treatments-xml/data/03/A0/E2/03A0E21C9853DA5FEAD3A26D5132DC6D.xml

<document id="E0A3FA8CFD0B6908C36C747F6080FE5C" ID-DOI="10.5281/zenodo.13244564" ID-ISSN="2345-7600" ID-Zenodo-Dep="13244564" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722970052887" checkinUser="felipe" docAuthor="Freitag, Hendrik &amp; Yeo, Darren C. J." docDate="2004" docId="03A0E21C9853DA5FEAD3A26D5132DC6D" docLanguage="en" docName="RafflesBZool.52.1.227-237.pdf" docOrigin="Raffles Bulletin of Zoology 52 (1)" docStyle="DocumentStyle:19380AF0FBA5B2F218DD253A34ABE57F.5:RafflesBZool.2003-2005.journal_article" docStyleId="19380AF0FBA5B2F218DD253A34ABE57F" docStyleName="RafflesBZool.2003-2005.journal_article" docStyleVersion="5" docTitle="Parathelphusa cabayugan Freitag &amp; Yeo, 2004, new species" docType="treatment" docVersion="2" lastPageNumber="232" masterDocId="FF999A649852DA5AE942A11D5465DB3D" masterDocTitle="Two New Species Of Parathelphusa H. Milne Edwards, 1853, From The Philippines (Crustacea: Decapoda: Brachyura: Parathelphusidae)" masterLastPageNumber="237" masterPageNumber="227" pageNumber="228" updateTime="1722974698588" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="EEDEEB6F66074E9353BD1CFBC4C24165">Two New Species Of Parathelphusa H. Milne Edwards, 1853, From The Philippines (Crustacea: Decapoda: Brachyura: Parathelphusidae)</mods:title>
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<mods:namePart id="EB950D1F2CB8CD0DB534773C197AAE48">Freitag, Hendrik</mods:namePart>
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<mods:namePart id="420F0DB0A8C5C299C3BB7C186676203E">Yeo, Darren C. J.</mods:namePart>
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<mods:title id="DC2A7F66AE57C197E771FDFB90610440">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="96D76695207BDCC7495E756997FD4CD7">2004</mods:date>
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<treatment id="03A0E21C9853DA5FEAD3A26D5132DC6D" LSID="urn:lsid:plazi:treatment:03A0E21C9853DA5FEAD3A26D5132DC6D" httpUri="http://treatment.plazi.org/id/03A0E21C9853DA5FEAD3A26D5132DC6D" lastPageId="5" lastPageNumber="232" pageId="1" pageNumber="228">
<subSubSection id="C31300819853DA5BEAD3A26D512DD8B5" box="[913,1352,880,904]" pageId="1" pageNumber="228" type="nomenclature">
<paragraph id="8BB6530A9853DA5BEAD3A26D512DD8B5" blockId="1.[913,1352,880,936]" box="[913,1352,880,904]" pageId="1" pageNumber="228">
<heading id="D0FEE4669853DA5BEAD3A26D512DD8B5" bold="true" box="[913,1352,880,904]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="228" reason="2">
<emphasis id="B97D8F189853DA5BEAD3A26D512DD8B5" bold="true" box="[913,1352,880,904]" pageId="1" pageNumber="228">
<taxonomicName id="4C0928899853DA5BEAD3A26D50D6D8B5" ID-CoL="75RRZ" authorityName="Freitag &amp; Yeo" authorityYear="2004" box="[913,1203,880,904]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="228" phylum="Arthropoda" rank="species" species="cabayugan" status="sp. nov.">
<emphasis id="B97D8F189853DA5BEAD3A26D50D6D8B5" bold="true" box="[913,1203,880,904]" italics="true" pageId="1" pageNumber="228">Parathelphusa cabayugan</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A24E32639853DA5BED80A26D512DD8B5" box="[1218,1352,880,904]" pageId="1" pageNumber="228" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31300819853DA5BED71A28C50C0D895" box="[1075,1189,913,936]" pageId="1" pageNumber="228" type="description">
<paragraph id="8BB6530A9853DA5BED71A28C50C0D895" blockId="1.[913,1352,880,936]" box="[1075,1189,913,936]" pageId="1" pageNumber="228">
(
<figureCitation id="13324F8F9853DA5BED79A28C50F9D895" box="[1083,1180,913,936]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="2.[136,171,2036,2057]" captionStartId-1="3.[113,148,2033,2054]" captionStartId-2="4.[136,171,2036,2057]" captionStartId-3="4.[837,872,2063,2084]" captionTargetBox-0="[165,1434,196,1966]" captionTargetBox-1="[154,1408,195,1984]" captionTargetBox-2="[136,776,870,2013]" captionTargetBox-3="[835,1475,1603,2045]" captionTargetId-0="figure-8@2.[109,1479,161,2011]" captionTargetId-1="figure-12@3.[135,1443,160,2008]" captionTargetId-2="figure-8@4.[136,779,869,2013]" captionTargetId-3="figure-10@4.[835,1477,1603,2045]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="4" captionText-0="Fig. 1. Parathelphusa cabayugan, new species, holotype male (32 by 25.3 mm) (PNM 17259). A, dorsal view of carapace; B, frontal view of carapace; C, right third ambulatory merus; D, right fourth ambulatory merus; E, left chela; F, abdominal segments 5-7. Scales = 10.0 mm." captionText-1="Fig. 2. Parathelphusa cabayugan, new species, left G1. A, B, E, F, holotype male (32 by 25.3 mm) (PNM 17259); C, D, G, H, paratype juvenile male (15.2 by 13.1 mm) (SPCP-ASTD). A, C, ventral view; B, D, dorsal view; E, G, ventral view of distal part; F, H, dorsal view of distal part. Scales = 1.0 mm." captionText-2="Fig. 3. Parathelphusa cabayugan, new species, holotype male (32 by 25.3 mm) (PNM 17259).A, dorsal view; B frontal view; C, ventral view." captionText-3="Fig. 4. Parathelphusa cabayugan, new species, paratype female (26.4 by 21.6 mm) (ZIH)." figureDoi-0="http://doi.org/10.5281/zenodo.13244566" figureDoi-1="http://doi.org/10.5281/zenodo.13244568" figureDoi-2="http://doi.org/10.5281/zenodo.13244570" figureDoi-3="http://doi.org/10.5281/zenodo.13244572" httpUri-0="https://zenodo.org/record/13244566/files/figure.png" httpUri-1="https://zenodo.org/record/13244568/files/figure.png" httpUri-2="https://zenodo.org/record/13244570/files/figure.png" httpUri-3="https://zenodo.org/record/13244572/files/figure.png" pageId="1" pageNumber="228">Figs. 1-4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31300819853DA5BEA6FA2D051C8DE18" pageId="1" pageNumber="228" type="materials_examined">
