399 lines
37 KiB
XML
399 lines
37 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/jhr.96.101873" ID-Pensoft-Pub="1314-2607-96-241" ID-Pensoft-UUID="A5722C06212C5BA7A2099B77C6A8DF54" ID-ZooBank="15A2B06B92F34E70AC8F6FEABF365E71" ModsDocID="1314-2607-96-241" checkinTime="1684817117864" checkinUser="pensoft" docAuthor="Wood, Thomas J." docDate="2023" docId="09000A93AC53372E78A678670C110D2D" docLanguage="en" docName="JourHymenoptRes 96: 241-484" docOrigin="Journal of Hymenoptera Research 96" docPubDate="2023-05-22" docSource="http://dx.doi.org/10.3897/jhr.96.101873" docTitle="Blandandrena fumida subsp. subgen. nov." docType="treatment" docVersion="1" id="A5722C06212C5BA7A2099B77C6A8DF54" lastPageNumber="241" masterDocId="A5722C06212C5BA7A2099B77C6A8DF54" masterDocTitle="The genus Andrena Fabricius, 1775 in the Iberian Peninsula (Hymenoptera, Andrenidae)" masterLastPageNumber="484" masterPageNumber="241" pageNumber="241" updateTime="1684817117864" updateUser="pensoft">
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<mods:mods id="7F52BC09E158483308DB9EAB0B7A0909" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="54C538B8B11DFBF10B7A52DD0D0D5EE8">
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<mods:title id="867E9A3CCCE6CBF7832397218A27AFF2">The genus Andrena Fabricius, 1775 in the Iberian Peninsula (Hymenoptera, Andrenidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="05267A01B7F463599ADD6D1615F0E1E7" type="personal">
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<mods:roleTerm id="49E76496836F67F5646AF3B29E86B0E6">Author</mods:roleTerm>
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<mods:namePart id="0926DD6216D7E147BCF355278A2110A2">Wood, Thomas J.</mods:namePart>
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<mods:nameIdentifier id="A704254789D0E1A495C6408E0CAECEC8" type="ORCID">https://orcid.org/0000-0001-5653-224X</mods:nameIdentifier>
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<mods:affiliation id="ED4F7DD09CD9E9079462488DEBD66C46">University of Mons, Research Institute for Biosciences, Laboratory of Zoology, Place du Parc 20, 7000, Mons, Belgium</mods:affiliation>
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<mods:nameIdentifier id="F9E6D1E4BBE7F8146297E3A3761DBE6F" type="email">thomasjames.wood@umons.ac.be</mods:nameIdentifier>
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<mods:typeOfResource id="61BB1BD355BABDF9C05A74BEF59F9F9B">text</mods:typeOfResource>
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<mods:titleInfo id="0F8E8DE4B453F9317489E0ECEBF261D9">
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<mods:title id="9B26E0A8545CEF938A6B8EEF81C6FADD">Journal of Hymenoptera Research</mods:title>
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</mods:titleInfo>
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<mods:part id="A910BF4A018989F9A29A9FD0F1469ED0">
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<mods:date id="B42878314D2A4D60D4F59D81E41BECA2">2023</mods:date>
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<mods:detail id="EC83EAB7B3E9C642AE8DB79988D7F132" type="pubDate">
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<mods:number id="481918DDD9F1017F50F8B01F5FB841F8">2023-05-22</mods:number>
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</mods:detail>
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<mods:detail id="3E77F0121456AB5C2EDA245094CD7E6C" type="volume">
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<mods:number id="65DFE39C088D9B435133AA863A5F8E6C">96</mods:number>
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<mods:start id="8D73A1BDB399220D1B48B1EB4B0A24F4">241</mods:start>
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<mods:end id="2A39EB8BDF09A8FDBDB3E96609D90F0A">484</mods:end>
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<mods:url id="76E140A90D54705E8F42B2EB14CAE7CE">http://dx.doi.org/10.3897/jhr.96.101873</mods:url>
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</mods:location>
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<mods:classification id="F9572609D1970CF04EAE4E2801583C6B">journal article</mods:classification>
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<mods:identifier id="A10467BC672562113DC7419AC3E558F1" type="DOI">http://dx.doi.org/10.3897/jhr.96.101873</mods:identifier>
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<mods:identifier id="3794A8613948BE2B4532E713E58C4AE7" type="Pensoft-Pub">1314-2607-96-241</mods:identifier>
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<treatment id="09000A93AC53372E78A678670C110D2D" LSID="urn:lsid:zoobank.org:act:E78F170A-60CC-4D38-9D56-9727B1347DDC" httpUri="http://treatment.plazi.org/id/09000A93AC53372E78A678670C110D2D" lastPageNumber="241" pageId="0" pageNumber="241">
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<subSubSection id="DC65AEBF21BC515DC621437498797504" pageId="0" pageNumber="241" type="nomenclature">
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<paragraph id="2C0ADBE2CE51165F3DB8A2B1324D4161" pageId="0" pageNumber="241">
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Subgenus
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<taxonomicName id="CAE3F6885B70889ECCE7F046ADF94DEC" LSID="https://zoobank.org/E78F170A-60CC-4D38-9D56-9727B1347DDC" family="Andrenidae" genus="Blandandrena" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Blandandrena subsp. subgen. nov." order="Hymenoptera" pageId="0" pageNumber="241" rank="subSpecies" species="fumida" subSpecies="subgen. nov.">Blandandrena subgen. nov.</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="9E38B28DD327D9E5836EABBA4374A63F" pageId="0" pageNumber="241" type="type species">
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<paragraph id="13826147DCDF52D3364F15EFB0E0C349" pageId="0" pageNumber="241">Type species.</paragraph>
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<paragraph id="7BA2CCEF160EE6503C93B18799B43064" pageId="0" pageNumber="241">
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<taxonomicName id="3DE99306A2D94E2479972827A835FD11" authorityName="Perez" authorityYear="1895" class="Insecta" family="Geraniaceae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena blanda" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="blanda">
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<emphasis id="C210AA4BEB7022600763C29F99942136" italics="true" pageId="0" pageNumber="241">Andrena blanda</emphasis>
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</taxonomicName>
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<normalizedToken id="ADAA2543A5E2F040A97534485D7D9390" originalValue="Pérez">Perez</normalizedToken>
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, 1895.
