" captionStartId="3.[151,264,1883,1906]" captionTargetBox="[317,1270,503,1849]" captionTargetId="figure@3.[306,1281,489,1862]" captionTargetPageId="3" captionText="FIGURES 6 – 8. Male genitalia of Luchoelmis spp: 6) aedeagus of L. kapenkemkensis; 7) detail of median lobe of L. kapenkemkensis; 8) aedeagus of L. cekalovici. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/226680/files/figure.png" pageId="1" pageNumber="564">Figs 6, 7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="89816513FF9BFFA566F0FBAAFBB9FE02" lastPageId="3" lastPageNumber="566" pageId="1" pageNumber="564" type="description">
<paragraph id="C1243698FF9BFFA766F0FBAAFB1FFBE1" blockId="1.[151,1436,762,2011]" box="[199,1162,1086,1111]" pageId="1" pageNumber="564">
<emphasis id="F3EFEA8AFF9BFFA766F0FBAAFECCFBE0" bold="true" box="[199,345,1086,1111]" pageId="1" pageNumber="564">Description.</emphasis>
<typeStatus id="1E20883AFF9BFFA76768FBAAFE59FBE0" box="[351,460,1086,1111]" pageId="1" pageNumber="564" type="holotype">Holotype</typeStatus>
: male, 
<quantity id="06639B7DFF9BFFA7642CFBABFDE5FBE1" box="[539,624,1087,1111]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="1" pageNumber="564" unit="mm" value="2.1">2.1 mm</quantity>
long; 
<quantity id="06639B7DFF9BFFA7648FFBABFC9BFBE1" box="[696,782,1087,1111]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="564" unit="mm" value="0.8">0.8 mm</quantity>
wide. Length range: 
<quantity id="06639B7DFF9BFFA765CBFBABFB13FBE1" box="[1020,1158,1087,1111]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.1" metricValueMin="1.9" pageId="1" pageNumber="564" unit="mm" value="2.0" valueMax="2.1" valueMin="1.9">1.9–2.1 mm</quantity>
.
</paragraph>
<paragraph id="C1243698FF9BFFA766F0FBF6FA88FB74" blockId="1.[151,1436,762,2011]" pageId="1" pageNumber="564">
Color: shiny, very dark brown except for antennae, femora and tibiae (except apex of femora and base of tibiae) and pretarsal claws reddish brown; each elytron with two reddish-brown markings (
<figureCitation id="59A02A1DFF9BFFA762FDFB12FA85FB28" box="[1226,1296,1158,1183]" captionStart="FIGURES 1 – 5" captionStartId="2.[151,264,1717,1740]" captionTargetBox="[327,1245,222,1679]" captionTargetId="figure@2.[281,1306,193,1696]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Adults of Luchoelmis: 1) habitus of L. kapenkemkensis; 2) abdominal ventrites of L. kapenkemkensis; 3) lateral view of L. kapenkemkensis; 4) habitus of L. cekalovici; 5) abdominal ventrites of L. cekalovici. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/226679/files/figure.png" pageId="1" pageNumber="564">Fig. 1</figureCitation>
), basal one smaller, close to inner margin, distal one larger, elongate, subapical on distal half close to outer margin.
</paragraph>
<paragraph id="C1243698FF9BFFA766F0FB5AFF20FA98" blockId="1.[151,1436,762,2011]" pageId="1" pageNumber="564">
Plastron on pronotal hypomeron and inner margin of epipleuron, sides of pro- meso- and metasternum, outer face of pro- and mesocoxa, surrounding basal half of profemur and mesofemur, and sides of ventrites I–III (
<figureCitation id="59A02A1DFF9BFFA76379FB66FF32FA98" captionStart="FIGURES 1 – 5" captionStartId="2.[151,264,1717,1740]" captionTargetBox="[327,1245,222,1679]" captionTargetId="figure@2.[281,1306,193,1696]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Adults of Luchoelmis: 1) habitus of L. kapenkemkensis; 2) abdominal ventrites of L. kapenkemkensis; 3) lateral view of L. kapenkemkensis; 4) habitus of L. cekalovici; 5) abdominal ventrites of L. cekalovici. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/226679/files/figure.png" pageId="1" pageNumber="564">Figs 2, 3</figureCitation>
).