<paragraph id="8BB6530A9853DA5BEA6FA2D057FADF20" blockId="1.[813,1452,973,1053]" pageId="1" pageNumber="228">
<emphasis id="B97D8F189853DA5BEA6FA2D05062D8DF" bold="true" box="[813,1031,973,994]" italics="true" pageId="1" pageNumber="228">Material examined. –</emphasis>
<materialsCitation id="3B6159579853DA5BED52A2D057FADF20" collectingDate="2001-06-05" collectionCode="PNM" collectorName="H. Freitag" location="Puerto Princesa, Palawan" pageId="1" pageNumber="228" specimenCode="PNM 17259" specimenCount="1" stateProvince="Puerto Princesa, Palawan" typeStatus="holotype">
<typeStatus id="54B2EDA89853DA5BED52A2D0500FD8DF" box="[1040,1130,973,994]" pageId="1" pageNumber="228" type="holotype">Holotype</typeStatus>
- male (32 by 
<quantity id="4CF1FEEF9853DA5BEC44A2D05105D8DF" box="[1286,1376,973,994]" metricMagnitude="-2" metricUnit="m" metricValue="2.53" pageId="1" pageNumber="228" unit="mm" value="25.3">25.3 mm</quantity>
) (
<specimenCode id="DBAFFB719853DA5BEC34A2D0570EDF3D" collectionCode="PNM" country="Philippines" httpUri="http://grbio.org/cool/mxaf-c5e2" name="Philippine National Museum" pageId="1" pageNumber="228">PNM 17259</specimenCode>
), 
<taxonomicName id="4C0928899853DA5BEA3BA2F6507ADF3D" authority="River" authorityName="River" box="[889,1055,1003,1024]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="228" phylum="Arthropoda" rank="species" species="cabayugan">Cabayugan River</taxonomicName>
, 
<collectingRegion id="49CD9DE89853DA5BED64A2F65176DF3D" box="[1062,1299,1003,1024]" isEnumeration="true" pageId="1" pageNumber="228">Puerto Princesa, Palawan</collectingRegion>
, coll. 
<collectorName id="26FC36DC9853DA5BEC0AA2F651CCDF3D" box="[1352,1449,1003,1024]" pageId="1" pageNumber="228">H. Freitag</collectorName>
, 
<date id="FFB775CA9853DA5BEA6FA51557FEDF20" box="[813,923,1032,1053]" pageId="1" pageNumber="228" value="2001-06-05">
<collectingDate id="EFF38C229853DA5BEA6FA51557FEDF20" box="[813,923,1032,1053]" pageId="1" pageNumber="228" value="2001-06-05">5 Jun.2001</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8BB6530A9853DA5BEA6FA55E516EDF8D" blockId="1.[813,1452,1091,1200]" pageId="1" pageNumber="228">
<typeStatus id="54B2EDA89853DA5BEA6FA55E57EFDF65" box="[813,906,1091,1112]" pageId="1" pageNumber="228" type="paratype">Paratypes</typeStatus>
- 
<specimenCount id="9D0F98839853DA5BEAE0A55E578BDF65" box="[930,1006,1091,1112]" count="2" pageId="1" pageNumber="228" type="male">2 males</specimenCount>
(larger 19.3 by 
<quantity id="4CF1FEEF9853DA5BEDCDA55E508FDF65" box="[1167,1258,1091,1112]" metricMagnitude="-2" metricUnit="m" metricValue="1.64" pageId="1" pageNumber="228" unit="mm" value="16.4">16.4 mm</quantity>
), 
<materialsCitation id="3B6159579853DA5BEDBFA55E504ADF48" collectionCode="ZRC" pageId="1" pageNumber="228" specimenCode="ZRC2004.0622" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D0F98839853DA5BEDBFA55E5134DF65" box="[1277,1361,1091,1112]" count="1" pageId="1" pageNumber="228" type="female">1 female</specimenCount>
(21.6 by 
<quantity id="4CF1FEEF9853DA5BEA6FA57D57E5DF48" box="[813,896,1120,1141]" metricMagnitude="-2" metricUnit="m" metricValue="1.78" pageId="1" pageNumber="228" unit="mm" value="17.8">17.8 mm</quantity>
) (
<specimenCode id="DBAFFB719853DA5BEAD0A57D504CDF48" box="[914,1065,1120,1141]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="228">ZRC2004.0622</specimenCode>
)
</materialsCitation>
, 
<materialsCitation id="3B6159579853DA5BED75A57D51CDDF48" box="[1079,1448,1120,1141]" collectionCode="ZIH" pageId="1" pageNumber="228" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="9D0F98839853DA5BED75A57D501ADF48" box="[1079,1151,1120,1141]" count="2" pageId="1" pageNumber="228" type="male">2 males</specimenCount>
(larger 20.7 by 
<quantity id="4CF1FEEF9853DA5BEC52A57D5101DF48" box="[1296,1380,1120,1141]" metricMagnitude="-2" metricUnit="m" metricValue="1.72" pageId="1" pageNumber="228" unit="mm" value="17.2">17.2 mm</quantity>
) (
<collectionCode id="ED18CBCF9853DA5BEC34A57D51C4DF48" box="[1398,1441,1120,1141]" pageId="1" pageNumber="228">ZIH</collectionCode>
)
</materialsCitation>
, 
<materialsCitation id="3B6159579853DA5BEA6FA560509EDFAF" box="[813,1275,1149,1170]" collectionCode="SPCP-ASTD" pageId="1" pageNumber="228" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="9D0F98839853DA5BEA6FA5605711DFAF" box="[813,884,1149,1170]" count="2" pageId="1" pageNumber="228" type="male">2 males</specimenCount>
(largest 17.2 by 
<quantity id="4CF1FEEF9853DA5BED4FA5605007DFAF" box="[1037,1122,1149,1170]" metricMagnitude="-2" metricUnit="m" metricValue="1.47" pageId="1" pageNumber="228" unit="mm" value="14.7">14.7 mm</quantity>
) (
<collectionCode id="ED18CBCF9853DA5BED31A5605090DFAF" box="[1139,1269,1149,1170]" pageId="1" pageNumber="228">SPCP-ASTD</collectionCode>
)
</materialsCitation>
, 
<taxonomicName id="4C0928899853DA5BEC41A56051CDDFAF" authority="River" authorityName="River" box="[1283,1448,1149,1170]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="228" phylum="Arthropoda" rank="species" species="cabayugan">Cabayugan River</taxonomicName>
, 
<collectingRegion id="49CD9DE89853DA5BEA6FA5865043DF8D" box="[813,1062,1179,1200]" isEnumeration="true" pageId="1" pageNumber="228">Puerto Princesa, Palawan</collectingRegion>
, coll. 