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</paragraph>
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</subSubSection>
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<subSubSection id="C0985AECC819674301C95D03EB98B4ED" pageId="0" pageNumber="241" type="diagnosis">
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<paragraph id="D23BCB31D06B22C5A61CAB1DC750DA2F" pageId="0" pageNumber="241">Diagnosis.</paragraph>
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<paragraph id="A2B023191F7C6924F4538840E2DDC97B" pageId="0" pageNumber="241">
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<taxonomicName id="61C185D953E2219BFFA6B8CBA4A80F39" family="Andrenidae" genus="Blandandrena" kingdom="Animalia" lsidName="Blandandrena" order="Hymenoptera" pageId="0" pageNumber="241" rank="genus">
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<emphasis id="5017BBDA12F257C453065BE297C94A41" italics="true" pageId="0" pageNumber="241">Blandandrena</emphasis>
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</taxonomicName>
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is monotypic, and hence diagnosis of
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<taxonomicName id="BDBAF65C641768065540688EDCD7C042" class="Insecta" family="Plantaginaceae" kingdom="Animalia" lsidName="A. blanda" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="blanda">
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<emphasis id="DDA388B59A32D7953ECA446F775E1FFF" italics="true" pageId="0" pageNumber="241">A. blanda</emphasis>
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</taxonomicName>
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(Fig.
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<figureCitation id="DC6C10EAFE3988DA099087C31F7C90FB" captionStart="Figure 40" captionStartId="F40" captionText="Figure 40. Andrena (Blandandrena) blanda Perez, 1895, female A profile B face, frontal view C propodeum, dorsal view D terga, dorsal view; male E profile F face, frontal view G terga, dorsal view H genital capsule." figureDoi="10.3897/jhr.96.101873.figure40" httpUri="https://binary.pensoft.net/fig/853432" pageId="0" pageNumber="241">40</figureCitation>
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) is
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<emphasis id="EC3971CA4F6C1264D3EB7996E72E4F4E" italics="true" pageId="0" pageNumber="241">de facto</emphasis>
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diagnosis of the subgenus. Through the combination of slightly upturned fore margin of the clypeus, fovea broad and occupying over
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<normalizedToken id="228DB0E10584EAED817998E3645B51F4" originalValue="½">1/2</normalizedToken>
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the space between the lateral ocellus and the inner margin of the compound eye, weak but distinct humeral angle, unmodified posterior face of the hind femur (without teeth, carinae, or spines), lack of squamous hairs, simple hind tibial spur (not broadened basally or medially), dark integument, black male clypeus, and essential absence of defining features it falls very close to members of the
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<taxonomicName id="69E9A25299AE406ED6DAC8713BB9B41B" authorityName="Warncke" authorityYear="1967" class="Insecta" family="Geraniaceae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena relata" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="relata">
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<emphasis id="32E4909DE1163936F3E90586C0DF6AC1" italics="true" pageId="0" pageNumber="241">Andrena relata</emphasis>
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</taxonomicName>
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-group and to
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<taxonomicName id="12C1D67C045B21AC4B1896852A10C7C7" class="Insecta" family="Geraniaceae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena (Ovandrena)" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="subGenus" subGenus="Ovandrena">Andrena Ovandrena</taxonomicName>
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subgen. nov. that were formerly lumped together under the subgenus
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<taxonomicName id="B863D355810412B69973154DC13F4F51" family="Andrenidae" genus="Ovandrena" kingdom="Animalia" lsidName="Ovandrena (Poliandrena)" order="Hymenoptera" pageId="0" pageNumber="241" rank="subGenus" subGenus="Poliandrena">Ovandrena Poliandrena</taxonomicName>
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(see above).
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<taxonomicName id="EDEDD8B811EE9DCA50037716215E79CE" authorityName="Perez" authorityYear="1895" class="Insecta" family="Geraniaceae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena blanda" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="blanda">
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<emphasis id="3FDF907365012D27F840E45730A8E047" italics="true" pageId="0" pageNumber="241">Andrena blanda</emphasis>
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</taxonomicName>
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females can provisionally be separated from the
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<taxonomicName id="39B2BC68AA9C5A2EBE5183840015B895" authorityName="Warncke" authorityYear="1967" class="Insecta" family="Geraniaceae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena relata" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="relata">
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<emphasis id="67371257294B2E268E323CECA7789784" italics="true" pageId="0" pageNumber="241">Andrena relata</emphasis>
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</taxonomicName>
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-group by the weakly punctate terga, punctures shallow and somewhat obscure, separated by 1-2 puncture diameters (Fig.