</paragraph>
<paragraph id="C1243698FF9BFFA766F0FAAEFC47FA2C" blockId="1.[151,1436,762,2011]" pageId="1" pageNumber="564">Head bent downwards, opisthognathous, covered by fine punctuation, punctures separated by 2–3 times their diameter. Fronto-clypeal suture evident, between antennal sockets. Labrum large, rectangular, wider than long. Antennae filiform, shorter than pronotum, distal antennomere the longest.</paragraph>
<paragraph id="C1243698FF9BFFA766F0FA32FDECF8D8" blockId="1.[151,1436,762,2011]" pageId="1" pageNumber="564">
Thorax. Pronotum widest behind middle, wider at base than at anterior margin; anterior margin convex, covering base of head capsule (
<figureCitation id="59A02A1DFF9BFFA767CEFA5EFDAEFA54" box="[505,571,1482,1507]" captionStart="FIGURES 1 – 5" captionStartId="2.[151,264,1717,1740]" captionTargetBox="[327,1245,222,1679]" captionTargetId="figure@2.[281,1306,193,1696]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Adults of Luchoelmis: 1) habitus of L. kapenkemkensis; 2) abdominal ventrites of L. kapenkemkensis; 3) lateral view of L. kapenkemkensis; 4) habitus of L. cekalovici; 5) abdominal ventrites of L. cekalovici. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/226679/files/figure.png" pageId="1" pageNumber="564">Fig. 1</figureCitation>
); lateral margins sinuate; posterior margin trisinuate. Disc of pronotum with large median gibbosity in posterior half; one strong central depression anterior to gibbosity, two smaller depressions on antero-lateral corners, two elongate small depressions behind central gibbosity joining together and forming V-shaped groove, two small central foveae at base, close to scutellar emargination. Surface covered by fine punctures, separated by 1–2 times their diameter. Prosternum in front of coxae short, prosternal process narrow, elongated and parallel-sided, with rounded apex, reaching base of mesosternum. Mesosternum with groove for reception of prosternal process, concave between mesocoxae. Metasternum large, convex on anterior half, with elongate triangular depression along midline on posterior half. Pro- and mesocoxa rounded, metacoxa transverse, subtriangular; outer margins of pro- and mesocoxa with plastron; femora as long as tibiae, pro- and mesofemur with plastron on basal half; tibiae slender, those of meso- and metathoracic legs with cleaning fringe on distal half (also present in females); tarsi longer than tibiae, apical tarsomere as long as preceding four tarsomeres combined; pretarsal claws simple, lacking basal tooth.
</paragraph>
<paragraph id="C1243698FF9BFFA766F0F8EEFC3BF86D" blockId="1.[151,1436,762,2011]" pageId="1" pageNumber="564">
Abdomen with five visible ventrites (
<figureCitation id="59A02A1DFF9BFFA76445F8EEFD2DF824" box="[626,696,1914,1939]" captionStart="FIGURES 1 – 5" captionStartId="2.[151,264,1717,1740]" captionTargetBox="[327,1245,222,1679]" captionTargetId="figure@2.[281,1306,193,1696]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Adults of Luchoelmis: 1) habitus of L. kapenkemkensis; 2) abdominal ventrites of L. kapenkemkensis; 3) lateral view of L. kapenkemkensis; 4) habitus of L. cekalovici; 5) abdominal ventrites of L. cekalovici. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/226679/files/figure.png" pageId="1" pageNumber="564">Fig. 2</figureCitation>
), convex laterally, without lobes; ventrites I–III with plastron on sides, disc of ventrites glabrous, with fine microsculpture; ventrites IV and V glabrous, microsculptured; distal margin of ventrite V evenly convex, margined with short golden setae.
</paragraph>
<caption id="95E46610FF98FFA466A0F921FAB2F95D" httpUri="https://zenodo.org/record/226679/files/figure.png" pageId="2" pageNumber="565" targetBox="[327,1245,222,1679]" targetPageId="2">
<paragraph id="C1243698FF98FFA466A0F921FAB2F95D" blockId="2.[151,1435,1717,1771]" pageId="2" pageNumber="565">
<emphasis id="F3EFEA8AFF98FFA466A0F921FEAFF97C" bold="true" box="[151,314,1717,1740]" pageId="2" pageNumber="565">FIGURES 1–5.</emphasis>
Adults of 
<taxonomicName id="069B4D1BFF98FFA46793F922FD8DF97B" box="[420,536,1718,1740]" class="Insecta" family="Elmidae" genus="Luchoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="565" phylum="Arthropoda" rank="genus">
<emphasis id="F3EFEA8AFF98FFA46793F922FD8DF97B" box="[420,536,1718,1740]" italics="true" pageId="2" pageNumber="565">Luchoelmis</emphasis>
</taxonomicName>
: 1) habitus of 
<taxonomicName id="069B4D1BFF98FFA4649DF923FCF2F97B" box="[682,871,1718,1740]" class="Insecta" family="Elmidae" genus="Luchoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="565" phylum="Arthropoda" rank="species" species="kapenkemkensis">
<emphasis id="F3EFEA8AFF98FFA4649DF923FCF2F97B" box="[682,871,1718,1740]" italics="true" pageId="2" pageNumber="565">L. kapenkemkensis</emphasis>
</taxonomicName>
; 2) abdominal ventrites of 
<taxonomicName id="069B4D1BFF98FFA46240F923FAA0F97B" box="[1143,1333,1718,1740]" class="Insecta" family="Elmidae" genus="Luchoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="565" phylum="Arthropoda" rank="species" species="kapenkemkensis">
<emphasis id="F3EFEA8AFF98FFA46240F923FAA0F97B" box="[1143,1333,1718,1740]" italics="true" pageId="2" pageNumber="565">L. kapenkemkensis</emphasis>
</taxonomicName>
; 3) lateral view of 
<taxonomicName id="069B4D1BFF98FFA466DDF942FE32F95D" box="[234,423,1748,1771]" class="Insecta" family="Elmidae" genus="Luchoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="565" phylum="Arthropoda" rank="species" species="kapenkemkensis">
<emphasis id="F3EFEA8AFF98FFA466DDF942FE32F95D" box="[234,423,1748,1771]" italics="true" pageId="2" pageNumber="565">L. kapenkemkensis</emphasis>
</taxonomicName>
; 4) habitus of 
<taxonomicName id="069B4D1BFF98FFA4640CF942FD28F95D" box="[571,701,1748,1771]" class="Insecta" family="Elmidae" genus="Luchoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="565" phylum="Arthropoda" rank="species" species="cekalovici">
<emphasis id="F3EFEA8AFF98FFA4640CF942FD28F95D" box="[571,701,1748,1771]" italics="true" pageId="2" pageNumber="565">L. cekalovici</emphasis>
</taxonomicName>
; 5) abdominal ventrites of 
<taxonomicName id="069B4D1BFF98FFA465F8F942FBC4F95D" box="[975,1105,1748,1771]" class="Insecta" family="Elmidae" genus="Luchoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="565" phylum="Arthropoda" rank="species" species="cekalovici">
<emphasis id="F3EFEA8AFF98FFA465F8F942FBC4F95D" box="[975,1105,1748,1771]" italics="true" pageId="2" pageNumber="565">L. cekalovici</emphasis>
</taxonomicName>
. Scale bars: 0.5 mm.