<collectorName id="26FC36DC9853DA5BED21A58650ADDF8D" box="[1123,1224,1179,1200]" pageId="1" pageNumber="228">H. Freitag</collectorName>
, 2001.
</paragraph>
<paragraph id="8BB6530A9853DA5BEA6FA5C851C8DE18" blockId="1.[813,1453,1237,1317]" pageId="1" pageNumber="228">
<materialsCitation id="3B6159579853DA5BEA6FA5C851CCDE18" collectionCode="SPCP-ASTD" pageId="1" pageNumber="228" specimenCount="166" specimenCount-female="28" specimenCount-juvenile="100" specimenCount-male="38">
Others (designated for ecological studies) - 
<specimenCount id="9D0F98839853DA5BED97A5C85149DFD7" box="[1237,1324,1237,1258]" count="38" pageId="1" pageNumber="228" type="male">38 males</specimenCount>
(largest 14.6 by 
<quantity id="4CF1FEEF9853DA5BEA08A5EE57C7DE35" box="[842,930,1267,1288]" metricMagnitude="-2" metricUnit="m" metricValue="1.24" pageId="1" pageNumber="228" unit="mm" value="12.4">12.4 mm</quantity>
), 
<specimenCount id="9D0F98839853DA5BEAF3A5EE507CDE35" box="[945,1049,1267,1288]" count="28" pageId="1" pageNumber="228" type="female">28 females</specimenCount>
(largest 26.4 by 
<quantity id="4CF1FEEF9853DA5BEDF5A5EE516ADE35" box="[1207,1295,1267,1288]" metricMagnitude="-2" metricUnit="m" metricValue="2.16" pageId="1" pageNumber="228" unit="mm" value="21.6">21.6 mm</quantity>
),&gt; 
<specimenCount id="9D0F98839853DA5BEC6EA5EE51C9DE35" box="[1324,1452,1267,1288]" count="100" pageId="1" pageNumber="228" type="juvenile">100 juveniles</specimenCount>
(smallest 5.0 by 
<quantity id="4CF1FEEF9853DA5BEA92A40D5079DE18" box="[976,1052,1296,1317]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="1" pageNumber="228" unit="mm" value="4.5">4.5 mm</quantity>
) (
<collectionCode id="ED18CBCF9853DA5BED72A40D50D1DE18" box="[1072,1204,1296,1317]" pageId="1" pageNumber="228">SPCP-ASTD</collectionCode>
), same data as paratypes
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31300819853DA5BEA6FA45651EBDD76" pageId="1" pageNumber="228" type="diagnosis">
<paragraph id="8BB6530A9853DA5BEA6FA45651EBDD76" blockId="1.[813,1452,1355,1611]" pageId="1" pageNumber="228">
<emphasis id="B97D8F189853DA5BEA6FA45657D8DE5E" bold="true" box="[813,957,1355,1379]" italics="true" pageId="1" pageNumber="228">Diagnosis. –</emphasis>
Carapace low, dorsal surface flat; cervical grooves distinct, deep, broad; epibranchial teeth well developed, sharp, directed forwards. Merus of ambulatory legs with well developed subdistal spine on dorsal margin. Male sixth abdominal segment elongated, relatively slender, longer than telson (segment 7). G1 gently sinuous, proximal part broader than distal part; tip truncate, gently curved outwards, with distinct subdistal notch on outer margin.
</paragraph>
</subSubSection>
<subSubSection id="C31300819853DA5BEA6FA76A511BDC87" pageId="1" pageNumber="228" type="etymology">
<paragraph id="8BB6530A9853DA5BEA6FA76A512EDDED" blockId="1.[813,1452,1655,1744]" pageId="1" pageNumber="228">
<emphasis id="B97D8F189853DA5BEA6FA76A57A4DDB2" bold="true" box="[813,961,1655,1679]" italics="true" pageId="1" pageNumber="228">Etymology. –</emphasis>
The species is named for the 
<typeStatus id="54B2EDA89853DA5BEC59A76A512CDDB3" box="[1307,1353,1655,1678]" pageId="1" pageNumber="228">type</typeStatus>
locality, 
<taxonomicName id="4C0928899853DA5BEA6FA7855157DD92" authority="River and Barangay Cabayugan" authorityName="River and Barangay Cabayugan" box="[813,1330,1688,1711]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="228" phylum="Arthropoda" rank="species" species="cabayugan">Cabayugan River and Barangay Cabayugan</taxonomicName>
of 
<collectingRegion id="49CD9DE89853DA5BEC23A78557A4DDED" country="Philippines" name="Puerto Princesa" pageId="1" pageNumber="228">Puerto Princesa City</collectingRegion>
. The name is a noun in apposition.
</paragraph>
<paragraph id="8BB6530A9853DA5BEA6FA7E1511BDC87" blockId="1.[813,1452,1788,1978]" pageId="1" pageNumber="228">
<emphasis id="B97D8F189853DA5BEA6FA7E157FCDC29" bold="true" box="[813,921,1788,1812]" italics="true" pageId="1" pageNumber="228">Colour. –</emphasis>
In life, dorsal carapace and walking legs dirty greyish brown, the latter with a slight cross-stripe pattern. Ventral parts (sternum and abdomen) a dirty-white colour. Eleven out of 
<specimenCount id="9D0F98839853DA5BEA8EA67D503ADC4A" box="[972,1119,1888,1911]" count="71" pageId="1" pageNumber="228" type="generic">71 specimens</specimenCount>
with blackened fingers of the chelipeds, while the rest with greyish brown fingers. In alcohol, all brown parts become a light orange-brown.