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<figureCitation id="2EE3BFC3E6EF5A199A840572F7E57CFE" captionStart="Figure 40" captionStartId="F40" captionText="Figure 40. Andrena (Blandandrena) blanda Perez, 1895, female A profile B face, frontal view C propodeum, dorsal view D terga, dorsal view; male E profile F face, frontal view G terga, dorsal view H genital capsule." figureDoi="10.3897/jhr.96.101873.figure40" httpUri="https://binary.pensoft.net/fig/853432" pageId="0" pageNumber="241">40D</figureCitation>
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) whereas in the
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<taxonomicName id="32CCA9BB459B0C334D295256DBF39A2E" authorityName="Warncke" authorityYear="1967" class="Insecta" family="Geraniaceae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena relata" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="relata">
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<emphasis id="7BA239766F30B6D68AFF6BE3A21E3DBB" italics="true" pageId="0" pageNumber="241">Andrena relata</emphasis>
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</taxonomicName>
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-group, tergal punctures are typically much stronger, clear and dense, separated by 1-2 puncture diameters but often by only 1 puncture diameter. This character works for West Mediterranean members of the
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<taxonomicName id="F0E48D8F12FB1EA70A45C64E73D2DF54" authorityName="Warncke" authorityYear="1967" class="Insecta" family="Geraniaceae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena relata" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="relata">
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<emphasis id="6C41ED8FA0AFBE6C5BFFE3C4A6C8DB26" italics="true" pageId="0" pageNumber="241">Andrena relata</emphasis>
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</taxonomicName>
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-group, but additional work is needed to define this subgenus in the east, and so it is not defined and described here.
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<taxonomicName id="FF2B648FF0FB59C49A5FCA60A2A7B1D8" authorityName="Perez" authorityYear="1895" class="Insecta" family="Geraniaceae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena blanda" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="blanda">
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<emphasis id="2AF89DEA309A7F4DB8058A88F3BAB026" italics="true" pageId="0" pageNumber="241">Andrena blanda</emphasis>
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</taxonomicName>
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females can be separated from the
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<taxonomicName id="4164B920B1DDF3CA211C93D7F87039DD" family="Andrenidae" genus="Ovandrena" kingdom="Animalia" lsidName="Ovandrena" order="Hymenoptera" pageId="0" pageNumber="241" rank="genus">
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<emphasis id="5D6B392ECD96F88088F9E1E8A9AFAC10" italics="true" pageId="0" pageNumber="241">Ovandrena</emphasis>
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</taxonomicName>
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by their simple scutal hairs (Fig.
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<figureCitation id="B4C7273B9E4FD4BFD7791CD2F9178894" captionStart="Figure 40" captionStartId="F40" captionText="Figure 40. Andrena (Blandandrena) blanda Perez, 1895, female A profile B face, frontal view C propodeum, dorsal view D terga, dorsal view; male E profile F face, frontal view G terga, dorsal view H genital capsule." figureDoi="10.3897/jhr.96.101873.figure40" httpUri="https://binary.pensoft.net/fig/853432" pageId="0" pageNumber="241">40A-C</figureCitation>
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; hairs semi-squamous in
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<taxonomicName id="6F99824779F95DE1B5CC33CC21E14D52" family="Andrenidae" genus="Ovandrena" kingdom="Animalia" lsidName="Ovandrena" order="Hymenoptera" pageId="0" pageNumber="241" rank="genus">
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<emphasis id="5FC4165EE9080BAEBF6C517FA24065D6" italics="true" pageId="0" pageNumber="241">Ovandrena</emphasis>
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</taxonomicName>
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) and poorly delineated propodeal triangle that lacks lateral carinae (Fig.
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<figureCitation id="305A743B595E6D4D3A0F51C5DCC588D3" captionStart="Figure 40" captionStartId="F40" captionText="Figure 40. Andrena (Blandandrena) blanda Perez, 1895, female A profile B face, frontal view C propodeum, dorsal view D terga, dorsal view; male E profile F face, frontal view G terga, dorsal view H genital capsule." figureDoi="10.3897/jhr.96.101873.figure40" httpUri="https://binary.pensoft.net/fig/853432" pageId="0" pageNumber="241">40C</figureCitation>
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; propodeal triangle clearly defined by presence of lateral carinae in
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<taxonomicName id="EFABF2D2B09DE05A041BEEEDC55F55CF" family="Andrenidae" genus="Ovandrena" kingdom="Animalia" lsidName="Ovandrena" order="Hymenoptera" pageId="0" pageNumber="241" rank="genus">
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<emphasis id="060DF0C535BFF4239702B758F69EC8A1" italics="true" pageId="0" pageNumber="241">Ovandrena</emphasis>
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</taxonomicName>
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; see below for additional detail).
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</paragraph>
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<caption id="6E7B20DB1F8FC2C1FBEF05710F5C156F" doi="10.3897/jhr.96.101873.figure40" httpUri="https://binary.pensoft.net/fig/853432" pageId="0" pageNumber="241" start="Figure 40" startId="F40">
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<paragraph id="EFD0B41B7655FB83183C2ED3BB1E9594" pageId="0" pageNumber="241">
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<emphasis id="DFE8257F09EFDBE22DC6424E09072B6A" bold="true" pageId="0" pageNumber="241">Figure 40.</emphasis>
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<taxonomicName id="0B61027C83586E822B74ACAF88D8AFFF" authorityName="Perez" authorityYear="1895" class="Insecta" family="Geraniaceae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena (Blandandrena) blanda" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="blanda" subGenus="Blandandrena">Andrena (Blandandrena) blanda</taxonomicName>
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<normalizedToken id="4A335E80E72EBBFA4F09B09F58B43C5B" originalValue="Pérez">Perez</normalizedToken>
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, 1895, female
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<emphasis id="5AEB86E46293AEC8DBC8B158ECD4934F" bold="true" pageId="0" pageNumber="241">A</emphasis>
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profile
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<emphasis id="D9AD1A6E9CE1600764AB3DA7058BC2AD" bold="true" pageId="0" pageNumber="241">B</emphasis>
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face, frontal view
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<emphasis id="D670F660B3C08CDED8EF9275DA987ABA" bold="true" pageId="0" pageNumber="241">C</emphasis>
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propodeum, dorsal view
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<emphasis id="BA8FD7F65FBA4512F99397B24AE29AF7" bold="true" pageId="0" pageNumber="241">D</emphasis>
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terga, dorsal view; male
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<emphasis id="99CB3146296AB623C74C3B43FEF41391" bold="true" pageId="0" pageNumber="241">E</emphasis>
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profile
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<emphasis id="1F270DEF18A6BA9917F868C526FE4AC6" bold="true" pageId="0" pageNumber="241">F</emphasis>
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face, frontal view
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<emphasis id="D82BB4EAA5A277E9FB1A590CF6DF76B5" bold="true" pageId="0" pageNumber="241">G</emphasis>
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terga, dorsal view
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<emphasis id="88CA3873FBA83EFDAAB94EDA07F02543" bold="true" pageId="0" pageNumber="241">H</emphasis>
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genital capsule.