</paragraph>
</caption>
<paragraph id="C1243698FF98FFA466F0F883FDEDF82B" blockId="2.[151,1436,1815,2023]" pageId="2" pageNumber="565">
Male genitalia: median lobe longer than parameres (
<figureCitation id="59A02A1DFF98FFA46527F883FCE0F898" box="[784,885,1815,1840]" captionStart="FIGURES 6 – 8" captionStartId="3.[151,264,1883,1906]" captionTargetBox="[317,1270,503,1849]" captionTargetId="figure@3.[306,1281,489,1862]" captionTargetPageId="3" captionText="FIGURES 6 – 8. Male genitalia of Luchoelmis spp: 6) aedeagus of L. kapenkemkensis; 7) detail of median lobe of L. kapenkemkensis; 8) aedeagus of L. cekalovici. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/226680/files/figure.png" pageId="2" pageNumber="565">Figs 6, 7</figureCitation>
), broader in middle portion, becoming slender in distal half, and constricted before rounded apex; phallotreme wide, almost at midlength of penis. Parameres broad at base, narrowing towards apex. Phallobase shorter than parameres and median lobe, longer than wide, asymmetrical, with three basal projections.
</paragraph>
<paragraph id="C1243698FF98FFA566F0F83EFBB9FE02" blockId="2.[151,1436,1815,2023]" lastBlockId="3.[151,1436,151,437]" lastPageId="3" lastPageNumber="566" pageId="2" pageNumber="565">
<emphasis id="F3EFEA8AFF98FFA466F0F83EFE2EF874" bold="true" box="[199,443,1962,1987]" pageId="2" pageNumber="565">Habitat and biology.</emphasis>
Nant y Fall stream is a 3rd order stream tributary of the Futaleufú river, and is located in a transitional mountain and piedmont area in the Northwest of Chubut province, 
<collectingCountry id="B98C7608FF98FFA46260F85AFB5FF850" box="[1111,1226,1998,2023]" name="Argentina" pageId="2" pageNumber="565">Argentina</collectingCountry>
. It belongs to the ecotone between the Subantarctic forest and the Patagonian steppe phytogeographical provinces. The substrate at the studied site is composed of gravel, cobbles, and boulders, with sand and smaller fractions at pools and depositional areas. Aquatic vegetation is abundant and diverse, with 
<taxonomicName id="069B4D1BFF99FFA565A3FF75FBDFFF40" box="[916,1098,224,248]" class="Lycopodiopsida" family="Isoetaceae" genus="Isoetes" kingdom="Plantae" order="Isoetales" pageId="3" pageNumber="566" phylum="Tracheophyta" rank="species" species="savatieri">
<emphasis id="F3EFEA8AFF99FFA565A3FF75FBDFFF40" box="[916,1098,224,248]" italics="true" pageId="3" pageNumber="566">Isoetes savatieri</emphasis>
</taxonomicName>
being the dominant species in riffles and 
<taxonomicName id="069B4D1BFF99FFA56724FE93FD82FEA8" box="[275,535,263,287]" class="Magnoliopsida" family="Haloragaceae" genus="Myriophyllum" kingdom="Plantae" order="Saxifragales" pageId="3" pageNumber="566" phylum="Tracheophyta" rank="species" species="quitense">
<emphasis id="F3EFEA8AFF99FFA56724FE93FD82FEA8" box="[275,535,263,287]" italics="true" pageId="3" pageNumber="566">Myriophyllum quitense</emphasis>
</taxonomicName>
and 
<taxonomicName id="069B4D1BFF99FFA56466FE93FCBDFEA8" box="[593,808,263,287]" class="Magnoliopsida" family="Plantaginaceae" genus="Callitriche" kingdom="Animalia" order="Lamiales" pageId="3" pageNumber="566" phylum="Tracheophyta" rank="species" species="lechleri">
<emphasis id="F3EFEA8AFF99FFA56466FE93FCBDFEA8" box="[593,808,263,287]" italics="true" pageId="3" pageNumber="566">Callitriche lechleri</emphasis>
</taxonomicName>
in pools. Current speed ranged from 0.22 to 1.16 ms -1, water temperature ranged from 4 to 18 °C, with an annual average of 11.6 °C; during the entire year water remains well oxygenated (9.95–13.88 mgl -1 O2) and above saturation (114.1–121.4 %) (Brand &amp; Miserendino 2012, Brand 
<emphasis id="F3EFEA8AFF99FFA566A0FEEDFF46FE27" box="[151,211,376,400]" italics="true" pageId="3" pageNumber="566">et al.</emphasis>
2012). Larvae assigned to 
<taxonomicName id="069B4D1BFF99FFA56425FEEDFD72FE27" box="[530,743,376,400]" class="Insecta" family="Elmidae" genus="Luchoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="566" phylum="Arthropoda" rank="species" species="kapenkemkensis">
<emphasis id="F3EFEA8AFF99FFA56425FEEDFD72FE27" box="[530,743,376,400]" italics="true" pageId="3" pageNumber="566">L. kapenkemkensis</emphasis>
</taxonomicName>
were found at all habitats, both erosional and depositional, while adults appeared to be most abundant in erosional boulder-pebble substrates.