</paragraph>
</subSubSection>
<subSubSection id="C31300819853DA5FEA6FA6FE5132DC6D" lastPageId="5" lastPageNumber="232" pageId="1" pageNumber="228" type="discussion">
<paragraph id="8BB6530A9853DA5EEA6FA6FE5561DAC5" blockId="1.[813,1452,2019,2107]" lastBlockId="4.[136,775,193,504]" lastPageId="4" lastPageNumber="231" pageId="1" pageNumber="228">
<emphasis id="B97D8F189853DA5BEA6FA6FE509ADCC6" box="[813,1279,2019,2043]" italics="true" pageId="1" pageNumber="228">
<emphasis id="B97D8F189853DA5BEA6FA6FE57DDDCC6" bold="true" box="[813,952,2019,2043]" italics="true" pageId="1" pageNumber="228">Remarks. –</emphasis>
<taxonomicName id="4C0928899853DA5BEA87A6F9509ADCC6" authorityName="Freitag &amp; Yeo" authorityYear="2004" box="[965,1279,2020,2043]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="228" phylum="Arthropoda" rank="species" species="cabayugan" status="sp. nov.">Parathelphusa cabayugan</taxonomicName>
</emphasis>
, 
<taxonomicNameLabel id="A24E32639853DA5BEC51A6FE51CDDCC7" box="[1299,1448,2019,2042]" pageId="1" pageNumber="228" rank="species">new species</taxonomicNameLabel>
, superficially resembles several known species of Philippine 
<taxonomicName id="4C0928899853DA5BEA6FA93957B4D306" authorityName="H. MILNE EDWARDS" authorityYear="1853" box="[813,977,2084,2107]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="228" phylum="Arthropoda" rank="genus">
<emphasis id="B97D8F189853DA5BEA6FA93957B4D306" box="[813,977,2084,2107]" italics="true" pageId="1" pageNumber="228">Parathelphusa</emphasis>
</taxonomicName>
. For example, the sharp, well-developed epibranchial teeth of 
<taxonomicName id="4C0928899856DA5EE838A1DC5676DBE5" authorityName="Freitag &amp; Yeo" authorityYear="2004" box="[378,531,193,216]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="231" phylum="Arthropoda" rank="species" species="cabayugan">
<emphasis id="B97D8F189856DA5EE838A1DC5676DBE5" box="[378,531,193,216]" italics="true" pageId="4" pageNumber="231">P. cabayugan</emphasis>
</taxonomicName>
resemble those of 
<taxonomicName id="4C0928899856DA5EEBB2A1DC55C1DBC5" authority="(Bott, 1969)" baseAuthorityName="Bott" baseAuthorityYear="1969" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="231" phylum="Arthropoda" rank="species" species="palawanensis">
<emphasis id="B97D8F189856DA5EEBB2A1DC557CDBC5" italics="true" pageId="4" pageNumber="231">P. palawanensis</emphasis>
(
<bibRefCitation id="EF982EFB9856DA5EE865A1FC55F9DBC5" author="Bott, R" box="[295,412,225,248]" pageId="4" pageNumber="231" pagination="359 - 366" refId="ref4975" refString="Bott, R., 1969. Flusskrabben aus Asien und ihre Klassifikation. (Crustacea, Decapoda). Senckenbergiana biologica, Frankfurt, 50 (5 / 6): 359 - 366." type="journal article" year="1969">Bott, 1969</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C0928899856DA5EE8ECA1FC5766DBC5" authority="Ng &amp; Takeda, 1993" authorityName="Ng &amp; Takeda" authorityYear="1993" box="[430,771,225,248]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="231" phylum="Arthropoda" rank="species" species="saginata">
<emphasis id="B97D8F189856DA5EE8ECA1FC564DDBC5" box="[430,552,225,248]" italics="true" pageId="4" pageNumber="231">P. saginata</emphasis>
Ng &amp; Takeda, 1993
</taxonomicName>
, 
<taxonomicName id="4C0928899856DA5EE9CAA01C55AADA25" authority="Ng &amp; Takeda, 1993" authorityName="Ng &amp; Takeda" authorityYear="1993" box="[136,463,257,280]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="231" phylum="Arthropoda" rank="species" species="mindoro">
<emphasis id="B97D8F189856DA5EE9CAA01C5499DA25" box="[136,252,257,280]" italics="true" pageId="4" pageNumber="231">P. mindoro</emphasis>
Ng &amp; Takeda, 1993
</taxonomicName>
, and 
<emphasis id="B97D8F189856DA5EEB40A01C56E3DA25" box="[514,646,257,280]" italics="true" pageId="4" pageNumber="231">
<taxonomicName id="4C0928899856DA5EEB40A01C56E7DA25" authorityName="Freitag &amp; Yeo" authorityYear="2004" box="[514,642,257,280]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="231" phylum="Arthropoda" rank="species" species="manguao" status="new species">P. manguao</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A24E32639856DA5EEBC9A01C5762DA25" box="[651,775,257,280]" pageId="4" pageNumber="231" rank="species">new species</taxonomicNameLabel>
(present study). In addition, the truncate tip of the G1 of 
<taxonomicName id="4C0928899856DA5EEBB2A03C5498DA65" authorityName="Freitag &amp; Yeo" authorityYear="2004" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="231" phylum="Arthropoda" rank="species" species="cabayugan">
<emphasis id="B97D8F189856DA5EEBB2A03C5498DA65" italics="true" pageId="4" pageNumber="231">P. cabayugan</emphasis>
</taxonomicName>
resembles that seen in 
<taxonomicName id="4C0928899856DA5EE8B2A05C56BEDA65" authority="(Bott, 1969)" baseAuthorityName="Bott" baseAuthorityYear="1969" box="[496,731,321,344]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="231" phylum="Arthropoda" rank="species" species="obtusa">
<emphasis id="B97D8F189856DA5EE8B2A05C5631DA65" box="[496,596,321,344]" italics="true" pageId="4" pageNumber="231">P. obtusa</emphasis>
(
<bibRefCitation id="EF982EFB9856DA5EEB23A05C56B6DA65" author="Bott, R" box="[609,723,321,344]" pageId="4" pageNumber="231" pagination="359 - 366" refId="ref4975" refString="Bott, R., 1969. Flusskrabben aus Asien und ihre Klassifikation. (Crustacea, Decapoda). Senckenbergiana biologica, Frankfurt, 50 (5 / 6): 359 - 366." type="journal article" year="1969">Bott, 1969</bibRefCitation>
)
</taxonomicName>
and 
<taxonomicName id="4C0928899856DA5EE9CAA07C55A7DA45" authority="Ng &amp; Takeda, 1993" authorityName="Ng &amp; Takeda" authorityYear="1993" box="[136,450,353,376]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="231" phylum="Arthropoda" rank="species" species="rasilis">
<emphasis id="B97D8F189856DA5EE9CAA07C548DDA45" box="[136,232,353,376]" italics="true" pageId="4" pageNumber="231">P. rasilis</emphasis>
Ng &amp; Takeda, 1993
</taxonomicName>
; and the overall form of its G1 is similar to that of 
<taxonomicName id="4C0928899856DA5EE8D1A09C5766DAA5" authority="Ng &amp; Takeda, 1993" authorityName="Ng &amp; Takeda" authorityYear="1993" box="[403,771,385,408]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="231" phylum="Arthropoda" rank="species" species="nana">
<emphasis id="B97D8F189856DA5EE8D1A09C5591DAA5" box="[403,500,385,408]" italics="true" pageId="4" pageNumber="231">P. nana</emphasis>
Ng &amp; Takeda, 1993
</taxonomicName>
. 