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</paragraph>
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</caption>
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<paragraph id="6A8D98272F57A88D42673D9D3E9704EC" pageId="0" pageNumber="241">
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<taxonomicName id="BC18182B81E5432AB0AFAD52899B79E0" authorityName="Perez" authorityYear="1895" class="Insecta" family="Geraniaceae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena blanda" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="blanda">
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<emphasis id="39CFB4BFA47913CF3A56E63FCFDEC29C" italics="true" pageId="0" pageNumber="241">Andrena blanda</emphasis>
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</taxonomicName>
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males can be separated by their black clypeus (Fig.
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<figureCitation id="3570247909E9F041938DD9B285A577E3" captionStart="Figure 40" captionStartId="F40" captionText="Figure 40. Andrena (Blandandrena) blanda Perez, 1895, female A profile B face, frontal view C propodeum, dorsal view D terga, dorsal view; male E profile F face, frontal view G terga, dorsal view H genital capsule." figureDoi="10.3897/jhr.96.101873.figure40" httpUri="https://binary.pensoft.net/fig/853432" pageId="0" pageNumber="241">40F</figureCitation>
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) in combination with their distinctive genital capsule (Fig.
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<figureCitation id="B92137405F011E25CD40E72E2F90295A" captionStart="Figure 40" captionStartId="F40" captionText="Figure 40. Andrena (Blandandrena) blanda Perez, 1895, female A profile B face, frontal view C propodeum, dorsal view D terga, dorsal view; male E profile F face, frontal view G terga, dorsal view H genital capsule." figureDoi="10.3897/jhr.96.101873.figure40" httpUri="https://binary.pensoft.net/fig/853432" pageId="0" pageNumber="241">40G</figureCitation>
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). Most members of the
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<taxonomicName id="1EBB66B44F47B4C7C30BF63BC972552F" authorityName="Warncke" authorityYear="1967" class="Insecta" family="Geraniaceae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena relata" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="relata">
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<emphasis id="1AF49DF9D43F33E024CA98477683CFD3" italics="true" pageId="0" pageNumber="241">Andrena relata</emphasis>
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</taxonomicName>
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-group have a yellow-marked clypeus, but for those with a black clypeus (e.g.
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|
<taxonomicName id="CA8292EE0D8C08497D419CD935B6CA53" class="Insecta" family="Plantaginaceae" kingdom="Animalia" lsidName="A. corax" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="corax">
|
|
<emphasis id="EF9894CF0D45CF751BD7BC47DC603D73" italics="true" pageId="0" pageNumber="241">A. corax</emphasis>
|
|
</taxonomicName>
|
|
), the genital capsule allows recognition. In
|
|
<taxonomicName id="4B8A6C9200AE9833E94F2DAFCF1C263D" class="Insecta" family="Plantaginaceae" kingdom="Animalia" lsidName="A. blanda" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="blanda">
|
|
<emphasis id="DE0B2F4A7DCEF45324678AEF7FC8406B" italics="true" pageId="0" pageNumber="241">A. blanda</emphasis>
|
|
</taxonomicName>
|
|
, the genital capsule is short and compact, more or less round, with gonocoxae with apically diverging inner margins and which are produced into short pointed teeth, and the gonostyli are apically strongly broadened and flattened, the disc being slightly broader than long. In the
|
|
<taxonomicName id="DF8E59287D411418922859246FDB869D" authorityName="Warncke" authorityYear="1967" class="Insecta" family="Geraniaceae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena relata" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="relata">
|
|
<emphasis id="3C7595A15C894A4A45B408C494D6C37F" italics="true" pageId="0" pageNumber="241">Andrena relata</emphasis>
|
|
</taxonomicName>
|
|
-group, the genital capsule is typically simple, without strongly apically broadened gonostyli, or when this is the case (e.g.
|
|
<taxonomicName id="E46392737C412C4D1CE61C94A01BB58C" class="Insecta" family="Plantaginaceae" kingdom="Animalia" lsidName="A. murana" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="murana">
|
|
<emphasis id="958B3E796AF05D94D94986C7BA68629B" italics="true" pageId="0" pageNumber="241">A. murana</emphasis>
|
|
</taxonomicName>
|
|
Warncke, 1975a) then the clypeus is yellow-marked and the gonocoxae are not produced into pointed teeth.
