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<document id="5C8D453C3556E6CE24F4AC762E8F4167" ID-CLB-Dataset="294579" ID-DOI="10.5194/fr-17-41-2014" ID-GBIF-Dataset="da2d7c61-e90d-49d9-abf6-3161806e8d57" ID-ISSN="2193-0074" ID-Zenodo-Dep="10965376" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1712925550276" checkinUser="felipe" docAuthor="Ghaderi, A., Leda, L., Schobben, M., Korn, D. &amp; Ashouri, A. R." docDate="2014" docId="107A8787FFA19364D10BFB9EFE30FE25" docLanguage="en" docName="FossRec.17.1.41-57.pdf" docOrigin="Fossil Record 17 (1)" docSource="http://dx.doi.org/10.5194/fr-17-41-2014" docStyle="DocumentStyle:1B3764FFAC190C6D25C29F616DE4358D.1:FossRec.2014-2021.journal_article" docStyleId="1B3764FFAC190C6D25C29F616DE4358D" docStyleName="FossRec.2014-2021.journal_article" docStyleVersion="1" docTitle="Arasella Ghaderi &amp; Leda &amp; Schobben &amp; Korn &amp; Ashouri 2014, n. gen." docType="treatment" docUuid="7B65D070-4D72-43F9-8138-5397A9BA10D5" docUuidSource="ZooBank" docVersion="4" lastPageNumber="55" masterDocId="EC43FFFFFFAC936AD238FF9EFF95FFC3" masterDocTitle="High-resolution stratigraphy of the Changhsingian (Late Permian) successions of NW Iran and the Transcaucasus based on lithological features, conodonts and ammonoids" masterLastPageNumber="57" masterPageNumber="41" pageNumber="54" updateTime="1713857359944" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="436F23794DB071C9184C5E0CA3ABA28C">High-resolution stratigraphy of the Changhsingian (Late Permian) successions of NW Iran and the Transcaucasus based on lithological features, conodonts and ammonoids</mods:title>
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<taxonomicName id="5FD34D12FFA19367D10BFB9EFC1AFBDB" ID-CoL="ZRR" authority="Ghaderi &amp; Leda &amp; Schobben &amp; Korn &amp; Ashouri, 2014" authorityName="Ghaderi &amp; Leda &amp; Schobben &amp; Korn &amp; Ashouri" authorityYear="2014" box="[819,911,1024,1048]" class="Cephalopoda" family="Xenodiscidae" genus="Arasella" kingdom="Animalia" order="Ceratitida" pageId="13" pageNumber="54" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="AAA7EA83FFA19367D10BFB9EFC1AFBDB" bold="true" box="[819,911,1024,1048]" italics="true" pageId="13" pageNumber="54">Arasella</emphasis>
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<taxonomicNameLabel id="B19457F8FFA19367D1AEFB9EFC75FBDB" box="[918,992,1024,1048]" pageId="13" pageNumber="54" rank="genus">n. gen.</taxonomicNameLabel>
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of name: after the
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, the locality of the
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material.
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species:
<taxonomicName id="5FD34D12FFA19367D182FB16FA8CFB5E" authority="Zakharov, 1983" authorityName="Zakharov" authorityYear="1983" box="[954,1305,1160,1181]" class="Cephalopoda" family="Tapashanitidae" genus="Sinoceltites" kingdom="Animalia" pageId="13" pageNumber="54" phylum="Mollusca" rank="species" species="minutus">
<emphasis id="AAA7EA83FFA19367D182FB16FBEEFB5E" box="[954,1147,1160,1181]" italics="true" pageId="13" pageNumber="54">Sinoceltites minutus</emphasis>
<bibRefCitation id="FC424B60FFA19367D6B8FB16FA8CFB5E" author="Zakharov, Y. D." box="[1152,1305,1160,1181]" pageId="13" pageNumber="54" pagination="150 - 157" refId="ref12700" refString="Zakharov, Y. D.: Ammonoidea, in: Kotlyar, G. V., Zakharov, Yu. D., Koczyrkevich, B. V., Kropatcheva, G. S., Rostovtsev, K. O., Chedia, I. O., Vuks, G. P., and Guseva, E. A.: Dzhul'finskii i Dorashamskii yarusy SSSR, edited by: Gramm, M. N. and Rostovtsev, K. O., Pozdnepermiskii etap evoliutsii organicheskogo mira, 150 - 157, 1983." type="book chapter" year="1983">Zakharov, 1983</bibRefCitation>
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.
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Diagnosis: genus of the
<taxonomicName id="5FD34D12FFA19367D614FB2DFB37FB0B" authorityName="Shevyrev" authorityYear="1965" box="[1068,1186,1203,1224]" class="Cephalopoda" family="Dzhulfitidae" kingdom="Animalia" pageId="13" pageNumber="54" phylum="Mollusca" rank="family">Dzhulfitidae</taxonomicName>
with small conch reaching
<quantity id="5F2B9B74FFA19367D10BFB4DFCE1FB2B" box="[819,884,1235,1256]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="13" pageNumber="54" unit="mm" value="30.0">30 mm</quantity>
diameter. Conch widely umbilicate with circular whorl cross section. Ornament with sharp ribs. Suture line with short external lobe, which reaches only 50 % of the depth of the adventive lobe. Adventive lobe and lateral lobe rounded and unserrated.
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Discussion:
<bibRefCitation id="FC424B60FFA19367D1FAFAC2FB17FAB0" author="Zakharov, Y. D." box="[962,1154,1372,1395]" pageId="13" pageNumber="54" pagination="150 - 157" refId="ref12700" refString="Zakharov, Y. D.: Ammonoidea, in: Kotlyar, G. V., Zakharov, Yu. D., Koczyrkevich, B. V., Kropatcheva, G. S., Rostovtsev, K. O., Chedia, I. O., Vuks, G. P., and Guseva, E. A.: Dzhul'finskii i Dorashamskii yarusy SSSR, edited by: Gramm, M. N. and Rostovtsev, K. O., Pozdnepermiskii etap evoliutsii organicheskogo mira, 150 - 157, 1983." type="book chapter" year="1983">Zakharov (1983)</bibRefCitation>
described the species as belonging to
<taxonomicName id="5FD34D12FFA19367D1FFFAE2FBD4FA50" authorityName="Zhao &amp; Liang" authorityYear="1974" box="[967,1089,1404,1427]" class="Cephalopoda" family="Tapashanitidae" genus="Sinoceltites" kingdom="Animalia" pageId="13" pageNumber="54" phylum="Mollusca" rank="genus">
<emphasis id="AAA7EA83FFA19367D1FFFAE2FBD4FA50" box="[967,1089,1404,1427]" italics="true" pageId="13" pageNumber="54">Sinoceltites</emphasis>
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, a genus belonging to the family
<taxonomicName id="5FD34D12FFA19367D10BFA02FC47FA70" box="[819,978,1436,1459]" class="Cephalopoda" family="Tapashanitidae" kingdom="Animalia" pageId="13" pageNumber="54" phylum="Mollusca" rank="family">Tapashanitidae</taxonomicName>
and known from occurrences in South
<collectingCountry id="E0C47601FFA19367D748FA02FA26FA70" box="[1392,1459,1436,1459]" name="China" pageId="13" pageNumber="54">China</collectingCountry>
. The species has a particular position in two respects: (1) it is obviously the stratigraphically youngest ammonoid to appear before the end-Permian mass extinction event in the NW Iranian sections, and (2) it has a morphology characterized by reduction of morphological characters such as the suture line and thus is difficult to interpret in terms of phylogeny. Two possible phylogenetic origins may be discussed:
</paragraph>
<footnote id="FBC82A9FFFA19367D10BF958FB6FF8DE" pageId="13" pageNumber="54">
<paragraph id="986C3691FFA19367D10BF958FB6FF8DE" blockId="13.[819,1463,1372,1821]" pageId="13" pageNumber="54">(1) A xenodiscid origin this would imply a ghost lineage ranging through the higher part of the Changhsingian. Evidence of a xenodiscid origin is lacking.</paragraph>
</footnote>
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(2) A paratirolitid origin this would imply a simplification of conch and suture morphology. Indeed, obvious phylogenetic relationships occur between the stratigraphically older
<taxonomicName id="5FD34D12FFA29364D284FE8EFED5FEE4" box="[188,320,272,295]" class="Cephalopoda" family="Dzhulfitidae" genus="Paratirolites" kingdom="Animalia" pageId="14" pageNumber="55" phylum="Mollusca" rank="genus">
<emphasis id="AAA7EA83FFA29364D284FE8EFED5FEE4" box="[188,320,272,295]" italics="true" pageId="14" pageNumber="55">Paratirolites</emphasis>
</taxonomicName>
towards the younger
<taxonomicName id="5FD34D12FFA29364D016FE8EFD05FEE4" authorityName="Shevyrev" authorityYear="1965" box="[558,656,272,295]" class="Cephalopoda" family="Dzhulfitidae" genus="Abichites" kingdom="Animalia" pageId="14" pageNumber="55" phylum="Mollusca" rank="genus">
<emphasis id="AAA7EA83FFA29364D016FE8EFD05FEE4" box="[558,656,272,295]" italics="true" pageId="14" pageNumber="55">Abichites</emphasis>
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in the reduction of size, the change from trapezoidal towards quadrate whorl cross sections, the disappearance of strong nodes in the sculpture and the reduction of sutural notching.
<taxonomicName id="5FD34D12FFA29364D043FEEEFD40FE44" authorityName="Ghaderi &amp; Leda &amp; Schobben &amp; Korn &amp; Ashouri" authorityYear="2014" box="[635,725,368,391]" class="Cephalopoda" family="Xenodiscidae" genus="Arasella" kingdom="Animalia" pageId="14" pageNumber="55" phylum="Mollusca" rank="genus">
<emphasis id="AAA7EA83FFA29364D043FEEEFD40FE44" box="[635,725,368,391]" italics="true" pageId="14" pageNumber="55">Arasella</emphasis>
</taxonomicName>
has some similarities with the stratigraphically younger species of
<taxonomicName id="5FD34D12FFA29364D2A3FE31FF68FE05" authorityName="Shevyrev" authorityYear="1965" box="[155,253,431,454]" class="Cephalopoda" family="Dzhulfitidae" genus="Abichites" kingdom="Animalia" pageId="14" pageNumber="55" phylum="Mollusca" rank="genus">
<emphasis id="AAA7EA83FFA29364D2A3FE31FF68FE05" box="[155,253,431,454]" italics="true" pageId="14" pageNumber="55">Abichites</emphasis>
</taxonomicName>
and may thus be an advanced but morphologically simplified descendent.
</paragraph>
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