<taxonomicName id="4C0928899856DA5EE9CAA0BC55B8DA85" authority="Ng, 1994" authorityName="Ng" authorityYear="1994" box="[136,477,417,440]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="231" phylum="Arthropoda" rank="species" species="ovum">
<emphasis id="B97D8F189856DA5EE9CAA0BC550FDA85" box="[136,362,417,440]" italics="true" pageId="4" pageNumber="231">Parathelphusa ovum</emphasis>
<bibRefCitation id="EF982EFB9856DA5EE836A0BC55B8DA85" author="Ng, P" box="[372,477,417,440]" pageId="4" pageNumber="231" pagination="73 - 92" refId="ref5492" refString="Ng, P. K. L., 1994. On a collection of freshwater decapod crustaceans from the Kinabatangan River, Sabah, Malaysia, with descriptions of three new species. Sabah Museum Journal, 1 (2): 73 - 92." type="journal article" year="1994">Ng, 1994</bibRefCitation>
</taxonomicName>
, from 
<collectingRegion id="49CD9DE89856DA5EEB68A0BC560BDA85" box="[554,622,417,440]" country="Malaysia" name="Sabah" pageId="4" pageNumber="231">Sabah</collectingRegion>
, Borneo, also possesses a G1 with a truncate tip that is similar to that of 
<taxonomicName id="4C0928899856DA5EEBB2A0DC5498DAC5" authorityName="Freitag &amp; Yeo" authorityYear="2004" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="231" phylum="Arthropoda" rank="species" species="cabayagan">
<emphasis id="B97D8F189856DA5EEBB2A0DC5498DAC5" italics="true" pageId="4" pageNumber="231">P. cabayagan</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="DF7603829850DA58E9CAA6E954D1D37F" ID-DOI="http://doi.org/10.5281/zenodo.13244566" ID-Zenodo-Dep="13244566" httpUri="https://zenodo.org/record/13244566/files/figure.png" pageId="2" pageNumber="229" startId="2.[136,171,2036,2057]" targetBox="[165,1434,196,1966]" targetPageId="2" targetType="figure">
<paragraph id="8BB6530A9850DA58E9CAA6E954D1D37F" blockId="2.[136,1476,2036,2114]" pageId="2" pageNumber="229">
Fig. 1. 
<emphasis id="B97D8F189850DA58E98FA6E955B5D334" box="[205,464,2036,2057]" italics="true" pageId="2" pageNumber="229">
<taxonomicName id="4C0928899850DA58E98FA6E955A8D334" authorityName="Freitag &amp; Yeo" authorityYear="2004" box="[205,461,2036,2057]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="229" phylum="Arthropoda" rank="species" species="cabayugan" status="sp. nov.">Parathelphusa cabayugan</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A24E32639850DA58E894A6E95628D334" box="[470,589,2036,2057]" pageId="2" pageNumber="229" rank="species">new species</taxonomicNameLabel>
, holotype male (32 by 25.3 mm) (PNM 17259). A, dorsal view of carapace; B, frontal view of carapace; C, right third ambulatory merus; D, right fourth ambulatory merus; E, left chela; F, abdominal segments 5-7. Scales = 10.0 mm.
</paragraph>
</caption>
<caption id="DF7603829851DA59E933A6EC55D5D37D" ID-DOI="http://doi.org/10.5281/zenodo.13244568" ID-Zenodo-Dep="13244568" httpUri="https://zenodo.org/record/13244568/files/figure.png" pageId="3" pageNumber="230" startId="3.[113,148,2033,2054]" targetBox="[154,1408,195,1984]" targetPageId="3" targetType="figure">
<paragraph id="8BB6530A9851DA59E933A6EC55D5D37D" blockId="3.[113,1452,2033,2112]" pageId="3" pageNumber="230">
Fig. 2. 
<emphasis id="B97D8F189851DA59E9FAA6EC55D8D33B" box="[184,445,2033,2054]" italics="true" pageId="3" pageNumber="230">
<taxonomicName id="4C0928899851DA59E9FAA6EC55DCD33B" authorityName="Freitag &amp; Yeo" authorityYear="2004" box="[184,441,2033,2054]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="230" phylum="Arthropoda" rank="species" species="cabayugan" status="sp. nov.">Parathelphusa cabayugan</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A24E32639851DA59E886A6EC565ED33B" box="[452,571,2033,2054]" pageId="3" pageNumber="230" rank="species">new species</taxonomicNameLabel>
, left G1. A, B, E, F, holotype male (32 by 25.3 mm) (PNM 17259); C, D, G, H, paratype juvenile male (15.2 by 13.1 mm) (SPCP-ASTD). A, C, ventral view; B, D, dorsal view; E, G, ventral view of distal part; F, H, dorsal view of distal part. Scales = 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BB6530A9856DA5EE9CAA33C56ADD985" blockId="4.[136,775,545,696]" pageId="4" pageNumber="231">
<taxonomicName id="4C0928899856DA5EE9CAA33C55CFD905" authorityName="Freitag &amp; Yeo" authorityYear="2004" box="[136,426,545,568]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="231" phylum="Arthropoda" rank="species" species="cabayugan">
<emphasis id="B97D8F189856DA5EE9CAA33C55CFD905" box="[136,426,545,568]" italics="true" pageId="4" pageNumber="231">Parathelphusa cabayugan</emphasis>
</taxonomicName>
, however, can be immediately separated from these and all other 
<taxonomicName id="4C0928899856DA5EE8B7A35C5683D965" box="[501,742,577,600]" pageId="4" pageNumber="231">
<emphasis id="B97D8F189856DA5EE8B7A35C56F7D965" box="[501,658,577,600]" italics="true" pageId="4" pageNumber="231">Parathelphusa</emphasis>
species
</taxonomicName>
by the presence of a distinct subdistal notch on the outer margin of the tip of its G1, in addition to a unique combination of external and other G1 characters (see Tables 1A, B).