|
|
<taxonomicName id="8CC9CFE2F938325D38B7585D743817A6" authorityName="Perez" authorityYear="1895" class="Insecta" family="Geraniaceae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena blanda" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="blanda">
|
|
<emphasis id="1D22BD0F7927D20F76B2E93E9A2857B4" italics="true" pageId="0" pageNumber="241">Andrena blanda</emphasis>
|
|
</taxonomicName>
|
|
can be separated from the
|
|
<taxonomicName id="663DECF4095E7E8C6FD532E4B402AF25" family="Andrenidae" genus="Ovandrena" kingdom="Animalia" lsidName="Ovandrena" order="Hymenoptera" pageId="0" pageNumber="241" rank="genus">
|
|
<emphasis id="D5BCEC5E82E3BB76A0321B7DB18FDC9A" italics="true" pageId="0" pageNumber="241">Ovandrena</emphasis>
|
|
</taxonomicName>
|
|
by the genital capsule, as in
|
|
<taxonomicName id="E721F74AE5479A9BB07230B3C3E181D8" family="Andrenidae" genus="Ovandrena" kingdom="Animalia" lsidName="Ovandrena" order="Hymenoptera" pageId="0" pageNumber="241" rank="genus">
|
|
<emphasis id="664878E6AF478F122FB478C5E0BC8784" italics="true" pageId="0" pageNumber="241">Ovandrena</emphasis>
|
|
</taxonomicName>
|
|
the gonostyli are apically flattened and spatulate, but the disc is always longer than broad. The gonocoxae are also produced into apically projecting teeth, but the inner margins of the gonocoxae are parallel and do not diverge. The propodeal triangle of
|
|
<taxonomicName id="6A91F1301446F2745FBE512DC0DA5766" family="Andrenidae" genus="Ovandrena" kingdom="Animalia" lsidName="Ovandrena" order="Hymenoptera" pageId="0" pageNumber="241" rank="genus">
|
|
<emphasis id="D1EAAF4B3E2D8E85F1136C38403CAF6E" italics="true" pageId="0" pageNumber="241">Ovandrena</emphasis>
|
|
</taxonomicName>
|
|
is also triangular and strongly defined by lateral carinae, whereas in
|
|
<taxonomicName id="5CA5E61DBAFC542A941901E7273213BA" class="Insecta" family="Plantaginaceae" kingdom="Animalia" lsidName="A. blanda" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="blanda">
|
|
<emphasis id="81E1B9DC524E2FA7ACB6A93901B340DF" italics="true" pageId="0" pageNumber="241">A. blanda</emphasis>
|
|
</taxonomicName>
|
|
it is poorly defined and lacks lateral carinae.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C71E84CA2424BDE2B49DA15F7FD69588" pageId="0" pageNumber="241" type="description">
|
|
<paragraph id="A26228AED048B90B132DCE95B54ADB54" pageId="0" pageNumber="241">Description.</paragraph>
|
|
<paragraph id="8889E7EAAD330AA7FBE7EB66659BFF35" pageId="0" pageNumber="241">Medium-sized bees (9-10 mm) with dark integument. Head broad, 1.4 times broader than long. Gena slightly exceeding width of compound eye; ocelloccipital distance 1.5-2 times diameter of lateral ocellus, slightly broader in male sex. Facial fovea broad, occupying almost entire distance between lateral ocellus and inner margin of compound eye. Mesosoma dorsally with moderately long light brown hairs, laterally with white hairs. Pronotum laterally with humeral angle. Dorsolateral surface of propodeum with obscure and finely raised rugosity; propodeal triangle broad, poorly delineated laterally, surface with fine granular reticulation, basally with weakly raised rugosity, propodeal triangle thus defined by change in surface sculpture. Forewing with nervulus antefurcal. Hind tibial spurs simple, not broadened basally or medially. Terga weakly and obscurely punctate, punctures separated by 1-2 puncture diameters. Male genital capsule rounded, more or less circular in outline, gonocoxae with inner margins apically diverging, produced into apically projecting short pointed teeth. Gonostyli apically broadened and flattened, apical disc slightly broader than long. Penis valves relatively narrow, occupying less than half space between gonostyli.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="62B170ECA0E138D41DE974466A4FD9B4" pageId="0" pageNumber="241" type="etymology">
|
|
<paragraph id="06279D5FF88512CFA4F9520B979F6DA5" pageId="0" pageNumber="241">Etymology.</paragraph>
|
|
<paragraph id="5D7E37921C7FB324CBAADBD1F6DBA26E" pageId="0" pageNumber="241">
|
|
The name is taken from the name of the type species
|
|
<taxonomicName id="DC2B4B567BF133E7F31F45C90B68E3EC" class="Insecta" family="Plantaginaceae" kingdom="Animalia" lsidName="A. blanda" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="blanda">
|
|
<emphasis id="84358213DAB7F0BFEB287E1EF1BF0F8B" italics="true" pageId="0" pageNumber="241">A. blanda</emphasis>
|
|
</taxonomicName>
|
|
, with
|
|
<taxonomicName id="E02B12526AD3CA1F67BC79885B24D3CD" authorityName="Perez" authorityYear="1895" class="Insecta" family="Geraniaceae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena blanda" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="blanda">
|
|