</paragraph>
</subSubSection>
<caption id="95E46610FF99FFA566A0F8CFFCADF827" httpUri="https://zenodo.org/record/226680/files/figure.png" pageId="3" pageNumber="566" targetBox="[317,1270,503,1849]" targetPageId="3">
<paragraph id="C1243698FF99FFA566A0F8CFFCADF827" blockId="3.[151,1437,1883,1936]" pageId="3" pageNumber="566">
<emphasis id="F3EFEA8AFF99FFA566A0F8CFFED5F8C6" bold="true" box="[151,320,1883,1906]" pageId="3" pageNumber="566">FIGURES 6–8.</emphasis>
Male genitalia of 
<taxonomicName id="069B4D1BFF99FFA56424F8C8FD1DF8C5" box="[531,648,1884,1906]" class="Insecta" family="Elmidae" genus="Luchoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="566" phylum="Arthropoda" rank="genus">
<emphasis id="F3EFEA8AFF99FFA56424F8C8FD1DF8C5" box="[531,648,1884,1906]" italics="true" pageId="3" pageNumber="566">Luchoelmis</emphasis>
</taxonomicName>
spp: 6) aedeagus of 
<taxonomicName id="069B4D1BFF99FFA56540F8C9FBAEF8C5" box="[887,1083,1884,1906]" class="Insecta" family="Elmidae" genus="Luchoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="566" phylum="Arthropoda" rank="species" species="kapenkemkensis">
<emphasis id="F3EFEA8AFF99FFA56540F8C9FBAEF8C5" box="[887,1083,1884,1906]" italics="true" pageId="3" pageNumber="566">L. kapenkemkensis</emphasis>
</taxonomicName>
; 7) detail of median lobe of 
<taxonomicName id="069B4D1BFF99FFA563B0F8C9FEAFF827" class="Insecta" family="Elmidae" genus="Luchoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="566" phylum="Arthropoda" rank="species" species="kapenkemkensis">
<emphasis id="F3EFEA8AFF99FFA563B0F8C9FEAFF827" italics="true" pageId="3" pageNumber="566">L. kapenkemkensis</emphasis>
</taxonomicName>
; 8) aedeagus of 
<taxonomicName id="069B4D1BFF99FFA567D6F8EFFDF7F827" box="[481,610,1914,1936]" class="Insecta" family="Elmidae" genus="Luchoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="566" phylum="Arthropoda" rank="species" species="cekalovici">
<emphasis id="F3EFEA8AFF99FFA567D6F8EFFDF7F827" box="[481,610,1914,1936]" italics="true" pageId="3" pageNumber="566">L. cekalovici</emphasis>
</taxonomicName>
. Scale bars: 0.1 mm.
</paragraph>
</caption>
<caption id="95E46610FF9EFFA266A0FD65FC30FC91" httpUri="https://zenodo.org/record/226681/files/figure.png" pageId="4" pageNumber="567" targetBox="[362,1243,227,697]" targetPageId="4">
<paragraph id="C1243698FF9EFFA266A0FD65FC30FC91" blockId="4.[151,1437,753,806]" pageId="4" pageNumber="567">
<emphasis id="F3EFEA8AFF9EFFA266A0FD65FEDCFCB0" bold="true" box="[151,329,753,776]" pageId="4" pageNumber="567">FIGURES 9–11.</emphasis>
Male genitalia of 
<taxonomicName id="069B4D1BFF9EFFA2643AFD66FD17FCBF" box="[525,642,754,776]" class="Insecta" family="Elmidae" genus="Luchoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="567" phylum="Arthropoda" rank="genus">
<emphasis id="F3EFEA8AFF9EFFA2643AFD66FD17FCBF" box="[525,642,754,776]" italics="true" pageId="4" pageNumber="567">Luchoelmis</emphasis>
</taxonomicName>
spp: 9) edeagus of 
<taxonomicName id="069B4D1BFF9EFFA26561FD67FC5FFCBF" box="[854,970,754,776]" class="Insecta" family="Elmidae" genus="Luchoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="567" phylum="Arthropoda" rank="species" species="aequalis">
<emphasis id="F3EFEA8AFF9EFFA26561FD67FC5FFCBF" box="[854,970,754,776]" italics="true" pageId="4" pageNumber="567">L. aequalis</emphasis>
</taxonomicName>
; 10) edeagus of 
<taxonomicName id="069B4D1BFF9EFFA26249FD67FB41FCBF" box="[1150,1236,755,776]" class="Insecta" family="Elmidae" genus="Luchoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="567" phylum="Arthropoda" rank="species" species="penai">
<emphasis id="F3EFEA8AFF9EFFA26249FD67FB41FCBF" box="[1150,1236,755,776]" italics="true" pageId="4" pageNumber="567">L. penai</emphasis>
</taxonomicName>
; 11) edeagus of 
<taxonomicName id="069B4D1BFF9EFFA263B0FD67FEBCFC91" class="Insecta" family="Elmidae" genus="Luchoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="567" phylum="Arthropoda" rank="species" species="magallanensis">
<emphasis id="F3EFEA8AFF9EFFA263B0FD67FEBCFC91" italics="true" pageId="4" pageNumber="567">L. magallanensis</emphasis>
</taxonomicName>
. (modified after Spangler &amp; Staines 2002). Scale bar: 0.1 mm.