</paragraph>
<paragraph id="8BB6530A9856DA5EE9CAA3FC57BFDBC5" blockId="4.[136,775,737,792]" lastBlockId="4.[835,1475,193,248]" pageId="4" pageNumber="231">
The diagnostic subdistal notch on the outer margin of the tip of the G1 and the sharp, well-developed epibranchial teeth are also present in juvenile specimens of 
<taxonomicName id="4C0928899856DA5EEDADA1DC51E5DBE5" authorityName="Freitag &amp; Yeo" authorityYear="2004" box="[1263,1408,193,216]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="231" phylum="Arthropoda" rank="species" species="cabayugan">
<emphasis id="B97D8F189856DA5EEDADA1DC51E5DBE5" box="[1263,1408,193,216]" italics="true" pageId="4" pageNumber="231">P. cabayugan</emphasis>
</taxonomicName>
(
<figureCitation id="13324F8F9856DA5EECCFA1DC57ABDBC5" captionStart="Fig" captionStartId="3.[113,148,2033,2054]" captionTargetBox="[154,1408,195,1984]" captionTargetId="figure-12@3.[135,1443,160,2008]" captionTargetPageId="3" captionText="Fig. 2. Parathelphusa cabayugan, new species, left G1. A, B, E, F, holotype male (32 by 25.3 mm) (PNM 17259); C, D, G, H, paratype juvenile male (15.2 by 13.1 mm) (SPCP-ASTD). A, C, ventral view; B, D, dorsal view; E, G, ventral view of distal part; F, H, dorsal view of distal part. Scales = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13244568" httpUri="https://zenodo.org/record/13244568/files/figure.png" pageId="4" pageNumber="231">Figs. 2C, D, G, H</figureCitation>
).
</paragraph>
<paragraph id="8BB6530A9856DA5EEA01A03C51F3D985" blockId="4.[835,1475,289,696]" pageId="4" pageNumber="231">
<emphasis id="B97D8F189856DA5EEA01A03C5071DA04" bold="true" box="[835,1044,289,313]" italics="true" pageId="4" pageNumber="231">Ecological notes. –</emphasis>
The habitats where 
<taxonomicName id="4C0928899856DA5EEDAFA03C51E5DA05" authorityName="Freitag &amp; Yeo" authorityYear="2004" box="[1261,1408,289,312]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="231" phylum="Arthropoda" rank="species" species="cabayugan">
<emphasis id="B97D8F189856DA5EEDAFA03C51E5DA05" box="[1261,1408,289,312]" italics="true" pageId="4" pageNumber="231">P. cabayugan</emphasis>
</taxonomicName>
occur are in the 
<taxonomicName id="4C0928899856DA5EEAE8A05C5007DA65" authority="River" authorityName="River" box="[938,1122,321,344]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="231" phylum="Arthropoda" rank="species" species="cabayugan">Cabayugan River</taxonomicName>
catchment (
<geoCoordinate id="EE3D35CD9856DA5EEDA1A05C5127DA65" box="[1251,1346,321,344]" degrees="10" direction="north" minutes="09" orientation="latitude" pageId="4" pageNumber="231" precision="925" value="10.15">10 09’N</geoCoordinate>
, 
<geoCoordinate id="EE3D35CD9856DA5EEC0EA05C51D0DA65" box="[1356,1461,321,344]" degrees="118" direction="east" minutes="51" orientation="longitude" pageId="4" pageNumber="231" precision="925" value="118.85">118 51’E</geoCoordinate>
), which has an area of approximately 3,683 ha (
<bibRefCitation id="EF982EFB9856DA5EEC11A07C5003DAA5" author="PAMB of Puerto Princesa City" pageId="4" pageNumber="231" pagination="1 - 9" refId="ref5539" refString="PAMB of Puerto Princesa City, 1998. St. Paul Subterranean National Park Management Plan. Pp. i-viii, 1 - 68, Figs. 1 - 9, unpublished." type="book chapter" year="1998">
PAMB of 
<collectingRegion id="49CD9DE89856DA5EEA01A09C5046DAA5" box="[835,1059,385,408]" country="Philippines" name="Puerto Princesa" pageId="4" pageNumber="231">Puerto Princesa City</collectingRegion>
, 1998
</bibRefCitation>
, unpublished). The catchment is surrounded by mountains up to 
<quantity id="4CF1FEEF9856DA5EED8AA0BC5147DA85" box="[1224,1314,417,440]" metricMagnitude="3" metricUnit="m" metricValue="1.028" pageId="4" pageNumber="231" unit="m" value="1028.0">1028 m</quantity>
high, but all permanent tributaries and sites sampled are lowland waters. The soils drained are yellowish to deep red clays on ultrabasic subsoils at the peripheral headwaters; alluvial soils on shale and sandstone at the upper to middle river course; and karst soils on limestone at the lower and subterranean course of Cabayugan/Underground River. This results in the pH being generally basic, ranging from 7.1 to 8.7, and the conductivity being relatively high, ranging from 140 to 740 S/cm.