<emphasis id="EE81AA2D3D37EAB4A32AD2099D4EE61F" italics="true" pageId="0" pageNumber="241">Andrena blanda</emphasis>
|
|
</taxonomicName>
|
|
being the feminine singular of the adjective
|
|
<emphasis id="CDA4EE5A8048A226B0FDDFB2800AE291" italics="true" pageId="0" pageNumber="241">blandus</emphasis>
|
|
which can mean pleasant, agreeable, smooth. It can be used to refer to the generally unremarkable nature of the species which has made it hard to assign to a particular group of species. The gender is feminine.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="4803D3B6F6D6E89CA06D9A5E11C9FA14" pageId="0" pageNumber="241" type="included species">
|
|
<paragraph id="6F034B0116AAE22D27263F6654CF150A" pageId="0" pageNumber="241">Included species.</paragraph>
|
|
<paragraph id="6223EBE5D3A53B489B56C94773203B5B" pageId="0" pageNumber="241">
|
|
<taxonomicName id="CC4152663FE5E22C36212C83C7782A31" authorityName="Perez" authorityYear="1895" class="Insecta" family="Geraniaceae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena blanda" order="Hymenoptera" pageId="0" pageNumber="241" phylum="Arthropoda" rank="species" species="blanda">
|
|
<emphasis id="AC19F949B78A7FDC57B20AD9DA76EEC0" italics="true" pageId="0" pageNumber="241">Andrena blanda</emphasis>
|
|
</taxonomicName>
|
|
(Spain, including mainland Spain and newly recorded for Fuerteventura), Morocco, Algeria, Tunisia;
|
|
<bibRefCitation id="BEF58F214A508BBC89CCC3FDD99F2466" author="Gusenleitner, F" journalOrPublisher="Entomofauna, Supplement" pageId="0" pageNumber="241" pagination="1 - 1280" refId="B38" refString="Gusenleitner, F, Schwarz, M, 2002. Weltweite Checkliste der Bienengattung Andrena mit Bemerkungen und Ergaenzungen zu palaearktischen Arten (Hymenoptera, Apidae, Andreninae, Andrena). Entomofauna, Supplement 10: 1 - 1280" title="Weltweite Checkliste der Bienengattung Andrena mit Bemerkungen und Ergaenzungen zu palaearktischen Arten (Hymenoptera, Apidae, Andreninae, Andrena)." volume="10" year="2002">Gusenleitner and Schwarz 2002</bibRefCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="E349F682FA6E94832BC0365AAE33B91A" pageId="0" pageNumber="241" type="materials_examined">
|
|
<paragraph id="3D9769F8BEE9E68815C51CF2D79E0563" pageId="0" pageNumber="241">Material examined (illustrative).</paragraph>
|
|
<paragraph id="94638F550E2C582F5DD9843EDE5EF55F" pageId="0" pageNumber="241">
|
|
<materialsCitation id="1BFD80A3E2F86BC60EA5E36FBDABAE64" collectingDate="1983-04-06" collectingDateMax="1983-04-07" collectingDateMin="1983-04-06" collectorName="R. Leys, Hurk" country="Algeria" latitude="34.86" location="Mecheria" longLatPrecision="6" longitude="5.6995" specimenCount="3" specimenCount-female="3" stateProvince="Biskra" typeStatus="lectotype">
|
|
<emphasis id="9D30476B583DD1CB83B1B6E663D547B1" bold="true" pageId="0" pageNumber="241">
|
|
<collectingCountry id="4FDFA94B12A67837AF6750B6D0407A44" name="Algeria">Algeria</collectingCountry>
|
|
</emphasis>
|
|
:
|
|
<collectingRegion id="4520F35EE94FB21B6EEEE9E4E87871A2" country="Algeria" name="Biskra">Biskra</collectingRegion>
|
|
[
|
|
<geoCoordinate id="BDB88B22040162031D2FE77835F5CE19" degrees="34.8600" direction="north" orientation="latitude" precision="5" value="34.86">34.8600°N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="F84A4DB4160023FB7F401AE2901E3D9D" degrees="5.6995" direction="east" orientation="longitude" precision="5" value="5.6995">5.6995°E</geoCoordinate>
|
|
],
|
|
<specimenCount id="12FBAB3812CEAAAC1FA2979E006198CA" type="female">1♀</specimenCount>
|
|
, MNHN (
|
|
<typeStatus id="892059FEA54A40AB651430AF8D7E4EF2">lectotype</typeStatus>
|
|
des.
|
|
<bibRefCitation id="156940160E0B2E2DF207E27C79B403A7" author="Warncke, K" journalOrPublisher="Eos" pageId="0" pageNumber="241" pagination="171 - 318" refId="B93" refString="Warncke, K, 1967. Beitrag zur Klaerung palaearktischer Andrena -Arten. Eos 43: 171 - 318" title="Beitrag zur Klaerung palaearktischer Andrena - Arten." volume="43" year="1967">Warncke 1967</bibRefCitation>
|
|
);
|
|
<quantity id="FB69AA0E87DD90A749DE527BEC2BD457" metricMagnitude="3" metricUnit="m" metricValue="5.0" unit="km" value="5.0">5 km</quantity>
|
|
N of
|
|
<location id="007C545A2A00EA3FEEF32E5E7892AE7B" LSID="urn:lsid:plazi:treatment:09000A93AC53372E78A678670C110D2D:007C545A2A00EA3FEEF32E5E7892AE7B" country="Algeria" latitude="34.86" longLatPrecision="6" longitude="5.6995" name="Mecheria" stateProvince="Biskra">Mecheria</location>
|
|
, st. 9,
|
|
<collectingDate id="DF124ABF63C16551854726622BCC4127" value="1983-04-07">7.iv.1983</collectingDate>
|
|
,
|
|
<specimenCount id="C54DDA95ED01E808C219F3603507AD73" type="female">1♀</specimenCount>
|
|
, leg.
|
|
<collectorName id="8B024B75A6A6F0434F83B14EF224B42A">R. Leys</collectorName>
|
|
& P. v. d.