</paragraph>
</caption>
<subSubSection id="89816513FF9EFFA266F0FCDAFF75FC65" pageId="4" pageNumber="567" type="etymology">
<paragraph id="C1243698FF9EFFA266F0FCDAFF75FC65" blockId="4.[151,1436,846,978]" pageId="4" pageNumber="567">
<emphasis id="F3EFEA8AFF9EFFA266F0FCDAFEDBFCD0" bold="true" box="[199,334,846,871]" pageId="4" pageNumber="567">Etymology.</emphasis>
From 
<emphasis id="F3EFEA8AFF9EFFA267A8FCDAFE77FCD1" box="[415,482,846,870]" italics="true" pageId="4" pageNumber="567">kapen</emphasis>
(Tehuelche, Boreal-Meridional= red) and 
<emphasis id="F3EFEA8AFF9EFFA265E7FCDAFBB1FCD1" box="[976,1060,846,870]" italics="true" pageId="4" pageNumber="567">kemken</emphasis>
(Tehuelche, Boreal-Meridional= back), for the reddish-brown spots on the elytra. The Tehuelche people, also known as “Patagones”, inhabited southern 
<collectingCountry id="B98C7608FF9EFFA26734FC02FEE7FC18" box="[259,370,918,943]" name="Argentina" pageId="4" pageNumber="567">Argentina</collectingCountry>
and 
<collectingCountry id="B98C7608FF9EFFA26786FC02FE64FC18" box="[433,497,918,943]" name="Chile" pageId="4" pageNumber="567">Chile</collectingCountry>
. Both Tehuelche words were taken from the Tehuelche dictionary (
<bibRefCitation id="A50A4B69FF9EFFA2633EFC02FF41FC65" author="Casamiquela" pageId="4" pageNumber="567" refString="Casamiquela, R. (2008) Diccionario Tehuelche. Patagonia Sur Libros, 112 pp." type="book" year="2008">Casamiquela 2008</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="89816513FF9EFFA266A0FBB2FA09FA55" pageId="4" pageNumber="567" type="key">
<paragraph id="C1243698FF9EFFA266A0FBB2FC71FB88" blockId="4.[151,996,1061,1087]" box="[151,996,1061,1087]" pageId="4" pageNumber="567">
<emphasis id="F3EFEA8AFF9EFFA266A0FBB2FD82FB89" bold="true" box="[151,535,1061,1087]" pageId="4" pageNumber="567">
Key to the species of 
<taxonomicName id="069B4D1BFF9EFFA267A7FBB1FD82FB89" ID-CoL="5H4V" box="[400,535,1061,1086]" class="Insecta" family="Elmidae" genus="Luchoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="567" phylum="Arthropoda" rank="genus">
<emphasis id="F3EFEA8AFF9EFFA267A7FBB1FD82FB89" bold="true" box="[400,535,1061,1086]" italics="true" pageId="4" pageNumber="567">Luchoelmis</emphasis>
</taxonomicName>
</emphasis>
(adapted from 
<bibRefCitation id="A50A4B69FF9EFFA264F4FBB2FC49FB88" author="Spangler" box="[707,988,1062,1087]" pageId="4" pageNumber="567" refString="Spangler, P. J. &amp; Staines, C. L. (2002) Luchoelmis, a new genus of Elmidae (Coleoptera) from Chile and Argentina. Insecta Mundi, 16, 215 - 220." type="journal article" year="2002">Spangler &amp; Staines 2002</bibRefCitation>
)
</paragraph>
<key id="B50A61A9FF9EFFA266A0FBF9FA09FA55" pageId="4" pageNumber="567">
<keyStep id="7A6F2E3DFF9EFFA266A0FBF9FA0EFB17" pageId="4" pageNumber="567">
<paragraph id="C1243698FF9EFFA266A0FBF9FA09FB35" blockId="4.[151,1437,1133,1506]" box="[151,1436,1133,1154]" pageId="4" pageNumber="567">
<keyLead id="7A6A95ADFF9EFFA266A0FBF9FA09FB35" box="[151,1436,1133,1154]" pageId="4" pageNumber="567">
1. Pronotum with basal margin as wide as anterior margin; male genitalia as in 
<figureCitation id="59A02A1DFF9EFFA26588FBF9FC6CFB35" box="[959,1017,1133,1154]" captionStart="FIGURES 9 – 11" captionStartId="4.[151,264,753,776]" captionTargetBox="[362,1243,227,697]" captionTargetId="figure@4.[304,1283,193,733]" captionTargetPageId="4" captionText="FIGURES 9 – 11. Male genitalia of Luchoelmis spp: 9) edeagus of L. aequalis; 10) edeagus of L. penai; 11) edeagus of L. magallanensis. (modified after Spangler &amp; Staines 2002). Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/226681/files/figure.png" pageId="4" pageNumber="567">Fig. 9</figureCitation>
.......... 