</paragraph>
<paragraph id="8BB6530A9856DA5EEA01A3FC57EADF65" blockId="4.[835,1474,737,1112]" pageId="4" pageNumber="231">
Highest abundance of 
<taxonomicName id="4C0928899856DA5EED68A3FC50DDD9C5" box="[1066,1208,737,760]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="231" phylum="Arthropoda" rank="species" species="cabayugan">
<emphasis id="B97D8F189856DA5EED68A3FC50DDD9C5" box="[1066,1208,737,760]" italics="true" pageId="4" pageNumber="231">P. cabayugan</emphasis>
</taxonomicName>
was found in undisturbed first to third order headwaters with moderate to low turbidity (&gt; 
<quantity id="4CF1FEEF9856DA5EEA20A23C57D7D805" box="[866,946,801,824]" metricMagnitude="0" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="231" unit="cm" value="120.0">120 cm</quantity>
light penetration). The specimens were collected from the river bed and in colonization baskets situated between stony and course organic substrates. Within such river sections, the crabs were found in different micro-habitats such as under small falls, in riffles, and in pools. Applying drift nets at all sites, several specimens were caught only at a spring brook running out off limestone rocks. Dissolved oxygen was occasionally found decreasing to 60% saturation in this site. All other sites sampled had generally higher values.
</paragraph>
<paragraph id="8BB6530A9856DA5FEA01A59C54D2DD4E" blockId="4.[835,1475,1153,1560]" lastBlockId="5.[113,753,1562,1651]" lastPageId="5" lastPageNumber="232" pageId="4" pageNumber="231">
Lower abundance of 
<taxonomicName id="4C0928899856DA5EED64A59C50DCDFA5" box="[1062,1209,1153,1176]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="231" phylum="Arthropoda" rank="species" species="cabayugan">
<emphasis id="B97D8F189856DA5EED64A59C50DCDFA5" box="[1062,1209,1153,1176]" italics="true" pageId="4" pageNumber="231">P. cabayugan</emphasis>
</taxonomicName>
was found in the middle river course that is influenced by anthropogenic activities (paddy fields, settlements). Those result in higher temperature fluctuations (26 C 3 C), raised turbidity and slightly increased BOD 
<subScript id="178D514F9856DA5EEAA5A413579ADE21" attach="left" box="[999,1023,1294,1308]" fontSize="6" pageId="4" pageNumber="231">2+5</subScript>
(max: 
<quantity id="4CF1FEEF9856DA5EED0EA41C50FCDE25" box="[1100,1177,1281,1304]" metricMagnitude="-6" metricUnit="kg" metricValue="2.6" pageId="4" pageNumber="231" unit="mg" value="2.6">2.6 mg</quantity>
/l, n=5). In paddy fields and their draining channels, no individuals of the species were found. This is in contrast to other parathelphusid crabs such as members of 
<taxonomicName id="4C0928899856DA5EED79A47C50C4DE45" authorityName="Naiyanetr" authorityYear="1994" box="[1083,1185,1377,1400]" class="Malacostraca" family="Gecarcinucidae" genus="Sayamia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="231" phylum="Arthropoda" rank="genus">
<emphasis id="B97D8F189856DA5EED79A47C50C4DE45" box="[1083,1185,1377,1400]" italics="true" pageId="4" pageNumber="231">Sayamia</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C0928899856DA5EED8AA47C511CDE45" authorityName="Naiyanetr" authorityYear="1994" box="[1224,1401,1377,1400]" class="Malacostraca" family="Gecarcinucidae" genus="Esanthelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="231" phylum="Arthropoda" rank="genus">
<emphasis id="B97D8F189856DA5EED8AA47C511CDE45" box="[1224,1401,1377,1400]" italics="true" pageId="4" pageNumber="231">Esanthelphusa</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0928899856DA5EEA01A49C5070DEA5" authorityName="Bott" authorityYear="1968" box="[835,1045,1409,1432]" class="Malacostraca" family="Gecarcinucidae" genus="Somanniathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="231" phylum="Arthropoda" rank="genus">
<emphasis id="B97D8F189856DA5EEA01A49C5070DEA5" box="[835,1045,1409,1432]" italics="true" pageId="4" pageNumber="231">Somanniathelphusa</emphasis>
</taxonomicName>
that occur in rice fields in many parts of Indochina and 
<collectingCountry id="F31E139A9856DA5EEAA3A4BC5044DE85" box="[993,1057,1441,1464]" name="China" pageId="4" pageNumber="231">China</collectingCountry>
(see Ng &amp; Naiyanetr, 1993; 
<bibRefCitation id="EF982EFB9856DA5EEC10A4BC5719DEE5" author="Naiyanetr, P" pageId="4" pageNumber="231" pagination="695 - 700" refId="ref5143" refString="Naiyanetr, P., 1994. On three new genera of Thai ricefield crabs allied to Somanniathelphusa Bott, 1968 (Crustacea: Decapoda: Brachyura: Parathelphusidae). Raffles Bulletin of Zoology, 42 (3): 695 - 700." type="journal article" year="1994">Naiyanetr, 1994</bibRefCitation>
; 
<bibRefCitation id="EF982EFB9856DA5EEAC9A4DC5792DEE5" author="Ng, P" box="[907,1015,1473,1496]" pageId="4" pageNumber="231" pagination="26 - 30" refId="ref5240" refString="Ng, P. K. L., 1995. Somanniathelphusa lacuvita, a new ricefield crab from Tonle Sap, Cambodia (Crustacea: Brachyura: Parathelphusidae). Asian Journal of Tropical Biology, 1: 26 - 30." type="journal article" year="1995">Ng, 1995</bibRefCitation>
; 
<bibRefCitation id="EF982EFB9856DA5EED44A4DC5012DEE5" author="Dai, A" box="[1030,1143,1473,1496]" pageId="4" pageNumber="231" refId="ref5011" refString="Dai, A. Y., 1999. Fauna Sinica (Arthropoda. Crustacea. Malacostraca. Decapoda. Parathelphusidae. Potamidae). Editorial Committee of Fauna Sinica, Academia Sinica, Science Press, Beijing, 501 pp, 238 figs., 30 pls." type="book" year="1999">Dai, 1999</bibRefCitation>
; Yeo &amp; Nguyen, 1999). It is possible that the absence or low abundance of 
<taxonomicName id="4C0928899856DA5EEC72A4FC51A7DEC5" box="[1328,1474,1505,1528]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="231" phylum="Arthropoda" rank="species" species="cabayugan">
<emphasis id="B97D8F189856DA5EEC72A4FC51A7DEC5" box="[1328,1474,1505,1528]" italics="true" pageId="4" pageNumber="231">P. cabayugan</emphasis>
</taxonomicName>
in the middle parts of the river may be due to natural physical/ environmental variables. However, the use of pesticides and fertilizers in the nearby paddy fields may also have played a part.
</paragraph>
<caption id="DF7603829856DA5EE9CAA6E954DBD37F" ID-DOI="http://doi.org/10.5281/zenodo.13244570" ID-Zenodo-Dep="13244570" httpUri="https://zenodo.org/record/13244570/files/figure.png" pageId="4" pageNumber="231" startId="4.[136,171,2036,2057]" targetBox="[136,776,870,2013]" targetPageId="4" targetType="figure">
<paragraph id="8BB6530A9856DA5EE9CAA6E954DBD37F" blockId="4.[136,776,2036,2114]" pageId="4" pageNumber="231">
Fig. 3. 
<emphasis id="B97D8F189856DA5EE98FA6E955B4D334" box="[205,465,2036,2057]" italics="true" pageId="4" pageNumber="231">
<taxonomicName id="4C0928899856DA5EE98FA6E955A8D334" authorityName="Freitag &amp; Yeo" authorityYear="2004" box="[205,461,2036,2057]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="231" phylum="Arthropoda" rank="species" species="cabayugan" status="sp. nov.">Parathelphusa cabayugan</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A24E32639856DA5EE895A6E9562BD334" box="[471,590,2036,2057]" pageId="4" pageNumber="231" rank="species">new species</taxonomicNameLabel>
, holotype male (32 by 25.3 mm) (PNM 17259). A, dorsal view; B frontal view; C, ventral view.
</paragraph>
</caption>
<caption id="DF7603829856DA5EEA07A9125021D37C" ID-DOI="http://doi.org/10.5281/zenodo.13244572" ID-Zenodo-Dep="13244572" httpUri="https://zenodo.org/record/13244572/files/figure.png" pageId="4" pageNumber="231" startId="4.[837,872,2063,2084]" targetBox="[835,1475,1603,2045]" targetPageId="4" targetType="figure">
<paragraph id="8BB6530A9856DA5EEA07A9125021D37C" blockId="4.[837,1476,2063,2113]" pageId="4" pageNumber="231">
Fig. 4. 
<emphasis id="B97D8F189856DA5EEACDA90D50F2D318" box="[911,1175,2064,2085]" italics="true" pageId="4" pageNumber="231">
<taxonomicName id="4C0928899856DA5EEACDA90D50F6D318" authorityName="Freitag &amp; Yeo" authorityYear="2004" box="[911,1171,2064,2085]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="231" phylum="Arthropoda" rank="species" species="cabayugan" status="sp. nov.">Parathelphusa cabayugan</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A24E32639856DA5EEDDDA912517CD319" box="[1183,1305,2063,2084]" pageId="4" pageNumber="231" rank="species">new species</taxonomicNameLabel>
, paratype female (26.4 by 21.6 mm) (ZIH).
</paragraph>
</caption>
<paragraph id="8BB6530A9857DA5FE933A78356FADDEB" blockId="5.[113,753,1694,1750]" pageId="5" pageNumber="232">The species was not detected in the other stretches of the river, viz., the subterranean course and the estuary.</paragraph>
<paragraph id="8BB6530A9857DA5FE933A61C54CFDCE2" blockId="5.[113,752,1793,2015]" pageId="5" pageNumber="232">The species appears to be primarily aquatic in habits, as majority of the specimens were collected from within the river. However, a single specimen caught on the river bank about 1 metre above the water surface, in an emergence trap for eclosing aquatic insects, suggests that the members of the species actively leave the water body for short times at least.</paragraph>
<paragraph id="8BB6530A9857DA5FE933A91757BBDDAD" blockId="5.[113,752,2058,2114]" lastBlockId="5.[813,1451,1561,1680]" pageId="5" pageNumber="232">
Juvenile stages (&lt;5 by 
<quantity id="4CF1FEEF9857DA5FE827A91755D2D31C" box="[357,439,2058,2081]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="5" pageNumber="232" unit="mm" value="4.5">4.5 mm</quantity>
) assigned to this species were found several times throughout the year at the same stream courses as the adults. 
<specimenCount id="9D0F98839857DA5FED62A70450FADD0D" box="[1056,1183,1561,1584]" count="1" pageId="5" pageNumber="232" type="female">One female</specimenCount>
(26.4 by 
<quantity id="4CF1FEEF9857DA5FEC4CA7045114DD0D" box="[1294,1393,1561,1584]" metricMagnitude="-2" metricUnit="m" metricValue="2.16" pageId="5" pageNumber="232" unit="mm" value="21.6">21.6 mm</quantity>
) was found carrying 
<specimenCount id="9D0F98839857DA5FEA92A7245034DD6D" box="[976,1105,1593,1616]" count="54" pageId="5" pageNumber="232" type="juvenile">54 juveniles</specimenCount>
of 2.0 by 
<quantity id="4CF1FEEF9857DA5FEDF9A7245169DD6D" box="[1211,1292,1593,1616]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="5" pageNumber="232" unit="mm" value="1.9">1.9 mm</quantity>
size. Juveniles of the same size were detected as smallest among the released free living ones.
</paragraph>
<paragraph id="8BB6530A9857DA5FEA6FA7A45132DC6D" blockId="5.[813,1452,1721,1872]" pageId="5" pageNumber="232">
The data from this semi-quantitative collection seems to indicate that 
<taxonomicName id="4C0928899857DA5FEA80A7C4503EDDCD" authorityName="Freitag &amp; Yeo" authorityYear="2004" box="[962,1115,1753,1776]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="232" phylum="Arthropoda" rank="species" species="cabayugan">
<emphasis id="B97D8F189857DA5FEA80A7C4503EDDCD" box="[962,1115,1753,1776]" italics="true" pageId="5" pageNumber="232">P. cabayugan</emphasis>
</taxonomicName>
has an affinity for small and undisturbed headwater streams, with clear and unpolluted water. The preferred habitats overlap in parts with those of the potamid genus 
<taxonomicName id="4C0928899857DA5FEABFA6245136DC6D" authority="Ng &amp; Takeda, 1992" authorityName="Ng &amp; Takeda" authorityYear="1992" box="[1021,1363,1849,1872]" class="Malacostraca" family="Potamidae" genus="Insulamon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="232" phylum="Arthropoda" rank="genus">
<emphasis id="B97D8F189857DA5FEABFA624500BDC6D" box="[1021,1134,1849,1872]" italics="true" pageId="5" pageNumber="232">Insulamon</emphasis>
Ng &amp; Takeda, 1992
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>