|
|
<collectorName id="441C099893F63B57793ED1AB15C273DC">Hurk</collectorName>
|
|
, RMNH;
|
|
<locationDeviation id="15B6B5340922D0C4DA5302653CCB2B49" location="Sfissifa">
|
|
<quantity id="F7EC55D16824A09F693F85C6DAAEB189" metricMagnitude="3" metricUnit="m" metricValue="5.0" unit="km" value="5.0">5 km</quantity>
|
|
SE of Sfissifa
|
|
</locationDeviation>
|
|
, st. 8,
|
|
<collectingDate id="510195E9517B8FC050CE6DB600D7EF1E" value="1983-04-06">6.iv.1983</collectingDate>
|
|
,
|
|
<specimenCount id="D73BB526874B9A2F7D35D0E7DCE65E2B" type="female">1♀</specimenCount>
|
|
, leg.
|
|
<collectorName id="DD88A2693FF939A545CD29E0DB28FF3D">R. Leys</collectorName>
|
|
& P. v. d.
|
|
<collectorName id="BE711662B4CE30E27424405B5BEB88A0">Hurk</collectorName>
|
|
, RMNH
|
|
</materialsCitation>
|
|
;
|
|
<materialsCitation id="E96116BB8E2926DD3A19454F73E1F694" collectingDate="1986-03-30" collectingDateMax="2022-04-15" collectingDateMin="1986-03-30" collectorName="T. J. Wood, Foum Zguid, M. Schwarz, Ifkern, Mi. Halada" country="Morocco" elevation="1600" location="Tazenakht" municipality="Draa-Tafilalet" specimenCount="28" specimenCount-female="27" specimenCount-male="1" typeStatus="lectotype">
|
|
<emphasis id="D190CA65D7E82564B58E0925019FE668" bold="true" pageId="0" pageNumber="241">
|
|
<collectingCountry id="CFC58BE193B65EEAE396835FE1AB3031" name="Morocco">Morocco</collectingCountry>
|
|
</emphasis>
|
|
:
|
|
<normalizedToken id="A983AD7236F09A22E821E150946F5177" originalValue="Drâa-Tafilalet">
|
|
<collectingMunicipality id="36CAE2BAFA1A94445EC3C7339500D848">Draa-Tafilalet</collectingMunicipality>
|
|
</normalizedToken>
|
|
,
|
|
<location id="53AD7205D0E703F5DB93C7A941A10156" LSID="urn:lsid:plazi:treatment:09000A93AC53372E78A678670C110D2D:53AD7205D0E703F5DB93C7A941A10156" country="Morocco" municipality="Draa-Tafilalet" name="Tazenakht">Tazenakht</location>
|
|
,
|
|
<locationDeviation id="FC1981CB52E54A49C5A9F0B61C96E75D" location="Anezal">
|
|
<quantity id="32EEDA187DBA6082F5DE5ECB764FBB64" metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="km" value="1.0">1 km</quantity>
|
|
W Anezal
|
|
</locationDeviation>
|
|
, c.
|
|
<quantity id="B04987D8980CEB7DDD8889E31EBAE116" metricMagnitude="3" metricUnit="m" metricValue="1.6" unit="m" value="1600.0">
|
|
<elevation id="04379039503B3B6B757CA9F188CC1E44" metricMagnitude="3" metricUnit="m" metricValue="1.6" unit="m" value="1600.0">1600 m</elevation>
|
|
</quantity>
|
|
,
|
|
<collectingDate id="8C002AE71734CFD2ABAD3704C12786BB" value="2022-04-15">15.iv.2022</collectingDate>
|
|
,
|
|
<specimenCount id="377D96DD368C8D2203C40377963D2092" type="female">1♀</specimenCount>
|
|
, leg.
|
|
<collectorName id="37410F428DC638F86668414C9F706E2F">T.J. Wood</collectorName>
|
|
, TJWC;
|
|
<collectorName id="D2406895E46396FA48C4111CAB4AEF1A">Foum Zguid</collectorName>
|
|
,
|
|
<quantity id="407733465075AE6DFC25EB2B00E95D88" metricMagnitude="4" metricUnit="m" metricValue="5.0" unit="km" value="50.0">50 km</quantity>
|
|
N,
|
|
<collectingDate id="C681B833211980919D2F8293ABB5326D" value="1986-03-30">30.iii.1986</collectingDate>
|
|
,
|
|
<specimenCount id="0A042743ACAAB9D40AD1AC3E3C91CB1B" type="male">1♂</specimenCount>
|
|
,
|
|
<specimenCount id="40FAB16B96FAEA18773FEE4976C37472" type="female">16♀</specimenCount>
|
|
, leg.
|
|
<collectorName id="ACDF6D48822D05132C02BF2EE1126536">M. Schwarz</collectorName>
|
|
, MSC;
|
|
<collectorName id="02B8ABFC0F0B930B85AB6AB3D7801267">Ifkern</collectorName>
|
|
,
|
|
<locationDeviation id="0DDEBAE2754B5AFE54B8D13237B3ABCA" location="Boulemane">
|
|
<quantity id="0883716DC42364484172A80EA3DDCA74" metricMagnitude="4" metricUnit="m" metricValue="2.5" unit="km" value="25.0">25 km</quantity>
|
|
E Boulemane
|
|
</locationDeviation>
|
|
,
|
|
<collectingDate id="6F9C67368EB91F06C392981F27D61350" value="1995-05-25">25.v.1995</collectingDate>
|
|
,
|
|
<specimenCount id="EA443E683B65E29C6EC580AC482D420E" type="female">10♀</specimenCount>
|
|
, leg.