<taxonomicName id="069B4D1BFF9EFFA26245FBF9FA09FB35" ID-CoL="72P9B" authority="Spangler &amp; Staines" authorityName="Spangler &amp; Staines" box="[1138,1436,1133,1154]" class="Insecta" family="Elmidae" genus="Luchoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="567" phylum="Arthropoda" rank="species" species="aequalis">
<emphasis id="F3EFEA8AFF9EFFA26245FBF9FB48FB35" box="[1138,1245,1133,1154]" italics="true" pageId="4" pageNumber="567">L. aequalis</emphasis>
Spangler &amp; Staines
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C1243698FF9EFFA266A0FB1FFA0EFB17" blockId="4.[151,1437,1133,1506]" box="[151,1435,1163,1184]" pageId="4" pageNumber="567">
<keyLead id="7A6A95ADFF9EFFA266A0FB1FFA0EFB17" box="[151,1435,1163,1184]" pageId="4" pageNumber="567">
- Pronotum with basal margin wider than anterior margin; male genitalia not as in 
<figureCitation id="59A02A1DFF9EFFA265D0FB1FFA0EFB17" box="[999,1435,1163,1184]" captionStart-0="FIGURES 9 – 11" captionStart-1="FIGURES 1 – 5" captionStartId-0="4.[151,264,753,776]" captionStartId-1="2.[151,264,1717,1740]" captionTargetBox-0="[362,1243,227,697]" captionTargetBox-1="[327,1245,222,1679]" captionTargetId-0="figure@4.[304,1283,193,733]" captionTargetId-1="figure@2.[281,1306,193,1696]" captionTargetPageId-0="4" captionTargetPageId-1="2" captionText-0="FIGURES 9 – 11. Male genitalia of Luchoelmis spp: 9) edeagus of L. aequalis; 10) edeagus of L. penai; 11) edeagus of L. magallanensis. (modified after Spangler &amp; Staines 2002). Scale bar: 0.1 mm." captionText-1="FIGURES 1 – 5. Adults of Luchoelmis: 1) habitus of L. kapenkemkensis; 2) abdominal ventrites of L. kapenkemkensis; 3) lateral view of L. kapenkemkensis; 4) habitus of L. cekalovici; 5) abdominal ventrites of L. cekalovici. Scale bars: 0.5 mm." httpUri-0="https://zenodo.org/record/226681/files/figure.png" httpUri-1="https://zenodo.org/record/226679/files/figure.png" pageId="4" pageNumber="567">Fig. 9.............................. 2</figureCitation>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="7A6F2E3DFF9EFFA266A0FB3CFA09FA85" pageId="4" pageNumber="567">
<paragraph id="C1243698FF9EFFA266A0FB3CFA0EFB4F" blockId="4.[151,1437,1133,1506]" pageId="4" pageNumber="567">
<keyLead id="7A6A95ADFF9EFFA266A0FB3CFA0EFB4F" pageId="4" pageNumber="567">
2. Each elytron with two reddish markings as in 
<figureCitation id="59A02A1DFF9EFFA26495FB3CFD49FB0A" box="[674,732,1192,1213]" captionStart="FIGURES 1 – 5" captionStartId="2.[151,264,1717,1740]" captionTargetBox="[327,1245,222,1679]" captionTargetId="figure@2.[281,1306,193,1696]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Adults of Luchoelmis: 1) habitus of L. kapenkemkensis; 2) abdominal ventrites of L. kapenkemkensis; 3) lateral view of L. kapenkemkensis; 4) habitus of L. cekalovici; 5) abdominal ventrites of L. cekalovici. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/226679/files/figure.png" pageId="4" pageNumber="567">Fig. 1</figureCitation>
. Abdominal ventrites I–III with plastron on sides (
<figureCitation id="59A02A1DFF9EFFA262FDFB3CFAB4FB0A" box="[1226,1313,1192,1213]" captionStart="FIGURES 1 – 5" captionStartId="2.[151,264,1717,1740]" captionTargetBox="[327,1245,222,1679]" captionTargetId="figure@2.[281,1306,193,1696]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Adults of Luchoelmis: 1) habitus of L. kapenkemkensis; 2) abdominal ventrites of L. kapenkemkensis; 3) lateral view of L. kapenkemkensis; 4) habitus of L. cekalovici; 5) abdominal ventrites of L. cekalovici. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/226679/files/figure.png" pageId="4" pageNumber="567">Figs 2, 3</figureCitation>
). Meso- and metatibiae with cleaning fringes on distal half. Male with median lobe of aedeagus strongly narrowing on distal half (
<figureCitation id="59A02A1DFF9EFFA2636FFB51FF7AFB4F" captionStart="FIGURES 6 – 8" captionStartId="3.[151,264,1883,1906]" captionTargetBox="[317,1270,503,1849]" captionTargetId="figure@3.[306,1281,489,1862]" captionTargetPageId="3" captionText="FIGURES 6 – 8. Male genitalia of Luchoelmis spp: 6) aedeagus of L. kapenkemkensis; 7) detail of median lobe of L. kapenkemkensis; 8) aedeagus of L. cekalovici. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/226680/files/figure.png" pageId="4" pageNumber="567">Figs 6, 7</figureCitation>
). Small species, 
<quantity id="06639B7DFF9EFFA267A7FB77FD96FB4F" box="[400,515,1251,1272]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.1" metricValueMin="1.9" pageId="4" pageNumber="567" unit="mm" value="2.0" valueMax="2.1" valueMin="1.9">1.9–2.1 mm</quantity>
.................................................. 
<taxonomicName id="069B4D1BFF9EFFA26258FB77FAB6FB4F" ID-CoL="3WBCC" box="[1135,1315,1251,1272]" class="Insecta" family="Elmidae" genus="Luchoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="567" phylum="Arthropoda" rank="species" species="kapenkemkensis" status="sp. nov.">
<emphasis id="F3EFEA8AFF9EFFA26258FB77FAB6FB4F" box="[1135,1315,1251,1272]" italics="true" pageId="4" pageNumber="567">L. kapenkemkensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8DC57F1FF9EFFA2631EFB77FA0EFB4F" box="[1321,1435,1251,1272]" pageId="4" pageNumber="567" rank="species">new species</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="C1243698FF9EFFA266A0FA94FA09FA85" blockId="4.[151,1437,1133,1506]" pageId="4" pageNumber="567">
<keyLead id="7A6A95ADFF9EFFA266A0FA94FA09FA85" pageId="4" pageNumber="567">
- Elytra reddish-brown to black, without markings (
<figureCitation id="59A02A1DFF9EFFA264F5FA94FD68FAA2" box="[706,765,1280,1301]" captionStart="FIGURES 1 – 5" captionStartId="2.[151,264,1717,1740]" captionTargetBox="[327,1245,222,1679]" captionTargetId="figure@2.[281,1306,193,1696]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Adults of Luchoelmis: 1) habitus of L. kapenkemkensis; 2) abdominal ventrites of L. kapenkemkensis; 3) lateral view of L. kapenkemkensis; 4) habitus of L. cekalovici; 5) abdominal ventrites of L. cekalovici. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/226679/files/figure.png" pageId="4" pageNumber="567">Fig. 4</figureCitation>
). Abdominal ventrites I–III without plastron on margins (
<figureCitation id="59A02A1DFF9EFFA26311FA94FAF4FAA2" box="[1318,1377,1280,1301]" captionStart="FIGURES 1 – 5" captionStartId="2.[151,264,1717,1740]" captionTargetBox="[327,1245,222,1679]" captionTargetId="figure@2.[281,1306,193,1696]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Adults of Luchoelmis: 1) habitus of L. kapenkemkensis; 2) abdominal ventrites of L. kapenkemkensis; 3) lateral view of L. kapenkemkensis; 4) habitus of L. cekalovici; 5) abdominal ventrites of L. cekalovici. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/226679/files/figure.png" pageId="4" pageNumber="567">Fig. 5</figureCitation>
). Tib- iae with or without cleaning fringes. Median lobe of aedeagus different. Larger species, 
<quantity id="06639B7DFF9EFFA2621AFA89FB35FA85" box="[1069,1184,1309,1330]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.4" metricValueMin="2.1" pageId="4" pageNumber="567" unit="mm" value="2.25" valueMax="2.4" valueMin="2.1">2.1–2.4 mm</quantity>
................... 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="7A6F2E3DFF9EFFA266A0FAAFFA0EFA1F" pageId="4" pageNumber="567">
<paragraph id="C1243698FF9EFFA266A0FAAFFA09FADA" blockId="4.[151,1437,1133,1506]" pageId="4" pageNumber="567">
<keyLead id="7A6A95ADFF9EFFA266A0FAAFFA09FADA" pageId="4" pageNumber="567">
3. No femora with cleaning fringe. Pronotum widest at about midlength; male genitalia as in 
<figureCitation id="59A02A1DFF9EFFA26273FAAFFB1EFAE7" box="[1092,1163,1339,1360]" captionStart="FIGURES 9 – 11" captionStartId="4.[151,264,753,776]" captionTargetBox="[362,1243,227,697]" captionTargetId="figure@4.[304,1283,193,733]" captionTargetPageId="4" captionText="FIGURES 9 – 11. Male genitalia of Luchoelmis spp: 9) edeagus of L. aequalis; 10) edeagus of L. penai; 11) edeagus of L. magallanensis. (modified after Spangler &amp; Staines 2002). Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/226681/files/figure.png" pageId="4" pageNumber="567">Fig. 10</figureCitation>
..................................................................................................... 
<taxonomicName id="069B4D1BFF9EFFA262B9FACCFA09FADA" ID-CoL="6QP9C" authority="Spangler &amp; Staines" authorityName="Spangler &amp; Staines" box="[1166,1436,1368,1389]" class="Insecta" family="Elmidae" genus="Luchoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="567" phylum="Arthropoda" rank="species" species="penai">
<emphasis id="F3EFEA8AFF9EFFA262B9FACCFB48FADA" box="[1166,1245,1368,1389]" italics="true" pageId="4" pageNumber="567">L. penai</emphasis>
Spangler &amp; Staines
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C1243698FF9EFFA266A0FAE1FA0EFA1F" blockId="4.[151,1437,1133,1506]" pageId="4" pageNumber="567">
<keyLead id="7A6A95ADFF9EFFA266A0FAE1FA0EFA1F" pageId="4" pageNumber="567">
- Pro- and mesofemora with cleaning fringe on some surfaces. Pronotum widest behind midlength; male genitalia not as in 
<figureCitation id="59A02A1DFF9EFFA26341FAE1FA0EFA1F" captionStart-0="FIGURES 9 – 11" captionStart-1="FIGURES 1 – 5" captionStartId-0="4.[151,264,753,776]" captionStartId-1="2.[151,264,1717,1740]" captionTargetBox-0="[362,1243,227,697]" captionTargetBox-1="[327,1245,222,1679]" captionTargetId-0="figure@4.[304,1283,193,733]" captionTargetId-1="figure@2.[281,1306,193,1696]" captionTargetPageId-0="4" captionTargetPageId-1="2" captionText-0="FIGURES 9 – 11. Male genitalia of Luchoelmis spp: 9) edeagus of L. aequalis; 10) edeagus of L. penai; 11) edeagus of L. magallanensis. (modified after Spangler &amp; Staines 2002). Scale bar: 0.1 mm." captionText-1="FIGURES 1 – 5. Adults of Luchoelmis: 1) habitus of L. kapenkemkensis; 2) abdominal ventrites of L. kapenkemkensis; 3) lateral view of L. kapenkemkensis; 4) habitus of L. cekalovici; 5) abdominal ventrites of L. cekalovici. Scale bars: 0.5 mm." httpUri-0="https://zenodo.org/record/226681/files/figure.png" httpUri-1="https://zenodo.org/record/226679/files/figure.png" pageId="4" pageNumber="567">Fig. 10................................................................................................. 4</figureCitation>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="7A6F2E3DFF9EFFA266A0FA24FA09FA55" pageId="4" pageNumber="567">
<paragraph id="C1243698FF9EFFA266A0FA24FA09FA72" blockId="4.[151,1437,1133,1506]" box="[151,1436,1456,1477]" pageId="4" pageNumber="567">
<keyLead id="7A6A95ADFF9EFFA266A0FA24FA09FA72" box="[151,1436,1456,1477]" pageId="4" pageNumber="567">
4. Female without cleaning fringe on front surface of profemora. Male genitalia as in 
<figureCitation id="59A02A1DFF9EFFA26234FA24FBA9FA72" box="[1027,1084,1456,1477]" captionStart="FIGURES 6 – 8" captionStartId="3.[151,264,1883,1906]" captionTargetBox="[317,1270,503,1849]" captionTargetId="figure@3.[306,1281,489,1862]" captionTargetPageId="3" captionText="FIGURES 6 – 8. Male genitalia of Luchoelmis spp: 6) aedeagus of L. kapenkemkensis; 7) detail of median lobe of L. kapenkemkensis; 8) aedeagus of L. cekalovici. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/226680/files/figure.png" pageId="4" pageNumber="567">Fig. 8</figureCitation>
.. 
<taxonomicName id="069B4D1BFF9EFFA26254FA24FA09FA72" ID-CoL="3WBCB" authority="Spangler &amp; Staines" authorityName="Spangler &amp; Staines" box="[1123,1436,1456,1477]" class="Insecta" family="Elmidae" genus="Luchoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="567" phylum="Arthropoda" rank="species" species="cekalovici">
<emphasis id="F3EFEA8AFF9EFFA26254FA24FB48FA72" box="[1123,1245,1456,1477]" italics="true" pageId="4" pageNumber="567">L. cekalovici</emphasis>
Spangler &amp; Staines
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C1243698FF9EFFA266A0FA59FA09FA55" blockId="4.[151,1437,1133,1506]" box="[151,1436,1485,1506]" pageId="4" pageNumber="567">
<keyLead id="7A6A95ADFF9EFFA266A0FA59FA09FA55" box="[151,1436,1485,1506]" pageId="4" pageNumber="567">
- Female with cleaning fringe on front surface of profemora. Male genitalia as in 
<figureCitation id="59A02A1DFF9EFFA265E9FA59FBBDFA55" box="[990,1064,1485,1506]" captionStart="FIGURES 9 – 11" captionStartId="4.[151,264,753,776]" captionTargetBox="[362,1243,227,697]" captionTargetId="figure@4.[304,1283,193,733]" captionTargetPageId="4" captionText="FIGURES 9 – 11. Male genitalia of Luchoelmis spp: 9) edeagus of L. aequalis; 10) edeagus of L. penai; 11) edeagus of L. magallanensis. (modified after Spangler &amp; Staines 2002). Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/226681/files/figure.png" pageId="4" pageNumber="567">Fig. 11</figureCitation>
.. 
<taxonomicName id="069B4D1BFF9EFFA2620EFA59FA09FA55" ID-CoL="3WBCD" authority="Spangler &amp; Staines" authorityName="Spangler &amp; Staines" box="[1081,1436,1485,1506]" class="Insecta" family="Elmidae" genus="Luchoelmis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="567" phylum="Arthropoda" rank="species" species="magallanensis">
<emphasis id="F3EFEA8AFF9EFFA2620EFA59FB4BFA55" box="[1081,1246,1485,1506]" italics="true" pageId="4" pageNumber="567">L. magallanensis</emphasis>
Spangler &amp; Staines
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>