|
|
<collectorName id="83D30504BB10BE6248A744E29491FC09">Mi. Halada</collectorName>
|
|
,
|
|
<normalizedToken id="49B4CF90EDF9088E00F1750BE1D11E6D" originalValue="OÖLM">OOELM</normalizedToken>
|
|
</materialsCitation>
|
|
;
|
|
<materialsCitation id="C17A2627633C673B786D928298A4D64D" collectingDate="1982-05-26" collectingDateMax="2015-04-01" collectingDateMin="1982-05-26" collectorName="A. Mueller, Fuerteventura, S, de Maria, J. Halada, R. Leys" country="Spain" county="Fuerteventura" location="Montana Pelada" municipality="Costa Calma" specimenCount="10" specimenCount-female="10" typeStatus="lectotype">
|
|
<emphasis id="080083FB16E8CC1E871E062F6E7696DE" bold="true" pageId="0" pageNumber="241">
|
|
<collectingCountry id="56D2B0DAB5B97FC01DCC9000DE1B19FA" name="Spain">Spain</collectingCountry>
|
|
</emphasis>
|
|
:
|
|
<collectingCounty id="BA9D0E624B4BA4301A350DF470FDA9C9">Fuerteventura</collectingCounty>
|
|
,
|
|
<collectingMunicipality id="D5729FE9D91F4532F754E770EC1617F2">Costa Calma</collectingMunicipality>
|
|
, ESE
|
|
<location id="1EC0334F12C92AA60876E19E7BC6D087" LSID="urn:lsid:plazi:treatment:09000A93AC53372E78A678670C110D2D:1EC0334F12C92AA60876E19E7BC6D087" country="Spain" county="Fuerteventura" municipality="Costa Calma" name="Montana Pelada">
|
|
<normalizedToken id="8C82AFD4D3205D9FAE16245FB1C6DC62" originalValue="Montaña">Montana</normalizedToken>
|
|
Pelada
|
|
</location>
|
|
,
|
|
<collectingDate id="9D00E19D8690E33829B53FE7CBC451D7" value="2015-03-29">29.iii.2015</collectingDate>
|
|
,
|
|
<specimenCount id="FC5D7999786A4177B243F309E908820B" type="female">3♀</specimenCount>
|
|
, leg.
|
|
<collectorName id="A2325F6DB923874A2B5BB3B396D85445">
|
|
A.
|
|
<normalizedToken id="7544B120F661DD16EEAFD63F6283D50C" originalValue="Müller">Mueller</normalizedToken>
|
|
</collectorName>
|
|
, AMC/TJWC;
|
|
<collectorName id="BDE3AF7FB8B5022827CC1C96970FCF03">Fuerteventura, S</collectorName>
|
|
Costa Calma,
|
|
<normalizedToken id="1BF069E5DFA03359F28E3AC272FCBDC6" originalValue="Montañeta">Montaneta</normalizedToken>
|
|
de los Verdes,
|
|
<collectingDate id="2784BF1835C92515FFFF72B6EF5F7444" value="2015-04-01">1.iv.2015</collectingDate>
|
|
,
|
|
<specimenCount id="358629932E65D501B3902FAAB09615A2" type="female">5♀</specimenCount>
|
|
, leg.
|
|
<collectorName id="667EA08049324B319D3EF42045F0DE55">
|
|
A.
|
|
<normalizedToken id="CF6CCC9A536C5CC564E2856CCE06899C" originalValue="Müller">Mueller</normalizedToken>
|
|
</collectorName>
|
|
, AMC;
|
|
<location id="5830DCB3347094D40A5743A76826B5A6" LSID="urn:lsid:plazi:treatment:09000A93AC53372E78A678670C110D2D:5830DCB3347094D40A5743A76826B5A6" country="Spain" county="Fuerteventura" municipality="Costa Calma" name="Sierra de Maria">
|
|
Sierra
|
|
<collectorName id="5B4964FDF9FC9758AF1DF25010CDE53A">
|
|
de
|
|
<normalizedToken id="04B18015828FA959E02CD2A82E05EC42" originalValue="María">Maria</normalizedToken>
|
|
</collectorName>
|
|
</location>
|
|
,
|
|
<locationDeviation id="EFA91C037D50BBE9929CF36BC4CF4A6E" location="Lorca">
|
|
<quantity id="6E3EBA185F5D13D31AABF2AA89BE95A6" metricMagnitude="4" metricUnit="m" metricValue="2.5" unit="km" value="25.0">25 km</quantity>
|
|
W Lorca
|
|
</locationDeviation>
|
|
,
|
|
<collectingDate id="E8A3B46E93DE29293DF53E637B031C62" value="2003-05-10">10.v.2003</collectingDate>
|
|
,
|
|
<specimenCount id="30B9F373C2B82BF02D4725C4E9E4C2D5" type="female">1♀</specimenCount>
|
|
, leg.
|
|
<collectorName id="BCB17E62D716D2AD061F9E2E546046EE">J. Halada</collectorName>
|
|
,
|
|
<normalizedToken id="9099A652739C9A21D44CB38FF887690B" originalValue="OÖLM">OOELM</normalizedToken>
|
|
; Granada, Pantano de los Bermejales,
|
|
<collectingDate id="C3764FFE93F1B67CAE31039B18572770" value="1982-05-26">26.v.1982</collectingDate>
|
|
,
|
|
<specimenCount id="2BD831E11FBB84F69F388DA6020F7D37" type="female">1♀</specimenCount>
|
|
, leg.
|
|
<collectorName id="B8C3234069D89C58DE942050A5260823">R. Leys</collectorName>
|
|
, RMNH
|
|
</materialsCitation>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |